229 lines
28 KiB
XML
229 lines
28 KiB
XML
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<mods:title id="C9B3E8C6E6D9B9AE2686584A54B72D7F">A new polydesmid milliped genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett and Gardner, 1968, and a summary of the family in Western North America (Polydesmida: Polydesmidae)</mods:title>
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<mods:namePart id="B93963975379FBB39EBD0B74D6990470">Shelley, Rowland M.</mods:namePart>
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<mods:date id="3F95F2C1F09827987FD2707E7B4EBF58">2003</mods:date>
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<treatment id="03A3096E3A1DFFFBF462FBEE4E16C6A3" ID-DOI="http://doi.org/10.5281/zenodo.5014405" ID-GBIF-Taxon="183531914" ID-Zenodo-Dep="5014405" LSID="urn:lsid:plazi:treatment:03A3096E3A1DFFFBF462FBEE4E16C6A3" httpUri="http://treatment.plazi.org/id/03A3096E3A1DFFFBF462FBEE4E16C6A3" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<subSubSection id="C310EBF33A1DFFFAF462FBEE4F1EC67B" box="[264,837,1044,1070]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BB5B8783A1DFFFAF462FBEE4F1EC67B" blockId="8.[264,837,1044,1070]" box="[264,837,1044,1070]" pageId="8" pageNumber="9">
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<heading id="D0FD0F143A1DFFFAF462FBEE4F1EC67B" bold="true" box="[264,837,1044,1070]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="B97E646A3A1DFFFAF462FBEE4F1EC67B" bold="true" box="[264,837,1044,1070]" pageId="8" pageNumber="9">
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Genus
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<taxonomicName id="4C0AC3FB3A1DFFFAF435FBEE4F1EC67B" ID-CoL="7NQQN" authority="Buckett and Gardner, 1968" authorityName="Buckett and Gardner" authorityYear="1968" box="[351,837,1044,1070]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B97E646A3A1DFFFAF435FBEE4DB0C67B" bold="true" box="[351,491,1044,1070]" italics="true" pageId="8" pageNumber="9">Bidentogon</emphasis>
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<bibRefCitation id="EF9BC5893A1DFFFAF499FBEE4F1EC67B" author="Buckett, J. S. & Gardner, M. R." box="[499,837,1044,1070]" pageId="8" pageNumber="9" pagination="198 - 202" refId="ref5008" refString="Buckett, J. S. & Gardner, M. R. (1968) A new genus and species of milliped from northern California (Polydesmida: Vanhoeffeniidae). Pan-Pacific Entomologist, 44, 198 - 202." type="journal article" year="1968">Buckett and Gardner, 1968</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C310EBF33A1DFFFAF462FBA54FFEC6C0" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="8BB5B8783A1DFFFAF462FBA54FFEC6C0" blockId="8.[264,1208,1118,1174]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="AB1A9F563A1DFFFAF462FBA54FFEC6C0" pageId="8" pageNumber="9">
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<treatmentCitation id="0AAB9E693A1DFFFAF462FBA54E80C620" author="Buckett, J. S. & Gardner, M. R." box="[264,731,1118,1142]" page="198" pageId="8" pageNumber="9" year="1968">
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<taxonomicName id="4C0AC3FB3A1DFFFAF462FBA54E80C620" ID-CoL="7NQQN" authority="Buckett and Gardner, 1968: 198" authorityName="Buckett and Gardner" authorityPageNumber="198" authorityYear="1968" box="[264,731,1118,1142]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B97E646A3A1DFFFAF462FBA54DD9C623" box="[264,386,1119,1142]" italics="true" pageId="8" pageNumber="9">Bidentogon</emphasis>
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<bibRefCitation id="EF9BC5893A1DFFFAF4E3FBA44E80C620" author="Buckett, J. S. & Gardner, M. R." box="[393,731,1118,1141]" pageId="8" pageNumber="9" pagination="198 - 202" refId="ref5008" refString="Buckett, J. S. & Gardner, M. R. (1968) A new genus and species of milliped from northern California (Polydesmida: Vanhoeffeniidae). Pan-Pacific Entomologist, 44, 198 - 202." type="journal article" year="1968">Buckett and Gardner, 1968:198</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<treatmentCitation id="0AAB9E693A1DFFFAF786FBA44F94C620" author="Shear, W. A." box="[748,975,1118,1141]" page="489" pageId="8" pageNumber="9" year="1972">
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<bibRefCitation id="EF9BC5893A1DFFFAF786FBA44F94C620" author="Shear, W. A." box="[748,975,1118,1141]" pageId="8" pageNumber="9" pagination="489 - 492" refId="ref5398" refString="Shear, W. A. (1972) The milliped genus Bidentogon (Diplopoda, Polydesmida, Trichopolydesmidae). Proceedings of the Biological Society of Washington, 85, 489 - 492." type="journal article" year="1972">Shear, 1972:489490</bibRefCitation>
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</treatmentCitation>
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||
.
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||
<treatmentCitation id="0AAB9E693A1DFFFAF6B5FBA448E8C620" author="Hoffman, R. L." box="[991,1203,1118,1141]" page="172" pageId="8" pageNumber="9" year="1980">
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<bibRefCitation id="EF9BC5893A1DFFFAF6B5FBA448E8C620" author="Hoffman, R. L." box="[991,1203,1118,1141]" pageId="8" pageNumber="9" refId="ref5165" refString="Hoffman, R. L. (1980) Classification of the Diplopoda. Museum d'Histoire Naturelle, Geneve, 237 pp." type="book" year="1980">Hoffman, 1980:172</bibRefCitation>
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||
</treatmentCitation>
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||
;
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<treatmentCitation id="0AAB9E693A1DFFFAF452FB844DFBC6C0" author="Hoffman, R. L." box="[312,416,1150,1173]" page="438" pageId="8" pageNumber="9" year="1999">
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||
<bibRefCitation id="EF9BC5893A1DFFFAF452FB844DFBC6C0" author="Hoffman, R. L." box="[312,416,1150,1173]" pageId="8" pageNumber="9" pagination="1 - 584" refId="ref5188" refString="Hoffman, R. L. (1999) Checklist of the Millipeds of North and Middle America. Virginia Mu s eum of Natural History Special Publication No. 8, 1 - 584." type="book chapter" year="1999">1999:438</bibRefCitation>
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</treatmentCitation>
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||
.
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<treatmentCitation id="0AAB9E693A1DFFFAF4DBFB844E96C6C0" author="Shelley, R. M. & Sierwald, P. & Kiser, S. B. & Golovatch, S. I." box="[433,717,1150,1174]" page="87" pageId="8" pageNumber="9" year="2000">
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<bibRefCitation id="EF9BC5893A1DFFFAF4DBFB844E96C6C0" author="Shelley, R. M. & Sierwald, P. & Kiser, S. B. & Golovatch, S. I." box="[433,717,1150,1174]" pageId="8" pageNumber="9" refId="ref5755" refString="Shelley, R. M., Sierwald, P., Kiser, S. B. & Golovatch, S. I. (2000) Nomenclator generum et familiarum Diplopodorum II. A List of the Genus and Family-Group Names in the Class Diplopoda from 1958 through 1999. Pensoft Publishers, Sofia-Moscow. 167 pp." type="book" year="2000">
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Shelley
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<emphasis id="B97E646A3A1DFFFAF76DFB854E66C6C3" box="[519,573,1151,1174]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
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, 2000:87–88
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</bibRefCitation>
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||
</treatmentCitation>
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||
.
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<treatmentCitation id="0AAB9E693A1DFFFAF7B7FB844FFAC6C0" author="Shelley, R. M." box="[733,929,1150,1173]" page="108" pageId="8" pageNumber="9" year="2002">
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||
<bibRefCitation id="EF9BC5893A1DFFFAF7B7FB844FFAC6C0" author="Shelley, R. M." box="[733,929,1150,1173]" pageId="8" pageNumber="9" pagination="90 - 115" refId="ref5719" refString="Shelley, R. M. (2002) Annotated checklist of the millipeds of California (Arthropoda: Diplopoda). Monographs of the Western North American Naturalist, 1, 90 - 115." type="journal article" year="2002">Shelley, 2002:108</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C310EBF33A1DFFFBF462FB3D4E16C6A3" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
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||
<paragraph id="8BB5B8783A1DFFFAF462FB3D48E7C6B4" blockId="8.[264,1212,1223,1249]" box="[264,1212,1223,1249]" pageId="8" pageNumber="9">
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||
<emphasis id="B97E646A3A1DFFFAF462FB3D4DF9C6B5" box="[264,418,1223,1248]" italics="true" pageId="8" pageNumber="9">
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||
<typeStatus id="54B106DA3A1DFFFAF462FB3D4D64C6B5" box="[264,319,1223,1248]" pageId="8" pageNumber="9">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C0AC3FB3A1DFFFAF4DBFB3D4FC2C6B4" authority="Buckett and Gardner, 1968" authorityName="Buckett and Gardner" authorityYear="1968" box="[433,921,1223,1249]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="helferorum">
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<emphasis id="B97E646A3A1DFFFAF4DBFB3D4E09C6B5" box="[433,594,1223,1248]" italics="true" pageId="8" pageNumber="9">B. helferorum</emphasis>
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<bibRefCitation id="EF9BC5893A1DFFFAF730FB3D4FC2C6B4" author="Buckett, J. S. & Gardner, M. R." box="[602,921,1223,1249]" pageId="8" pageNumber="9" pagination="198 - 202" refId="ref5008" refString="Buckett, J. S. & Gardner, M. R. (1968) A new genus and species of milliped from northern California (Polydesmida: Vanhoeffeniidae). Pan-Pacific Entomologist, 44, 198 - 202." type="journal article" year="1968">Buckett and Gardner, 1968</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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||
<paragraph id="8BB5B8783A1DFFFAF462FAEE4FB0C72B" blockId="8.[264,1323,1300,1886]" pageId="8" pageNumber="9">
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<emphasis id="B97E646A3A1DFFFAF462FAEE4DDCC778" box="[264,391,1300,1325]" italics="true" pageId="8" pageNumber="9">Diagnosis.</emphasis>
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A genus of minute
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<taxonomicName id="4C0AC3FB3A1DFFFAF716FAEE4F7BC77B" authorityName="Leach" authorityYear="1815" box="[636,800,1300,1326]" class="Diplopoda" family="Polydesmidae" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Polydesmidae</taxonomicName>
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without distinct pulvillus but with scattered hairs along caudal surface of gonopodal telopodite; division point located distad, tibiotarsus and endomerite widely separated, subparallel to each other.
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</paragraph>
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<paragraph id="8BB5B8783A1DFFFAF452FA764D81C7F3" blockId="8.[264,1323,1300,1886]" box="[312,474,1420,1446]" pageId="8" pageNumber="9">
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||
<emphasis id="B97E646A3A1DFFFAF452FA764DCCC7F0" box="[312,407,1420,1445]" italics="true" pageId="8" pageNumber="9">Species.</emphasis>
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Two
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</paragraph>
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<paragraph id="8BB5B8783A1DFFFAF452FA4E4F4FC7A3" blockId="8.[264,1323,1300,1886]" pageId="8" pageNumber="9">
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||
<emphasis id="B97E646A3A1DFFFAF452FA4E4D94C798" box="[312,463,1460,1485]" italics="true" pageId="8" pageNumber="9">Distribution.</emphasis>
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||
The coast of
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<collectingRegion id="49CE769A3A1DFFFAF719FA4E4EB1C79B" box="[627,746,1460,1486]" country="United States of America" name="California" pageId="8" pageNumber="9">California</collectingRegion>
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from Mendocino to Santa Cruz counties, extending inland to Sacramento County (
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<figureCitation id="1331A4FD3A1DFFFAF7F1FA264E84C7A3" box="[667,735,1500,1526]" captionStart="FIGURE 1" captionStartId="2.[264,368,1327,1351]" captionTargetBox="[470,1120,597,1316]" captionTargetId="figure-214@2.[464,1126,597,1317]" captionTargetPageId="2" captionText="FIGURE 1. Occurrence of the Polydesmidae in western North America; 1, Scytonotus; 2, Calianotus; 3, Utadesmus; 4, Speodesmus; 5, Bidentogon; 6, Retrorsia." figureDoi="http://doi.org/10.5281/zenodo.5014429" httpUri="https://zenodo.org/record/5014429/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
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[5]).
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</paragraph>
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<paragraph id="8BB5B8783A1DFFFBF452F9FE48E4C16B" blockId="8.[264,1323,1300,1886]" lastBlockId="9.[264,1323,284,1270]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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||
<emphasis id="B97E646A3A1DFFFAF452F9FE4DC5C448" box="[312,414,1540,1565]" italics="true" pageId="8" pageNumber="9">Remarks</emphasis>
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||
.
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<bibRefCitation id="EF9BC5893A1DFFFAF4D9F9FE4EC8C44B" author="Chamberlin, R. V." box="[435,659,1540,1566]" pageId="8" pageNumber="9" pagination="9 - 11" refId="ref5079" refString="Chamberlin, R. V. (1918) Four new western diplopods. Pomona College Journal of Entomology and Zoology, 10, 9 - 11." type="journal article" year="1918">Chamberlin (1918)</bibRefCitation>
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||
based his description of
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<taxonomicName id="4C0AC3FB3A1DFFFAF6DFF9FE48A8C448" authorityName="Shelley" authorityYear="2003" box="[949,1267,1540,1565]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="californicus">
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||
<emphasis id="B97E646A3A1DFFFAF6DFF9FE48A8C448" box="[949,1267,1540,1565]" italics="true" pageId="8" pageNumber="9">Brachydesmus californicus</emphasis>
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||
</taxonomicName>
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on a sample from Stanford, Santa Clara County, containing only
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<specimenCount id="9D0C73F13A1DFFFAF6BCF9D6481DC413" box="[982,1094,1580,1606]" pageId="8" pageNumber="9" type="male">one male</specimenCount>
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||
, which is therefore fixed as the
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<typeStatus id="54B106DA3A1DFFFAF4FDF9AE4DA5C43B" box="[407,510,1620,1646]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
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||
by monotypy (Article 73.1.2 of the Code); the date of collection was not provided. During the next 46 years,
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||
<bibRefCitation id="EF9BC5893A1DFFFAF798F9864FFAC4C3" author="Attems, C." box="[754,929,1660,1686]" pageId="8" pageNumber="9" pagination="1 - 576" refId="ref4897" refString="Attems, C. (1940) Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich, Lief. 70, 1 - 576." type="journal article" year="1940">Attems (1940)</bibRefCitation>
|
||
,
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||
<bibRefCitation id="EF9BC5893A1DFFFAF6C5F986497DC4C3" author="Chamberlin, R. V. & Hoffman, R. L." box="[943,1318,1660,1686]" pageId="8" pageNumber="9" pagination="1 - 236" refId="ref5129" refString="Chamberlin, R. V., & Hoffman, R. L. (1958) Checklist of the millipeds of North America. United States National Museum Bulletin 212, 1 - 236." type="journal article" year="1958">Chamberlin & Hoffman (1958)</bibRefCitation>
|
||
, and
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||
<bibRefCitation id="EF9BC5893A1DFFFAF456F95E4DB4C4EB" author="Buckett, J. S." box="[316,495,1700,1726]" pageId="8" pageNumber="9" refId="ref4979" refString="Buckett, J. S. (1964) Annotated list of the Diplopoda of California. Simmons Publishing Co., Davis, California, 34 pp." type="book" year="1964">Buckett (1964)</bibRefCitation>
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all recorded Stanford as the type locality, but
|
||
<bibRefCitation id="EF9BC5893A1DFFFAF149F95E48E5C4EB" author="Shear, W. A." box="[1059,1214,1700,1726]" pageId="8" pageNumber="9" pagination="489 - 492" refId="ref5398" refString="Shear, W. A. (1972) The milliped genus Bidentogon (Diplopoda, Polydesmida, Trichopolydesmidae). Proceedings of the Biological Society of Washington, 85, 489 - 492." type="journal article" year="1972">Shear (1972)</bibRefCitation>
|
||
reported discovering the type sample at the MCZ, which contained an unspecified number of male and female
|
||
<typeStatus id="54B106DA3A1DFFFAF4C8F90E4E50C55B" box="[418,523,1780,1806]" pageId="8" pageNumber="9" type="syntype">syntypes</typeStatus>
|
||
, was collected in
|
||
<date id="FFB49EB83A1DFFFAF795F90E4FFFC55B" box="[767,932,1780,1806]" pageId="8" pageNumber="9" value="1914-01">January 1914</date>
|
||
from Sacramento, Sacramento County, and was labeled “ ‘
|
||
<typeStatus id="54B106DA3A1DFFFAF730F8E64EA5C563" box="[602,766,1820,1846]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
|
||
.’ ” Subsequently,
|
||
<bibRefCitation id="EF9BC5893A1DFFFAF6BDF8E648CCC563" author="Hoffman, R. L." box="[983,1175,1820,1846]" pageId="8" pageNumber="9" pagination="1 - 584" refId="ref5188" refString="Hoffman, R. L. (1999) Checklist of the Millipeds of North and Middle America. Virginia Mu s eum of Natural History Special Publication No. 8, 1 - 584." type="book chapter" year="1999">Hoffman (1999)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF9BC5893A1DFFFAF1B8F8E64D0DC50B" author="Shelley, R. M." pageId="8" pageNumber="9" pagination="90 - 115" refId="ref5719" refString="Shelley, R. M. (2002) Annotated checklist of the millipeds of California (Arthropoda: Diplopoda). Monographs of the Western North American Naturalist, 1, 90 - 115." type="journal article" year="2002">Shelley (2002)</bibRefCitation>
|
||
reported Sacramento as the type locality. I have examined this sample and found it to contain
|
||
<specimenCount id="9D0C73F13A1CFFFBF4E8FEE64DA2C363" box="[386,505,284,310]" pageId="9" pageNumber="10" type="male">two males</specimenCount>
|
||
and
|
||
<specimenCount id="9D0C73F13A1CFFFBF758FEE64E95C363" box="[562,718,284,310]" pageId="9" pageNumber="10" type="female">three females</specimenCount>
|
||
with a label stating “
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF6AFFEE64958C360" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[965,1283,284,309]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="californicus" status="sp. nov.">
|
||
<emphasis id="B97E646A3A1CFFFBF6AFFEE64958C360" box="[965,1283,284,309]" italics="true" pageId="9" pageNumber="10">Brachydesmus californicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DD9113A1CFFFBF063FEE64D45C30B" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
” instead of one stating “
|
||
<typeStatus id="54B106DA3A1CFFFBF72CFEBE4EB1C30B" box="[582,746,324,350]" pageId="9" pageNumber="10" type="holotype">HOLOTYPE</typeStatus>
|
||
.” Evidently, Chamberlin or somebody else mistakenly placed the “ sp. n.” label with the Sacramento sample because it does not match the locality or number of specimens reported by
|
||
<bibRefCitation id="EF9BC5893A1CFFFBF656FE6E4847C3FB" author="Chamberlin, R. V." box="[828,1052,404,430]" pageId="9" pageNumber="10" pagination="9 - 11" refId="ref5079" refString="Chamberlin, R. V. (1918) Four new western diplopods. Pomona College Journal of Entomology and Zoology, 10, 9 - 11." type="journal article" year="1918">Chamberlin (1918)</bibRefCitation>
|
||
in the original description. There is a vial with
|
||
<specimenCount id="9D0C73F13A1CFFFBF741FE464ECEC383" box="[555,661,444,470]" pageId="9" pageNumber="10" type="male">one male</specimenCount>
|
||
at the NMNH that was collected at Stanford in 1910 and matches the requirements for the type, but the possibility cannot be discounted that there might have been another pre1918 sample from Stanford that is now lost. As both the Sacramento sample and that taken at Stanford in 1910 were collected prior to Chamberlin’s description (1918) of
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF76DFDA64EE8C020" authorityName="Shelley" authorityYear="2003" box="[519,691,604,629]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="californicus">
|
||
<emphasis id="B97E646A3A1CFFFBF76DFDA64EE8C020" box="[519,691,604,629]" italics="true" pageId="9" pageNumber="10">B. californicus</emphasis>
|
||
</taxonomicName>
|
||
, one can conjecture that he had both samples on hand when preparing the account and placed the label in the wrong vial himself. We will never know what happened, but Chamberlin’s clear intent, as evidenced by the published account, was to apply this name to the form at Stanford, the designated type locality. I have therefore placed a label in this vial stating “
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF63BFD064D1EC16B" authority="Chamberlin, 1918" authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="californicus">
|
||
<emphasis id="B97E646A3A1CFFFBF63BFD0648CBC140" box="[849,1168,764,789]" italics="true" pageId="9" pageNumber="10">Brachydesmus californicus</emphasis>
|
||
<bibRefCitation id="EF9BC5893A1CFFFBF1F3FD064D1EC16B" author="Chamberlin, R. V." pageId="9" pageNumber="10" pagination="9 - 11" refId="ref5079" refString="Chamberlin, R. V. (1918) Four new western diplopods. Pomona College Journal of Entomology and Zoology, 10, 9 - 11." type="journal article" year="1918">Chamberlin, 1918</bibRefCitation>
|
||
</taxonomicName>
|
||
, PROBABLE
|
||
<typeStatus id="54B106DA3A1CFFFBF498FCDE4ECBC16B" box="[498,656,804,830]" pageId="9" pageNumber="10" type="holotype">HOLOTYPE</typeStatus>
|
||
" and deposited it in the NMNH type collection.
|
||
</paragraph>
|
||
<paragraph id="8BB5B8783A1CFFFBF452FCB64E16C6A3" blockId="9.[264,1323,284,1270]" pageId="9" pageNumber="10">
|
||
<bibRefCitation id="EF9BC5893A1CFFFBF452FCB64D94C133" author="Shear, W. A." box="[312,463,844,870]" pageId="9" pageNumber="10" pagination="489 - 492" refId="ref5398" refString="Shear, W. A. (1972) The milliped genus Bidentogon (Diplopoda, Polydesmida, Trichopolydesmidae). Proceedings of the Biological Society of Washington, 85, 489 - 492." type="journal article" year="1972">Shear (1972)</bibRefCitation>
|
||
noted that the endomerite in the males from Sacramento is apically broad whereas that of the
|
||
<typeStatus id="54B106DA3A1CFFFBF49FFC8E4E7CC1DB" box="[501,551,884,910]" pageId="9" pageNumber="10">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF738FC8E4F38C1D8" authorityName="Buckett and Gardner" authorityYear="1968" box="[594,867,884,909]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="helferorum">
|
||
<emphasis id="B97E646A3A1CFFFBF738FC8E4F38C1D8" box="[594,867,884,909]" italics="true" pageId="9" pageNumber="10">Bidentogon helferorum</emphasis>
|
||
</taxonomicName>
|
||
, from
|
||
<quantity id="4CF2159D3A1CFFFBF6DCFC8E4FB6C1DB" box="[950,1005,884,910]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="9" pageNumber="10" unit="mi" value="1.0">1 mi</quantity>
|
||
(1.6 km) north of Mendocino, Mendocino County, is narrow and acuminate. I have examined most of the available males of
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF41CFC3E4DA6C188" authorityName="Buckett and Gardner" authorityYear="1968" box="[374,509,964,989]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A1CFFFBF41CFC3E4DA6C188" box="[374,509,964,989]" italics="true" pageId="9" pageNumber="10">Bidentogon</emphasis>
|
||
</taxonomicName>
|
||
and found that all those collected along the coast and San Francisco Bay, including Stanford, exhibit the latter condition; the only sample in which the tibiotarsus is broad is that from Sacramento. Consequently, as the true
|
||
<typeStatus id="54B106DA3A1CFFFBF14BFBEE4808C67B" box="[1057,1107,1044,1070]" pageId="9" pageNumber="10">type</typeStatus>
|
||
localities of both nominal species occur along the coast, its name must be changed to
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF12FFBC6488AC600" authorityName="Chamberlin" authorityYear="1918" box="[1093,1233,1084,1109]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="californicus">
|
||
<emphasis id="B97E646A3A1CFFFBF12FFBC6488AC600" box="[1093,1233,1084,1109]" italics="true" pageId="9" pageNumber="10">californicus</emphasis>
|
||
</taxonomicName>
|
||
, which holds priority over
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF49EFB9E4E2EC628" authorityName="Buckett and Gardner" authorityYear="1968" box="[500,629,1124,1149]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="helferorum">
|
||
<emphasis id="B97E646A3A1CFFFBF49EFB9E4E2EC628" box="[500,629,1124,1149]" italics="true" pageId="9" pageNumber="10">helferorum</emphasis>
|
||
</taxonomicName>
|
||
by 50 years. No name is available for that from Sacramento, so I propose
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF49DFB764EAAC6F0" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[503,753,1164,1189]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="expansus" status="sp. nov.">
|
||
<emphasis id="B97E646A3A1CFFFBF49DFB764EAAC6F0" box="[503,753,1164,1189]" italics="true" pageId="9" pageNumber="10">Bidentogon expansus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24DD9113A1CFFFBF793FB764F6CC6F3" box="[761,823,1164,1190]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
I have placed the
|
||
<typeStatus id="54B106DA3A1CFFFBF17CFB764813C6F3" box="[1046,1096,1164,1190]" pageId="9" pageNumber="10">type</typeStatus>
|
||
label for
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF1D0FB764D76C698" authorityName="Shelley" authorityYear="2003" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="expansus">
|
||
<emphasis id="B97E646A3A1CFFFBF1D0FB764D76C698" italics="true" pageId="9" pageNumber="10">B. expansus</emphasis>
|
||
</taxonomicName>
|
||
in this vial along with one stating that Stanford is the true
|
||
<typeStatus id="54B106DA3A1CFFFBF136FB4E48D5C69B" box="[1116,1166,1204,1230]" pageId="9" pageNumber="10">type</typeStatus>
|
||
locality for
|
||
<taxonomicName id="4C0AC3FB3A1CFFFBF462FB264E1DC6A0" authorityName="Chamberlin" authorityYear="1918" box="[264,582,1244,1269]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="californicus">
|
||
<emphasis id="B97E646A3A1CFFFBF462FB264E1DC6A0" box="[264,582,1244,1269]" italics="true" pageId="9" pageNumber="10">Brachydesmus californicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |