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<document id="A5AF1E8036B4FA94E0EB9F51B1E6BE66" ID="10.11646/zootaxa.4267.1.1SLASH11021" ID-CLB-Dataset="33061" ID-DOI="10.5281/zenodo.575641" ID-GBIF-Dataset="810b55e4-b8ba-4368-bf9c-8517ee33e7d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="575641" ID-ZooBank="8920893E-E5F8-482A-A60D-7A248D2B0DCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494577389872" checkinUser="plazi" docAuthor="Skelley, Paul, Xu, Guang, Tang, William, Lindström, Anders J., Marler, Thomas, Khuraijam, Jibankumar Singh, Singh, Rita &amp; Rich, Stephen" docDate="2017" docId="03A38781FFCEFFA5FF70FB25FC06C299" docLanguage="en" docName="zootaxa.4267.1.1SLASH11021.pdf" docOrigin="Zootaxa 4267 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cycadophila (Cycadophila) papua Skelley, Xu &amp; Tang, new species" docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="FF9AFFF9FFE6FF8FFFE7FFEBFFD5C639" masterDocTitle="Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species" masterLastPageNumber="63" masterPageNumber="1" pageNumber="41" updateTime="1698439510239" updateUser="ExternalLinkService">
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<mods:title id="7E6B9B7C8703AAC14E12D9F725C5B9D6">Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species</mods:title>
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<mods:namePart id="8D82709F138EEB2D7C2B533B03BA1725">Xu, Guang</mods:namePart>
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<mods:namePart id="69F031C7B189C150D48E76E02367369B">Tang, William</mods:namePart>
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<mods:namePart id="A58579E8FFE4071F07005C173AB4ECC1">Lindström, Anders J.</mods:namePart>
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<mods:namePart id="42E7BE02A82E36234AB46A58ABB8C1E7">Marler, Thomas</mods:namePart>
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<mods:namePart id="1072A0C7806203D8B1326A88472D0298">Khuraijam, Jibankumar Singh</mods:namePart>
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<mods:namePart id="B4ABAB2719A3764047EAFE25C55843D6">Singh, Rita</mods:namePart>
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<mods:namePart id="DBB011D9D8054D7D6AC9F36F57E08C4E">Rich, Stephen</mods:namePart>
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<treatment id="03A38781FFCEFFA5FF70FB25FC06C299" ID-DOI="http://doi.org/10.5281/zenodo.5999219" ID-GBIF-Taxon="130809504" ID-Zenodo-Dep="5999219" LSID="urn:lsid:plazi:treatment:03A38781FFCEFFA5FF70FB25FC06C299" httpUri="http://treatment.plazi.org/id/03A38781FFCEFFA5FF70FB25FC06C299" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="C310651CFFCEFFA7FF70FB25FEF1C330" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8BB53697FFCEFFA7FF70FB25FC1FC2D1" blockId="40.[151,970,1230,1290]" box="[151,970,1230,1256]" pageId="40" pageNumber="41">
<heading id="D0FD81FBFFCEFFA7FF70FB25FC1FC2D1" bold="true" box="[151,970,1230,1256]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="B97EEA85FFCEFFA7FF70FB25FC1FC2D1" bold="true" box="[151,970,1230,1256]" pageId="40" pageNumber="41">
<taxonomicName id="4C0A4D14FFCEFFA7FF70FB25FCF8C2D1" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[151,813,1230,1256]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="papua" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFCEFFA7FF70FB25FEFBC2D1" bold="true" box="[151,302,1230,1256]" italics="true" pageId="40" pageNumber="41">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFCEFFA7FED8FB25FE03C2D1" bold="true" box="[319,470,1230,1256]" italics="true" pageId="40" pageNumber="41">Cycadophila</emphasis>
)
<emphasis id="B97EEA85FFCEFFA7FE0FFB25FDE6C2D1" bold="true" box="[488,563,1230,1256]" italics="true" pageId="40" pageNumber="41">papua</emphasis>
Skelley, Xu &amp; Tang
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,
<taxonomicNameLabel id="A24D57FEFFCEFFA7FCDDFB25FC1FC2D1" box="[826,970,1230,1256]" pageId="40" pageNumber="41" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BB53697FFCEFFA7FF70FB1AFEF1C330" blockId="40.[151,970,1230,1290]" box="[151,292,1265,1290]" pageId="40" pageNumber="41">
<figureCitation id="13312A12FFCEFFA7FF70FB1AFF3AC333" box="[151,239,1265,1290]" captionStart="FIGURE 13" captionStartId="41.[151,250,1569,1591]" captionTargetBox="[151,1436,409,1548]" captionTargetId="figure@41.[151,1436,409,1548]" captionTargetPageId="41" captionText="FIGURE 13. Photographs of Cycadophila (Cycadophila) papua, holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia." httpUri="https://zenodo.org/record/575654/files/figure.png" pageId="40" pageNumber="41">Figs. 13</figureCitation>
AE
</paragraph>
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<subSubSection id="C310651CFFCEFFA7FF70FAD2FB5CC048" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8BB53697FFCEFFA7FF70FAD2FCF8C3A3" blockId="40.[151,1436,1337,2010]" pageId="40" pageNumber="41">
<emphasis id="B97EEA85FFCEFFA7FF70FAD2FE89C36B" bold="true" box="[151,348,1337,1362]" pageId="40" pageNumber="41">Adult diagnosis.</emphasis>
A member of
<taxonomicName id="4C0A4D14FFCEFFA7FDF6FAD2FC98C36B" box="[529,845,1337,1362]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFCEFFA7FDF6FAD2FD75C36B" box="[529,672,1337,1362]" italics="true" pageId="40" pageNumber="41">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFCEFFA7FD52FAD2FC91C36B" box="[693,836,1337,1362]" italics="true" pageId="40" pageNumber="41">Cycadophila</emphasis>
)
</taxonomicName>
<taxonomicName id="4C0A4D14FFCEFFA7FCBFFAD1FC4EC368" box="[856,923,1338,1361]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="papua">papua</taxonomicName>
species group with long maxillary palpi as discussed for the group, recognized by its narrow orange mark on the elytra, prosternal process with coxal lines diverging posteriorly, and occurring in
<collectingCountry id="F31D7607FFCEFFA7FDAAFA69FCFCC3A3" box="[589,809,1409,1434]" name="Papua New Guinea" pageId="40" pageNumber="41">
<taxonomicName id="4C0A4D14FFCEFFA7FDAAFA69FD47C3A0" box="[589,658,1410,1433]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="43" phylum="Arthropoda" rank="species" species="papua">
<collectingRegion id="49CEF875FFCEFFA7FDAAFA69FD47C3A0" box="[589,658,1410,1433]" country="Indonesia" name="Papua Barat" pageId="40" pageNumber="41">Papua</collectingRegion>
</taxonomicName>
New Guinea
</collectingCountry>
.
</paragraph>
<paragraph id="8BB53697FFCEFFA7FF20FA4EFB5CC048" blockId="40.[151,1436,1337,2010]" pageId="40" pageNumber="41">
<emphasis id="B97EEA85FFCEFFA7FF20FA4EFE74C387" bold="true" box="[199,417,1445,1470]" pageId="40" pageNumber="41">Adult description.</emphasis>
Length
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, width
<quantity id="4CF29B72FFCEFFA7FD19FA4DFC73C384" box="[766,934,1446,1470]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="40" pageNumber="41" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.702.80 mm</quantity>
. Body in dorsal view elongate, sides evenly arcuate, greatest width at middle of elytra; in lateral view weakly convex dorsally. General body and appendage color solid reddish-brown, with nearly black base of head, pronotal disc, and sutural and lateral margins of elytra, remaining red-orange mark on elytra broad, at base from striae IVVI; dorsal surface distinctly punctate, surface glossy, dorsally appearing glabrous, with short procumbent hairs associated with punctation behind eyes and along lateral sides and declivity of elytra, ventrally mostly covered with short procumbent setae.
</paragraph>
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<subSubSection id="C310651CFFCEFFA5FF20F996FACCC52C" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="description">
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<emphasis id="B97EEA85FFCEFFA7FF20F996FED0C0AF" bold="true" box="[199,261,1661,1686]" italics="true" pageId="40" pageNumber="41">Head</emphasis>
in dorsal view conical, gradually narrowed apically, surface flat to slightly convex, distinctly punctured; width
<quantity id="4CF29B72FFCEFFA7FF39F949FE51C080" box="[222,388,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="1.135" metricValueMax="1.17" metricValueMin="1.1" pageId="40" pageNumber="41" unit="mm" value="1.135" valueMax="1.17" valueMin="1.1">1.101.17 mm</quantity>
; dorsal interocular distance
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, head width/dorsal interocular distance ratio 1.70 1.77, ventral interocular distance
<quantity id="4CF29B72FFCEFFA7FDC4F92DFD19C0E4" box="[547,716,1734,1758]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="40" pageNumber="41" unit="mm" value="0.45499999999999996" valueMax="0.48" valueMin="0.43">0.430.48 mm</quantity>
, head width/ventral interocular distance ratio 2.442.56. Eye globular; bordered dorsally with supraocular stria; projected laterally, with large black facets and short, inconspicuous interfacetal setae. Frons with short, dark, subcuticular suture on each side, approximately 1/4 width of frons in length, extending obliquely to lateral margin above and anterior to antennal insertion. Antennal length about 0.9 0 times pronotal width, and longer than head width, antennomere relative lengths from base to end approximately 16:12:14:10:10:10:10:10:14:14:18; antennomere I (scape) fairly large, slightly elongate, antennomere II slightly smaller than III; IVVIII small, width equals length; club fairly large, IXXI similar in length, XI globular. Clypeus truncate anteriorly, anterior margin with long setae; densely punctate; somewhat emarginate, with narrow margin. Maxillary palpomeres II and III longer than wide; IV elongate, apex densely papillate; relative lengths of IIIV approximately 4:4:8. Labium with mentum pentagonal with carina forming triangular plate projecting medially, with small lateral pockets; labial palpomere III elongate-oval.
<collectingRegion id="49CEF875FFCFFFA6FAECFF0BFABFC6CE" box="[1291,1386,224,247]" country="Myanmar" name="Chin" pageId="41" pageNumber="42">Mentum</collectingRegion>
and submentum with moderate punctation and setation, setae short. Gular area smooth, without setae or punctation, except near suture with submentum where a shallow U-shaped fovea is present, deepest laterally, bearing setose punctures.
</paragraph>
<caption id="DF75661FFFCFFFA6FF70F9CAFD22C063" httpUri="https://zenodo.org/record/575654/files/figure.png" pageId="41" pageNumber="42" targetBox="[151,1436,409,1548]" targetPageId="41">
<paragraph id="8BB53697FFCFFFA6FF70F9CAFD22C063" blockId="41.[151,1435,1569,1626]" pageId="41" pageNumber="42">
<emphasis id="B97EEA85FFCFFFA6FF70F9CAFEF7C00E" bold="true" box="[151,290,1569,1591]" pageId="41" pageNumber="42">FIGURE 13.</emphasis>
Photographs of
<taxonomicName id="4C0A4D14FFCFFFA6FE37F9CAFCE1C00E" box="[464,820,1569,1591]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="papua" subGenus="Cycadophila">
<emphasis id="B97EEA85FFCFFFA6FE37F9CAFD87C00E" box="[464,594,1569,1591]" italics="true" pageId="41" pageNumber="42">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFCFFFA6FD84F9CAFD31C00E" box="[611,740,1569,1591]" italics="true" pageId="41" pageNumber="42">Cycadophila</emphasis>
)
<emphasis id="B97EEA85FFCFFFA6FD12F9C9FCE1C00E" box="[757,820,1570,1591]" italics="true" pageId="41" pageNumber="42">papua</emphasis>
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, holotype: A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia.
</paragraph>
</caption>
<paragraph id="8BB53697FFCFFFA5FF20F961FC86C75D" blockId="41.[151,1436,1674,2023]" lastBlockId="42.[151,1437,151,1184]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<emphasis id="B97EEA85FFCFFFA6FF20F961FEC8C09A" bold="true" box="[199,285,1674,1699]" italics="true" pageId="41" pageNumber="42">Thorax</emphasis>
with pronotum transverse in dorsal view, with marginal beads on all sides; length/width ratio 0.58 0.61, moderately convex, sides evenly arcuate entire length; anterior angles rounded; lateral carina with fine bead, bead bearing row of fine setose punctures; posterior angle exteriorly angulate; posterior margin projecting medially; with narrow, short longitudinal furrow posterolaterally, beginning at pore on posterior margin, extending forward and ending abruptly, length of furrows relative to pronotal length 0.170.19. Prosternum with anterior margin slightly emarginate, finely denticulate with fringe of long, anteriorly directed setae; posterior prosternal process round and convex; coxal lines diverging posteriorly. Hypomeron with coarse punctures laterally, medially with longitudinal striations. Scutellar shield transverse, posterior margin projected, pentagonal. Elytra elongate, evenly gradually arcuate laterally, convex; length/width 3.383.43, greatest width near midlength; with marginal bead basally, scutellary striole with 10 punctures, intervals of striae with distinct fine punctures. All elytral punctures bearing a single short seta; seta length barely reaching outside of puncture. Punctation on meso- and metaventrite distinct. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over half metaventral length. Legs narrow, similar in length; femora compressed laterally; tibiae shorter than femora, weakly dilated to narrow apex. Protibia weakly dilated at apex, apical and lateral margin lacking stout spinules, 23 small stout spines present on apical-lateral angle giving angle a sharper appearance. Meso- and metatibia less dilated, nearly parallel-sided at apex.
</paragraph>
<paragraph id="8BB53697FFCCFFA5FF20FE84FD7DC405" blockId="42.[151,1437,151,1184]" pageId="42" pageNumber="43">
<emphasis id="B97EEA85FFCCFFA5FF20FE84FEE0C7B1" bold="true" box="[199,309,367,392]" italics="true" pageId="42" pageNumber="43">Abdomen</emphasis>
with 5 ventrites bearing fine punctures; anterior margin with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I lacking subcoxal line, slightly longer medially than II; IIV each with pair short hairs located adjacent to midline (often abraded); IIIV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex bearing row of short appressed, densely arranged setae.
</paragraph>
<paragraph id="8BB53697FFCCFFA5FF20FDACFACCC52C" blockId="42.[151,1437,151,1184]" pageId="42" pageNumber="43">
<emphasis id="B97EEA85FFCCFFA5FF20FDACFEBBC459" bold="true" box="[199,366,583,608]" italics="true" pageId="42" pageNumber="43">Male genitalia</emphasis>
(tegmen and median lobe of aedeagus) twisted approximately 45° toward the left side of animal and resting on side. Tegmen sclerotized, triangular, gently twisted; with anterior region ring-like, posterior region sheath-like, posterior margin slightly arcuate; lateral margins gradually converging posteriorly; dorsally with 2 large, elongate, somewhat spatula-shaped parameres with oblique apex. Parameres with long setae apically; length/ width ratio 2:1. Aedeagus with median lobe short, slender, highly sclerotized, apex acuminate. Length of median lobe to penile strut ratio is around 1:4.
<emphasis id="B97EEA85FFCCFFA5FDAEFD17FD75C52C" bold="true" box="[585,672,764,789]" italics="true" pageId="42" pageNumber="43">Female</emphasis>
similar to male; pro- and mesotibia narrower than male.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFCCFFA5FF20FCF4FECCC5D5" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8BB53697FFCCFFA5FF20FCF4FD65C501" blockId="42.[151,1437,151,1184]" box="[199,688,799,824]" pageId="42" pageNumber="43">
<materialsCitation id="3B623CCAFFCCFFA5FF20FCF4FD65C501" ID-GBIF-Occurrence="1502685021" box="[199,688,799,824]" country="Papua New Guinea" location="Papua New Guinea" pageId="42" pageNumber="43" specimenCount="1" typeStatus="holotype">
<emphasis id="B97EEA85FFCCFFA5FF20FCF4FEB0C501" bold="true" box="[199,357,799,824]" pageId="42" pageNumber="43">
<typeStatus id="54B18835FFCCFFA5FF20FCF4FED7C501" box="[199,258,799,824]" pageId="42" pageNumber="43">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D7607FFCCFFA5FE8BFCCBFD92C501" box="[364,583,799,824]" name="Papua New Guinea" pageId="42" pageNumber="43">
<taxonomicName id="4C0A4D14FFCCFFA5FE8BFCCBFE65C50E" box="[364,432,800,823]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="papua">Papua</taxonomicName>
New Guinea
</collectingCountry>
, Nadzad.
</materialsCitation>
</paragraph>
<paragraph id="8BB53697FFCCFFA5FF20FCAFFECCC5D5" blockId="42.[151,1437,151,1184]" pageId="42" pageNumber="43">
<emphasis id="B97EEA85FFCCFFA5FF20FCAFFDF6C564" bold="true" box="[199,547,836,861]" pageId="42" pageNumber="43">Material examined. Holotype</emphasis>
(by designation) male with the following labels: 1) [rectangular; white; printed in black ink] “Nadzab, Markam R. Val. NGuinea, Au, K. V. Krombein”; 2) [rectangular; white; printed in black ink] “ExColl KVKrombein”; 3) [rectangular; red; printed in black ink] “HOLOTYPE ♂
<taxonomicName id="4C0A4D14FFCCFFA5FB78FC67FE2BC5F1" authority="P. Skelley, G. Xu &amp; W. Tang 2017" authorityName="P. Skelley, G. Xu &amp; W. Tang" authorityYear="2017" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="papua">
<emphasis id="B97EEA85FFCCFFA5FB78FC67FAABC59D" box="[1183,1406,908,933]" italics="true" pageId="42" pageNumber="43">Cycadophila papua</emphasis>
P. Skelley, G. Xu &amp; W. Tang 2017
</taxonomicName>
”. Deposited in the USNM.
<materialsCitation id="3B623CCAFFCCFFA5FCDBFC44FEC0C5D5" ID-GBIF-Occurrence="1502685027" collectionCode="USNM" collectorName="Deposited" pageId="42" pageNumber="43" specimenCount="1" typeStatus="allotype">
<emphasis id="B97EEA85FFCCFFA5FCDBFC44FC74C5F1" bold="true" box="[828,929,943,968]" pageId="42" pageNumber="43">
<typeStatus id="54B18835FFCCFFA5FCDBFC44FC74C5F1" box="[828,929,943,968]" pageId="42" pageNumber="43" type="allotype">Allotype</typeStatus>
</emphasis>
female, same data as holotype.
<collectorName id="26FF5341FFCCFFA5FAEAFC44FAAAC5F1" box="[1293,1407,943,968]" pageId="42" pageNumber="43">Deposited</collectorName>
in the
<collectionCode id="ED1BAE52FFCCFFA5FF27FC3FFEC0C5D5" box="[192,277,980,1004]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFCCFFA5FF20FC1CFA55C229" box="[199,1408,1015,1040]" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8BB53697FFCCFFA5FF20FC1CFA55C229" blockId="42.[151,1437,151,1184]" box="[199,1408,1015,1040]" pageId="42" pageNumber="43">
<emphasis id="B97EEA85FFCCFFA5FF20FC1CFE9BC229" bold="true" box="[199,334,1015,1040]" pageId="42" pageNumber="43">Etymology.</emphasis>
Named for the region of New
<collectingCountry id="F31D7607FFCCFFA5FD41FC1CFD22C229" box="[678,759,1015,1040]" name="Guinea-Bissau" pageId="42" pageNumber="43">Guinea</collectingCountry>
the
<typeStatus id="54B18835FFCCFFA5FCC0FC13FCB7C229" box="[807,866,1016,1040]" pageId="42" pageNumber="43">types</typeStatus>
are from,
<taxonomicName id="4C0A4D14FFCCFFA5FC31FC13FBCFC236" box="[982,1050,1016,1039]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="papua">
<collectingRegion id="49CEF875FFCCFFA5FC31FC13FBCFC236" box="[982,1050,1016,1039]" country="Indonesia" name="Papua Barat" pageId="42" pageNumber="43">Papua</collectingRegion>
</taxonomicName>
is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFCCFFA5FF20FBF7FC06C299" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BB53697FFCCFFA5FF20FBF7FC06C299" blockId="42.[151,1437,151,1184]" pageId="42" pageNumber="43">
<emphasis id="B97EEA85FFCCFFA5FF20FBF7FEEEC20C" bold="true" box="[199,315,1052,1077]" pageId="42" pageNumber="43">Remarks.</emphasis>
Based on the fact that the closely related species,
<taxonomicName id="4C0A4D14FFCCFFA5FC9BFBF7FBBDC20D" box="[892,1128,1052,1077]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="samara">
<emphasis id="B97EEA85FFCCFFA5FC9BFBF7FBBDC20D" box="[892,1128,1052,1077]" italics="true" pageId="42" pageNumber="43">Cycadophila samara</emphasis>
</taxonomicName>
, in this species group, was collected on cycads, it is anticipated that
<taxonomicName id="4C0A4D14FFCCFFA5FD99FBABFD3DC261" box="[638,744,1088,1112]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="papua">
<emphasis id="B97EEA85FFCCFFA5FD99FBABFD3DC261" box="[638,744,1088,1112]" italics="true" pageId="42" pageNumber="43">C. papua</emphasis>
</taxonomicName>
also has a cycad association. The
<typeStatus id="54B18835FFCCFFA5FB64FBABFB66C261" box="[1155,1203,1088,1112]" pageId="42" pageNumber="43">type</typeStatus>
locality, Nadzab, is situated in the highlands of New
<collectingCountry id="F31D7607FFCCFFA5FDE9FB8FFD8AC244" box="[526,607,1124,1149]" name="Guinea-Bissau" pageId="42" pageNumber="43">Guinea</collectingCountry>
and is habitat for
<taxonomicName id="4C0A4D14FFCCFFA5FCC9FB8EFB65C245" authority="Hill 1994a" authorityName="Hill" authorityYear="1994" box="[814,1200,1124,1149]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="schumanniana">
<emphasis id="B97EEA85FFCCFFA5FCC9FB8EFBCBC244" box="[814,1054,1124,1149]" italics="true" pageId="42" pageNumber="43">Cycas schumanniana</emphasis>
(
<bibRefCitation id="EF9B4B66FFCCFFA5FBCAFB8FFB7DC245" author="Hill" box="[1069,1192,1124,1149]" pageId="42" pageNumber="43" refString="Hill, K. D. (1994 a) The Cycas media group (Cycadaceae) in New Guinea. Australian Systematic Botany, 7, 527 - 541. https: // doi. org / 10.1071 / SB 9940527" type="journal article" year="1994" yearSuffix="a">Hill 1994a</bibRefCitation>
)
</taxonomicName>
and another species,
<taxonomicName id="4C0A4D14FFCCFFA5FF70FB63FEC5C299" box="[151,272,1159,1184]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="rumphii">
<emphasis id="B97EEA85FFCCFFA5FF70FB63FEC5C299" box="[151,272,1159,1184]" italics="true" pageId="42" pageNumber="43">C. rumphii</emphasis>
</taxonomicName>
, occurs along coastal habitats at lower elevations (
<bibRefCitation id="EF9B4B66FFCCFFA5FCAEFB6CFC12C299" author="Hill" box="[841,967,1159,1184]" pageId="42" pageNumber="43" refString="Hill, K. D. (1994 b) The Cycas rumphii complex (Cycadaceae) in New Guinea and the Western Pacific. Australian Systematic Botany, 7, 543 - 567." type="journal article" year="1994" yearSuffix="b">Hill 1994b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>