225 lines
26 KiB
XML
225 lines
26 KiB
XML
<document id="7653857B3DC9BE218C46CD4D53AEB8A9" ID-CLB-Dataset="5496" ID-DOI="10.11646/zootaxa.296.1.1" ID-GBIF-Dataset="3ddea826-544c-452c-994e-01c1a4973a4b" ID-ISSN="11755334" ID-Zenodo-Dep="5014425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624382618413" checkinUser="felipe" docAuthor="Shelley, Rowland M." docDate="2003" docId="03A3096E3A17FFF6F462FA3E4E25C1DB" docLanguage="en" docName="zt00296.pdf" docOrigin="Zootaxa 296 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.296.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Retrorsia Shelley 2003, new genus" docType="treatment" docVersion="6" lastPageNumber="5" masterDocId="FF9A71163A15FFF2F56AFFFA4C5BC255" masterDocTitle="A new polydesmid milliped genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett and Gardner, 1968, and a summary of the family in Western North America (Polydesmida: Polydesmidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1699020097571" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8A8B61D853A416676CFB120AF35349F7">A new polydesmid milliped genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett and Gardner, 1968, and a summary of the family in Western North America (Polydesmida: Polydesmidae)</mods:title>
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<mods:namePart id="A0EA2861AC6C80CF0FF9085F9454A587">Shelley, Rowland M.</mods:namePart>
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<mods:title id="5E695D93E5C2315E28FBB1CCA8D7E6C0">Zootaxa</mods:title>
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<mods:date id="28DCA29EC167B71E0988317F2572EABD">2003</mods:date>
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<mods:number id="5459D6FA5CF5D539544902E3A62FE384">2003-09-16</mods:number>
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<treatment id="03A3096E3A17FFF6F462FA3E4E25C1DB" ID-DOI="http://doi.org/10.5281/zenodo.5014401" ID-GBIF-Taxon="183531918" ID-Zenodo-Dep="5014401" LSID="urn:lsid:plazi:treatment:03A3096E3A17FFF6F462FA3E4E25C1DB" httpUri="http://treatment.plazi.org/id/03A3096E3A17FFF6F462FA3E4E25C1DB" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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<subSubSection id="C310EBF33A17FFF0F462FA3E4E52C78B" box="[264,521,1476,1502]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BB5B8783A17FFF0F462FA3E4E52C78B" blockId="2.[264,521,1476,1502]" box="[264,521,1476,1502]" pageId="2" pageNumber="3">
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<heading id="D0FD0F143A17FFF0F462FA3E4E52C78B" bold="true" box="[264,521,1476,1502]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<taxonomicName id="4C0AC3FB3A17FFF0F462FA3E4D23C78B" ID-CoL="7PXJN" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[264,376,1476,1502]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B97E646A3A17FFF0F462FA3E4D23C78B" bold="true" box="[264,376,1476,1502]" italics="true" pageId="2" pageNumber="3">Retrorsia</emphasis>
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</taxonomicName>
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,
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<emphasis id="B97E646A3A17FFF0F4EDFA3E4E52C78B" bold="true" box="[391,521,1476,1502]" pageId="2" pageNumber="3">
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<taxonomicNameLabel id="A24DD9113A17FFF0F4EDFA3E4E52C78B" box="[391,521,1476,1502]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C310EBF33A17FFF1F462F9EF4F62C18B" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8BB5B8783A17FFF0F462F9EF4F7FC47B" blockId="2.[264,804,1556,1582]" box="[264,804,1556,1582]" pageId="2" pageNumber="3">
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<emphasis id="B97E646A3A17FFF0F462F9EF4DC1C47B" box="[264,410,1557,1582]" italics="true" pageId="2" pageNumber="3">
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<typeStatus id="54B106DA3A17FFF0F462F9EF4D64C47B" box="[264,319,1557,1582]" pageId="2" pageNumber="3">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="4C0AC3FB3A17FFF0F4C0F9EF4EDBC47B" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[426,640,1557,1582]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="leonardi" status="sp. nov.">
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<emphasis id="B97E646A3A17FFF0F4C0F9EF4EDBC47B" box="[426,640,1557,1582]" italics="true" pageId="2" pageNumber="3">Retrorsia leonardi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24DD9113A17FFF0F7E5F9EE4F7BC47B" box="[655,800,1556,1582]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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.
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</paragraph>
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<paragraph id="8BB5B8783A17FFF1F462F9984DB2C073" blockId="2.[264,1323,1634,1900]" lastBlockId="3.[264,1323,284,990]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<emphasis id="B97E646A3A17FFF0F462F9984DDDC42E" box="[264,390,1634,1659]" italics="true" pageId="2" pageNumber="3">Diagnosis.</emphasis>
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A genus of minute, pallid
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<taxonomicName id="4C0AC3FB3A17FFF0F7A0F9984F35C429" authorityName="Leach" authorityYear="1815" box="[714,878,1634,1660]" class="Diplopoda" family="Polydesmidae" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Polydesmidae</taxonomicName>
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with 19 segments including epiproct, adult body dimensions ca. 5.6 mm in length and 0.7 mm in width; body noticeably broader with wider metaterga from segments 10–14. 6th antennomere distinctly swollen. Collum slightly narrower than succeeding tergites, not covering epicranium, setae scattered subuniformly across surface. Dorsum slightly convex, metaterga lightly papillate with dense coverings of parallelsided setae, latter arranged in four rows on segments 2–9, five rows on segments 10–17, and four rows again on segments 18–19; paranota present on all segments, margins variably dentate. Sterna without modifications. Legs with prefemora noticeably swollen through segment 10, becoming progressively narrower caudad; tarsal claws inconspicuous. Gonopodal telopodite relatively long, curving ventrad for most of length, divided distad into tibiotarsus and endomerite, with variable expansion proximal to division point and rounded or oblong pulvillus proximal to expansion; tibiotarsus curving variably dorsad, countering axis of stem; endomerite located laterad, length and configuration variable.
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</paragraph>
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<paragraph id="8BB5B8783A16FFF1F452FDCE4D9AC023" blockId="3.[264,1323,284,990]" pageId="3" pageNumber="4">
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<emphasis id="B97E646A3A16FFF1F452FDCE4DCBC018" box="[312,400,564,589]" italics="true" pageId="3" pageNumber="4">Species</emphasis>
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. Two are known,
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<taxonomicName id="4C0AC3FB3A16FFF1F704FDCE4EA9C018" authorityName="Shelley" authorityYear="2003" box="[622,754,564,589]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="leonardi">
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<emphasis id="B97E646A3A16FFF1F704FDCE4EA9C018" box="[622,754,564,589]" italics="true" pageId="3" pageNumber="4">R. leonardi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0AC3FB3A16FFF1F647FDCE4F96C018" authorityName="Shelley" authorityYear="2003" box="[813,973,564,589]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="benedictae">
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<emphasis id="B97E646A3A16FFF1F647FDCE4F96C018" box="[813,973,564,589]" italics="true" pageId="3" pageNumber="4">R. benedictae</emphasis>
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</taxonomicName>
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, n. spp., but others may exist in nearby areas.
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</paragraph>
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<paragraph id="8BB5B8783A16FFF1F452FD7E4E9CC093" blockId="3.[264,1323,284,990]" pageId="3" pageNumber="4">
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<emphasis id="B97E646A3A16FFF1F452FD7E4D9CC0C8" box="[312,455,644,669]" italics="true" pageId="3" pageNumber="4">Distribution</emphasis>
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. Western
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<collectingRegion id="49CE769A3A16FFF1F726FD7E4EFFC0CB" box="[588,676,644,670]" country="United States of America" name="Oregon" pageId="3" pageNumber="4">Oregon</collectingRegion>
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and
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<collectingRegion id="49CE769A3A16FFF1F789FD7E4F2AC0CB" box="[739,881,644,670]" country="United States of America" name="Washington" pageId="3" pageNumber="4">Washington</collectingRegion>
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,
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<collectingCountry id="F31DF8E83A16FFF1F615FD7E4FE6C0CB" box="[895,957,644,670]" name="United States of America" pageId="3" pageNumber="4">USA</collectingCountry>
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, most sites being in the lower Columbia River Valley (
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<figureCitation id="1331A4FD3A16FFF1F74CFD564E2FC093" box="[550,628,684,710]" captionStart="FIGURE 1" captionStartId="2.[264,368,1327,1351]" captionTargetBox="[470,1120,597,1316]" captionTargetId="figure-214@2.[464,1126,597,1317]" captionTargetPageId="2" captionText="FIGURE 1. Occurrence of the Polydesmidae in western North America; 1, Scytonotus; 2, Calianotus; 3, Utadesmus; 4, Speodesmus; 5, Bidentogon; 6, Retrorsia." figureDoi="http://doi.org/10.5281/zenodo.5014429" httpUri="https://zenodo.org/record/5014429/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
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[6], 2).
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</paragraph>
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<paragraph id="8BB5B8783A16FFF1F452FD2E4E78C143" blockId="3.[264,1323,284,990]" pageId="3" pageNumber="4">
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<emphasis id="B97E646A3A16FFF1F452FD2E4DE8C0B8" box="[312,435,724,749]" italics="true" pageId="3" pageNumber="4">Etymology</emphasis>
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. The generic name denotes the retrorse curvature of the tibiotarsus, the most distinctive feature.
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</paragraph>
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<paragraph id="8BB5B8783A16FFF1F452FCDE4F62C18B" blockId="3.[264,1323,284,990]" pageId="3" pageNumber="4">
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<emphasis id="B97E646A3A16FFF1F452FCDE4DC5C168" box="[312,414,804,829]" italics="true" pageId="3" pageNumber="4">Remarks</emphasis>
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. The species of
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<taxonomicName id="4C0AC3FB3A16FFF1F71FFCDE4EB8C168" authorityName="Shelley" authorityYear="2003" box="[629,739,804,829]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B97E646A3A16FFF1F71FFCDE4EB8C168" box="[629,739,804,829]" italics="true" pageId="3" pageNumber="4">Retrorsia</emphasis>
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</taxonomicName>
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are the smallest known polydesmids in western North America, smaller than even the species of
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<taxonomicName id="4C0AC3FB3A16FFF1F60FFCB64FB7C130" authorityName="Buckett and Gardner" authorityYear="1968" box="[869,1004,844,869]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B97E646A3A16FFF1F60FFCB64FB7C130" box="[869,1004,844,869]" italics="true" pageId="3" pageNumber="4">Bidentogon</emphasis>
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</taxonomicName>
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, which are over
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<quantity id="4CF2159D3A16FFF1F1ABFCB64952C133" box="[1217,1289,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="4" unit="mm" value="6.0">6 mm</quantity>
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in length (
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<bibRefCitation id="EF9BC5893A16FFF1F408FC8E4EF2C1DB" author="Buckett, J. S. & Gardner, M. R." box="[354,681,884,910]" pageId="3" pageNumber="4" pagination="198 - 202" refId="ref5008" refString="Buckett, J. S. & Gardner, M. R. (1968) A new genus and species of milliped from northern California (Polydesmida: Vanhoeffeniidae). Pan-Pacific Entomologist, 44, 198 - 202." type="journal article" year="1968">Buckett and Gardner, 1968</bibRefCitation>
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). They are thus comparable in size to representatives of the
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<taxonomicName id="4C0AC3FB3A16FFF1F432FC664E57C1E3" authorityName="Silvestri" authorityYear="1896" box="[344,524,924,950]" class="Diplopoda" family="Pyrgodesmidae" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Pyrgodesmidae</taxonomicName>
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and
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<taxonomicName id="4C0AC3FB3A16FFF1F720FC664F18C1E3" authorityName="Brolemann" authorityYear="1916" box="[586,835,924,950]" class="Diplopoda" family="Fuhrmannodesmidae" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Fuhrmannodesmidae</taxonomicName>
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, and their gonopods are only microns in length, mere specks in a watchglass of alcohol.
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</paragraph>
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</subSubSection>
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<caption id="DF75E8F03A16FFF1F462F8F04D9EC51C" ID-DOI="http://doi.org/10.5281/zenodo.5014433" ID-Zenodo-Dep="5014433" httpUri="https://zenodo.org/record/5014433/files/figure.png" pageId="3" pageNumber="4" startId="3.[264,367,1802,1826]" targetBox="[521,1074,1077,1791]" targetPageId="3">
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<paragraph id="8BB5B8783A16FFF1F462F8F04D9EC51C" blockId="3.[264,1322,1802,1865]" pageId="3" pageNumber="4">
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<emphasis id="B97E646A3A16FFF1F462F8F04DDDC577" bold="true" box="[264,390,1802,1826]" pageId="3" pageNumber="4">FIGURE 2</emphasis>
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. Distribution of
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<emphasis id="B97E646A3A16FFF1F723F8F14EEDC577" box="[585,694,1803,1826]" italics="true" pageId="3" pageNumber="4">
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<taxonomicName id="4C0AC3FB3A16FFF1F723F8F14EF4C577" authorityName="Shelley" authorityYear="2003" box="[585,687,1803,1826]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Retrorsia</taxonomicName>
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;
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</emphasis>
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dots,
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<taxonomicName id="4C0AC3FB3A16FFF1F791F8F14F2FC577" authorityName="Shelley" authorityYear="2003" box="[763,884,1803,1826]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="leonardi">
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<emphasis id="B97E646A3A16FFF1F791F8F14F2FC577" box="[763,884,1803,1826]" italics="true" pageId="3" pageNumber="4">R. leonardi</emphasis>
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</taxonomicName>
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; triangles,
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<taxonomicName id="4C0AC3FB3A16FFF1F69BF8F148DEC577" authorityName="Shelley" authorityYear="2003" box="[1009,1157,1803,1826]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="benedictae">
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<emphasis id="B97E646A3A16FFF1F69BF8F148DEC577" box="[1009,1157,1803,1826]" italics="true" pageId="3" pageNumber="4">R. benedictae</emphasis>
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</taxonomicName>
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. OR, Oregon; WA, Washington.
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</paragraph>
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</caption>
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<subSubSection id="C310EBF33A11FFF6F452FEE64E25C1DB" pageId="4" pageNumber="5" type="reference_group">
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<paragraph id="8BB5B8783A11FFF6F452FEE64E25C1DB" blockId="4.[264,1323,284,910]" pageId="4" pageNumber="5">
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<taxonomicName id="4C0AC3FB3A11FFF6F452FEE64DFDC360" ID-CoL="7PXJN" authorityName="Shelley" authorityYear="2003" box="[312,422,284,309]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B97E646A3A11FFF6F452FEE64DFDC360" box="[312,422,284,309]" italics="true" pageId="4" pageNumber="5">Retrorsia</emphasis>
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</taxonomicName>
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is related to
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<taxonomicName id="4C0AC3FB3A11FFF6F750FEE64EE7C360" ID-CoL="7Q5ZD" authorityName="Chamberlin and Hoffman" authorityYear="1950" box="[570,700,284,309]" class="Diplopoda" family="Polydesmidae" genus="Utadesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B97E646A3A11FFF6F750FEE64EE7C360" box="[570,700,284,309]" italics="true" pageId="4" pageNumber="5">Utadesmus</emphasis>
|
||
</taxonomicName>
|
||
, whose closest known locality is in the Henry Mountains, Garfield County,
|
||
<collectingRegion id="49CE769A3A11FFF6F74FFEBE4E05C30B" box="[549,606,324,350]" country="United States of America" name="Utah" pageId="4" pageNumber="5">Utah</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EF9BC5893A11FFF6F71BFEBE4F42C30B" author="Shelley, R. M." box="[625,793,324,350]" pageId="4" pageNumber="5" pagination="9 - 16" refId="ref5641" refString="Shelley, R. M. (1996) The milliped genus Utadesmus Chamberlin and Hoffman (Polydesmida: Polydesmidae). Myriapodologica, 4, 9 - 16." type="journal article" year="1996">Shelley, 1996</bibRefCitation>
|
||
), some
|
||
<quantity id="4CF2159D3A11FFF6F611FEBE4F8AC30B" box="[891,977,324,350]" metricMagnitude="6" metricUnit="m" metricValue="1.2391948799999999" pageId="4" pageNumber="5" unit="mi" value="770.0">770 mi</quantity>
|
||
(
|
||
<quantity id="4CF2159D3A11FFF6F68EFEBE4802C30B" box="[996,1113,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.232" pageId="4" pageNumber="5" unit="km" value="1.232">1,232 km</quantity>
|
||
) to the southeast (
|
||
<figureCitation id="1331A4FD3A11FFF6F47BFE964D0FC3D3" box="[273,340,364,390]" captionStart="FIGURE 1" captionStartId="2.[264,368,1327,1351]" captionTargetBox="[470,1120,597,1316]" captionTargetId="figure-214@2.[464,1126,597,1317]" captionTargetPageId="2" captionText="FIGURE 1. Occurrence of the Polydesmidae in western North America; 1, Scytonotus; 2, Calianotus; 3, Utadesmus; 4, Speodesmus; 5, Bidentogon; 6, Retrorsia." figureDoi="http://doi.org/10.5281/zenodo.5014429" httpUri="https://zenodo.org/record/5014429/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
|
||
[3]). The telopodal division point is distal in both genera, and the endomerites arise there along with long, terminal tibiotarsi. This configuration contrasts with that exhibited by
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F444FE464DF6C380" ID-CoL="7PZ9Y" authorityName="C. L. Koch" authorityYear="1847" box="[302,429,444,469]" class="Diplopoda" family="Polydesmidae" genus="Scytonotus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A11FFF6F444FE464DF6C380" box="[302,429,444,469]" italics="true" pageId="4" pageNumber="5">Scytonotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F48DFE464E33C380" ID-CoL="7NRK8" authorityName="Shelley" authorityYear="1997" box="[487,616,444,469]" class="Diplopoda" family="Polydesmidae" genus="Calianotus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A11FFF6F48DFE464E33C380" box="[487,616,444,469]" italics="true" pageId="4" pageNumber="5">Calianotus</emphasis>
|
||
</taxonomicName>
|
||
, in which the endomerites arise basally and are subequal in lengths to the tibiotarsi (
|
||
<bibRefCitation id="EF9BC5893A11FFF6F749FE1E4E9DC3AB" author="Shelley, R. M." box="[547,710,484,510]" pageId="4" pageNumber="5" pagination="1 - 60" refId="ref5611" refString="Shelley, R. M. (1993) Revision of the milliped genus Scytonotus Koch (Polydesmida: Polydes-midae). Brimleyana, 19, 1 - 60." type="journal article" year="1993">Shelley, 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9BC5893A11FFF6F7B8FE1E4F4AC3AB" author="Shelley, R. M." box="[722,785,484,510]" pageId="4" pageNumber="5" pagination="59 - 67" refId="ref5671" refString="Shelley, R. M. (1997) The identity of Polydesmus sastianus Chamberlin, proposal of a new milliped genus, and remarks on the identity of Phreatodesmus hastingsus (Chamberlin) (Polydesmida: Polydesmidae). Myriapodologica, 4, 59 - 67." type="journal article" year="1997">1997</bibRefCitation>
|
||
). The tibiotarsus is coaxial with the telopodal stem in
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F40FFDF64DBCC070" ID-CoL="7Q5ZD" authorityName="Chamberlin and Hoffman" authorityYear="1950" box="[357,487,524,549]" class="Diplopoda" family="Polydesmidae" genus="Utadesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A11FFF6F40FFDF64DBCC070" box="[357,487,524,549]" italics="true" pageId="4" pageNumber="5">Utadesmus</emphasis>
|
||
</taxonomicName>
|
||
, whereas it curves dorsad and counters the general curvature in
|
||
<emphasis id="B97E646A3A11FFF6F1BFFDF64D6AC018" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F1BFFDF64D76C018" ID-CoL="7PXJN" authorityName="Shelley" authorityYear="2003" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Retrorsia</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The pulvillus in
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F761FDCE4ED6C018" ID-CoL="7Q5ZD" authorityName="Chamberlin and Hoffman" authorityYear="1950" box="[523,653,564,589]" class="Diplopoda" family="Polydesmidae" genus="Utadesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A11FFF6F761FDCE4ED6C018" box="[523,653,564,589]" italics="true" pageId="4" pageNumber="5">Utadesmus</emphasis>
|
||
</taxonomicName>
|
||
is a cupulate, perpendicular projection located distad at the bases of the endomerite and tibiotarsus; it is located near 1/3 length in
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F112FDA648BEC020" ID-CoL="7PXJN" authorityName="Shelley" authorityYear="2003" box="[1144,1253,604,629]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A11FFF6F112FDA648BEC020" box="[1144,1253,604,629]" italics="true" pageId="4" pageNumber="5">Retrorsia</emphasis>
|
||
</taxonomicName>
|
||
. The oblong structure in
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F487FD7E4ED7C0C8" ID-CoL="6X47F" authorityName="Shelley" authorityYear="2003" box="[493,652,644,669]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="benedictae">
|
||
<emphasis id="B97E646A3A11FFF6F487FD7E4ED7C0C8" box="[493,652,644,669]" italics="true" pageId="4" pageNumber="5">R. benedictae</emphasis>
|
||
</taxonomicName>
|
||
is intermediate between the condition in
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F11DFD7E48A2C0C8" ID-CoL="7Q5ZD" authorityName="Chamberlin and Hoffman" authorityYear="1950" box="[1143,1273,644,669]" class="Diplopoda" family="Polydesmidae" genus="Utadesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A11FFF6F11DFD7E48A2C0C8" box="[1143,1273,644,669]" italics="true" pageId="4" pageNumber="5">Utadesmus</emphasis>
|
||
</taxonomicName>
|
||
and the small, rounded structure in
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F714FD564F58C090" ID-CoL="78RVF" authorityName="Shelley" authorityYear="2003" box="[638,771,684,709]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="leonardi">
|
||
<emphasis id="B97E646A3A11FFF6F714FD564F58C090" box="[638,771,684,709]" italics="true" pageId="4" pageNumber="5">R. leonardi</emphasis>
|
||
</taxonomicName>
|
||
. This affinity between
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F14EFD5648CAC090" ID-CoL="7PXJN" authorityName="Shelley" authorityYear="2003" box="[1060,1169,684,709]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A11FFF6F14EFD5648CAC090" box="[1060,1169,684,709]" italics="true" pageId="4" pageNumber="5">Retrorsia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0AC3FB3A11FFF6F1A4FD564D6CC0B8" ID-CoL="7Q5ZD" authorityName="Chamberlin and Hoffman" authorityYear="1950" class="Diplopoda" family="Polydesmidae" genus="Utadesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97E646A3A11FFF6F1A4FD564D6CC0B8" italics="true" pageId="4" pageNumber="5">Utadesmus</emphasis>
|
||
</taxonomicName>
|
||
suggests that isolated populations of related polydesmids may inhabit intervening areas in the Great Basin Physiographic Province, particularly the Blue Mountains in eastern
|
||
<collectingRegion id="49CE769A3A11FFF6F45EFCDE4DD4C16B" box="[308,399,804,830]" country="United States of America" name="Oregon" pageId="4" pageNumber="5">Oregon</collectingRegion>
|
||
, the Ruby and Shell Creek Mountains and Wheeler Peak (Great Basin National Park),
|
||
<collectingRegion id="49CE769A3A11FFF6F43FFCB64DE9C133" box="[341,434,844,870]" country="United States of America" name="Nevada" pageId="4" pageNumber="5">Nevada</collectingRegion>
|
||
, and the Deep Creek Mountains in western
|
||
<collectingRegion id="49CE769A3A11FFF6F6C6FCB64FB3C133" box="[940,1000,844,870]" country="United States of America" name="Utah" pageId="4" pageNumber="5">Utah</collectingRegion>
|
||
, which are forested and relatively moist at high elevations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |