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<document id="AD2C14524EB043D6087658153DC4FC38" ID-CLB-Dataset="54139" ID-DOI="10.11646/zootaxa.5243.1.1" ID-GBIF-Dataset="1476f0e1-3886-401f-bf56-c294a0a99962" ID-ISSN="1175-5326" ID-Zenodo-Dep="7642400" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676448183279" checkinUser="plazi" docAuthor="Tian, Mingyi, Huang, Sunbin &amp; Jia, Xinyang" docDate="2023" docId="039587ADBF10FF85FF7F478F230CF9D8" docLanguage="en" docName="zootaxa.5243.1.1.pdf" docOrigin="Zootaxa 5243 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5243.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dongodytes (Dongodytodes) pingjingae Tian &amp; Huang &amp; Jia 2023, sp. n." docType="treatment" docVersion="5" lastPageNumber="45" masterDocId="FFACFFD5BF38FFA9FFE841652260FFF5" masterDocTitle="A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="41" updateTime="1699475196432" updateUser="ExternalLinkService">
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<mods:title id="91459E720C05525D41F5A3B728B01188">A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini)</mods:title>
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<mods:namePart id="51D973E00383B5E944A6CC8D547D4A28">Tian, Mingyi</mods:namePart>
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<mods:namePart id="1AC7B2CDFB1D793C7D37E7DADD5A9BD7">Huang, Sunbin</mods:namePart>
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<mods:namePart id="4C30C3718C28AAE6690719594CD129F1">Jia, Xinyang</mods:namePart>
<mods:nameIdentifier id="8C72BBF8C8610662DFB33F630D867BAD" type="ORCID">0000-0003-1936-5926</mods:nameIdentifier>
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<mods:title id="4B029F86E930F3BB524F2C719D9CCEEC">Zootaxa</mods:title>
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<mods:date id="24DEAE23AE926BEB6C596A0CF3671891">2023</mods:date>
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<mods:number id="37F65BC348BEB08F6080FA22B330DEFF">2023-02-15</mods:number>
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<mods:number id="3EBEB47DE7DFC08C42B72B261F68C794">5243</mods:number>
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<mods:classification id="4A0A4C05554C0B3C8D261D2F34C74F7D">journal article</mods:classification>
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<treatment id="039587ADBF10FF85FF7F478F230CF9D8" ID-DOI="http://doi.org/10.5281/zenodo.7647042" ID-GBIF-Taxon="209296920" ID-Zenodo-Dep="7647042" LSID="urn:lsid:plazi:treatment:039587ADBF10FF85FF7F478F230CF9D8" httpUri="http://treatment.plazi.org/id/039587ADBF10FF85FF7F478F230CF9D8" lastPageId="44" lastPageNumber="45" pageId="40" pageNumber="41">
<subSubSection id="C3266530BF10FF81FF7F478F20DDF8F0" box="[151,701,1770,1797]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8B8336BBBF10FF81FF7F478F20DDF8F0" blockId="40.[151,701,1770,1833]" box="[151,701,1770,1797]" pageId="40" pageNumber="41">
<heading id="D0CB81D7BF10FF81FF7F478F20DDF8F0" bold="true" box="[151,701,1770,1797]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="4">
<emphasis id="B948EAA9BF10FF81FF7F478F20DDF8F0" bold="true" box="[151,701,1770,1797]" pageId="40" pageNumber="41">
<taxonomicName id="4C3C4D38BF10FF81FF7F478F2013F8F0" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[151,627,1770,1797]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pingjingae" status="sp. nov." subGenus="Dongodytodes">
<emphasis id="B948EAA9BF10FF81FF7F478F2013F8F0" bold="true" box="[151,627,1770,1797]" italics="true" pageId="40" pageNumber="41">Dongodytes (Dongodytodes) pingjingae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57D2BF10FF81FD92478E20DDF8F0" box="[634,701,1771,1797]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF10FF81FF7F466A23F9F8DC" box="[151,409,1807,1833]" pageId="40" pageNumber="41" type="description">
<paragraph id="8B8336BBBF10FF81FF7F466A23F9F8DC" blockId="40.[151,701,1770,1833]" box="[151,409,1807,1833]" pageId="40" pageNumber="41">
<figureCitation id="13072A3EBF10FF81FF7F466A2369F8DC" box="[151,265,1807,1833]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="40" pageNumber="41">Figures 1i</figureCitation>
,
<figureCitation id="13072A3EBF10FF81FEFE466A2354F8DC" box="[278,308,1807,1833]" captionStart="FIGURE 26" captionStartId="41.[151,250,1942,1966]" captionTargetId="figure-20@41.[330,1256,181,1919]" captionTargetPageId="41" captionText="FIGURE 26. Habitus of Dongodytes (Dongodytodes) pingjingae sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7647032" httpUri="https://zenodo.org/record/7647032/files/figure.png" pageId="40" pageNumber="41">26</figureCitation>
,
<figureCitation id="13072A3EBF10FF81FEA8466A2311F8DC" box="[320,369,1807,1833]" captionStart="FIGURE 27" captionStartId="42.[151,250,1965,1989]" captionTargetBox="[280,1306,453,1927]" captionTargetId="figure-129@42.[280,1307,453,1942]" captionTargetPageId="42" captionText="FIGURE 27. Left elytra (white dots indicating the chaetotaxy) of Dongodytes species (white dots indicating the chaetotaxy). A. Dongodytes (Dongodytodes) pingjingae sp. n.; B. Dongodytes (s. str.) mingruoae n. sp." figureDoi="http://doi.org/10.5281/zenodo.7642446" httpUri="https://zenodo.org/record/7642446/files/figure.png" pageId="40" pageNumber="41">27B</figureCitation>
,
<figureCitation id="13072A3EBF10FF81FE95466A23F9F8DC" box="[381,409,1807,1833]" captionStart="FIGURE 28" captionStartId="43.[151,250,1717,1741]" captionTargetBox="[163,1401,220,1692]" captionTargetId="figure-20@43.[163,1424,181,1693]" captionTargetPageId="43" captionText="FIGURE 28. Dongodytes (Dongodytodes) pingjingae sp. n. A. median lobe, lateral view; B. apical lobe, dorsal view; C. an active individual in cave." figureDoi="http://doi.org/10.5281/zenodo.7642448" httpUri="https://zenodo.org/record/7642448/files/figure.png" pageId="40" pageNumber="41">28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF10FF81FF7F463321ADF84C" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8B8336BBBF10FF81FF7F463321ADF84C" blockId="40.[151,1437,1878,1977]" pageId="40" pageNumber="41">
<emphasis id="B948EAA9BF10FF81FF7F46332366F885" bold="true" box="[151,262,1878,1904]" pageId="40" pageNumber="41">Material.</emphasis>
<materialsCitation id="3B543CE6BF10FF81FEFA463223E9F84C" ID-GBIF-Occurrence="4030741311" collectingDate="2016-07-09" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Pingjing Yang &amp; Mingruo Tang &amp; Jingli Cheng" county="Dahua Yao Autonomous County" elevation="240" latitude="23.9" location="Guhe" longLatPrecision="752" longitude="107.74" municipality="Xiannu Dong" pageId="40" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi Zhuang Autonomous Region" typeStatus="holotype">
<typeStatus id="54878819BF10FF81FEFA4632231AF884" box="[274,378,1879,1905]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3AFD32BF10FF81FE6D463223DEF884" box="[389,446,1879,1905]" pageId="40" pageNumber="41" type="male">male</specimenCount>
,
<collectingMunicipality id="6BE7ACC1BF10FF81FE264632200DF884" box="[462,621,1879,1905]" pageId="40" pageNumber="41">Xiannü Dong</collectingMunicipality>
,
<location id="8EE36060BF10FF81FD95463220DCF884" LSID="urn:lsid:plazi:treatment:039587ADBF10FF85FF7F478F230CF9D8:8EE36060BF10FF81FD95463220DCF884" box="[637,700,1879,1905]" county="Dahua Yao Autonomous County" latitude="23.9" longLatPrecision="752" longitude="107.74" municipality="Xiannu Dong" name="Guhe" pageId="40" pageNumber="41" stateProvince="Guangxi Zhuang Autonomous Region">Guhe</location>
,
<collectingCounty id="62E24E37BF10FF81FD2446322626F884" box="[716,1094,1879,1905]" pageId="40" pageNumber="41">Dahua Yao Autonomous County</collectingCounty>
,
<location id="8EE36060BF10FF81FBBD463226ABF884" LSID="urn:lsid:plazi:treatment:039587ADBF10FF85FF7F478F230CF9D8:8EE36060BF10FF81FBBD463226ABF884" box="[1109,1227,1879,1905]" county="Dahua Yao Autonomous County" latitude="23.9" longLatPrecision="752" longitude="107.74" municipality="Xiannu Dong" name="Hechi Shi" pageId="40" pageNumber="41" stateProvince="Guangxi Zhuang Autonomous Region">Hechi Shi</location>
,
<collectingRegion id="49F8F859BF10FF81FB33463223E2F860" pageId="40" pageNumber="41">Guangxi Zhuang Autonomous Region</collectingRegion>
,
<geoCoordinate id="EE08507CBF10FF81FE64461E2389F860" box="[396,489,1915,1941]" degrees="23.90" direction="north" orientation="latitude" pageId="40" pageNumber="41" precision="555" value="23.9">23.90 N</geoCoordinate>
,
<geoCoordinate id="EE08507CBF10FF81FE1B461E2005F860" box="[499,613,1915,1941]" degrees="107.74" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="555" value="107.74">107.74° E</geoCoordinate>
,
<quantity id="4CC49B5EBF10FF81FD98461E20D9F860" box="[624,697,1915,1941]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="40" pageNumber="41" unit="m" value="240.0">
<elevation id="0011D188BF10FF81FD98461E20D9F860" box="[624,697,1915,1941]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="40" pageNumber="41" unit="m" value="240.0">240 m</elevation>
</quantity>
,
<date id="FF82107BBF10FF81FD2C461E212EF860" box="[708,846,1915,1941]" pageId="40" pageNumber="41" value="2016-07-09">
<collectingDate id="EFC6E993BF10FF81FD2C461E212EF860" box="[708,846,1915,1941]" pageId="40" pageNumber="41" value="2016-07-09">VII. 9. 2016</collectingDate>
</date>
,
<collectorName id="26C9536DBF10FF81FCB1461E2186F860" box="[857,998,1915,1941]" pageId="40" pageNumber="41">Mingyi Tian</collectorName>
,
<collectorName id="26C9536DBF10FF81FC19461E26F2F860" box="[1009,1170,1915,1941]" pageId="40" pageNumber="41">Pingjing Yang</collectorName>
,
<collectorName id="26C9536DBF10FF81FB74461E275CF860" box="[1180,1340,1915,1941]" pageId="40" pageNumber="41">Mingruo Tang</collectorName>
&amp;
<collectorName id="26C9536DBF10FF81FAB6461E2280F84C" pageId="40" pageNumber="41">Jingli Cheng</collectorName>
leg., in
<collectionCode id="ED2DAE7EBF10FF81FED346FA23E9F84C" box="[315,393,1951,1977]" collectionName="SCAU" pageId="40" pageNumber="41">SCAU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B543CE6BF10FF81FE7C46FA2132F84C" ID-GBIF-Occurrence="4030741313" box="[404,850,1951,1977]" collectionCode="IBID" pageId="40" pageNumber="41" specimenCount="12" specimenCount-female="9" specimenCount-male="3" stateProvince="Guangxi" typeStatus="paratype">
<typeStatus id="54878819BF10FF81FE7C46FA2065F84C" box="[404,517,1951,1977]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3AFD32BF10FF81FDF846FA2006F84C" box="[528,614,1951,1977]" pageId="40" pageNumber="41" type="male">3 males</specimenCount>
and
<specimenCount id="9D3AFD32BF10FF81FD7446FA216BF84C" box="[668,779,1951,1977]" pageId="40" pageNumber="41" type="female">9 females</specimenCount>
,
<collectionCode id="ED2DAE7EBF10FF81FCFE46FA2132F84C" box="[790,850,1951,1977]" pageId="40" pageNumber="41">IBID</collectionCode>
</materialsCitation>
, in
<collectionCode id="ED2DAE7EBF10FF81FC9346FA21A9F84C" box="[891,969,1951,1977]" pageId="40" pageNumber="41">SCAU</collectionCode>
.
</paragraph>
</subSubSection>
<caption id="DF436633BF11FF80FF7F46F32676F85A" ID-DOI="http://doi.org/10.5281/zenodo.7647032" ID-Zenodo-Dep="7647032" box="[151,1046,1942,1967]" httpUri="https://zenodo.org/record/7647032/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,1942,1966]" targetBox="[342,1240,193,1914]" targetPageId="41">
<paragraph id="8B8336BBBF11FF80FF7F46F32676F85A" blockId="41.[151,1046,1942,1967]" box="[151,1046,1942,1967]" pageId="41" pageNumber="42">
<emphasis id="B948EAA9BF11FF80FF7F46F32340F85B" bold="true" box="[151,288,1942,1966]" pageId="41" pageNumber="42">FIGURE 26.</emphasis>
Habitus of
<taxonomicName id="4C3C4D38BF11FF80FE7F46F2214CF85B" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[407,812,1943,1967]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="pingjingae" status="sp. nov." subGenus="Dongodytodes">
<emphasis id="B948EAA9BF11FF80FE7F46F2214CF85B" box="[407,812,1943,1967]" italics="true" pageId="41" pageNumber="42">Dongodytes (Dongodytodes) pingjingae</emphasis>
</taxonomicName>
<emphasis id="B948EAA9BF11FF80FCDA46F2210BF85A" bold="true" box="[818,875,1943,1967]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A27B57D2BF11FF80FCDA46F2210BF85A" box="[818,875,1943,1967]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male, holotype.
</paragraph>
</caption>
<subSubSection id="C3266530BF12FF83FF2F41F321C2FF20" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8B8336BBBF12FF83FF2F41F321C2FF20" blockId="42.[151,1437,150,429]" pageId="42" pageNumber="43">
<emphasis id="B948EAA9BF12FF83FF2F41F32322FF45" bold="true" box="[199,322,150,176]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
Medium-sized for
<taxonomicName id="4C3C4D38BF12FF83FDFF41F220D9FF44" authorityName="Tian" authorityYear="2011" box="[535,697,151,177]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="45" phylum="Arthropoda" rank="subGenus" subGenus="Dongodytodes">
<emphasis id="B948EAA9BF12FF83FDFF41F220D9FF44" box="[535,697,151,177]" italics="true" pageId="42" pageNumber="43">Dongodytodes</emphasis>
</taxonomicName>
species, whole surface pubescent, head hardly expanded, fore part (head including mandibles plus prothorax) slightly longer than elytra.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF12FF85FF2F41BB2604FB88" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43" type="description">
<paragraph id="8B8336BBBF12FF83FF2F41BB2623FF0C" blockId="42.[151,1437,150,429]" box="[199,1091,222,249]" pageId="42" pageNumber="43">
<emphasis id="B948EAA9BF12FF83FF2F41BB2339FF0D" bold="true" box="[199,345,222,248]" pageId="42" pageNumber="43">Description.</emphasis>
Length:
<quantity id="4CC49B5EBF12FF83FE5641BA2028FF0C" box="[446,584,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="7.9" metricValueMin="7.7" pageId="42" pageNumber="43" unit="mm" value="7.8" valueMax="7.9" valueMin="7.7">7.7–7.9 mm</quantity>
; width:
<quantity id="4CC49B5EBF12FF83FD4A41BA214CFF0C" box="[674,812,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="42" pageNumber="43" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
. Habitus as in
<figureCitation id="13072A3EBF12FF83FC2741BA265FFF0C" box="[975,1087,223,249]" captionStart="FIGURE 26" captionStartId="41.[151,250,1942,1966]" captionTargetId="figure-20@41.[330,1256,181,1919]" captionTargetPageId="41" captionText="FIGURE 26. Habitus of Dongodytes (Dongodytodes) pingjingae sp. n., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7647032" httpUri="https://zenodo.org/record/7647032/files/figure.png" pageId="42" pageNumber="43">Figure 26</figureCitation>
.
</paragraph>
<paragraph id="8B8336BBBF12FF83FF2F4066204EFE58" blockId="42.[151,1437,150,429]" pageId="42" pageNumber="43">Brown, but antennae, mouthparts palps and tarsi pale. Microsculptural engraved meshes moderately or transverse on head, strongly transverse on pronotum, invisible on elytra. Body wholly pubescent, sparser on head and pronotum, denser on elytra; underside of head and thorax sparsely covered with longer hairs; ventrites densely and shortly pubescent, propleura glabrous but with a few short setae anteriorly. Head and pronotum smooth, elytra vaguely punctate. Moderately shiny.</paragraph>
<caption id="DF436633BF12FF83FF7F46C8266FF81C" ID-DOI="http://doi.org/10.5281/zenodo.7642446" ID-Zenodo-Dep="7642446" httpUri="https://zenodo.org/record/7642446/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1965,1989]" targetBox="[280,1306,453,1927]" targetPageId="42">
<paragraph id="8B8336BBBF12FF83FF7F46C8266FF81C" blockId="42.[151,1437,1965,2025]" pageId="42" pageNumber="43">
<emphasis id="B948EAA9BF12FF83FF7F46C8237FF830" bold="true" box="[151,287,1965,1989]" pageId="42" pageNumber="43">FIGURE 27.</emphasis>
Left elytra (white dots indicating the chaetotaxy) of
<taxonomicName id="4C3C4D38BF12FF83FCD846C82197F830" box="[816,1015,1965,1989]" pageId="42" pageNumber="43">
<emphasis id="B948EAA9BF12FF83FCD846C821C9F830" box="[816,937,1965,1989]" italics="true" pageId="42" pageNumber="43">Dongodytes</emphasis>
species
</taxonomicName>
(white dots indicating the chaetotaxy). A.
<taxonomicName id="4C3C4D38BF12FF83FF7F46B4204CF81C" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[151,556,2001,2025]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pingjingae" status="sp. nov." subGenus="Dongodytodes">
<emphasis id="B948EAA9BF12FF83FF7F46B4204CF81C" box="[151,556,2001,2025]" italics="true" pageId="42" pageNumber="43">Dongodytes (Dongodytodes) pingjingae</emphasis>
</taxonomicName>
<emphasis id="B948EAA9BF12FF83FDDB46B4200CF81C" bold="true" box="[563,620,2001,2025]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="A27B57D2BF12FF83FDDB46B4200CF81C" box="[563,620,2001,2025]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; B.
<taxonomicName id="4C3C4D38BF12FF83FD7F46B421B0F81C" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[663,976,2001,2025]" class="Insecta" family="Carabidae" genus="Dongodytes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="mingruoae" status="sp. nov.">
<emphasis id="B948EAA9BF12FF83FD7F46B421B0F81C" box="[663,976,2001,2025]" italics="true" pageId="42" pageNumber="43">Dongodytes (s. str.) mingruoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B57D2BF12FF83FC3E46B4266FF81C" box="[982,1039,2001,2025]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF436633BF13FF82FF7F47D023F7F904" ID-DOI="http://doi.org/10.5281/zenodo.7642448" ID-Zenodo-Dep="7642448" httpUri="https://zenodo.org/record/7642448/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1717,1741]" targetBox="[163,1401,220,1692]" targetPageId="43">
<paragraph id="8B8336BBBF13FF82FF7F47D023F7F904" blockId="43.[151,1436,1717,1777]" pageId="43" pageNumber="44">
<emphasis id="B948EAA9BF13FF82FF7F47D02342F938" bold="true" box="[151,290,1717,1741]" pageId="43" pageNumber="44">FIGURE 28.</emphasis>
<taxonomicName id="4C3C4D38BF13FF82FEC147D020A2F938" authority="Tian &amp; Huang &amp; Jia, 2023" authorityName="Tian &amp; Huang &amp; Jia" authorityYear="2023" box="[297,706,1717,1741]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pingjingae" status="sp. nov." subGenus="Dongodytodes">
<emphasis id="B948EAA9BF13FF82FEC147D020A2F938" box="[297,706,1717,1741]" italics="true" pageId="43" pageNumber="44">Dongodytes (Dongodytodes) pingjingae</emphasis>
</taxonomicName>
<emphasis id="B948EAA9BF13FF82FD2247D32165F93B" bold="true" box="[714,773,1718,1742]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="A27B57D2BF13FF82FD2247D32165F93B" box="[714,773,1718,1742]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A. median lobe, lateral view; B. apical lobe, dorsal view; C. an active individual in cave.
</paragraph>
</caption>
<paragraph id="8B8336BBBF13FF85FF2F464523C9FE7C" blockId="43.[151,1437,1824,2030]" lastBlockId="44.[151,1437,151,1581]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
Head elongate, much longer than wide, HLm/HW=3.0–3.1; widest at about 1/3 excluding mandibles, as long as pronotum excluding mandibles; genae hardly expanded, gradually narrowed toward base, suddenly constricted before the collar-shaped neck which is about 2/5 as wide as head; frons flat, vertex convex; clypeus transverse, 6-setose; labrum nearly straight at frontal margin, 6-setose; two pairs of supraorbital setae present, anterior and posterior ones at about 1/3 and 7/9 from labrum to neck constriction respectively; a pair of suborbital setae present, mentum and submentum well separated by a clear labial suture, mentum with two pairs of setae, at base of tooth and sides of basal concave respectively; mental tooth short and bluntly bifid at apex; submentum 12- to 14-setose; mouthparts palps glabrous except 2
<superScript id="7C499BF3BF14FF85FDD841DF2020FF3D" attach="left" box="[560,576,186,200]" fontSize="6" pageId="44" pageNumber="45">nd</superScript>
labial palpomere bisetose on inner margin; 2
<superScript id="7C499BF3BF14FF85FBB841DF2600FF3D" attach="left" box="[1104,1120,186,200]" fontSize="6" pageId="44" pageNumber="45">nd</superScript>
labial palpomere 1.4 times longer than 3
<superScript id="7C499BF3BF14FF85FED841BB235EFF19" attach="none" box="[304,318,222,236]" fontSize="6" pageId="44" pageNumber="45">rd</superScript>
, 3
<superScript id="7C499BF3BF14FF85FEB341BB2309FF19" attach="left" box="[347,361,222,236]" fontSize="6" pageId="44" pageNumber="45">rd</superScript>
maxillary palpomere 1.1 times longer than 4
<superScript id="7C499BF3BF14FF85FC9941BB211EFF19" attach="none" box="[881,894,222,236]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
; ligula 10-setose; two pairs of suborbital pores present; antennae filiform, slender long, extending at about apical 1/7 of elytra; 1
<superScript id="7C499BF3BF14FF85FBC94066264CFEE4" attach="left" box="[1057,1068,259,273]" fontSize="6" pageId="44" pageNumber="45">st</superScript>
antennomere much more dilated than others, slightly longer than 2
<superScript id="7C499BF3BF14FF85FDFE40432046FEC1" attach="left" box="[534,550,294,308]" fontSize="6" pageId="44" pageNumber="45">nd</superScript>
which is shortest; the comparative length of each antennomere from the 1
<superScript id="7C499BF3BF14FF85FA9B4042271EFEC0" attach="left" box="[1395,1406,295,309]" fontSize="6" pageId="44" pageNumber="45">st</superScript>
to 11
<superScript id="7C499BF3BF14FF85FF5A402F22DFFEAD" attach="left" box="[178,191,330,344]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
in
<typeStatus id="54878819BF14FF85FF09402E2323FE90" box="[225,323,331,357]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
as: 1
<superScript id="7C499BF3BF14FF85FE95402E23E8FEAC" attach="left" box="[381,392,331,345]" fontSize="6" pageId="44" pageNumber="45">st</superScript>
(1.05) / 2
<superScript id="7C499BF3BF14FF85FE1C402F2064FEAD" attach="left" box="[500,516,330,344]" fontSize="6" pageId="44" pageNumber="45">nd</superScript>
(1.00) / 3
<superScript id="7C499BF3BF14FF85FD98402F201EFEAD" attach="left" box="[624,638,330,344]" fontSize="6" pageId="44" pageNumber="45">rd</superScript>
(2.47) / 4
<superScript id="7C499BF3BF14FF85FD02402F2097FEAD" attach="left" box="[746,759,330,344]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
(2.37) / 5
<superScript id="7C499BF3BF14FF85FC8A402F210FFEAD" attach="left" box="[866,879,330,344]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
(2.37) / 6
<superScript id="7C499BF3BF14FF85FC33402F2188FEAD" attach="left" box="[987,1000,330,344]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
(2.32) / 7
<superScript id="7C499BF3BF14FF85FBBC402F2601FEAD" attach="left" box="[1108,1121,330,344]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
(2.00) / 8
<superScript id="7C499BF3BF14FF85FB25402F26BAFEAD" attach="left" box="[1229,1242,330,344]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
(1.58) / 9
<superScript id="7C499BF3BF14FF85FAAE402F2733FEAD" attach="left" box="[1350,1363,330,344]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
(1.58) / 10
<superScript id="7C499BF3BF14FF85FF2A400B22AFFE89" attach="left" box="[194,207,366,380]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
(1.26) / 11
<superScript id="7C499BF3BF14FF85FEA2400B2337FE89" attach="left" box="[330,343,366,380]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
(1.53).
</paragraph>
<paragraph id="8B8336BBBF14FF85FF2F40F62085FD70" blockId="44.[151,1437,151,1581]" pageId="44" pageNumber="45">Prothorax long, propleura moderately tumid, and well visible from above, slightly wider than head, PrW/HW = 1.12–1.18; distinctly wider than pronotum, PrW/PnW = 0.76–0.85 (mean 0.81); widest at about 1/3 from base. Pronotum elongate, much longer than wide, PrL/PrW =1.88–1.94; slightly narrower than head, PnW/HW= 0.87– 0.92; sub-parallel-sided in most parts, slightly sinuate just before hind angles; front distinctly narrower than base, PfW/PbW =0.69–0.77; front angles bluntly right, hind angles roundly obtuse; two pairs of latero-marginal setae present, located at about 4/7 from base and a little before hind angles respectively; posterior transversal impression well-marked. Scutellum quite large, inverse triangle.</paragraph>
<paragraph id="8B8336BBBF14FF85FF2F43EA266BFCA8" blockId="44.[151,1437,151,1581]" pageId="44" pageNumber="45">
Elytra elongate, long than wide, EL/EW= 2.05–2.13; much wider than prothorax, EW/PrW=1.87–2.10; slightly shorter than head including mandibles and pronotum combined, EL/(HLm+PnL) = 0.94–1.00; base rather narrow, lateral margins smooth and glabrous, widest at about apical 4/7 of elytra; disc strongly convex though rather flat near base; striae not traceable; chaetotaxy (
<figureCitation id="13072A3EBF14FF85FDD6439E20A4FCE0" box="[574,708,763,789]" captionStart="FIGURE 27" captionStartId="42.[151,250,1965,1989]" captionTargetBox="[280,1306,453,1927]" captionTargetId="figure-129@42.[280,1307,453,1942]" captionTargetPageId="42" captionText="FIGURE 27. Left elytra (white dots indicating the chaetotaxy) of Dongodytes species (white dots indicating the chaetotaxy). A. Dongodytes (Dongodytodes) pingjingae sp. n.; B. Dongodytes (s. str.) mingruoae n. sp." figureDoi="http://doi.org/10.5281/zenodo.7642446" httpUri="https://zenodo.org/record/7642446/files/figure.png" pageId="44" pageNumber="45">Figure 27A</figureCitation>
): only a dorsal pores present on the site of 3
<superScript id="7C499BF3BF14FF85FB41439F26D7FCFD" attach="left" box="[1193,1207,762,776]" fontSize="6" pageId="44" pageNumber="45">rd</superScript>
stria, at about apical 4/7, preapical pore present, at about apical 1/8; humeral group of the marginal umbilicate pores not aggregated, 4
<superScript id="7C499BF3BF14FF85FA78427B27FDFCD9" attach="none" box="[1424,1437,798,812]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
backwardly shifted, closer to 5
<superScript id="7C499BF3BF14FF85FE184227239DFCA5" attach="left" box="[496,509,834,848]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
pore than to 3
<superScript id="7C499BF3BF14FF85FD76422720CCFCA5" attach="none" box="[670,684,834,848]" fontSize="6" pageId="44" pageNumber="45">rd</superScript>
; the anguloapical pore present.
</paragraph>
<paragraph id="8B8336BBBF14FF85FF2F42022676FC50" blockId="44.[151,1437,151,1581]" pageId="44" pageNumber="45">
Legs thin and slender, tibiae not longitudinally sulcate; the 1
<superScript id="7C499BF3BF14FF85FC8642022119FC80" attach="left" box="[878,889,871,885]" fontSize="6" pageId="44" pageNumber="45">st</superScript>
tarsomere slightly shorter, slightly longer and as long as 2
<superScript id="7C499BF3BF14FF85FF1442EF236CFC6D" attach="left" box="[252,268,906,920]" fontSize="6" pageId="44" pageNumber="45">nd</superScript>
– 4
<superScript id="7C499BF3BF14FF85FEDF42EF2324FC6D" attach="left" box="[311,324,906,920]" fontSize="6" pageId="44" pageNumber="45">th</superScript>
tarsomeres combined in fore, middle and hind legs respectively.
</paragraph>
<paragraph id="8B8336BBBF14FF85FF2F42CA208AFC3C" blockId="44.[151,1437,151,1581]" box="[199,746,943,969]" pageId="44" pageNumber="45">Ventrite VII bisetose in male, 4-setose in female.</paragraph>
<paragraph id="8B8336BBBF14FF85FF2F42B62604FB88" blockId="44.[151,1437,151,1581]" pageId="44" pageNumber="45">
Male genitalia (
<figureCitation id="13072A3EBF14FF85FE9642B6204BFC18" box="[382,555,979,1005]" captionStart="FIGURE 28" captionStartId="43.[151,250,1717,1741]" captionTargetBox="[163,1401,220,1692]" captionTargetId="figure-20@43.[163,1424,181,1693]" captionTargetPageId="43" captionText="FIGURE 28. Dongodytes (Dongodytodes) pingjingae sp. n. A. median lobe, lateral view; B. apical lobe, dorsal view; C. an active individual in cave." figureDoi="http://doi.org/10.5281/zenodo.7642448" httpUri="https://zenodo.org/record/7642448/files/figure.png" pageId="44" pageNumber="45">Figure 28A, B</figureCitation>
): Male genitalia well-sclerotized, median lobe very elongate and thin, basal orifice comparatively small, but with a large sagittal aileron; base quite large, strongly curved ventrally in apical half, apical lobe roundly blunt; inner sac with a weakly-marked copulatory piece about 1/4 as long as median lobe; in dorsal view, apical lobe distinctly narrowed and nearly parallel-sided before the rounded apex; parameres slender, base much widened, apical part distinctly narrowed, each with four long setae apically.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF14FF85FF2F45E2264FFAC4" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8B8336BBBF14FF85FF2F45E2264FFAC4" blockId="44.[151,1437,151,1581]" pageId="44" pageNumber="45">
<emphasis id="B948EAA9BF14FF85FF2F45E2235BFB54" bold="true" box="[199,315,1159,1185]" pageId="44" pageNumber="45">Remarks.</emphasis>
This new species differs from other congeners of the subgenus
<taxonomicName id="4C3C4D38BF14FF85FBEE45E22746FB54" authority="Tian, 2011" authorityName="Tian" authorityYear="2011" box="[1030,1318,1159,1185]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subGenus" subGenus="Dongodytodes">
<emphasis id="B948EAA9BF14FF85FBEE45E226C8FB54" box="[1030,1192,1159,1185]" italics="true" pageId="44" pageNumber="45">Dongodytodes</emphasis>
Tian, 2011
</taxonomicName>
by having a well-defined labial suture. It may be close to
<taxonomicName id="4C3C4D38BF14FF85FD2245CE2697FB30" authority="(Tian et al. 2014)" baseAuthorityName="Tian" baseAuthorityYear="2014" box="[714,1271,1195,1221]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="yaophilus" subGenus="Dongodytodes">
<emphasis id="B948EAA9BF14FF85FD2245CE2085FB31" box="[714,741,1195,1220]" italics="true" pageId="44" pageNumber="45">D.</emphasis>
(
<emphasis id="B948EAA9BF14FF85FD1145CE267CFB30" box="[761,1052,1195,1221]" italics="true" pageId="44" pageNumber="45">Dongodytodes) yaophilus</emphasis>
(
<bibRefCitation id="EFAD4B4ABF14FF85FBC745CE268FFB30" author="Tian, M. Y. &amp; Yin, H. M. &amp; Huang, S. B." box="[1071,1263,1195,1221]" pageId="44" pageNumber="45" pagination="69 - 107" refId="ref28960" refString="Tian, M. Y., Yin, H. M. &amp; Huang, S. B. (2014) Du'an Karst of Guangxi: a kingdom of the cavernicolous genus Dongodytes Deuve (Coleoptera: Carabidae: Trechinae). ZooKeys, 454, 69 - 107. https: // doi. org / 10.3897 / zookeys. 454.7269" type="journal article" year="2014">
Tian
<emphasis id="B948EAA9BF14FF85FB8645CE26CBFB30" box="[1134,1195,1195,1221]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2014
</bibRefCitation>
)
</taxonomicName>
which is also recorded from Dahua County, and the localities (Xiannü Dong and Qiaoxu Dong) of both species are distant from only 20 kilometres in a straight line. However, the male genitalia is slenderer, with a more widened apical lobe, and parameres are narrowed at apices instead of broadly rounded in
<taxonomicName id="4C3C4D38BF14FF85FC8C44722648FAC4" box="[868,1064,1303,1329]" class="Insecta" family="Carabidae" genus="Dongodytes" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="yaophilus" subGenus="Dongodytodes">
<emphasis id="B948EAA9BF14FF85FC8C447221E0FAC5" box="[868,896,1303,1328]" italics="true" pageId="44" pageNumber="45">D.</emphasis>
(
<emphasis id="B948EAA9BF14FF85FC7844722648FAC4" box="[912,1064,1303,1329]" italics="true" pageId="44" pageNumber="45">D.) yaophilus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF14FF85FF2F445E2077FA8C" pageId="44" pageNumber="45" type="etymology">
<paragraph id="8B8336BBBF14FF85FF2F445E2077FA8C" blockId="44.[151,1437,151,1581]" pageId="44" pageNumber="45">
<emphasis id="B948EAA9BF14FF85FF2F445E232EFAA0" bold="true" box="[199,334,1339,1365]" pageId="44" pageNumber="45">Etymology.</emphasis>
This beautiful species is named after Ms. Pingjing Yang (a member of SCAU caving team), one of the collectors of the
<typeStatus id="54878819BF14FF85FE94443A23CCFA8C" box="[380,428,1375,1401]" pageId="44" pageNumber="45">type</typeStatus>
material.
</paragraph>
</subSubSection>
<subSubSection id="C3266530BF14FF85FF2F44E7230CF9D8" pageId="44" pageNumber="45" type="distribution">
<paragraph id="8B8336BBBF14FF85FF2F44E7277BFA68" blockId="44.[151,1437,151,1581]" box="[199,1307,1410,1437]" pageId="44" pageNumber="45">
<emphasis id="B948EAA9BF14FF85FF2F44E72301FA69" bold="true" box="[199,353,1410,1436]" pageId="44" pageNumber="45">Distribution.</emphasis>
<collectingCountry id="F32B762BBF14FF85FE8F44E623CAFA68" box="[359,426,1411,1437]" name="China" pageId="44" pageNumber="45">China</collectingCountry>
(
<collectingRegion id="49F8F859BF14FF85FE5144E62040FA68" box="[441,544,1411,1437]" country="China" name="Guangxi" pageId="44" pageNumber="45">Guangxi</collectingRegion>
). Known only from the
<typeStatus id="54878819BF14FF85FCC344E6213BFA68" box="[811,859,1411,1437]" pageId="44" pageNumber="45">type</typeStatus>
locality, cave Xiannü Dong (
<figureCitation id="13072A3EBF14FF85FB4B44E6276BFA68" box="[1187,1291,1411,1437]" captionStart="FIGURE 1" captionStartId="3.[152,255,1900,1924]" captionTargetBox="[151,1432,955,1874]" captionTargetId="figure-296@3.[151,1436,955,1874]" captionTargetPageId="3" captionText="FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong." figureDoi="http://doi.org/10.5281/zenodo.7642402" httpUri="https://zenodo.org/record/7642402/files/figure.png" pageId="44" pageNumber="45">Figure 1i</figureCitation>
).
</paragraph>
<paragraph id="8B8336BBBF14FF85FF2F44C2230CF9D8" blockId="44.[151,1437,151,1581]" pageId="44" pageNumber="45">
Xiannü Dong is a disused limestone show cave, about 2 kilometres long. There are several chambers along the main passage. Apart from the cave beetles, crickets, millipedes, woodlice, snails and a white mouse were also observed inside the cave. The beetles were wandering on walls and on the ground in several wet places in the dark zone (
<figureCitation id="13072A3EBF14FF85FF334776233FF9D8" box="[219,351,1555,1581]" captionStart="FIGURE 28" captionStartId="43.[151,250,1717,1741]" captionTargetBox="[163,1401,220,1692]" captionTargetId="figure-20@43.[163,1424,181,1693]" captionTargetPageId="43" captionText="FIGURE 28. Dongodytes (Dongodytodes) pingjingae sp. n. A. median lobe, lateral view; B. apical lobe, dorsal view; C. an active individual in cave." figureDoi="http://doi.org/10.5281/zenodo.7642448" httpUri="https://zenodo.org/record/7642448/files/figure.png" pageId="44" pageNumber="45">Figure 28C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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