treatments-xml/data/03/A3/87/03A387AEFFFCFF9AFF79FBC8A63AE959.xml

<document id="8281D4715623210BAF24E610323A9F81" ID-DOI="10.11646/zootaxa.3957.2.4" ID-GBIF-Dataset="1bead3a5-b5c1-4562-b3be-1cd40fe7bafb" ID-ISSN="1175-5326" ID-Zenodo-Dep="240263" ID-ZooBank="116FFA26-515E-49E8-9180-719B9B58F758" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461060847470" checkinUser="plazi" docAuthor="Turvey, Samuel T., Hansford, James, Kennerley, Rosalind J., Nuñez-Miño, José M., Brocca, Jorge L. &amp; Young, Richard P." docDate="2015" docId="03A387AEFFFCFF9AFF79FBC8A63AE959" docLanguage="en" docName="zt03957p214.pdf" docOrigin="Zootaxa 3957 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Plagiodontia aedium subsp. bondi Turvey, Hansford, Kennerley, Nuñez-Miño, Brocca &amp; Young, 2015, subsp. nov." docType="treatment" docVersion="8" lastPageNumber="210" masterDocId="FF9AFFD6FFFAFF93FFEEFFBFA526EA4D" masterDocTitle="A new subspecies of hutia (Plagiodontia, Capromyidae, Rodentia) from southern Hispaniola" masterLastPageNumber="214" masterPageNumber="201" pageNumber="207" updateTime="1698582131793" updateUser="plazi">
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<mods:title id="73429EA9364B708364D9A9685C2A0983">A new subspecies of hutia (Plagiodontia, Capromyidae, Rodentia) from southern Hispaniola</mods:title>
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<mods:namePart id="45A3528D95B1B464BB74FE26F020BCF1">Hansford, James</mods:namePart>
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<mods:namePart id="0ADDB63A09EA4C6F2A71EF02DDBE5EE5">Kennerley, Rosalind J.</mods:namePart>
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<mods:namePart id="C6D3F796519BCF9A5CEA6D299B0B22B8">Nuñez-Miño, José M.</mods:namePart>
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<mods:namePart id="6C2207072D64304FA5EAA017B37BAA43">Brocca, Jorge L.</mods:namePart>
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<mods:namePart id="2FFA04B4DCAB07CCC5F7E700D2A920B0">Young, Richard P.</mods:namePart>
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<treatment id="03A387AEFFFCFF9AFF79FBC8A63AE959" ID-DOI="http://doi.org/10.5281/zenodo.5682450" ID-GBIF-Taxon="119614093" ID-Zenodo-Dep="5682450" LSID="urn:lsid:plazi:treatment:03A387AEFFFCFF9AFF79FBC8A63AE959" httpUri="http://treatment.plazi.org/id/03A387AEFFFCFF9AFF79FBC8A63AE959" lastPageId="9" lastPageNumber="210" pageId="6" pageNumber="207">
<subSubSection id="C3106533FFFCFF95FF79FBC8A659EEDF" box="[151,895,1143,1170]" pageId="6" pageNumber="207" type="nomenclature">
<paragraph id="8BB536B8FFFCFF95FF79FBC8A659EEDF" blockId="6.[151,895,1143,1170]" box="[151,895,1143,1170]" pageId="6" pageNumber="207">
<heading id="D0FD81D4FFFCFF95FF79FBC8A659EEDF" bold="true" box="[151,895,1143,1170]" fontSize="11" level="1" pageId="6" pageNumber="207" reason="1">
<emphasis id="B97EEAAAFFFCFF95FF79FBC8A659EEDF" bold="true" box="[151,895,1143,1170]" pageId="6" pageNumber="207">
<taxonomicName id="4C0A4D3BFFFCFF95FF79FBC8A4F9EEDC" box="[151,479,1143,1169]" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="207" phylum="Chordata" rank="subSpecies" species="aedium" status="subsp. nov." subSpecies="bondi">
<emphasis id="B97EEAAAFFFCFF95FF79FBC8A4F9EEDC" bold="true" box="[151,479,1143,1169]" italics="true" pageId="6" pageNumber="207">Plagiodontia aedium bondi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57D1FFFCFF95FE0BFBC7A749EEDF" box="[485,623,1144,1170]" pageId="6" pageNumber="207" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
(James Bond’s hutia)
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BB536B8FFFCFF95FF79FB01A14CEEB6" blockId="6.[151,1436,1214,2031]" pageId="6" pageNumber="207">
<emphasis id="B97EEAAAFFFCFF95FF79FB01A42DEE9A" bold="true" box="[151,267,1214,1239]" pageId="6" pageNumber="207">
<typeStatus id="54B1881AFFFCFF95FF79FB01A420EE9A" box="[151,262,1214,1239]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
.
</emphasis>
LDUCZ-Z2807, complete skeleton of adult individual (
<figureCitation id="13312A3DFFFCFF95FC92FB01A6F9EE9A" box="[892,991,1214,1239]" captionStart="FIGURE 4" captionStartId="8.[151,250,1549,1571]" captionTargetBox="[154,1433,879,1523]" captionTargetId="figure@8.[151,1435,877,1527]" captionTargetPageId="8" captionText="FIGURE 4. Holotype skull and mandible of Plagiodontia aedium bondi (LDUCZ-Z 2807): a, dorsal view of skull; b, ventral view of skull; c, lateral view of skull; d, external view of mandible. Scale bar = 1 cm." httpUri="https://zenodo.org/record/240267/files/figure.png" pageId="6" pageNumber="207">Figure 4</figureCitation>
). Found dead in ravine, apparently after fatal fall from tree branch; collected by Ramon “Moncho” Espinal on 
<date id="FFB41078FFFCFF95FC46FB5DA141EEB6" box="[936,1127,1250,1275]" pageId="6" pageNumber="207" value="2011-10-25">25 October 2011</date>
.
</paragraph>
<paragraph id="8BB536B8FFFCFF95FF29FAB9A473EF2A" blockId="6.[151,1436,1214,2031]" pageId="6" pageNumber="207">
<emphasis id="B97EEAAAFFFCFF95FF29FAB9A41DEF52" bold="true" box="[199,315,1286,1311]" pageId="6" pageNumber="207">
<typeStatus id="54B1881AFFFCFF95FF29FAB9A410EF52" box="[199,310,1286,1311]" pageId="6" pageNumber="207" type="paratype">Paratype</typeStatus>
.
</emphasis>
LDUCZ-Z2808, complete skeleton of young adult with full tooth eruption and substantial but not complete closure of ectocranial sutures; possibly offspring of 
<typeStatus id="54B1881AFFFCFF95FCBAFA95A690EF0E" box="[852,950,1322,1347]" pageId="6" pageNumber="207" type="holotype">holotype</typeStatus>
individual. Collected at same location and date as 
<typeStatus id="54B1881AFFFCFF95FF05FAF1A477EF2A" box="[235,337,1358,1383]" pageId="6" pageNumber="207" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3106533FFFCFF95FF29FACDA648EC2E" pageId="6" pageNumber="207" type="etymology">
<paragraph id="8BB536B8FFFCFF95FF29FACDA46CEF9F" blockId="6.[151,1436,1214,2031]" pageId="6" pageNumber="207">
<emphasis id="B97EEAAAFFFCFF95FF29FACDA468EFC6" bold="true" box="[199,334,1394,1419]" pageId="6" pageNumber="207">Etymology.</emphasis>
Named after the ornithologist James Bond (
<date id="FFB41078FFFCFF95FCA1FACCA6F5EFC6" box="[847,979,1395,1419]" pageId="6" pageNumber="207" value="1900" valueMax="1989">1900–1989</date>
), to acknowledge his recognition of the biogeographic line in southern Hispaniola that represents the allopatric barrier between the new subspecies and 
<taxonomicName id="4C0A4D3BFFFCFF95FA69FA28A464EF9E" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="207" phylum="Chordata" rank="subSpecies" species="aedium" subSpecies="aedium">
<emphasis id="B97EEAAAFFFCFF95FA69FA28A464EF9E" italics="true" pageId="6" pageNumber="207">P. aedium aedium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB536B8FFFCFF95FF29FA61A648EC2E" blockId="6.[151,1436,1214,2031]" pageId="6" pageNumber="207">
<emphasis id="B97EEAAAFFFCFF95FF29FA61A441EFBA" bold="true" box="[199,359,1502,1527]" pageId="6" pageNumber="207">Local names.</emphasis>
This subspecies is referred to by local informants in southeastern 
<collectingCountry id="F31D7628FFFCFF95FB87FA61A187EFBA" box="[1129,1185,1502,1527]" name="Haiti" pageId="6" pageNumber="207">Haiti</collectingCountry>
and the southwestern 
<collectingCountry id="F31D7628FFFCFF95FF79F9BDA4AFEC56" box="[151,393,1538,1563]" name="Dominican Republic" pageId="6" pageNumber="207">Dominican Republic</collectingCountry>
by several names, including 
<emphasis id="B97EEAAAFFFCFF95FCEFF9BCA613EC57" box="[769,821,1539,1562]" italics="true" pageId="6" pageNumber="207">jutía</emphasis>
(commonest name used in 
<collectingCountry id="F31D7628FFFCFF95FB77F9BDA0B7EC56" box="[1177,1425,1538,1563]" name="Dominican Republic" pageId="6" pageNumber="207">Dominican Republic</collectingCountry>
), 
<emphasis id="B97EEAAAFFFCFF95FF79F999A438EC72" box="[151,286,1574,1599]" italics="true" pageId="6" pageNumber="207">kochondenn</emphasis>
(commonest name used in 
<collectingCountry id="F31D7628FFFCFF95FDBDF999A7A9EC72" box="[595,655,1574,1599]" name="Haiti" pageId="6" pageNumber="207">Haiti</collectingCountry>
), 
<emphasis id="B97EEAAAFFFCFF95FD4DF999A630EC72" box="[675,790,1574,1599]" italics="true" pageId="6" pageNumber="207">solenodon</emphasis>
, 
<emphasis id="B97EEAAAFFFCFF95FCCAF999A65FEC72" box="[804,889,1574,1599]" italics="true" pageId="6" pageNumber="207">kombee</emphasis>
, and 
<emphasis id="B97EEAAAFFFCFF95FC56F998A6C0EC73" box="[952,998,1575,1598]" italics="true" pageId="6" pageNumber="207">rata</emphasis>
, reflecting confusion with other native and introduced co-occurring land mammals (
<bibRefCitation id="EF9B4B49FFFCFF95FD60F9F4A644EC2E" author="Turvey" box="[654,866,1610,1635]" pageId="6" pageNumber="207" refString="Turvey, S. T., Fernandez-Secades, C., Nunez-Mino, J. M., Hart, T., Martinez, P., Brocca, J. L. &amp; Young, R. P. (2014) Is local ecological knowledge a useful conservation tool for small mammals in a Caribbean multicultural landscape? Biological Conservation, 169, 189 - 197. http: // dx. doi. org / 10.1016 / j. biocon. 2013.11.018" type="journal article" year="2014">
Turvey 
<emphasis id="B97EEAAAFFFCFF95FD08F9F3A630EC2E" box="[742,790,1610,1635]" italics="true" pageId="6" pageNumber="207">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3106533FFFCFF95FF29F9D1A632ECE7" pageId="6" pageNumber="207" type="materials_examined">
<paragraph id="8BB536B8FFFCFF95FF29F9D1A632ECE7" blockId="6.[151,1436,1214,2031]" pageId="6" pageNumber="207">
<emphasis id="B97EEAAAFFFCFF95FF29F9D1A443ECCA" bold="true" box="[199,357,1646,1671]" pageId="6" pageNumber="207">
<typeStatus id="54B1881AFFFCFF95FF29F9D1A424ECCA" box="[199,258,1646,1671]" pageId="6" pageNumber="207">Type</typeStatus>
locality.
</emphasis>
Vicinity of Mencia village, Pedernales Province, 
<collectingCountry id="F31D7628FFFCFF95FC61F9D1A151ECCA" box="[911,1143,1646,1671]" name="Dominican Republic" pageId="6" pageNumber="207">Dominican Republic</collectingCountry>
(18°10'10N, 71°44'25W), in area of primary broadleaf forest at elevation of 
<specimenCount id="9D0CFD31FFFCFF95FD28F92DA632ECE7" box="[710,788,1682,1707]" pageId="6" pageNumber="207" type="generic">
<quantity id="4CF29B5DFFFCFF95FD28F92DA636ECE7" box="[710,784,1682,1707]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="6" pageNumber="207" unit="m" value="460.0">460 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C3106533FFFCFF95FF29F909A07BED12" pageId="6" pageNumber="207" type="distribution">
<paragraph id="8BB536B8FFFCFF95FF29F909A07BED12" blockId="6.[151,1436,1214,2031]" pageId="6" pageNumber="207">
<emphasis id="B97EEAAAFFFCFF95FF29F909A446EC82" bold="true" box="[199,352,1718,1743]" pageId="6" pageNumber="207">Distribution.</emphasis>
The southern palaeo-island of Hispaniola south of the Neiba Valley and east of the Jacmel– Fauché depression; found in both southeastern 
<collectingCountry id="F31D7628FFFCFF95FD0DF965A63DECBE" box="[739,795,1754,1779]" name="Haiti" pageId="6" pageNumber="207">Haiti</collectingCountry>
(departments Ouest and Sud-Est) and southwestern 
<collectingCountry id="F31D7628FFFCFF95FF79F941A4A3ED5A" box="[151,389,1790,1815]" name="Dominican Republic" pageId="6" pageNumber="207">Dominican Republic</collectingCountry>
(Barahona, Independencia and Pedernales provinces). Hutias are known to occur in the Massif de la Selle (
<collectingCountry id="F31D7628FFFCFF95FE6AF89DA4E7ED76" box="[388,449,1826,1851]" name="Haiti" pageId="6" pageNumber="207">Haiti</collectingCountry>
), and the Sierra de Bahoruco mountain range and Barahona 
<taxonomicName id="4C0A4D3BFFFCFF95FB73F89DA02CED76" box="[1181,1290,1826,1851]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="212" phylum="Mollusca" rank="species" species="peninsula">Peninsula</taxonomicName>
(
<collectingCountry id="F31D7628FFFCFF95FAF0F89DA5D8ED12" name="Dominican Republic" pageId="6" pageNumber="207">Dominican Republic</collectingCountry>
) (
<bibRefCitation id="EF9B4B49FFFCFF95FEFAF8F9A493ED12" author="Sullivan" box="[276,437,1862,1887]" pageId="6" pageNumber="207" refString="Sullivan, C. P. (1983) Status and distribution of Plagiodontia aedium in the Dominican Republic. Unpublished MSc thesis, University of Florida." type="book" year="1983">Sullivan 1983</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFFCFF95FE2FF8F9A7A9ED12" author="Woods" box="[449,655,1862,1887]" pageId="6" pageNumber="207" refString="Woods, C. A., Ottenwalder, J. A. &amp; Oliver, W. L. R. (1985) Lost mammals of the Greater Antilles: the summarised findings of a ten weeks field survey in the Dominican Republic, Haiti and Puerto Rico. Dodo, 22, 23 - 42." type="journal article" year="1985">
Woods 
<emphasis id="B97EEAAAFFFCFF95FDFBF8F8A763ED12" box="[533,581,1862,1887]" italics="true" pageId="6" pageNumber="207">et al</emphasis>
. 1985
</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFFCFF95FD75F8F9A60CED12" author="Woods" box="[667,810,1862,1887]" pageId="6" pageNumber="207" refString="Woods, C. A. (1986) Mammals of the national parks of Haiti. Unpublished report, prepared for USAID / Haiti under contract number 521 - 0169 - C- 00 - 3083 - 00, pp. 1 - 80." type="book chapter" year="1986">Woods 1986</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFFCFF95FCDBF8F9A155ED12" author="Woods" box="[821,1139,1862,1887]" pageId="6" pageNumber="207" refString="Woods, C. A. &amp; Ottenwalder, J. A. (1992) The natural history of southern Haiti. Florida Museum of Natural History, Gainesville, FL, 221 pp." type="book" year="1992">Woods &amp; Ottenwalder 1992</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFFCFF95FB91F8F8A077ED12" author="Turvey" box="[1151,1361,1862,1887]" pageId="6" pageNumber="207" refString="Turvey, S. T., Fernandez-Secades, C., Nunez-Mino, J. M., Hart, T., Martinez, P., Brocca, J. L. &amp; Young, R. P. (2014) Is local ecological knowledge a useful conservation tool for small mammals in a Caribbean multicultural landscape? Biological Conservation, 169, 189 - 197. http: // dx. doi. org / 10.1016 / j. biocon. 2013.11.018" type="journal article" year="2014">
Turvey 
<emphasis id="B97EEAAAFFFCFF95FB3BF8F8A020ED12" box="[1237,1286,1862,1887]" italics="true" pageId="6" pageNumber="207">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3106533FFFCFF94FF29F8D5A15FEAB5" lastPageId="7" lastPageNumber="208" pageId="6" pageNumber="207" type="diagnosis">
<paragraph id="8BB536B8FFFCFF94FF29F8D5A15FEAB5" blockId="6.[151,1436,1214,2031]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="208" pageId="6" pageNumber="207">
<emphasis id="B97EEAAAFFFCFF95FF29F8D5A467EDCE" bold="true" box="[199,321,1898,1923]" pageId="6" pageNumber="207">Diagnosis.</emphasis>
Subspecies of Hispaniolan hutia which is significantly larger than the closely related 
<taxonomicName id="4C0A4D3BFFFCFF95FAE4F8D5A46BEDEA" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="6" pageNumber="207" phylum="Chordata" rank="subSpecies" species="aedium" subSpecies="aedium">
<emphasis id="B97EEAAAFFFCFF95FAE4F8D5A46BEDEA" italics="true" pageId="6" pageNumber="207">Plagiodontia aedium aedium</emphasis>
</taxonomicName>
in the following craniodental characters: maximum interorbital width, minimum anterior interorbital width, length and width of incisive foramina, external width across bony palate at PM4, crown width of PM4, M1 and M2, crown length of M3, crown width of pm4 and m1, coronoid–condyle length, and coronoid– angular length. Significantly larger than 
<taxonomicName id="4C0A4D3BFFFDFF94FDB3FF26A617EAFD" box="[605,817,151,176]" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="208" phylum="Chordata" rank="subSpecies" species="aedium" subSpecies="hylaeum">
<emphasis id="B97EEAAAFFFDFF94FDB3FF26A74DEAFD" box="[605,619,153,176]" italics="true" pageId="7" pageNumber="208">P</emphasis>
. 
<emphasis id="B97EEAAAFFFDFF94FD97FF28A617EAFD" box="[633,817,151,176]" italics="true" pageId="7" pageNumber="208">aedium hylaeum</emphasis>
</taxonomicName>
in width of incisive foramina and crown length of M2, and somewhat larger than this subspecies in crown length of M3, height of toothrow at pm4, and coronoid–condyle length. Postcranial and soft-tissue differences between all three subspecies are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3106533FFFDFF9AFF29FEBBA63AE959" lastPageId="9" lastPageNumber="210" pageId="7" pageNumber="208" type="description">
<paragraph id="8BB536B8FFFDFF94FF29FEBBA679E98A" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">
<emphasis id="B97EEAAAFFFDFF94FF29FEBBA47FEB50" bold="true" box="[199,345,260,285]" pageId="7" pageNumber="208">Description.</emphasis>
Craniodental anatomy of 
<taxonomicName id="4C0A4D3BFFFDFF94FD7EFEBBA6A6EB50" box="[656,896,260,285]" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="aedium">
<emphasis id="B97EEAAAFFFDFF94FD7EFEBBA6A6EB50" box="[656,896,260,285]" italics="true" pageId="7" pageNumber="208">Plagiodontia aedium</emphasis>
</taxonomicName>
, including material from Pedernales Province referable to the new subspecies, described in detail by 
<bibRefCitation id="EF9B4B49FFFDFF94FCCAFE98A17DEB0D" author="Woods" box="[804,1115,295,320]" pageId="7" pageNumber="208" refString="Woods, C. A. &amp; Howland, E. B. (1979) Adaptive radiation of capromyid rodents: anatomy of the masticatory apparatus. Journal of Mammalogy, 60, 95 - 116. http: // dx. doi. org / 10.2307 / 1379762" type="journal article" year="1979">
Woods &amp; 
<collectingCountry id="F31D7628FFFDFF94FC71FE98A120EB0D" box="[927,1030,295,320]" name="United States Pacific Island Wildlife Refuges" pageId="7" pageNumber="208">Howland</collectingCountry>
(1979)
</bibRefCitation>
and Woods (2001). Nasals slightly anteriorly inflated. No supraorbital ridge; slight postorbital process. Zygomatic arch broad, with greatest width opposite M3; well-defined lateral jugal fossa present, restricted to level of molar toothrow. Superior zygomatic root of maxillary broad, inferior zygomatic root narrow, with well-developed masseteric tuberosity on ventral surface; posterior zygomatic root formed by squamosal. Incisive foramen distinct, relatively short and broad; septum formed mainly by premaxillary. Alisphenoid canal forms crescent-shaped fissure in rear wall of orbit, with optic foramen small and deep within this fissure; no sphenopalatine foramen on medial canal wall. Posterior margin of palate with V-shaped mesopterygoid fossa extending opposite M3, and with long, delicate lateral pterygoid processes. Large pterygoid plate extends medially posterior to M3, contacting base of pterygoid process; masticatory foramen in middle of pterygoid plate; lateral pterygoid ridge absent. Pterygoid fossa restricted by pterygoid plate; medial to M3. Paroccipital process long and broad; closely associated with short, spike-like lateral process that is independent from bullae. Incisors not procumbent; upper incisors strongly curved; alveolar capsule of lower incisors terminates below m1. Molariform cheek teeth hypsodont and rootless; upper and lower toothrows both parallel. Cheek teeth with enamel ridges forming deeply overlapping cement-filled oblique reentrant folds oriented at 45° angle to body axis; upper cheek teeth with one labial re-entrant fold and one lingual reentrant fold; lower cheek teeth with one labial re-entrant fold and two lingual re-entrant folds. Anterior margin of upper cheek teeth sigmoid; lower cheek teeth not sigmoid. Mandible with well-developed coronoid process, and angular process with masseteric crest and broad, flat-bottomed pterygoid shelf; mental foramen absent; retromolar fossa present as large pit with mandibular foramen in its centre.
</paragraph>
<paragraph id="8BB536B8FFFDFF94FF29FC6AA4BCEE15" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">
New subspecies distinguished phenotypically from other subspecies of 
<taxonomicName id="4C0A4D3BFFFDFF94FC03FC6AA17BE9A0" box="[1005,1117,980,1005]" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="aedium">
<emphasis id="B97EEAAAFFFDFF94FC03FC6AA6DDE9A1" box="[1005,1019,981,1004]" italics="true" pageId="7" pageNumber="208">P</emphasis>
. 
<emphasis id="B97EEAAAFFFDFF94FBE4FC6BA17BE9A0" box="[1034,1117,980,1005]" italics="true" pageId="7" pageNumber="208">aedium</emphasis>
</taxonomicName>
in this study on the basis of morphometric analysis using quantitative character measurements; no qualitative differences in craniodental character states are recognized. Craniodental measurements are as follows, taken from both individuals in 
<typeStatus id="54B1881AFFFDFF94FA82FBA2A0BAEE78" box="[1388,1436,1053,1077]" pageId="7" pageNumber="208">type</typeStatus>
series unless indicated:
</paragraph>
<paragraph id="8BB536B8FFFDFF94FF29FBDBA079ECAD" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="208">
Skull: greatest occipitonasal length, 
<quantity id="4CF29B5DFFFDFF94FD8EFBDBA633EE31" box="[608,789,1124,1149]" metricMagnitude="-2" metricUnit="m" metricValue="7.587999999999999" metricValueMax="7.76" metricValueMin="7.4159999999999995" pageId="7" pageNumber="208" unit="mm" value="75.88" valueMax="77.6" valueMin="74.16">74.16–77.6 mm</quantity>
; height of skull above palate, 
<quantity id="4CF29B5DFFFDFF94FB86FBDBA00DEE31" box="[1128,1323,1124,1149]" metricMagnitude="-2" metricUnit="m" metricValue="2.0409999999999995" metricValueMax="2.1239999999999997" metricValueMin="1.9579999999999997" pageId="7" pageNumber="208" unit="mm" value="20.409999999999997" valueMax="21.24" valueMin="19.58">19.58–21.24 mm</quantity>
; height of braincase, 
<quantity id="4CF29B5DFFFDFF94FEE0FB37A4F6EEED" box="[270,464,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="1.766" metricValueMax="1.812" metricValueMin="1.72" pageId="7" pageNumber="208" unit="mm" value="17.66" valueMax="18.12" valueMin="17.2">17.20–18.12 mm</quantity>
; width across braincase, 
<quantity id="4CF29B5DFFFDFF94FD09FB38A68FEEED" box="[743,937,1159,1184]" metricMagnitude="-2" metricUnit="m" metricValue="2.251" metricValueMax="2.316" metricValueMin="2.186" pageId="7" pageNumber="208" unit="mm" value="22.509999999999998" valueMax="23.16" valueMin="21.86">21.86–23.16 mm</quantity>
; width of occipital region, 
<quantity id="4CF29B5DFFFDFF94FB38FB38A0BEEEED" box="[1238,1432,1159,1184]" metricMagnitude="-2" metricUnit="m" metricValue="2.799" metricValueMax="2.822" metricValueMin="2.776" pageId="7" pageNumber="208" unit="mm" value="27.99" valueMax="28.22" valueMin="27.76">27.76–28.22 mm</quantity>
; internal width across occipital condyle, 
<quantity id="4CF29B5DFFFDFF94FD8FFB13A602EE89" box="[609,804,1196,1221]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" metricValueMax="1.15" metricValueMin="1.03" pageId="7" pageNumber="208" unit="mm" value="10.9" valueMax="11.5" valueMin="10.3">10.30–11.50 mm</quantity>
; width across paroccipital processes, 
<quantity id="4CF29B5DFFFDFF94FB3AFB13A0BEEE89" box="[1236,1432,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.735" metricValueMax="2.738" metricValueMin="2.732" pageId="7" pageNumber="208" unit="mm" value="27.35" valueMax="27.38" valueMin="27.32">27.32–27.38 mm</quantity>
; height of occipital region excluding paroccipital processes, 
<quantity id="4CF29B5DFFFDFF94FCD2FB70A6D9EEA5" box="[828,1023,1231,1256]" metricMagnitude="-2" metricUnit="m" metricValue="1.7579999999999998" metricValueMax="1.7600000000000002" metricValueMin="1.7559999999999998" pageId="7" pageNumber="208" unit="mm" value="17.58" valueMax="17.6" valueMin="17.56">17.56–17.60 mm</quantity>
; height of occipital region including paroccipital processes, 
<quantity id="4CF29B5DFFFDFF94FE70FB4BA744EF41" box="[414,610,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="2.6625" metricValueMax="2.7399999999999998" metricValueMin="2.585" pageId="7" pageNumber="208" unit="mm" value="26.625" valueMax="27.4" valueMin="25.85">25.85–27.40 mm</quantity>
; width across auditory bullae, 
<quantity id="4CF29B5DFFFDFF94FC2EFB4BA1A2EF41" box="[960,1156,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="2.8895" metricValueMax="2.905" metricValueMin="2.8739999999999997" pageId="7" pageNumber="208" unit="mm" value="28.895" valueMax="29.05" valueMin="28.74">28.74–29.05 mm</quantity>
; width across zygomatic arches, 41.00– 
<quantity id="4CF29B5DFFFDFF94FED1FAA8A49FEF7D" box="[319,441,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="4.164" pageId="7" pageNumber="208" unit="mm" value="41.64">41.64 mm</quantity>
; minimum breadth of zygomatic plate, 
<quantity id="4CF29B5DFFFDFF94FC7CFAA7A118EF7D" box="[914,1086,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="4.63" metricValueMax="4.88" metricValueMin="4.38" pageId="7" pageNumber="208" unit="mm" value="4.63" valueMax="4.88" valueMin="4.38">4.38–4.88 mm</quantity>
; maximum interorbital width, 24.18–25.00 mm; minimum anterior interorbital width, 
<quantity id="4CF29B5DFFFDFF94FCF6FA83A6FBEF19" box="[792,989,1340,1365]" metricMagnitude="-2" metricUnit="m" metricValue="2.007" metricValueMax="2.05" metricValueMin="1.964" pageId="7" pageNumber="208" unit="mm" value="20.07" valueMax="20.5" valueMin="19.64">19.64–20.50 mm</quantity>
; minimum posterior interorbital width, 
<quantity id="4CF29B5DFFFDFF94FF79FADFA47BEF35" box="[151,349,1376,1400]" metricMagnitude="-2" metricUnit="m" metricValue="1.925" metricValueMax="1.978" metricValueMin="1.8719999999999999" pageId="7" pageNumber="208" unit="mm" value="19.25" valueMax="19.78" valueMin="18.72">18.72–19.78 mm</quantity>
; maximum width of frontals, 
<quantity id="4CF29B5DFFFDFF94FD2EFAE0A6A0EF35" box="[704,902,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="2.4539999999999997" metricValueMax="2.55" metricValueMin="2.3579999999999997" pageId="7" pageNumber="208" unit="mm" value="24.54" valueMax="25.5" valueMin="23.58">23.58–25.50 mm</quantity>
; length of nasals, 
<quantity id="4CF29B5DFFFDFF94FB8FFAE0A1FEEF35" box="[1121,1240,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="7" pageNumber="208" unit="mm" value="25.1">25.10 mm</quantity>
(LDUCZ-Z2807 only); width of nasals, 
<quantity id="4CF29B5DFFFDFF94FE70FA3BA760EFD1" box="[414,582,1412,1437]" metricMagnitude="-3" metricUnit="m" metricValue="9.399999999999999" metricValueMax="9.6" metricValueMin="9.2" pageId="7" pageNumber="208" unit="mm" value="9.399999999999999" valueMax="9.6" valueMin="9.2">9.20–9.60 mm</quantity>
; width of single incisor, 
<quantity id="4CF29B5DFFFDFF94FC8FFA3BA12FEFD1" box="[865,1033,1412,1437]" metricMagnitude="-3" metricUnit="m" metricValue="2.915" metricValueMax="2.95" metricValueMin="2.88" pageId="7" pageNumber="208" unit="mm" value="2.915" valueMax="2.95" valueMin="2.88">2.88–2.95 mm</quantity>
; length of diastema (from posterior incisor margin), 
<quantity id="4CF29B5DFFFDFF94FEBAFA18A73EEF8D" box="[340,536,1447,1472]" metricMagnitude="-2" metricUnit="m" metricValue="1.823" metricValueMax="1.936" metricValueMin="1.71" pageId="7" pageNumber="208" unit="mm" value="18.23" valueMax="19.36" valueMin="17.1">17.10–19.36 mm</quantity>
; length of incisive foramina, 7.00– 
<quantity id="4CF29B5DFFFDFF94FC44FA17A134EF8D" box="[938,1042,1448,1472]" metricMagnitude="-3" metricUnit="m" metricValue="8.34" pageId="7" pageNumber="208" unit="mm" value="8.34">8.34 mm</quantity>
; width of incisive foramina, 
<quantity id="4CF29B5DFFFDFF94FAB3FA17A427EFA9" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="7" pageNumber="208" unit="mm" value="3.05" valueMax="3.3" valueMin="2.8">2.80– 3.30 mm</quantity>
; internal width across bony palate at PM4, 
<quantity id="4CF29B5DFFFDFF94FD13FA73A683EFA9" box="[765,933,1484,1509]" metricMagnitude="-3" metricUnit="m" metricValue="2.57" metricValueMax="2.6" metricValueMin="2.54" pageId="7" pageNumber="208" unit="mm" value="2.57" valueMax="2.6" valueMin="2.54">2.54–2.60 mm</quantity>
; external width across bony palate at PM4, 
<quantity id="4CF29B5DFFFDFF94FF79FA4FA42CEC45" box="[151,266,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.474" pageId="7" pageNumber="208" unit="mm" value="14.74">14.74 mm</quantity>
(both specimens); internal width across bony palate at M3, 
<quantity id="4CF29B5DFFFDFF94FC59FA50A178EC45" box="[951,1118,1519,1544]" metricMagnitude="-3" metricUnit="m" metricValue="6.76" metricValueMax="7.04" metricValueMin="6.48" pageId="7" pageNumber="208" unit="mm" value="6.76" valueMax="7.04" valueMin="6.48">6.48–7.04 mm</quantity>
; external width across bony palate at M3, 
<quantity id="4CF29B5DFFFDFF94FEDCF9ABA4D2EC61" box="[306,500,1556,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.5755" metricValueMax="1.596" metricValueMin="1.555" pageId="7" pageNumber="208" unit="mm" value="15.755" valueMax="15.96" valueMin="15.55">15.55–15.96 mm</quantity>
; length from 
<taxonomicName id="4C0A4D3BFFFDFF94FD65F9ABA7FBEC60" box="[651,733,1556,1581]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="208" phylum="Tracheophyta" rank="species" species="interior">interior</taxonomicName>
alveolus of PM4 to back of palate, 
<quantity id="4CF29B5DFFFDFF94FB81F9ABA014EC61" box="[1135,1330,1556,1580]" metricMagnitude="-2" metricUnit="m" metricValue="2.071" metricValueMax="2.1399999999999997" metricValueMin="2.002" pageId="7" pageNumber="208" unit="mm" value="20.71" valueMax="21.4" valueMin="20.02">20.02–21.40 mm</quantity>
; width of mesopterygoid fossa, 
<quantity id="4CF29B5DFFFDFF94FE60F988A710EC1D" box="[398,566,1591,1616]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.56" metricValueMin="4.24" pageId="7" pageNumber="208" unit="mm" value="4.4" valueMax="4.56" valueMin="4.24">4.24–4.56 mm</quantity>
; postpalatal length (to occipital foramen), 
<quantity id="4CF29B5DFFFDFF94FBF6F988A1FAEC1D" box="[1048,1244,1591,1616]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" metricValueMax="2.466" metricValueMin="2.314" pageId="7" pageNumber="208" unit="mm" value="23.9" valueMax="24.66" valueMin="23.14">23.14–24.66 mm</quantity>
; crown length of cheek tooth row, 
<quantity id="4CF29B5DFFFDFF94FEB6F9E3A73CEC39" box="[344,538,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.0465" metricValueMax="2.077" metricValueMin="2.016" pageId="7" pageNumber="208" unit="mm" value="20.465" valueMax="20.77" valueMin="20.16">20.16–20.77 mm</quantity>
; alveolar length of cheek tooth row, 
<quantity id="4CF29B5DFFFDFF94FC5AF9E3A150EC39" box="[948,1142,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="2.2435" metricValueMax="2.245" metricValueMin="2.242" pageId="7" pageNumber="208" unit="mm" value="22.435" valueMax="22.45" valueMin="22.42">22.42–22.45 mm</quantity>
; maximum alveolar width of cheek tooth row, 
<quantity id="4CF29B5DFFFDFF94FE91F9C0A701ECD5" box="[383,551,1663,1688]" metricMagnitude="-3" metricUnit="m" metricValue="6.26" metricValueMax="6.58" metricValueMin="5.94" pageId="7" pageNumber="208" unit="mm" value="6.26" valueMax="6.58" valueMin="5.94">5.94–6.58 mm</quantity>
; crown length of PM4, 
<quantity id="4CF29B5DFFFDFF94FCD4F9C0A6C4ECD5" box="[826,994,1663,1688]" metricMagnitude="-3" metricUnit="m" metricValue="6.22" metricValueMax="6.26" metricValueMin="6.18" pageId="7" pageNumber="208" unit="mm" value="6.22" valueMax="6.26" valueMin="6.18">6.18–6.26 mm</quantity>
; crown width of PM4, 
<quantity id="4CF29B5DFFFDFF94FB1EF9C0A0BEECD5" box="[1264,1432,1663,1688]" metricMagnitude="-3" metricUnit="m" metricValue="5.605" metricValueMax="5.85" metricValueMin="5.36" pageId="7" pageNumber="208" unit="mm" value="5.605" valueMax="5.85" valueMin="5.36">5.36–5.85 mm</quantity>
; crown length of M1, 
<quantity id="4CF29B5DFFFDFF94FE7FF91BA71CECF1" box="[401,570,1700,1725]" metricMagnitude="-3" metricUnit="m" metricValue="5.035" metricValueMax="5.14" metricValueMin="4.93" pageId="7" pageNumber="208" unit="mm" value="5.035" valueMax="5.14" valueMin="4.93">4.93–5.14 mm</quantity>
; crown width of M1, 
<quantity id="4CF29B5DFFFDFF94FCD3F91BA6C0ECF1" box="[829,998,1700,1725]" metricMagnitude="-3" metricUnit="m" metricValue="5.855" metricValueMax="5.96" metricValueMin="5.75" pageId="7" pageNumber="208" unit="mm" value="5.855" valueMax="5.96" valueMin="5.75">5.75–5.96 mm</quantity>
; crown length of M2, 
<quantity id="4CF29B5DFFFDFF94FB01F91BA0BEECF1" box="[1263,1432,1700,1724]" metricMagnitude="-3" metricUnit="m" metricValue="4.845" metricValueMax="4.99" metricValueMin="4.7" pageId="7" pageNumber="208" unit="mm" value="4.845" valueMax="4.99" valueMin="4.7">4.70–4.99 mm</quantity>
; crown width of M2, 
<quantity id="4CF29B5DFFFDFF94FE90F978A702ECAD" box="[382,548,1735,1760]" metricMagnitude="-3" metricUnit="m" metricValue="5.46" metricValueMax="5.62" metricValueMin="5.3" pageId="7" pageNumber="208" unit="mm" value="5.46" valueMax="5.62" valueMin="5.3">5.30–5.62 mm</quantity>
; crown length of M3, 
<quantity id="4CF29B5DFFFDFF94FCF2F978A6E4ECAD" box="[796,962,1735,1760]" metricMagnitude="-3" metricUnit="m" metricValue="4.93" metricValueMax="5.02" metricValueMin="4.84" pageId="7" pageNumber="208" unit="mm" value="4.93" valueMax="5.02" valueMin="4.84">4.84–5.02 mm</quantity>
; crown width of M3, 
<quantity id="4CF29B5DFFFDFF94FB5BF978A07DECAD" box="[1205,1371,1735,1760]" metricMagnitude="-3" metricUnit="m" metricValue="4.609999999999999" metricValueMax="4.64" metricValueMin="4.58" pageId="7" pageNumber="208" unit="mm" value="4.609999999999999" valueMax="4.64" valueMin="4.58">4.58–4.64 mm</quantity>
.
</paragraph>
<paragraph id="8BB536B8FFFDFF9BFF29F953A008EB72" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1436,151,824]" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">
Mandible: mandible length (incisor base to condyle), 
<quantity id="4CF29B5DFFFDFF94FCA8F953A128ED49" box="[838,1038,1772,1797]" metricMagnitude="-2" metricUnit="m" metricValue="5.081" metricValueMax="5.098" metricValueMin="5.064" pageId="7" pageNumber="208" unit="mm" value="50.81" valueMax="50.98" valueMin="50.64">50.64–50.98 mm</quantity>
; mandible length (incisor base to angular process), 
<quantity id="4CF29B5DFFFDFF94FEB3F8B0A739ED65" box="[349,543,1807,1832]" metricMagnitude="-2" metricUnit="m" metricValue="5.346" metricValueMax="5.534" metricValueMin="5.1579999999999995" pageId="7" pageNumber="208" unit="mm" value="53.46" valueMax="55.34" valueMin="51.58">51.58–55.34 mm</quantity>
; width of incisor, 
<quantity id="4CF29B5DFFFDFF94FD06F8AFA6A8ED65" box="[744,910,1808,1832]" metricMagnitude="-3" metricUnit="m" metricValue="2.74" metricValueMax="2.78" metricValueMin="2.7" pageId="7" pageNumber="208" unit="mm" value="2.74" valueMax="2.78" valueMin="2.7">2.70–2.78 mm</quantity>
; length of symphysis, 
<quantity id="4CF29B5DFFFDFF94FB67F8B0A06DED65" box="[1161,1355,1807,1832]" metricMagnitude="-2" metricUnit="m" metricValue="2.4045" metricValueMax="2.416" metricValueMin="2.393" pageId="7" pageNumber="208" unit="mm" value="24.045" valueMax="24.16" valueMin="23.93">23.93–24.16 mm</quantity>
; length of diastema, 
<quantity id="4CF29B5DFFFDFF94FEC7F88BA4CAED01" box="[297,492,1844,1869]" metricMagnitude="-2" metricUnit="m" metricValue="1.359" metricValueMax="1.426" metricValueMin="1.292" pageId="7" pageNumber="208" unit="mm" value="13.59" valueMax="14.26" valueMin="12.92">12.92–14.26 mm</quantity>
; length from incisor base to posterior bulb of incisor root, 
<quantity id="4CF29B5DFFFDFF94FB68F88BA06CED01" box="[1158,1354,1844,1869]" metricMagnitude="-2" metricUnit="m" metricValue="2.193" metricValueMax="2.234" metricValueMin="2.152" pageId="7" pageNumber="208" unit="mm" value="21.93" valueMax="22.34" valueMin="21.52">21.52–22.34 mm</quantity>
; crown length of cheek tooth row, 
<quantity id="4CF29B5DFFFDFF94FE36F8E8A7B8ED3D" box="[472,670,1879,1904]" metricMagnitude="-2" metricUnit="m" metricValue="1.9625" metricValueMax="2.016" metricValueMin="1.909" pageId="7" pageNumber="208" unit="mm" value="19.625" valueMax="20.16" valueMin="19.09">19.09–20.16 mm</quantity>
; alveolar length of cheek tooth row, 
<quantity id="4CF29B5DFFFDFF94FBBBF8E8A03DED3D" box="[1109,1307,1879,1904]" metricMagnitude="-2" metricUnit="m" metricValue="2.194" metricValueMax="2.196" metricValueMin="2.192" pageId="7" pageNumber="208" unit="mm" value="21.94" valueMax="21.96" valueMin="21.92">21.92–21.96 mm</quantity>
; maximum alveolar width of cheek tooth row, 
<quantity id="4CF29B5DFFFDFF94FDDCF8C3A7FDEDD9" box="[562,731,1916,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.99" metricValueMax="6.08" metricValueMin="5.9" pageId="7" pageNumber="208" unit="mm" value="5.99" valueMax="6.08" valueMin="5.9">5.90–6.08 mm</quantity>
; crown length of pm4, 
<quantity id="4CF29B5DFFFDFF94FC01F8C3A1BEEDD9" box="[1007,1176,1916,1941]" metricMagnitude="-3" metricUnit="m" metricValue="6.07" metricValueMax="6.6" metricValueMin="5.54" pageId="7" pageNumber="208" unit="mm" value="6.07" valueMax="6.6" valueMin="5.54">5.54–6.60 mm</quantity>
; crown width of pm4, 
<quantity id="4CF29B5DFFFDFF94FF79F81FA41AEDF5" box="[151,316,1952,1977]" metricMagnitude="-3" metricUnit="m" metricValue="4.91" metricValueMax="5.06" metricValueMin="4.76" pageId="7" pageNumber="208" unit="mm" value="4.91" valueMax="5.06" valueMin="4.76">4.76–5.06 mm</quantity>
; crown length of m1, 4.76–5.00 mm; crown width of m1, 
<quantity id="4CF29B5DFFFDFF94FC2AF81FA14CEDF5" box="[964,1130,1952,1977]" metricMagnitude="-3" metricUnit="m" metricValue="5.82" metricValueMax="5.88" metricValueMin="5.76" pageId="7" pageNumber="208" unit="mm" value="5.82" valueMax="5.88" valueMin="5.76">5.76–5.88 mm</quantity>
; crown length of m2, 
<quantity id="4CF29B5DFFFDFF94FAB3F81FA5D9ED91" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.68" metricValueMin="4.32" pageId="7" pageNumber="208" unit="mm" value="4.5" valueMax="4.68" valueMin="4.32">4.32– 4.68 mm</quantity>
; crown width of m2, 
<quantity id="4CF29B5DFFFDFF94FE1CF87BA7BFED91" box="[498,665,1988,2013]" metricMagnitude="-3" metricUnit="m" metricValue="5.54" metricValueMax="5.62" metricValueMin="5.46" pageId="7" pageNumber="208" unit="mm" value="5.54" valueMax="5.62" valueMin="5.46">5.46–5.62 mm</quantity>
; crown length of m3, 4.00– 
<quantity id="4CF29B5DFFFDFF94FC3FF87BA11FED91" box="[977,1081,1988,2013]" metricMagnitude="-3" metricUnit="m" metricValue="4.36" pageId="7" pageNumber="208" unit="mm" value="4.36">4.36 mm</quantity>
; crown width of m3, 
<quantity id="4CF29B5DFFFDFF9BFAC2F87BA5E0EAE2" lastPageId="8" lastPageNumber="209" metricMagnitude="-3" metricUnit="m" metricValue="4.82" metricValueMax="4.94" metricValueMin="4.7" pageId="7" pageNumber="208" unit="mm" value="4.82" valueMax="4.94" valueMin="4.7">4.70–4.94 mm</quantity>
; maximum width of entire mandibular body, 21.00– 
<quantity id="4CF29B5DFFF2FF9BFCE1FF27A6A2EAE2" box="[783,900,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.209" pageId="8" pageNumber="209" unit="mm" value="22.09">22.09 mm</quantity>
; maximum width of angular process, 
<quantity id="4CF29B5DFFF2FF9BFAC2FF28A5E0EA99" metricMagnitude="-3" metricUnit="m" metricValue="6.484999999999999" metricValueMax="6.95" metricValueMin="6.02" pageId="8" pageNumber="209" unit="mm" value="6.484999999999999" valueMax="6.95" valueMin="6.02">6.02–6.95 mm</quantity>
; internal height of mandible (base to alveolar lip), 
<quantity id="4CF29B5DFFF2FF9BFD10FF03A6E6EA99" box="[766,960,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.387" metricValueMax="1.424" metricValueMin="1.35" pageId="8" pageNumber="209" unit="mm" value="13.87" valueMax="14.24" valueMin="13.5">13.50–14.24 mm</quantity>
; internal height of mandible (base to tooth crown), 
<quantity id="4CF29B5DFFF2FF9BFF18FF60A49DEABA" box="[246,443,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.6164999999999998" metricValueMax="1.627" metricValueMin="1.6059999999999999" pageId="8" pageNumber="209" unit="mm" value="16.165" valueMax="16.27" valueMin="16.06">16.06–16.27 mm</quantity>
; height of toothrow at pm4 (internal aspect), 
<quantity id="4CF29B5DFFF2FF9BFC29FF5FA149EABA" box="[967,1135,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="3.82" metricValueMax="4.2" metricValueMin="3.44" pageId="8" pageNumber="209" unit="mm" value="3.82" valueMax="4.2" valueMin="3.44">3.44–4.20 mm</quantity>
; height of toothrow at m3 (internal aspect), 
<quantity id="4CF29B5DFFF2FF9BFEB5FEBBA727EB51" box="[347,513,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" metricValueMax="1.2" metricValueMin="1.12" pageId="8" pageNumber="209" unit="mm" value="1.16" valueMax="1.2" valueMin="1.12">1.12–1.20 mm</quantity>
; breadth across distal end of condyle, 
<quantity id="4CF29B5DFFF2FF9BFC5EFEBBA154EB51" box="[944,1138,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.0345" metricValueMax="1.037" metricValueMin="1.032" pageId="8" pageNumber="209" unit="mm" value="10.344999999999999" valueMax="10.37" valueMin="10.32">10.32–10.37 mm</quantity>
; coronoid–condyle length, 
<quantity id="4CF29B5DFFF2FF9BFF79FE98A47FEB72" box="[151,345,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.705" metricValueMax="1.8359999999999999" metricValueMin="1.574" pageId="8" pageNumber="209" unit="mm" value="17.05" valueMax="18.36" valueMin="15.74">15.74–18.36 mm</quantity>
; coronoid–angular length, 
<quantity id="4CF29B5DFFF2FF9BFD6BFE97A661EB72" box="[645,839,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.063" metricValueMax="3.114" metricValueMin="3.012" pageId="8" pageNumber="209" unit="mm" value="30.63" valueMax="31.14" valueMin="30.12">30.12–31.14 mm</quantity>
; condyle–angular length, 
<quantity id="4CF29B5DFFF2FF9BFB86FE98A00CEB72" box="[1128,1322,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.631" metricValueMax="2.67" metricValueMin="2.592" pageId="8" pageNumber="209" unit="mm" value="26.31" valueMax="26.7" valueMin="25.92">25.92–26.70 mm</quantity>
.
</paragraph>
<paragraph id="8BB536B8FFF2FF9BFF29FEF3A039E975" blockId="8.[151,1436,151,824]" pageId="8" pageNumber="209">
Detailed description of soft-tissue characters for the two specimens reported here is unfortunately not possible, as they were almost completely decomposed by the time they became available for study. General external morphology of hutias from southern 
<collectingCountry id="F31D7628FFF2FF9BFDBAFE2BA7A9EBE0" box="[596,655,404,429]" name="Haiti" pageId="8" pageNumber="209">Haiti</collectingCountry>
, representing individuals of both 
<taxonomicName id="4C0A4D3BFFF2FF9BFBC8FE2AA1DCEBE0" box="[1062,1274,404,429]" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="209" phylum="Chordata" rank="subSpecies" species="aedium" subSpecies="aedium">
<emphasis id="B97EEAAAFFF2FF9BFBC8FE2AA112EBE1" box="[1062,1076,405,428]" italics="true" pageId="8" pageNumber="209">P</emphasis>
. 
<emphasis id="B97EEAAAFFF2FF9BFBA6FE2BA1DCEBE0" box="[1096,1274,404,429]" italics="true" pageId="8" pageNumber="209">aedium aedium</emphasis>
</taxonomicName>
and the new subspecies, was described by 
<bibRefCitation id="EF9B4B49FFF2FF9BFE1BFE08A677EB9D" author="Woods" box="[501,849,439,464]" pageId="8" pageNumber="209" refString="Woods, C. A. &amp; Ottenwalder, J. A. (1992) The natural history of southern Haiti. Florida Museum of Natural History, Gainesville, FL, 221 pp." type="book" year="1992">Woods &amp; Ottenwalder (1992)</bibRefCitation>
, and live individuals encountered in the Sierra de Bahoruco during fieldwork by the authors in 
<date id="FFB41078FFF2FF9BFD5BFE63A614EBB9" box="[693,818,476,500]" pageId="8" pageNumber="209" value="2009" valueMax="2013">2009–2013</date>
match the broad description given by these authors (
<figureCitation id="13312A3DFFF2FF9BFF70FE40A426E855" box="[158,256,511,536]" captionStart="FIGURE 5" captionStartId="9.[151,250,1369,1391]" captionTargetBox="[154,1432,847,1343]" captionTargetId="figure@9.[151,1436,844,1348]" captionTargetPageId="9" captionText="FIGURE 5. Living individuals of Plagiodontia aedium bondi: a, typical individual, photographed on 5 November 2009 near Mencia (18 ° 10 ' 49 &quot; N, 71 ° 44 ' 40 &quot; W); b, albino individual, photographed on 29 November 2009 at Fondo Paradi, Parque Nacional Jaragua (17 ° 47 ' 36 &quot; N, 71 ° 27 ' 57 &quot; W)." httpUri="https://zenodo.org/record/240268/files/figure.png" pageId="8" pageNumber="209">Figure 5</figureCitation>
a). However, previously published information on supposed differences in external body proportions and pelage characteristics between northern and southwestern Hispaniolan hutia populations (
<bibRefCitation id="EF9B4B49FFF2FF9BFB4DFD9BA017E871" author="Miller" box="[1187,1329,548,573]" pageId="8" pageNumber="209" refString="Miller, G. S. Jr. (1928) The rodents of the genus Plagiodontia. Proceedings of the United States National Museum, 72, 1 - 8. http: // dx. doi. org / 10.5479 / si. 00963801.72 - 2712.1" type="journal article" year="1928">Miller 1928</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFF2FF9BFAAEFD9BA5F4E82D" author="Johnson" pageId="8" pageNumber="209" refString="Johnson, D. H. (1948) A rediscovered Haitian rodent, Plagiodontia aedium, with a synopsis of related species. Proceedings of the Biological Society of Washington, 61, 69 - 76." type="journal article" year="1948">Johnson 1948</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFF2FF9BFF33FDF8A4AAE82D" author="Anderson" box="[221,396,583,608]" pageId="8" pageNumber="209" refString="Anderson, S. (1965) Conspecificity of Plagiodontia aedium and P. hylaeum (Rodentia). Proceedings of the Biological Society of Washington, 78, 95 - 98." type="journal article" year="1965">Anderson 1965</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFF2FF9BFE79FDF8A78BE82D" author="Woods" box="[407,685,583,608]" pageId="8" pageNumber="209" refString="Woods, C. A. &amp; Howland, E. B. (1979) Adaptive radiation of capromyid rodents: anatomy of the masticatory apparatus. Journal of Mammalogy, 60, 95 - 116. http: // dx. doi. org / 10.2307 / 1379762" type="journal article" year="1979">
Woods &amp; 
<collectingCountry id="F31D7628FFF2FF9BFDE9FDF8A748E82D" box="[519,622,583,608]" name="United States Pacific Island Wildlife Refuges" pageId="8" pageNumber="209">Howland</collectingCountry>
1979
</bibRefCitation>
) is conflicting and of uncertain taxonomic value, and identification of systematic soft-tissue differences between different subspecies requires further study. Seventeen adult hutias were captured and released around Mencia in 
<date id="FFB41078FFF2FF9BFD49FD2FA602E8E5" box="[679,804,656,680]" pageId="8" pageNumber="209" value="2011" valueMax="2013">2011–2013</date>
during field research into ranging behavior and habitat use; adult males have a mean body mass of 
<quantity id="4CF29B5DFFF2FF9BFD7AFD0BA7DEE880" box="[660,760,692,717]" metricMagnitude="0" metricUnit="kg" metricValue="1.3229" pageId="8" pageNumber="209" unit="g" value="1322.9">1322.9 g</quantity>
(n=12, range= 
<quantity id="4CF29B5DFFF2FF9BFC4DFD0BA11FE880" box="[931,1081,692,717]" metricMagnitude="0" metricUnit="kg" metricValue="1.4175" metricValueMax="1.795" metricValueMin="1.04" pageId="8" pageNumber="209" unit="g" value="1417.5" valueMax="1795.0" valueMin="1040.0">
<date id="FFB41078FFF2FF9BFC4DFD0BA139E880" box="[931,1055,692,717]" pageId="8" pageNumber="209" value="1040" valueMax="1795">1040–1795</date>
g
</quantity>
, SD=280.4), and adult females have a mean body mass of 1355.0 g (n=5, range= 
<quantity id="4CF29B5DFFF2FF9BFD30FD68A653E8BD" box="[734,885,727,752]" metricMagnitude="0" metricUnit="kg" metricValue="1.355" metricValueMax="1.53" metricValueMin="1.18" pageId="8" pageNumber="209" unit="g" value="1355.0" valueMax="1530.0" valueMin="1180.0">1180–1530 g</quantity>
, SD=157.9), with total mean body mass for all individuals of 1335.0 g. These values are higher than the mean body mass of 
<quantity id="4CF29B5DFFF2FF9BFBA8FD43A1BEE958" box="[1094,1176,764,789]" metricMagnitude="0" metricUnit="kg" metricValue="1.018" pageId="8" pageNumber="209" unit="g" value="1018.0">1018 g</quantity>
for eight wild-caught individuals of 
<taxonomicName id="4C0A4D3BFFF2FF9BFED7FC9EA726E975" box="[313,512,799,824]" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="209" phylum="Chordata" rank="subSpecies" species="aedium" subSpecies="aedium">
<emphasis id="B97EEAAAFFF2FF9BFED7FC9EA461E975" box="[313,327,801,824]" italics="true" pageId="8" pageNumber="209">P</emphasis>
. 
<emphasis id="B97EEAAAFFF2FF9BFEBAFCA0A726E975" box="[340,512,799,824]" italics="true" pageId="8" pageNumber="209">aedium aedium</emphasis>
</taxonomicName>
from the Massif de la Hotte reported by 
<bibRefCitation id="EF9B4B49FFF2FF9BFC24FCA0A03CE975" author="Woods" box="[970,1306,799,824]" pageId="8" pageNumber="209" refString="Woods, C. A. &amp; Ottenwalder, J. A. (1992) The natural history of southern Haiti. Florida Museum of Natural History, Gainesville, FL, 221 pp." type="book" year="1992">Woods &amp; Ottenwalder (1992)</bibRefCitation>
.
</paragraph>
<caption id="DF756630FFF2FF9BFF79F9B2A6DEEC0F" httpUri="https://zenodo.org/record/240267/files/figure.png" pageId="8" pageNumber="209" targetBox="[154,1433,879,1523]" targetPageId="8">
<paragraph id="8BB536B8FFF2FF9BFF79F9B2A6DEEC0F" blockId="8.[151,1435,1549,1602]" pageId="8" pageNumber="209">
<emphasis id="B97EEAAAFFF2FF9BFF79F9B2A432EC6F" bold="true" box="[151,276,1549,1571]" pageId="8" pageNumber="209">FIGURE 4.</emphasis>
Holotype skull and mandible of 
<taxonomicName id="4C0A4D3BFFF2FF9BFD87F9B2A6A7EC6E" box="[617,897,1549,1571]" class="Mammalia" family="Capromyidae" genus="Plagiodontia" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="209" phylum="Chordata" rank="subSpecies" species="aedium" subSpecies="bondi">
<emphasis id="B97EEAAAFFF2FF9BFD87F9B2A6A7EC6E" box="[617,897,1549,1571]" italics="true" pageId="8" pageNumber="209">Plagiodontia aedium bondi</emphasis>
</taxonomicName>
(LDUCZ-Z2807): a, dorsal view of skull; b, ventral view of skull; c, lateral view of skull; d, external view of mandible. Scale bar = 1 cm.
</paragraph>
</caption>
<paragraph id="8BB536B8FFF2FF9AFF29F9D6A4F9EA98" blockId="8.[151,1437,1641,2026]" lastBlockId="9.[151,1437,151,789]" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209">
<emphasis id="B97EEAAAFFF2FF9BFF29F9D6A708ECCF" bold="true" box="[199,558,1641,1666]" pageId="8" pageNumber="209">Biological and ecological data.</emphasis>
Hispaniolan hutias are considered to be predominantly arboreal when in goodquality forest (
<bibRefCitation id="EF9B4B49FFF2FF9BFED2F933A4E8ECE8" author="Woods" box="[316,462,1676,1701]" pageId="8" pageNumber="209" refString="Woods, C. A. (1981) Last endemic mammals in Hispaniola. Oryx, 16, 146 - 152. http: // dx. doi. org / 10.1017 / S 0030605300017105" type="journal article" year="1981">Woods 1981</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFF2FF9BFE35F933A75AECE8" author="Sullivan" box="[475,636,1676,1701]" pageId="8" pageNumber="209" refString="Sullivan, C. P. (1983) Status and distribution of Plagiodontia aedium in the Dominican Republic. Unpublished MSc thesis, University of Florida." type="book" year="1983">Sullivan 1983</bibRefCitation>
), but during the course of the 
<date id="FFB41078FFF2FF9BFC3AF932A177ECE8" box="[980,1105,1677,1701]" pageId="8" pageNumber="209" value="2011" valueMax="2013">2011–2013</date>
study some individuals were caught on the ground, and signs of feeding on saplings on the ground were often observed. Females caught on 
<date id="FFB41078FFF2FF9BFA60F90EA40BECA0" pageId="8" pageNumber="209" value="2013-01-06">6 January 2013</date>
and 
<date id="FFB41078FFF2FF9BFE8DF96AA70CECA0" box="[355,554,1749,1774]" pageId="8" pageNumber="209" value="2013-02-15">15 February 2013</date>
were both pregnant, and a female caught on 
<date id="FFB41078FFF2FF9BFBCFF96AA1DDECA0" box="[1057,1275,1749,1774]" pageId="8" pageNumber="209" value="2012-09-14">14 September 2012</date>
was lactating. Field observations and camera trap photos recorded young juveniles (
<quantity id="4CF29B5DFFF2FF9BFC55F946A111ED5F" box="[955,1079,1785,1810]" metricMagnitude="-1" metricUnit="kg" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="8" pageNumber="209" unit="g" value="275.0" valueMax="350.0" valueMin="200.0">200–350 g</quantity>
body mass) in separate family groups on 
<date id="FFB41078FFF2FF9BFEF8F8A2A4FAED78" box="[278,476,1821,1846]" pageId="8" pageNumber="209" value="2012-01-15">15 January 2012</date>
, 
<date id="FFB41078FFF2FF9BFE02F8A2A797ED78" box="[492,689,1821,1846]" pageId="8" pageNumber="209" value="2012-01-27">27 January 2012</date>
, 
<date id="FFB41078FFF2FF9BFD2FF8A2A685ED78" box="[705,931,1821,1846]" pageId="8" pageNumber="209" value="2012-11-22">22 November 2012</date>
and 
<date id="FFB41078FFF2FF9BFC0CF8A2A181ED78" box="[994,1191,1821,1846]" pageId="8" pageNumber="209" value="2013-01-11">11 January 2013</date>
, suggesting probable parturition during the autumn, with larger juveniles (
<quantity id="4CF29B5DFFF2FF9BFD13F8FEA65FED17" box="[765,889,1857,1882]" metricMagnitude="-1" metricUnit="kg" metricValue="4.6" metricValueMax="5.0" metricValueMin="4.2" pageId="8" pageNumber="209" unit="g" value="460.0" valueMax="500.0" valueMin="420.0">420–500 g</quantity>
body mass) recorded on 
<date id="FFB41078FFF2FF9BFB48F8FEA064ED14" box="[1190,1346,1857,1882]" pageId="8" pageNumber="209" value="2012-05-12">12 May 2012</date>
and 
<date id="FFB41078FFF2FF9BFA6EF8FEA474ED30" pageId="8" pageNumber="209" value="2012-09-13">13 September 2012</date>
. Feeding signs detected during fieldwork and local reports suggest that hutias in the southeastern 
<collectingCountry id="F31D7628FFF2FF9BFF79F836A4A4EDEF" box="[151,386,1929,1954]" name="Dominican Republic" pageId="8" pageNumber="209">Dominican Republic</collectingCountry>
feed upon a relatively wide variety of plants, including 
<taxonomicName id="4C0A4D3BFFF2FF9BFBFBF836A01EEDEC" box="[1045,1336,1929,1954]" class="Magnoliopsida" family="Apocynaceae" genus="Rauvolfia" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFBFBF836A1A4EDEF" box="[1045,1154,1929,1954]" italics="true" pageId="8" pageNumber="209">Rauvolfia</emphasis>
(Apocynaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9BFAA8F836A410ED8B" class="Magnoliopsida" family="Cleomaceae" genus="Cleome" kingdom="Plantae" order="Capparales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFAA8F836A0BDEDEF" box="[1350,1435,1929,1954]" italics="true" pageId="8" pageNumber="209">Cleome</emphasis>
(Cleomaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9BFEA7F812A717ED8B" box="[329,561,1965,1990]" class="Insecta" family="Clusiidae" genus="Clusia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFEA7F812A4B5ED8B" box="[329,403,1965,1990]" italics="true" pageId="8" pageNumber="209">Clusia</emphasis>
(Clusiaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9BFDA8F811A6A4ED8B" box="[582,898,1965,1990]" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" kingdom="Plantae" order="Solanales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFDA8F811A78EED88" box="[582,680,1966,1989]" italics="true" pageId="8" pageNumber="209">Ipomoea</emphasis>
(Convulvulaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9BFC78F811A18BED8B" box="[918,1197,1965,1990]" class="Magnoliopsida" family="Euphorbiaceae" genus="Tragia" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFC78F811A6F9ED88" box="[918,991,1966,1989]" italics="true" pageId="8" pageNumber="209">Tragia</emphasis>
(Euphorbiaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9BFB2FF812A029ED89" box="[1217,1295,1965,1988]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFB2FF812A029ED89" box="[1217,1295,1965,1988]" italics="true" pageId="8" pageNumber="209">Acacia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D3BFFF2FF9BFAB9F812A433EDA7" class="Magnoliopsida" family="Fabaceae" genus="Senna" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFAB9F812A0BAED89" box="[1367,1436,1965,1988]" italics="true" pageId="8" pageNumber="209">Senna</emphasis>
(Fabaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9BFEDCF86DA702EDA4" box="[306,548,2001,2025]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFEDCF86DA4A5EDA4" box="[306,387,2002,2025]" italics="true" pageId="8" pageNumber="209">Ocotea</emphasis>
(Lauraceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9BFDAFF86DA7B0EDA4" box="[577,662,2002,2025]" class="Magnoliopsida" family="Meliaceae" genus="Guarea" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFDAFF86DA7B0EDA4" box="[577,662,2002,2025]" italics="true" pageId="8" pageNumber="209">Guarea</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D3BFFF2FF9BFD1EF86EA6D5EDA7" box="[752,1011,2001,2026]" class="Magnoliopsida" family="Meliaceae" genus="Trichilia" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFD1EF86EA669EDA7" box="[752,847,2001,2026]" italics="true" pageId="8" pageNumber="209">Trichilia</emphasis>
(Meliaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9BFBFEF86DA1CDEDA4" box="[1040,1259,2001,2025]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="209" phylum="Mollusca" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFBFEF86DA168EDA4" box="[1040,1102,2002,2025]" italics="true" pageId="8" pageNumber="209">Ficus</emphasis>
(Moraceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF2FF9AFAE6F86EA47FEAFD" class="Magnoliopsida" family="Phytolaccaceae" genus="Trichostigma" kingdom="Plantae" lastPageId="9" lastPageNumber="210" order="Caryophyllales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF2FF9BFAE6F86EA0BAEDA7" box="[1288,1436,2001,2026]" italics="true" pageId="8" pageNumber="209">Trichostigma</emphasis>
(Phytolaccaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF3FF9AFE8BFF27A4E1EAE2" box="[357,455,152,175]" class="Actinopterygii" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B97EEAAAFFF3FF9AFE8BFF27A4E1EAE2" box="[357,455,152,175]" italics="true" pageId="9" pageNumber="210">Gouania</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D3BFFF3FF9AFE11FF28A67EEAFD" box="[511,856,151,176]" class="Magnoliopsida" family="Rhamnaceae" genus="Krugiodendron" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="210" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF3FF9AFE11FF28A78DEAFD" box="[511,683,151,176]" italics="true" pageId="9" pageNumber="210">Krugiodendron</emphasis>
(Rhamnaceae)
</taxonomicName>
, 
<taxonomicName id="4C0A4D3BFFF3FF9AFC8AFF28A129EAFD" box="[868,1039,151,176]" class="Magnoliopsida" family="Sapotaceae" genus="Chrysophyllum" kingdom="Plantae" order="Ericales" pageId="9" pageNumber="210" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF3FF9AFC8AFF28A129EAFD" box="[868,1039,151,176]" italics="true" pageId="9" pageNumber="210">Chrysophyllum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0A4D3BFFF3FF9AFBA9FF28A04EEAFD" box="[1095,1384,151,176]" class="Magnoliopsida" family="Sapotaceae" genus="Sideroxylon" kingdom="Plantae" order="Ericales" pageId="9" pageNumber="210" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF3FF9AFBA9FF28A1EAEAFD" box="[1095,1228,151,176]" italics="true" pageId="9" pageNumber="210">Sideroxylon</emphasis>
(Sapotaceae)
</taxonomicName>
, and 
<taxonomicName id="4C0A4D3BFFF3FF9AFF79FF02A4FAEA98" box="[151,476,188,213]" class="Magnoliopsida" family="Zygophyllaceae" genus="Guaiacum" kingdom="Plantae" order="Zygophyllales" pageId="9" pageNumber="210" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEAAAFFF3FF9AFF79FF02A42AEA99" box="[151,268,189,212]" italics="true" pageId="9" pageNumber="210">Guaiacum</emphasis>
(Zygophyllaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BB536B8FFF3FF9AFF29FF60A0BBE870" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="210">
The relatively limited existing literature on previous field studies of Hispaniolan hutias provides some further information on habitat preferences and ecological requirements of the new subspecies. 
<bibRefCitation id="EF9B4B49FFF3FF9AFB8FFEBBA029EB51" author="Sullivan" box="[1121,1295,260,285]" pageId="9" pageNumber="210" refString="Sullivan, C. P. (1983) Status and distribution of Plagiodontia aedium in the Dominican Republic. Unpublished MSc thesis, University of Florida." type="book" year="1983">Sullivan (1983)</bibRefCitation>
reported the presence of hutias referable to this subspecies in low-elevation (0–234 masl) dry subtropical forest around Laguna Oviedo on the Barahona 
<taxonomicName id="4C0A4D3BFFF3FF9AFE56FEF3A70EEB28" box="[440,552,332,357]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="212" phylum="Mollusca" rank="species" species="peninsula">Peninsula</taxonomicName>
, where they have also been observed by the current authors (
<figureCitation id="13312A3DFFF3FF9AFB04FEF3A06AEB28" box="[1258,1356,332,357]" captionStart="FIGURE 5" captionStartId="9.[151,250,1369,1391]" captionTargetBox="[154,1432,847,1343]" captionTargetId="figure@9.[151,1436,844,1348]" captionTargetPageId="9" captionText="FIGURE 5. Living individuals of Plagiodontia aedium bondi: a, typical individual, photographed on 5 November 2009 near Mencia (18 ° 10 ' 49 &quot; N, 71 ° 44 ' 40 &quot; W); b, albino individual, photographed on 29 November 2009 at Fondo Paradi, Parque Nacional Jaragua (17 ° 47 ' 36 &quot; N, 71 ° 27 ' 57 &quot; W)." httpUri="https://zenodo.org/record/240268/files/figure.png" pageId="9" pageNumber="210">Figure 5</figureCitation>
b), and also in humid subtropical forest in the Sierra de Bahoruco across an elevational gradient from mixed hardwood forest (500–900 masl) to deciduous vegetation in ravines within pine forest (1300–1500 masl), with all sites associated with rocky limestone substrate that provides suitable cavities and crevices for shelter. Hutias have also been recorded from the Massif de la Selle in southeastern 
<collectingCountry id="F31D7628FFF3FF9AFCC6FE63A646EBB8" box="[808,864,476,501]" name="Haiti" pageId="9" pageNumber="210">Haiti</collectingCountry>
in undisturbed broad-leaved forest (locally known as “rak bwa”) on the steep north slope of the main ridge (2160 masl) and along the ravine of the Riviere Blanche (1730 masl) (
<bibRefCitation id="EF9B4B49FFF3FF9AFEC5FD9BA4DFE870" author="Woods" box="[299,505,548,573]" pageId="9" pageNumber="210" refString="Woods, C. A., Ottenwalder, J. A. &amp; Oliver, W. L. R. (1985) Lost mammals of the Greater Antilles: the summarised findings of a ten weeks field survey in the Dominican Republic, Haiti and Puerto Rico. Dodo, 22, 23 - 42." type="journal article" year="1985">
Woods 
<emphasis id="B97EEAAAFFF3FF9AFE6EFD9AA497E870" box="[384,433,548,573]" italics="true" pageId="9" pageNumber="210">et al</emphasis>
. 1985
</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFF3FF9AFDEAFD9BA7B2E870" author="Woods" box="[516,660,548,573]" pageId="9" pageNumber="210" refString="Woods, C. A. (1986) Mammals of the national parks of Haiti. Unpublished report, prepared for USAID / Haiti under contract number 521 - 0169 - C- 00 - 3083 - 00, pp. 1 - 80." type="book chapter" year="1986">Woods 1986</bibRefCitation>
; 
<bibRefCitation id="EF9B4B49FFF3FF9AFD70FD9BA6FDE871" author="Woods" box="[670,987,548,573]" pageId="9" pageNumber="210" refString="Woods, C. A. &amp; Ottenwalder, J. A. (1992) The natural history of southern Haiti. Florida Museum of Natural History, Gainesville, FL, 221 pp." type="book" year="1992">Woods &amp; Ottenwalder 1992</bibRefCitation>
), again near areas of exposed limestone.
</paragraph>
<paragraph id="8BB536B8FFF3FF9AFF3FFDF8A63AE959" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="210">
Interestingly, albinism has been documented in a relatively large number of individuals in this subspecies. Two separate albino individuals were caught during fieldwork around Mencia in 
<date id="FFB41078FFF3FF9AFBA6FDD3A1DEE8C9" box="[1096,1272,620,645]" pageId="9" pageNumber="210" value="2011" valueMax="2012">2011 and 2012</date>
, and a further albino individual was observed in Parque Nacional Jaragua in 2009 (
<figureCitation id="13312A3DFFF3FF9AFC58FD30A13CE8E5" box="[950,1050,655,680]" captionStart="FIGURE 5" captionStartId="9.[151,250,1369,1391]" captionTargetBox="[154,1432,847,1343]" captionTargetId="figure@9.[151,1436,844,1348]" captionTargetPageId="9" captionText="FIGURE 5. Living individuals of Plagiodontia aedium bondi: a, typical individual, photographed on 5 November 2009 near Mencia (18 ° 10 ' 49 &quot; N, 71 ° 44 ' 40 &quot; W); b, albino individual, photographed on 29 November 2009 at Fondo Paradi, Parque Nacional Jaragua (17 ° 47 ' 36 &quot; N, 71 ° 27 ' 57 &quot; W)." httpUri="https://zenodo.org/record/240268/files/figure.png" pageId="9" pageNumber="210">Figure 5</figureCitation>
b). A further “partially albinistic” individual “with pink eyes and unpigmented hands, feet and tail” from the “mountains of southern 
<collectingCountry id="F31D7628FFF3FF9AFB1EFD0BA00CE880" box="[1264,1322,692,717]" name="Haiti" pageId="9" pageNumber="210">Haiti</collectingCountry>
” (specific location unknown), which could refer either to the Massif de la Hotte in southwestern 
<collectingCountry id="F31D7628FFF3FF9AFB6EFD68A19EE8BD" box="[1152,1208,727,752]" name="Haiti" pageId="9" pageNumber="210">Haiti</collectingCountry>
or the Massif de la Selle in southeastern 
<collectingCountry id="F31D7628FFF3FF9AFE68FD43A4E7E958" box="[390,449,764,789]" name="Haiti" pageId="9" pageNumber="210">Haiti</collectingCountry>
, was described by 
<bibRefCitation id="EF9B4B49FFF3FF9AFD7DFD42A63FE959" author="Tate" box="[659,793,764,788]" pageId="9" pageNumber="210" refString="Tate, G. H. H. (1948) Notes on the Hispaniolan hutia, Plagiodontia, and extinct related genera. Journal of Mammalogy, 29, 176 - 178. http: // dx. doi. org / 10.2307 / 1375245" type="journal article" year="1948">Tate (1948)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>