160 lines
18 KiB
XML
160 lines
18 KiB
XML
<document id="CA84B7446D14ED4DB54729EEDD60F367" ID-CLB-Dataset="4204" ID-DOI="10.11646/palaeoentomology.4.5.9" ID-GBIF-Dataset="d21ee62b-62da-4e85-8c9d-9bae52962a16" ID-ISSN="2624-2834" ID-Zenodo-Dep="5530327" ID-ZooBank="16401A1E-5D42-44BD-8DE8-E37599F34875" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632741517667" checkinUser="plazi" docAuthor="SZADZIEWSKI, RYSZARD, SONTAG, ELŻBIETA & PANKOWSKI, MADELINE V." docDate="2021" docId="03A387A3FFDBFF9532624E2DEF1D1EA7" docLanguage="en" docName="palaeoentomology.4.5.9.pdf" docOrigin="Palaeoentomology 4 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.5.9" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Forcipomyia rasnitsyni SZADZIEWSKI & SONTAG & PANKOWSKI 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="443" masterDocId="FF9AFFDBFFDBFF9731484A27EE6F1F74" masterDocTitle="A new biting midge of the genus Forcipomyia Meigen, 1818 from Miocene Ethiopian amber (Diptera: Ceratopogonidae)" masterLastPageNumber="444" masterPageNumber="441" pageNumber="441" updateTime="1699450254848" updateUser="ExternalLinkService">
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<mods:title id="F6EA8A105B0606BD80CB6D964A166FB8">A new biting midge of the genus Forcipomyia Meigen, 1818 from Miocene Ethiopian amber (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="45E8D0EE3E7AA22C6A6A2F138DAE0863">SZADZIEWSKI, RYSZARD</mods:namePart>
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<mods:namePart id="87AAB20266D4DD9739A0E9DF008490FF">SONTAG, ELŻBIETA</mods:namePart>
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<mods:namePart id="C7A453BD9F2774164DAD40BFFADF7436">PANKOWSKI, MADELINE V.</mods:namePart>
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<mods:title id="3E4582A87CC0B16BAC283D7AEBC31327">Palaeoentomology</mods:title>
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<mods:part id="20164D919A7B8C54F32D64E42238E50D">
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<mods:date id="1C56604A55D289F7D3B5592F28066A65">2021</mods:date>
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<subSubSection id="C310653EFFDBFF9732624E2DEAEA1B57" box="[810,1157,1034,1059]" pageId="0" pageNumber="441" type="nomenclature">
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<paragraph id="8BB536B5FFDBFF9732624E2DEAEA1B57" blockId="0.[810,1157,1034,1095]" box="[810,1157,1034,1059]" pageId="0" pageNumber="441">
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<heading id="D0FD81D9FFDBFF9732624E2DEAEA1B57" bold="true" box="[810,1157,1034,1059]" fontSize="10" level="2" pageId="0" pageNumber="441" reason="3">
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<emphasis id="B97EEAA7FFDBFF9732624E2DEAEA1B57" bold="true" box="[810,1157,1034,1059]" pageId="0" pageNumber="441">
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<taxonomicName id="4C0A4D36FFDBFF9732624E2DEA461B56" ID-CoL="9J2QS" authority="SZADZIEWSKI & SONTAG & PANKOWSKI, 2021" authorityName="SZADZIEWSKI & SONTAG & PANKOWSKI" authorityYear="2021" box="[810,1065,1034,1058]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="441" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
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<emphasis id="B97EEAA7FFDBFF9732624E2DEA461B56" bold="true" box="[810,1065,1034,1058]" italics="true" pageId="0" pageNumber="441">Forcipomyia rasnitsyni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24D57DCFFDBFF9735784E2CEAEA1B57" box="[1072,1157,1035,1059]" pageId="0" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C310653EFFDBFF9732624E09EDC31B33" box="[810,940,1070,1095]" pageId="0" pageNumber="441" type="description">
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<paragraph id="8BB536B5FFDBFF9732624E09EDC31B33" blockId="0.[810,1157,1034,1095]" box="[810,940,1070,1095]" pageId="0" pageNumber="441">
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(
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<figureCitation id="13312A30FFDBFF97327A4E09EDCB1B33" box="[818,932,1070,1095]" captionStart="FIGURE 1" captionStartId="1.[136,235,2038,2062]" captionTargetBox="[257,1330,189,2015]" captionTargetId="figure-19@1.[257,1330,189,2015]" captionTargetPageId="1" captionText="FIGURE 1. Forcipomyia rasnitsyni sp. nov., female holotype. A, Total habitus. B, Right flagellum. C, Proboscis with palpi." figureDoi="http://doi.org/10.5281/zenodo.5530333" httpUri="https://zenodo.org/record/5530333/files/figure.png" pageId="0" pageNumber="441">Fig. 1A–C</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C310653EFFDBFF9732624E50EA8F1A6B" pageId="0" pageNumber="441" type="materials_examined">
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<paragraph id="8BB536B5FFDBFF9732624E50EA8F1A6B" blockId="0.[810,1452,1142,2092]" pageId="0" pageNumber="441">
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<emphasis id="B97EEAA7FFDBFF9732624E50EDBE1BFA" bold="true" box="[810,977,1142,1167]" pageId="0" pageNumber="441">Type material.</emphasis>
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<materialsCitation id="3B623CE8FFDBFF9732904E50EA8F1A6B" ID-GBIF-Occurrence="3358416301" county="Ethiopian" location="Museum of Amber Inclusions" municipality="International Amber Association" pageId="0" pageNumber="441" specimenCount="1" specimenCount-worker="1" typeStatus="holotype">
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<typeStatus id="54B18817FFDBFF9732904E50EA541BFB" box="[984,1083,1143,1167]" pageId="0" pageNumber="441" type="holotype">Holotype</typeStatus>
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female, deposited in the
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<location id="8ED5606EFFDBFF9734064E50EA7B1BC7" LSID="urn:lsid:plazi:treatment:03A387A3FFDBFF9532624E2DEF1D1EA7:8ED5606EFFDBFF9734064E50EA7B1BC7" county="Ethiopian" municipality="International Amber Association" name="Museum of Amber Inclusions" pageId="0" pageNumber="441">Museum of Amber Inclusions</location>
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,
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<location id="8ED5606EFFDBFF97356E4EBCEB731BC7" LSID="urn:lsid:plazi:treatment:03A387A3FFDBFF9532624E2DEF1D1EA7:8ED5606EFFDBFF97356E4EBCEB731BC7" box="[1062,1308,1179,1203]" county="Ethiopian" municipality="International Amber Association" name="University of Gdansk" pageId="0" pageNumber="441">University of Gdańsk</location>
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, MAI 6697, infrared spectroscopy certificate from the
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<collectingMunicipality id="6BD1ACCFFFDBFF97346B4E98ED951B8F" pageId="0" pageNumber="441">International Amber Association</collectingMunicipality>
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no. 22251.
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<collectingCounty id="62D44E39FFDBFF9735CB4EC4EA841B8F" box="[1155,1259,1251,1275]" pageId="0" pageNumber="441">Ethiopian</collectingCounty>
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amber, Miocene. Syninclusion:
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<specimenCount id="9D0CFD3CFFDBFF97328B4F2FEA531A6B" box="[963,1084,1287,1311]" pageId="0" pageNumber="441" type="worker">one worker</specimenCount>
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of
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<taxonomicName id="4C0A4D36FFDBFF9735174F20EAB31A6B" box="[1119,1244,1287,1311]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="441" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
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.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C310653EFFDBFF9732174F0CEDD51AA7" pageId="0" pageNumber="441" type="etymology">
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<paragraph id="8BB536B5FFDBFF9732174F0CEDD51AA7" blockId="0.[810,1452,1142,2092]" pageId="0" pageNumber="441">
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<emphasis id="B97EEAA7FFDBFF9732174F0CED8F1A37" bold="true" box="[863,992,1323,1347]" pageId="0" pageNumber="441">Etymology.</emphasis>
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This new species is named in honour of professor Alexandr P. Rasnitsyn of the Paleontological Institute of the Russian Academy of Sciences in Moscow in recognition of his important contributions to the study of fossil insects.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C310653EFFDBFF9732174FF8EB7119FC" pageId="0" pageNumber="441" type="diagnosis">
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<paragraph id="8BB536B5FFDBFF9732174FF8EB7119FC" blockId="0.[810,1452,1142,2092]" pageId="0" pageNumber="441">
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<emphasis id="B97EEAA7FFDBFF9732174FF8EDBB1A83" bold="true" box="[863,980,1503,1527]" pageId="0" pageNumber="441">Diagnosis.</emphasis>
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The only fossil species of
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<taxonomicName id="4C0A4D36FFDBFF97346B4FC7EBC31A83" authorityName="Meigen" authorityYear="1818" box="[1315,1452,1504,1527]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="441" phylum="Arthropoda" rank="genus">
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<emphasis id="B97EEAA7FFDBFF97346B4FC7EBC31A83" box="[1315,1452,1504,1527]" italics="true" pageId="0" pageNumber="441">Forcipomyia</emphasis>
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</taxonomicName>
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with a high tarsal ratio (2.3–2.6), very short distal female flagellomeres (AR 0.8), short and separated palpal segments 4 and 5, claws with single apices, both first radial cells present, wing length
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<quantity id="4CF29B50FFDBFF9735414C57EA0319F3" box="[1033,1132,1648,1672]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="0" pageNumber="441" unit="mm" value="0.81">0.81 mm</quantity>
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. Male unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C310653EFFDBFF9532174CB4EF1D1EA7" lastPageId="2" lastPageNumber="443" pageId="0" pageNumber="441" type="description">
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<paragraph id="8BB536B5FFDBFF9532174CB4EFA51EC4" blockId="0.[810,1452,1142,2092]" lastBlockId="2.[136,771,159,467]" lastPageId="2" lastPageNumber="443" pageId="0" pageNumber="441">
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<emphasis id="B97EEAA7FFDBFF9732174CB4ED8519DF" bold="true" box="[863,1002,1683,1707]" pageId="0" pageNumber="441">Description.</emphasis>
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Female. The specimen is well preserved, complete (
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<figureCitation id="13312A30FFDBFF9732EC4C91EA6C19BA" box="[932,1027,1718,1742]" captionStart="FIGURE 1" captionStartId="1.[136,235,2038,2062]" captionTargetBox="[257,1330,189,2015]" captionTargetId="figure-19@1.[257,1330,189,2015]" captionTargetPageId="1" captionText="FIGURE 1. Forcipomyia rasnitsyni sp. nov., female holotype. A, Total habitus. B, Right flagellum. C, Proboscis with palpi." figureDoi="http://doi.org/10.5281/zenodo.5530333" httpUri="https://zenodo.org/record/5530333/files/figure.png" pageId="0" pageNumber="441">Fig. 1A</figureCitation>
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). Body hairy, total length (without antennae)
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<quantity id="4CF29B50FFDBFF9732D34CFEED9C1984" box="[923,1011,1753,1777]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="0" pageNumber="441" unit="mm" value="1.4">1.4 mm</quantity>
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. Flagellum with 13 flagellomeres, total length
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<quantity id="4CF29B50FFDBFF9732304CDBEDB11867" box="[888,990,1788,1812]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="0" pageNumber="441" unit="mm" value="0.56">0.56 mm</quantity>
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, combined length of flagellomeres 9–13 smaller than combined length of flagellomeres 1–8, AR 0.81 (
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<figureCitation id="13312A30FFDBFF9732204D65EDD3182E" box="[872,956,1858,1882]" captionStart="FIGURE 1" captionStartId="1.[136,235,2038,2062]" captionTargetBox="[257,1330,189,2015]" captionTargetId="figure-19@1.[257,1330,189,2015]" captionTargetPageId="1" captionText="FIGURE 1. Forcipomyia rasnitsyni sp. nov., female holotype. A, Total habitus. B, Right flagellum. C, Proboscis with palpi." figureDoi="http://doi.org/10.5281/zenodo.5530333" httpUri="https://zenodo.org/record/5530333/files/figure.png" pageId="0" pageNumber="441">Fig. 1B</figureCitation>
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). Microsetae between ommatidia not visible. Proboscis short. Palpus 5-segmented, fourth and fifth palpal segments separated, short and broad, of similar size, combined length 4+5 about
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<quantity id="4CF29B50FFDBFF97351A4D8CEAAE18B7" box="[1106,1217,1963,1987]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="441" unit="mm" value="0.05">0.050 mm</quantity>
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; third palpal segment slightly swollen on basal 2/3, length about
|
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<quantity id="4CF29B50FFDBFF9734614DE9EBCF1892" box="[1321,1440,1998,2022]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="0" pageNumber="441" unit="mm" value="0.06">0.060 mm</quantity>
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||
, sensory pit not visible (
|
||
<figureCitation id="13312A30FFDBFF9735604DD6EAF0177D" box="[1064,1183,2033,2057]" captionStart="FIGURE 1" captionStartId="1.[136,235,2038,2062]" captionTargetBox="[257,1330,189,2015]" captionTargetId="figure-19@1.[257,1330,189,2015]" captionTargetPageId="1" captionText="FIGURE 1. Forcipomyia rasnitsyni sp. nov., female holotype. A, Total habitus. B, Right flagellum. C, Proboscis with palpi." figureDoi="http://doi.org/10.5281/zenodo.5530333" httpUri="https://zenodo.org/record/5530333/files/figure.png" pageId="0" pageNumber="441">Fig. 1A, C</figureCitation>
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||
). Anterior anepisternum narrow, weakly B-shaped. Wing length measured from basal arculus
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||
<quantity id="4CF29B50FFD9FF9531A94AB8EF271FC3" box="[225,328,159,183]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="2" pageNumber="443" unit="mm" value="0.81">0.81 mm</quantity>
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, costal ratio CR 0.51; wing membrane densely covered by long slender macrotrichia; both first radial cells present, first one slit-like, second one distinct, rectangular. Legs slender, tibiae without scales, claws simple, without modifications, empodium large. Tarsal ratio TR: hind leg TR(3) 2.28, midleg TR(2) 2.64, foreleg TR(1) 2.61. Abdomen without modifications, cerci short, seminal capsules not visible (
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||
<figureCitation id="13312A30FFD9FF9530204BB0EFD21EC4" box="[360,445,407,432]" captionStart="FIGURE 1" captionStartId="1.[136,235,2038,2062]" captionTargetBox="[257,1330,189,2015]" captionTargetId="figure-19@1.[257,1330,189,2015]" captionTargetPageId="1" captionText="FIGURE 1. Forcipomyia rasnitsyni sp. nov., female holotype. A, Total habitus. B, Right flagellum. C, Proboscis with palpi." figureDoi="http://doi.org/10.5281/zenodo.5530333" httpUri="https://zenodo.org/record/5530333/files/figure.png" pageId="2" pageNumber="443">Fig. 1A</figureCitation>
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).
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||
</paragraph>
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<caption id="DF75663DFFDAFF9631C04DD1EB17177A" ID-DOI="http://doi.org/10.5281/zenodo.5530333" ID-Zenodo-Dep="5530333" box="[136,1400,2038,2062]" httpUri="https://zenodo.org/record/5530333/files/figure.png" pageId="1" pageNumber="442" startId="1.[136,235,2038,2062]" targetBox="[257,1330,189,2015]" targetPageId="1">
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<paragraph id="8BB536B5FFDAFF9631C04DD1EB17177A" blockId="1.[136,1400,2038,2062]" box="[136,1400,2038,2062]" pageId="1" pageNumber="442">
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<emphasis id="B97EEAA7FFDAFF9631C04DD1EF6B177A" bold="true" box="[136,260,2038,2062]" pageId="1" pageNumber="442">FIGURE 1.</emphasis>
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<taxonomicName id="4C0A4D36FFDAFF9630424DD0EF98177A" authority="SZADZIEWSKI & SONTAG & PANKOWSKI, 2021" authorityName="SZADZIEWSKI & SONTAG & PANKOWSKI" authorityYear="2021" box="[266,503,2039,2062]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
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<emphasis id="B97EEAA7FFDAFF9630424DD0EF98177A" box="[266,503,2039,2062]" italics="true" pageId="1" pageNumber="442">Forcipomyia rasnitsyni</emphasis>
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</taxonomicName>
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<emphasis id="B97EEAA7FFDAFF9630B54DD1EC22177A" bold="true" box="[509,589,2038,2062]" pageId="1" pageNumber="442">
|
||
<taxonomicNameLabel id="A24D57DCFFDAFF9630B54DD1EC22177A" box="[509,589,2038,2062]" pageId="1" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype.
|
||
<emphasis id="B97EEAA7FFDAFF9632434DD1ED72177A" bold="true" box="[779,797,2038,2062]" pageId="1" pageNumber="442">A</emphasis>
|
||
, Total habitus.
|
||
<emphasis id="B97EEAA7FFDAFF9632F14DD1EDA5177A" bold="true" box="[953,970,2038,2062]" pageId="1" pageNumber="442">B</emphasis>
|
||
, Right flagellum.
|
||
<emphasis id="B97EEAA7FFDAFF9635CB4DD1EAFA177A" bold="true" box="[1155,1173,2038,2062]" pageId="1" pageNumber="442">C</emphasis>
|
||
, Proboscis with palpi.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536B5FFD9FF9531F54B9CEF1D1EA7" blockId="2.[136,771,159,467]" box="[189,370,443,467]" pageId="2" pageNumber="443">Male. Unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |