treatments-xml/data/03/95/08/039508639B4DFFEEFF7711D8FB24EFAE.xml

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<mods:title id="0EEA9B907B7F07846E0E0F1AAEFD0FD7">A new species of Raphidopus Stimpson, 1858, from the Persian Gulf (Crustacea: Decapoda: Anomura: Porcellanidae)</mods:title>
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<mods:namePart id="A52C57F7CD4B114DA07F2378CBEB29AD">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="54BD011A0E5A87F31566846F3BA40A2D">Safaie, Mohsen</mods:namePart>
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<paragraph id="8B83B9759B4DFFECFF7711D8FD9EEEA0" blockId="0.[151,554,1720,1779]" box="[151,554,1720,1746]" pageId="0" pageNumber="54">
<heading id="D0CB0E199B4DFFECFF7711D8FD9EEEA0" bold="true" box="[151,554,1720,1746]" fontSize="11" level="1" pageId="0" pageNumber="54" reason="1">
<emphasis id="B94865679B4DFFECFF7711D8FD9EEEA0" bold="true" box="[151,554,1720,1746]" pageId="0" pageNumber="54">
<taxonomicName id="4C3CC2F69B4DFFECFF7711D8FE26EEA0" ID-CoL="206e7acb-1620-4fbb-9520-6b99322c03b9" box="[151,402,1720,1746]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="54" phylum="Arthropoda" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B94865679B4DFFECFF7711D8FE26EEA0" bold="true" box="[151,402,1720,1746]" italics="true" pageId="0" pageNumber="54">Raphidopus persicus</emphasis>
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<taxonomicNameLabel id="A27BD81C9B4DFFECFE7A11D8FD9EEEA0" box="[410,554,1720,1746]" pageId="0" pageNumber="54" rank="species">new species</taxonomicNameLabel>
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(
<figureCitation id="1307A5F09B4DFFECFF7F11BBFF59EE81" box="[159,237,1755,1779]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="0" pageNumber="54">Figs. 1</figureCitation>
, 
<figureCitation id="1307A5F09B4DFFECFF1A11BBFEBDEE81" box="[250,265,1755,1779]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="0" pageNumber="54">2</figureCitation>
)
</paragraph>
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<subSubSection id="C326EAFE9B4DFFECFF771043FEC1EF04" pageId="0" pageNumber="54" type="reference_group">
<paragraph id="8B83B9759B4DFFECFF771043FEC1EF04" blockId="0.[151,1435,1827,1910]" pageId="0" pageNumber="54">
<treatmentCitationGroup id="AB2C9E5B9B4DFFECFF771043FEC1EF04" pageId="0" pageNumber="54">
<taxonomicName id="4C3CC2F69B4DFFECFF771043FEEBEF4B" ID-CoL="445049fb-994e-4886-ba7b-e2521935c222" box="[151,351,1827,1849]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="54" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4DFFECFF771043FEEBEF4B" box="[151,351,1827,1849]" italics="true" pageId="0" pageNumber="54">Raphidopus indicus</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0A9D9F649B4DFFECFE991043FDB8EF4B" author="Haig" box="[377,524,1827,1849]" page="62" pageId="0" pageNumber="54" year="1966">
<bibRefCitation id="EFADC4849B4DFFECFE991043FE52EF4B" author="Haig" box="[377,486,1827,1849]" pageId="0" pageNumber="54" refString="Haig, J. (1966) The Porcellanidae (Crustacea Anomura) of the Iranian Gulf and Gulf of Oman. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 129, 49 - 65." type="journal article" year="1966">Haig 1966</bibRefCitation>
: 62
</treatmentCitation>
, fig. 6; 
<treatmentCitation id="0A9D9F649B4DFFECFDB91043FD5DEF4B" author="Apel" box="[601,745,1827,1849]" page="43" pageId="0" pageNumber="54" year="2001">
<bibRefCitation id="EFADC4849B4DFFECFDB91043FD72EF4B" author="Apel" box="[601,710,1827,1849]" pageId="0" pageNumber="54" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes. Unpublished Ph. D. thesis, Johann Wolfgang Goethe-Universitat, Frankfurt am Main, pp. 1 - 268." type="book chapter" year="2001">Apel 2001</bibRefCitation>
: 43
</treatmentCitation>
(not 
<taxonomicName id="4C3CC2F69B4DFFECFCFF1043FB2BEF48" ID-CoL="445049fb-994e-4886-ba7b-e2521935c222" authority="Henderson, 1893" authorityName="Henderson" authorityYear="1893" box="[799,1183,1827,1850]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="54" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4DFFECFCFF1043FC53EF4B" box="[799,999,1827,1849]" italics="true" pageId="0" pageNumber="54">Raphidopus indicus</emphasis>
<bibRefCitation id="EFADC4849B4DFFECFC0E1043FB2BEF48" author="Henderson" box="[1006,1183,1827,1850]" pageId="0" pageNumber="54" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnean Society of London, Zoology, (2) 5, 325 - 458, pls. 36 - 40." type="journal article" year="1893">Henderson, 1893</bibRefCitation>
</taxonomicName>
). 
<taxonomicName id="4C3CC2F69B4DFFECFF771021FED7EF25" ID-CoL="5c8de6e7-6ae2-4bcc-aa45-cf04be1da142" box="[151,355,1857,1879]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="54" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4DFFECFF771021FED7EF25" box="[151,355,1857,1879]" italics="true" pageId="0" pageNumber="54">Raphidopus ciliatus</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0A9D9F649B4DFFECFE9C1021FD3AEF25" author="Tirmizi" box="[380,654,1857,1879]" page="71" pageId="0" pageNumber="54" year="1994">
<bibRefCitation id="EFADC4849B4DFFECFE9C1021FDDDEF25" author="Tirmizi" box="[380,617,1857,1879]" pageId="0" pageNumber="54" refString="Tirmizi, N. M. &amp; Ghani, N. (1994) An Indian Ocean record for a porcellanid crab Raphidopus ciliatus Stimpson, 1858. Pakistan Journal of Marine Sciences, 3 (1), 69 - 72, fig. 1." type="journal article" year="1994">Tirmizi &amp; Ghani 1994</bibRefCitation>
: 71
</treatmentCitation>
(part), fig. 1J; 
<treatmentCitation id="0A9D9F649B4DFFECFCD11021FBE7EF25" author="Siddiqui" box="[817,1107,1857,1879]" page="88" pageId="0" pageNumber="54" year="2003">
<bibRefCitation id="EFADC4849B4DFFECFCD11021FB9AEF25" author="Siddiqui" box="[817,1070,1857,1879]" pageId="0" pageNumber="54" refString="Siddiqui, F. A. &amp; Kazmi, Q. B. (2003) A checklist of marine anomurans (Crustacea: Decapoda) of Pakistan, northern Arabian Sea. Memoirs of Museum Victoria, 60 (1), 87 - 89." type="journal article" year="2003">Siddiqui &amp; Kazmi 2003</bibRefCitation>
: 88
</treatmentCitation>
(part) (not 
<taxonomicName id="4C3CC2F69B4DFFECFB301021FEDEEF04" ID-CoL="5c8de6e7-6ae2-4bcc-aa45-cf04be1da142" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="54" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4DFFECFB301021FA2FEF25" box="[1232,1435,1857,1879]" italics="true" pageId="0" pageNumber="54">Raphidopus ciliatus</emphasis>
<bibRefCitation id="EFADC4849B4DFFECFF271000FEDEEF04" author="Stimpson" box="[199,362,1888,1910]" pageId="0" pageNumber="54" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum quoe in Expeditione ad Oceanum Pacificum eptentrionalem a Republica Federata Missa, Cadwaladaro Ringgold et Johann Rodgers Ducibus, observavit et descripsit - Part VII, Crustacea Anomura. Proceedings of the Academy of Natural Sciences, Philadelphia, 10, 225 - 252." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
).
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</paragraph>
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<paragraph id="8B83B9759B4DFFEDFF7710C2FE68E8A6" blockId="0.[151,1436,1954,2015]" lastBlockId="1.[151,1437,151,2012]" lastPageId="1" lastPageNumber="55" pageId="0" pageNumber="54">
<emphasis id="B94865679B4DFFECFF7710C2FE36EFC9" bold="true" box="[151,386,1954,1979]" pageId="0" pageNumber="54">Material examined.</emphasis>
<typeStatus id="548707D79B4DFFECFE6C10C3FE4CEFC9" box="[396,504,1955,1979]" pageId="0" pageNumber="54" type="holotype">Holotype</typeStatus>
: male (8.5 × 
<quantity id="4CC414909B4DFFECFD7310C3FD58EFC9" box="[659,748,1955,1979]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="0" pageNumber="54" unit="mm" value="6.8">6.8 mm</quantity>
) (
<collectionCode id="ED2D21B09B4DFFECFCE610C3FCECEFC9" box="[774,856,1955,1979]" pageId="0" pageNumber="54">ZMSB</collectionCode>
), west of Hormozgan, 
<collectingCountry id="F32BF9E59B4DFFECFBBF10C3FB1BEFC9" box="[1119,1199,1955,1979]" name="Iran" pageId="0" pageNumber="54">Qeshm</collectingCountry>
Island, 
<collectingCountry id="F32BF9E59B4DFFECFAED10C4FA89EFC9" box="[1293,1341,1956,1979]" name="Iran" pageId="0" pageNumber="54">Iran</collectingCountry>
, Persian Gulf, 
<geoCoordinate id="EE08DFB29B4DFFECFF3B10A7FE83EFAD" box="[219,311,1991,2015]" direction="north" orientation="latitude" pageId="0" pageNumber="54" precision="925" value="27.016666">27°01'N</geoCoordinate>
<geoCoordinate id="EE08DFB29B4DFFECFEA210A7FE28EFAD" box="[322,412,1991,2015]" direction="east" orientation="longitude" pageId="0" pageNumber="54" precision="925" value="56.133335">56°08'E</geoCoordinate>
- 
<geoCoordinate id="EE08DFB29B4DFFECFE5910A7FDA1EFAD" box="[441,533,1991,2015]" direction="north" orientation="latitude" pageId="0" pageNumber="54" precision="925" value="27.016666">27°01'N</geoCoordinate>
<geoCoordinate id="EE08DFB29B4DFFECFDC010A7FD31EFAD" box="[544,645,1991,2015]" direction="east" orientation="longitude" pageId="0" pageNumber="54" precision="925" value="56.2">56°12' E</geoCoordinate>
, 
<quantity id="4CC414909B4DFFECFD7510A7FD45EFAD" box="[661,753,1991,2015]" metricMagnitude="1" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="0" pageNumber="54" unit="m" value="18.0" valueMax="19.0" valueMin="17.0">17–19m</quantity>
depth, coll. shrimp bottom trawl, M. Safaei, 
<date id="FF829FB59B4DFFEDFAE310A7FF67E8DD" lastPageId="1" lastPageNumber="55" pageId="0" pageNumber="54" value="2010-12-20">20 December 2010</date>
. 
<typeStatus id="548707D79B4CFFEDFF0017F8FEF2E8C2" box="[224,326,152,176]" pageId="1" pageNumber="55" type="paratype">Paratype</typeStatus>
: 
<specimenCount id="9D3A72FC9B4CFFEDFEB417F7FE16E8DD" box="[340,418,151,175]" pageId="1" pageNumber="55" type="male">1 male</specimenCount>
(7.5 × 
<quantity id="4CC414909B4CFFEDFE1D17F7FDEDE8DD" box="[509,601,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="55" unit="mm" value="5.8">5.8 mm</quantity>
) (
<collectionCode id="ED2D21B09B4CFFEDFD9217F7FD1AE8DD" box="[626,686,151,175]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="55">ZRC</collectionCode>
), northwestern Persian Gulf, 
<collectingCountry id="F32BF9E59B4CFFEDFBE217F8FB86E8DD" box="[1026,1074,152,175]" name="Iraq" pageId="1" pageNumber="55">Iraq</collectingCountry>
, 
<geoCoordinate id="EE08DFB29B4CFFEDFBA017F7FB5CE8DD" box="[1088,1256,151,175]" direction="north" orientation="latitude" pageId="1" pageNumber="55" precision="1" value="29.86224">29°51'44.06&quot;N</geoCoordinate>
<geoCoordinate id="EE08DFB29B4CFFEDFB1117F7FA23E8DD" box="[1265,1431,151,175]" direction="east" orientation="longitude" pageId="1" pageNumber="55" precision="1" value="48.7315">48°43'53.40&quot;E</geoCoordinate>
, cοll• đređge, Μ• Naser, 2OII•
</paragraph>
</subSubSection>
<subSubSection id="C326EAFE9B4CFFEDFF2717BFFDCFEAD5" pageId="1" pageNumber="55" type="diagnosis">
<paragraph id="8B83B9759B4CFFEDFF2717BFFDCFEAD5" blockId="1.[151,1437,151,2012]" pageId="1" pageNumber="55">
<emphasis id="B94865679B4CFFEDFF2717BFFEF5E88A" bold="true" box="[199,321,223,248]" pageId="1" pageNumber="55">Diagnosis.</emphasis>
Carapace ovoid, dorsal surfaces almost glabrous, without pubescence (
<figureCitation id="1307A5F09B4CFFEDFB8017BFFB1EE885" box="[1120,1194,223,247]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A); anterolateral, posterolateral margins strongly arcuate (
<figureCitation id="1307A5F09B4CFFEDFD5E1664FCB8E96E" box="[702,780,260,284]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A); regions indistinct, demarcated by shallow grooves (
<figureCitation id="1307A5F09B4CFFEDFF1E1647FE8AE94D" box="[254,318,295,319]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Fig. 1</figureCitation>
A, B, D); frontal margin trilobate from dorsal view (
<figureCitation id="1307A5F09B4CFFEDFC681647FC65E94D" box="[904,977,295,319]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A); lateral margin divided into 2 parts by small fissure just after beginning of cervical groove; anterolateral margin behind cleft granulated, no trace of lateral spines (
<figureCitation id="1307A5F09B4CFFEDFE7D160FFE5CE9F5" box="[413,488,367,391]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A); third antennal article relatively short, about 2 times length of fourth article (
<figureCitation id="1307A5F09B4CFFEDFF0E16F4FE8FE9DE" box="[238,315,404,428]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A); third maxilliped with merus longer than ischium, inner margin of ischium auriculiform, exopod foliaceous, length 2.1 times width (
<figureCitation id="1307A5F09B4CFFEDFD2C16D7FCA5E9BD" box="[716,785,439,463]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">Fig. 2</figureCitation>
D); anterior margin of thoracic sternum with median part prominently convex towards buccal cavity (
<figureCitation id="1307A5F09B4CFFEDFD6316BCFD73E986" box="[643,711,476,500]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">Fig. 2</figureCitation>
B); merus of cheliped with distinct spine on inner ventral margin (
<figureCitation id="1307A5F09B4CFFEDFF7F169FFF57EA65" box="[159,227,511,535]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">
Fig. 
<date id="FF829FB59B4CFFEDFF34169FFF57EA65" box="[212,227,511,535]" pageId="1" pageNumber="55">2</date>
</figureCitation>
I, J); outer margin of carpus of cheliped with 3 or 4 prominent spines on distal third (
<figureCitation id="1307A5F09B4CFFEDFB76169FFB6AEA65" box="[1174,1246,511,535]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A–C, 2J); dorsal (inner) margin of palm cristate with rounded ridge, sometimes lined with short oblique ridges (
<figureCitation id="1307A5F09B4CFFEDFB351544FAABEA4E" box="[1237,1311,548,572]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A–C, 2K); cutting edge of dactylus of larger chela with 2 strong inwardly-directed subproximal teeth, pollex with 1 subdistal, 1 subproximal teeth (
<figureCitation id="1307A5F09B4CFFEDFE6F150CFE62EAF6" box="[399,470,620,644]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">Fig. 2</figureCitation>
K); ambulatory legs relatively long; length of first to third meri 3.9, 4.7, 3.8 times maximum width, respectively (
<figureCitation id="1307A5F09B4CFFEDFE1415EFFD87EAD5" box="[500,563,655,679]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Fig. 1</figureCitation>
A, B).
</paragraph>
</subSubSection>
<subSubSection id="C326EAFE9B4CFFEEFF2715D3FBEAEA4E" lastPageId="2" lastPageNumber="56" pageId="1" pageNumber="55" type="description">
<paragraph id="8B83B9759B4CFFEDFF2715D3FDE3EC9A" blockId="1.[151,1437,151,2012]" pageId="1" pageNumber="55">
<emphasis id="B94865679B4CFFEDFF2715D3FD85EABE" bold="true" box="[199,561,691,716]" pageId="1" pageNumber="55">
Description of 
<typeStatus id="548707D79B4CFFEDFE9F15D4FE52EABE" box="[383,486,692,716]" pageId="1" pageNumber="55" type="holotype">holotype</typeStatus>
male.
</emphasis>
Carapace ovoid, dorsal surfaces smooth, almost glabrous, without trace of pubescence (
<figureCitation id="1307A5F09B4CFFEDFECC15B7FECFEA9D" box="[300,379,727,751]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A); anterolateral, posterolateral margins strongly arcuate; branchiostegites with dense coat of long, soft setae that do not completely obscure dorsal surface or margins; postfrontal region depressed, forming 2 transverse grooves, medially separated by fissure (
<figureCitation id="1307A5F09B4CFFEDFC37147FFB90EB45" box="[983,1060,799,823]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A); regions indistinct, demarcated by shallow grooves (
<figureCitation id="1307A5F09B4CFFEDFDE71424FDF2EB2E" box="[519,582,836,860]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Fig. 1</figureCitation>
A, B, D). Frontal margin slightly deflexed downwards, trilobate from dorsal view, lobes rounded, median lobe most prominent (
<figureCitation id="1307A5F09B4CFFEDFD091407FC82EB0D" box="[745,822,871,895]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A). Lateral margin divided into 2 parts by small fissure just after beginning of cervical groove; posterior margin cleft subcristate, lined with sharp to rounded granules but not spinulate, no trace of lateral spines (
<figureCitation id="1307A5F09B4CFFEDFD1414CFFC8AEBB5" box="[756,830,943,967]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A). Eyes small, clearly visible from dorsal view (
<figureCitation id="1307A5F09B4CFFEDFF3F14B4FE98EB9E" box="[223,300,980,1004]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A). Basal antennular segment with well developed triangular submedian tooth on gently granulated anterior margin (
<figureCitation id="1307A5F09B4CFFEDFE301497FDA7EC7D" box="[464,531,1015,1039]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">Fig. 2</figureCitation>
E), outer surface with indistinct, very low, weakly granulated oblique ridge. First antennal article relatively broad, fused to carapace; third antennal article relatively short, about 2 times length of fourth article (
<figureCitation id="1307A5F09B4CFFEDFED7135FFE34EC25" box="[311,384,1087,1111]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, D, 2A). Third maxilliped relatively narrow, inner margins with long setae; merus longer than ischium; inner margin of ischium auriculiform; merus narrow, proximal inner margin with triangular auriculiform structure; exopod broad, foliaceous, length 2.1 times width, not reaching distal edge of merus, with well developed flagellum (
<figureCitation id="1307A5F09B4CFFEDFE7013CCFE62ECB6" box="[400,470,1196,1220]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">Fig. 2</figureCitation>
D). Anterior margin of thoracic sternum distinctly sinuous, median part prominently convex towards buccal cavity (
<figureCitation id="1307A5F09B4CFFEDFE1213B0FD82EC9A" box="[498,566,1232,1256]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">Fig. 2</figureCitation>
B).
</paragraph>
<paragraph id="8B83B9759B4CFFEDFF291394FBE5EF3E" blockId="1.[151,1437,151,2012]" pageId="1" pageNumber="55">
Chelae stout, asymmetrical; outer surface with dense coat of long, soft setae on lower part, especially along ventral margin, completely obscuring margin (
<figureCitation id="1307A5F09B4CFFEDFD491278FD41ED42" box="[681,757,1304,1328]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A–C); inner surfaces of merus, carpus more densely setose than outer surfaces, lower surfaces almost glabrous (
<figureCitation id="1307A5F09B4CFFEDFD03125CFC95ED26" box="[739,801,1340,1364]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Fig. 1</figureCitation>
A–C). Surfaces, margins of palm, merus, carpus, fingers of larger cheliped slightly rugose ventral margins, lower surfaces of segments gently granulose (
<figureCitation id="1307A5F09B4CFFEDFB151200FAF6ED0A" box="[1269,1346,1376,1400]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
C, 2K). Outer, lower surfaces of merus, carpus gently convex, inner surfaces distinctly concave; indistinctly cristate along outer margins (
<figureCitation id="1307A5F09B4CFFEDFEA312C8FE39EDB2" box="[323,397,1448,1472]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A–C, 
<date id="FF829FB59B4CFFEDFE3612C8FE5BEDB2" box="[470,495,1448,1472]" pageId="1" pageNumber="55">2I</date>
, J). Merus subquadrate, with distinct spine on ventral margin, outer margin almost smooth (
<figureCitation id="1307A5F09B4CFFEDFF1912ACFE89ED96" box="[249,317,1484,1508]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">
Fig. 
<date id="FF829FB59B4CFFEDFECE12ACFE89ED96" box="[302,317,1484,1508]" pageId="1" pageNumber="55">2</date>
</figureCitation>
I, J). Dorsal margin of carpus gently granular; outer margin with 3 prominent spines on distal third, rest of margin entire; dorsal surface with low longitudinal median ridge (
<figureCitation id="1307A5F09B4CFFEDFC2D1290FBA2EE7A" box="[973,1046,1520,1544]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A–C, 2J). Outer surfaces of palms gently convex, with submedian longitudinal ridge; dorsal (inner) margin cristate with rounded ridge, sometimes with lined with short oblique ridges (
<figureCitation id="1307A5F09B4CFFEDFDD11158FDCDEE22" box="[561,633,1592,1616]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A–C, 2K); outer margin rounded, with some denticles on proximal half. Inner, outer surfaces immediately adjacent to base of fingers (including cutting edges) covered with long setae that partially obscure structures (
<figureCitation id="1307A5F09B4CFFEDFE3611E0FDA1EEEA" box="[470,533,1664,1688]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Fig. 1</figureCitation>
C). Cutting edge of dactylus of larger chela almost blade-like throughout length, with 2 strong, inwardly-directed subproximal teeth (inner tooth smaller); pollex with proximal part of cutting edge flattened, area relatively broad; with single subdistal, subproximal teeth, margin between teeth lined with denticles (
<figureCitation id="1307A5F09B4CFFEDFF7F118CFF50EF76" box="[159,228,1772,1796]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="1" pageNumber="55">Fig. 2</figureCitation>
K). Cutting edges of fingers of smaller chela with several subproximal teeth, numerous denticles, tooth on pollex flattened proximally like that of larger chela but flattened area relatively less broad (
<figureCitation id="1307A5F09B4CFFEDFB461070FB52EF5A" box="[1190,1254,1808,1832]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Fig. 1</figureCitation>
C). Dorsal margins of dactylus subcristate, with shallow median longitudinal groove (
<figureCitation id="1307A5F09B4CFFEDFC501054FC4EEF3E" box="[944,1018,1844,1868]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
C, 2K).
</paragraph>
<paragraph id="8B83B9759B4CFFEEFF291038FB20E8A6" blockId="1.[151,1437,151,2012]" lastBlockId="2.[151,1437,151,2012]" lastPageId="2" lastPageNumber="56" pageId="1" pageNumber="55">
Ambulatory legs relatively long, second leg longest; margins of segments lined with long plumose setae which partially obscure surfaces (
<figureCitation id="1307A5F09B4CFFEDFE26101CFDA4EFE6" box="[454,528,1916,1940]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, 2F–H). Merus long, slender, length of first to third meri 3.9, 4.7, 3.8 times maximum width, respectively; margins unarmed; outer surface, margins of merus, carpus, propodus densely covered with long, soft setae which obscure surface; inner surfaces smooth, glabrous (
<figureCitation id="1307A5F09B4CFFEDFBAA10A4FB21EFAE" box="[1098,1173,1988,2012]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="1" pageNumber="55">Figs. 1</figureCitation>
A, B, 2F–H). Dactylus long, slender, styliform, almost straight, margins of distal half partcularly setose, forming a brush-like structure; length of first to third dactyli 7.5, 5.7, 5.6 times maximum width, respectively (
<figureCitation id="1307A5F09B4FFFEEFBEF17DCFBE5E8A6" box="[1039,1105,188,212]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="2" pageNumber="56">Fig. 2</figureCitation>
F–H).
</paragraph>
<paragraph id="8B83B9759B4FFFEEFF2917BFFAF6E96E" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="56">
Abdomen with 6 free somites and telson, relatively broad, subrectangular in shape, covering entire thoracic sternum; telson with 7 plates, with pair of well developed pleopods on second abdominal somite (
<figureCitation id="1307A5F09B4FFFEEFB3D1664FA94E96E" box="[1245,1312,260,284]" captionStart="FIGURE 2" captionStartId="5.[151,250,1866,1889]" captionTargetBox="[210,1329,209,1807]" captionTargetId="figure@5.[199,1367,193,1824]" captionTargetPageId="5" captionText="FIGURE 2. Raphidopus persicus new species, holotype male (8.5 × 6.8 mm) (ZMSB), Iran. A, left side of carapace; B, anterior thoracic sternum; C, male telson and somite 6; D, left third maxilliped; E, left basal antennular article; F, right first ambulatory leg; G, right second ambulatory leg; H, right third ambulatory leg; I, ventral view of merus of left cheliped; J, dorsal view of merus and carpus of left cheliped; K, outer view of left major chela. All structures denuded. Scales: A – D, F – K = 1.0 mm; E = 0.5 mm." httpUri="https://zenodo.org/record/215482/files/figure.png" pageId="2" pageNumber="56">Fig. 2</figureCitation>
C).
</paragraph>
<paragraph id="8B83B9759B4FFFEEFF271648FBEAEA4E" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="56">
<emphasis id="B94865679B4FFFEEFF271648FEC1E932" bold="true" box="[199,373,296,320]" pageId="2" pageNumber="56">
<typeStatus id="548707D79B4FFFEEFF271648FE80E932" box="[199,308,296,320]" pageId="2" pageNumber="56" type="paratype">Paratype</typeStatus>
male
</emphasis>
. The smaller 
<typeStatus id="548707D79B4FFFEEFDF21648FDC6E932" box="[530,626,296,320]" pageId="2" pageNumber="56" type="paratype">paratype</typeStatus>
male (
<figureCitation id="1307A5F09B4FFFEEFD221647FCB6E94D" box="[706,770,295,319]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="2" pageNumber="56">Fig. 1</figureCitation>
B) differs from the 
<typeStatus id="548707D79B4FFFEEFC031647FBF1E94D" box="[995,1093,295,319]" pageId="2" pageNumber="56" type="holotype">holotype</typeStatus>
male in several aspects of the chelipeds, none of which are important at the species level. The right carpus has four prominent spines followed by several denticles on the outer margin, but these denticles do not reach beyond half the length of the segment. The left carpus has been damaged and regenerated, and is unarmed on the outer margin. However, the regenerated distal third of the margin is unevenly crenulated and indicates there were three spines originally at this position. The inner ridge on the merus is also relatively more crenulated and granular than the 
<typeStatus id="548707D79B4FFFEEFC1616BCFBECE986" box="[1014,1112,476,500]" pageId="2" pageNumber="56" type="holotype">holotype</typeStatus>
and the outer surface of the chelae is relatively more granular and uneven, with the submedian longitudinal ridge stronger. The outer surfaces of both chelipeds of the 
<typeStatus id="548707D79B4FFFEEFE461545FDB2EA4F" box="[422,518,549,573]" pageId="2" pageNumber="56" type="paratype">paratype</typeStatus>
male are relatively more rugose than in the 
<typeStatus id="548707D79B4FFFEEFC141544FBEEEA4E" box="[1012,1114,548,572]" pageId="2" pageNumber="56" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C326EAFE9B4FFFEEFF271528FB93EA2D" box="[199,1063,583,608]" pageId="2" pageNumber="56" type="etymology">
<paragraph id="8B83B9759B4FFFEEFF271528FB93EA2D" blockId="2.[151,1437,151,2012]" box="[199,1063,583,608]" pageId="2" pageNumber="56">
<emphasis id="B94865679B4FFFEEFF271528FEFAEA12" bold="true" box="[199,334,584,608]" pageId="2" pageNumber="56">Etymology.</emphasis>
The species is named after the region it occurs, the Persian Gulf.
</paragraph>
</subSubSection>
<subSubSection id="C326EAFE9B4FFFEEFF27150CFA28EFCA" pageId="2" pageNumber="56" type="discussion">
<paragraph id="8B83B9759B4FFFEEFF27150CFDE7EBD6" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="56">
<emphasis id="B94865679B4FFFEEFF27150CFE8FEAF6" bold="true" box="[199,315,620,644]" pageId="2" pageNumber="56">Remarks.</emphasis>
Ng &amp; 
<bibRefCitation id="EFADC4849B4FFFEEFE6C150CFDF9EAF6" box="[396,589,620,644]" pageId="2" pageNumber="56" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella Sankarankutty, 1961, with description of a new mangrove species, Raphidopus johnsoni from Singapore (Decapoda, Anomura). Crustaceana, 66 (1), 1 - 21." type="journal article">Nakasone (1994)</bibRefCitation>
redescribed 
<taxonomicName id="4C3CC2F69B4FFFEEFD3F150CFC74EAF6" box="[735,960,620,644]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFD3F150CFC74EAF6" box="[735,960,620,644]" italics="true" pageId="2" pageNumber="56">Raphidopus ciliatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3CC2F69B4FFFEEFC18150DFBDEEAF6" box="[1016,1130,620,644]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFC18150DFBDEEAF6" box="[1016,1130,620,644]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
in detail and provided figures of both species. The two species are very different, and they even suggested that they can perhaps be separated into two genera. They also described a new species (
<taxonomicName id="4C3CC2F69B4FFFEEFD0415D5FCD0EABE" box="[740,868,692,716]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B94865679B4FFFEEFD0415D5FCD0EABE" box="[740,868,692,716]" italics="true" pageId="2" pageNumber="56">R. johnsoni</emphasis>
</taxonomicName>
) from 
<collectingCountry id="F32BF9E59B4FFFEEFC5215D4FB97EABE" box="[946,1059,692,716]" name="Singapore" pageId="2" pageNumber="56">Singapore</collectingCountry>
that closely resembled 
<taxonomicName id="4C3CC2F69B4FFFEEFACC15D5FA28EABE" box="[1324,1436,692,716]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFACC15D5FA28EABE" box="[1324,1436,692,716]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
but differed mainly in the structure of the third maxillipeds, shape of the chelipeds and relative proportions of the ambulatory legs. The major diagnostic characters of 
<taxonomicName id="4C3CC2F69B4FFFEEFD0C159DFCDDEB66" box="[748,873,765,788]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B94865679B4FFFEEFD0C159DFCDDEB66" box="[748,873,765,788]" italics="true" pageId="2" pageNumber="56">R. persicus</emphasis>
</taxonomicName>
<emphasis id="B94865679B4FFFEEFC91159CFC4FEB66" bold="true" box="[881,1019,763,788]" pageId="2" pageNumber="56">
<taxonomicNameLabel id="A27BD81C9B4FFFEEFC91159CFC4FEB66" box="[881,1019,763,788]" pageId="2" pageNumber="56" rank="species">new species</taxonomicNameLabel>
</emphasis>
, are a lateral portion the of carapace that is rounded and not armed with spines, relatively short third antennal article, form and proportion of the exopod of the third maxilliped, presence of three or four spines on the outer margin of the carpus of the cheliped; relatively long ambulatory legs, and the median part of the thoracic sternum is strongly convex. The differences between the four species are summarised in Table 1.
</paragraph>
<paragraph id="8B83B9759B4FFFEEFF2914CFFF4FEE22" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="56">
The carapace of 
<taxonomicName id="4C3CC2F69B4FFFEEFE6514D1FDB7EBBA" box="[389,515,945,968]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="B94865679B4FFFEEFE6514D1FDB7EBBA" box="[389,515,945,968]" italics="true" pageId="2" pageNumber="56">R. persicus</emphasis>
</taxonomicName>
is superficially very similar to that of 
<taxonomicName id="4C3CC2F69B4FFFEEFC5714D1FB9CEBBA" box="[951,1064,944,968]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFC5714D1FB9CEBBA" box="[951,1064,944,968]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
. However, in the morphology of the chelipeds (especially the serrated carpus) and the elongated ambulatory legs, 
<taxonomicName id="4C3CC2F69B4FFFEEFBCD14B5FB1EEB9E" box="[1069,1194,981,1004]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="B94865679B4FFFEEFBCD14B5FB1EEB9E" box="[1069,1194,981,1004]" italics="true" pageId="2" pageNumber="56">R. persicus</emphasis>
</taxonomicName>
is more like 
<taxonomicName id="4C3CC2F69B4FFFEEFAA114B5FF0CEC62" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFAA114B5FF0CEC62" italics="true" pageId="2" pageNumber="56">R. ciliatus</emphasis>
</taxonomicName>
. In 
<taxonomicName id="4C3CC2F69B4FFFEEFF051499FEE2EC62" box="[229,342,1016,1040]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" sensu="stricto" species="indicus">
<emphasis id="B94865679B4FFFEEFF051499FEE2EC62" box="[229,342,1016,1040]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BD81C9B4FFFEEFEBE1499FE2EEC62" box="[350,410,1017,1040]" pageId="2" pageNumber="56" sensu="stricto">s. str.</taxonomicNameLabel>
(like in 
<taxonomicName id="4C3CC2F69B4FFFEEFE1B1499FDC8EC62" box="[507,636,1016,1040]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B94865679B4FFFEEFE1B1499FDC8EC62" box="[507,636,1016,1040]" italics="true" pageId="2" pageNumber="56">R. johnsoni</emphasis>
</taxonomicName>
), the outer margin of the carpus of the cheliped is entire and unarmed. The ambulatory legs are in addition relatively short in 
<taxonomicName id="4C3CC2F69B4FFFEEFD16137DFCD1EC46" box="[758,869,1052,1076]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFD16137DFCD1EC46" box="[758,869,1052,1076]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
but are proportionately longer in 
<taxonomicName id="4C3CC2F69B4FFFEEFB3D137DFAFAEC46" box="[1245,1358,1052,1076]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFB3D137DFAFAEC46" box="[1245,1358,1052,1076]" italics="true" pageId="2" pageNumber="56">R. ciliatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3CC2F69B4FFFEEFA64137DFF43EC2A" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B94865679B4FFFEEFA64137DFF43EC2A" italics="true" pageId="2" pageNumber="56">R. johnsoni</emphasis>
</taxonomicName>
. As such, in view of the relatively brief description of 
<taxonomicName id="4C3CC2F69B4FFFEEFC921321FC50EC2A" box="[882,996,1088,1112]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFC921321FC50EC2A" box="[882,996,1088,1112]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFADC4849B4FFFEEFBF2135FFF5CEC0E" author="Henderson" pageId="2" pageNumber="56" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnean Society of London, Zoology, (2) 5, 325 - 458, pls. 36 - 40." type="journal article" year="1893">Henderson (1893: 427, pl. 39 figs. 19–22)</bibRefCitation>
, it is easy to confuse 
<taxonomicName id="4C3CC2F69B4FFFEEFE0D1305FDD9EC0E" box="[493,621,1125,1148]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="B94865679B4FFFEEFE0D1305FDD9EC0E" box="[493,621,1125,1148]" italics="true" pageId="2" pageNumber="56">R. persicus</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C3CC2F69B4FFFEEFD531305FC93EC0E" box="[691,807,1124,1148]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFD531305FC93EC0E" box="[691,807,1124,1148]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
. It should be noted that when 
<bibRefCitation id="EFADC4849B4FFFEEFB721304FA28EC0E" author="Haig" box="[1170,1436,1124,1148]" pageId="2" pageNumber="56" refString="Haig, J. (1966) The Porcellanidae (Crustacea Anomura) of the Iranian Gulf and Gulf of Oman. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 129, 49 - 65." type="journal article" year="1966">Haig (1966: 62, fig. 6)</bibRefCitation>
reported and figured the carapace of “ 
<taxonomicName id="4C3CC2F69B4FFFEEFDA613E9FD03ECD2" box="[582,695,1160,1184]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFDA613E9FD03ECD2" box="[582,695,1160,1184]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
” from Kharg I. and Bushire (= Bandar Bushehr) from 
<collectingCountry id="F32BF9E59B4FFFEEFAC713E9FAE0ECD2" box="[1319,1364,1161,1184]" name="Iran" pageId="2" pageNumber="56">Iran</collectingCountry>
in the Persian Gulf, she compared the species at length with 
<taxonomicName id="4C3CC2F69B4FFFEEFD1E13CDFCC4ECB6" box="[766,880,1196,1220]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFD1E13CDFCC4ECB6" box="[766,880,1196,1220]" italics="true" pageId="2" pageNumber="56">R. ciliatus</emphasis>
</taxonomicName>
but made no mention of differences in the structure of the carpus of the cheliped and proportions of the ambulatory legs. This suggests the specimens she had of both species had similar chelipeds and ambulatory legs, and indicates her specimens of “ 
<taxonomicName id="4C3CC2F69B4FFFEEFB611395FB45ED7E" box="[1153,1265,1268,1292]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFB611395FB45ED7E" box="[1153,1265,1268,1292]" italics="true" pageId="2" pageNumber="56">R. indicus</emphasis>
</taxonomicName>
” were actually 
<taxonomicName id="4C3CC2F69B4FFFEEFF771279FEA0ED42" box="[151,276,1305,1328]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="B94865679B4FFFEEFF771279FEA0ED42" box="[151,276,1305,1328]" italics="true" pageId="2" pageNumber="56">R. persicus</emphasis>
</taxonomicName>
instead. The record of 
<taxonomicName id="4C3CC2F69B4FFFEEFDC01279FD27ED42" box="[544,659,1304,1328]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFDC01279FD27ED42" box="[544,659,1304,1328]" italics="true" pageId="2" pageNumber="56">R. ciliatus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFADC4849B4FFFEEFD211278FC66ED42" author="Tirmizi" box="[705,978,1304,1328]" pageId="2" pageNumber="56" refString="Tirmizi, N. M. &amp; Ghani, N. (1994) An Indian Ocean record for a porcellanid crab Raphidopus ciliatus Stimpson, 1858. Pakistan Journal of Marine Sciences, 3 (1), 69 - 72, fig. 1." type="journal article" year="1994">Tirmizi &amp; Ghani (1994)</bibRefCitation>
from Karachi in 
<collectingCountry id="F32BF9E59B4FFFEEFB7A1278FB4DED42" box="[1178,1273,1304,1328]" name="Pakistan" pageId="2" pageNumber="56">Pakistan</collectingCountry>
seems to be a composite. Their description and figures of most of the specimens (
<bibRefCitation id="EFADC4849B4FFFEEFC7B125CFA88ED26" author="Tirmizi" box="[923,1340,1340,1364]" pageId="2" pageNumber="56" refString="Tirmizi, N. M. &amp; Ghani, N. (1994) An Indian Ocean record for a porcellanid crab Raphidopus ciliatus Stimpson, 1858. Pakistan Journal of Marine Sciences, 3 (1), 69 - 72, fig. 1." type="journal article" year="1994">Tirmizi &amp; Ghani 1994: 62 fig. 1A–I</bibRefCitation>
) closely resemble 
<taxonomicName id="4C3CC2F69B4FFFEEFEE41201FEC2ED0A" box="[260,374,1376,1400]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFEE41201FEC2ED0A" box="[260,374,1376,1400]" italics="true" pageId="2" pageNumber="56">R. ciliatus</emphasis>
</taxonomicName>
as currently defined (see Ng &amp; 
<bibRefCitation id="EFADC4849B4FFFEEFD051200FC2CED0A" author="Nakasone" box="[741,920,1376,1400]" pageId="2" pageNumber="56" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella Sankarankutty, 1961, with description of a new mangrove species, Raphidopus johnsoni from Singapore (Decapoda, Anomura). Crustaceana, 66 (1), 1 - 21." type="journal article" year="1994">Nakasone 1994</bibRefCitation>
), but one specimen is problematic. 
<bibRefCitation id="EFADC4849B4FFFEEFACB1200FEECEDEE" author="Tirmizi" pageId="2" pageNumber="56" refString="Tirmizi, N. M. &amp; Ghani, N. (1994) An Indian Ocean record for a porcellanid crab Raphidopus ciliatus Stimpson, 1858. Pakistan Journal of Marine Sciences, 3 (1), 69 - 72, fig. 1." type="journal article" year="1994">Tirmizi &amp; Ghani (1994: 71)</bibRefCitation>
comment that “One female (cl. 
<quantity id="4CC414909B4FFFEEFD2212E4FCAEEDEE" box="[706,794,1412,1436]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="56" unit="mm" value="6.5">6.5 mm</quantity>
) collected on 
<date id="FF829FB59B4FFFEEFC5812E4FB82EDEE" box="[952,1078,1412,1436]" pageId="2" pageNumber="56" value="1992-02-23">23.02.1992</date>
differs from all the other specimens in being less pilose. The carapace and chelipeds have fewer granules. The lateral margins and the oblique posterolateral ridge of the carapace are unarmed (
<figureCitation id="1307A5F09B4FFFEEFD2712ACFCBCED96" box="[711,776,1484,1508]" captionStart="FIGURE 1" captionStartId="4.[151,250,1977,2000]" captionTargetBox="[180,1383,195,1905]" captionTargetId="figure@4.[180,1385,195,1906]" captionTargetPageId="4" captionText="FIGURE 1. Raphidopus persicus new species. A, C, D, holotype male (8.5 × 6.8 mm) (ZMSB), Iran; B, paratype male (7.5 × 5.8 mm) (ZRC), Iraq. A, B, overall dorsal views; C, outer view of right smaller chela; D, dorsal view of carapace (denuded)." httpUri="https://zenodo.org/record/215481/files/figure.png" pageId="2" pageNumber="56">Fig. 1</figureCitation>
J). The specimen has characters which are more like those of 
<taxonomicName id="4C3CC2F69B4FFFEEFF581290FDD3EE7A" authority="Henderson, 1893" authorityName="Henderson" authorityYear="1893" box="[184,615,1520,1544]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFF581290FE2CEE7A" box="[184,408,1520,1544]" italics="true" pageId="2" pageNumber="56">Raphidopus indicus</emphasis>
<bibRefCitation id="EFADC4849B4FFFEEFE421290FDD3EE7A" author="Henderson" box="[418,615,1520,1544]" pageId="2" pageNumber="56" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnean Society of London, Zoology, (2) 5, 325 - 458, pls. 36 - 40." type="journal article" year="1893">Henderson, 1893</bibRefCitation>
</taxonomicName>
. However, we are placing it under 
<taxonomicName id="4C3CC2F69B4FFFEEFBE01291FA84EE7A" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[1024,1328,1520,1544]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFBE01291FBC0EE7A" box="[1024,1140,1520,1544]" italics="true" pageId="2" pageNumber="56">R. ciliatus</emphasis>
<bibRefCitation id="EFADC4849B4FFFEEFB9E1290FA84EE7A" author="Stimpson" box="[1150,1328,1520,1544]" pageId="2" pageNumber="56" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum quoe in Expeditione ad Oceanum Pacificum eptentrionalem a Republica Federata Missa, Cadwaladaro Ringgold et Johann Rodgers Ducibus, observavit et descripsit - Part VII, Crustacea Anomura. Proceedings of the Academy of Natural Sciences, Philadelphia, 10, 225 - 252." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
till more specimens are available.” Their figure of the carapace of this specimen agrees well with what is here defined as 
<taxonomicName id="4C3CC2F69B4FFFEEFA641175FF40EE22" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="B94865679B4FFFEEFA641175FF40EE22" italics="true" pageId="2" pageNumber="56">R. persicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B83B9759B4FFFEEFF29113CFA28EFCA" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="56">
The distribution patterns of the four species are noteworthy. 
<taxonomicName id="4C3CC2F69B4FFFEEFC8C113CFBF8EE06" box="[876,1100,1628,1652]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFC8C113CFBF8EE06" box="[876,1100,1628,1652]" italics="true" pageId="2" pageNumber="56">Raphidopus ciliatus</emphasis>
</taxonomicName>
has only been reliably known from the Indo-West Pacific, reaching as far south as Western 
<collectingCountry id="F32BF9E59B4FFFEEFC82111FFC78EEE5" box="[866,972,1663,1687]" name="Australia" pageId="2" pageNumber="56">Australia</collectingCountry>
, and eastwards to Queensland (
<bibRefCitation id="EFADC4849B4FFFEEFADA111FFEECEECE" author="Grant" pageId="2" pageNumber="56" refString="Grant, F. E. &amp; McCulloch, A. R. (1906) On a collection of Crustacea from the Port Curtis district, Queensland. Proceedings of the Linnean Society of New South Wales, 1906, 2 - 53, pls. 1 - 4." type="proceedings paper" year="1906">Grant &amp; McCulloch 1906</bibRefCitation>
: 42; 
<bibRefCitation id="EFADC4849B4FFFEEFE6F11C4FDBDEECE" author="Haig" box="[399,521,1700,1724]" pageId="2" pageNumber="56" refString="Haig, J. (1965) The Porcellanidae (Crustacea, Anomura) of Western Australia with description of four new Australian species. Journal of the Royal Society of Western Australia, 48, 97 - 117." type="journal article" year="1965">Haig 1965</bibRefCitation>
: 116; Ng &amp; 
<bibRefCitation id="EFADC4849B4FFFEEFD7311C4FCF1EECE" author="Nakasone" box="[659,837,1700,1724]" pageId="2" pageNumber="56" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella Sankarankutty, 1961, with description of a new mangrove species, Raphidopus johnsoni from Singapore (Decapoda, Anomura). Crustaceana, 66 (1), 1 - 21." type="journal article" year="1994">Nakasone 1994</bibRefCitation>
: 3; 
<bibRefCitation id="EFADC4849B4FFFEEFC8E11C4FBEAEECE" author="Osawa" box="[878,1118,1700,1724]" pageId="2" pageNumber="56" refString="Osawa, M. &amp; Chan, T. - Y. (2010) Part III. Porcellanidae (Porcelain crabs). In: Crustacean Fauna of Taiwan: Crab-like anomurans (Hippoidea, Lithodoidea and Porcellanidae). T. - Y. Chan (Ed.), National Taiwan Ocean University, pp. 67 - 182." type="book chapter" year="2010">Osawa &amp; Chan 2010</bibRefCitation>
: 180, Fig. 141, 142). 
<taxonomicName id="4C3CC2F69B4FFFEEFAB011C4FE86EE92" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94865679B4FFFEEFAB011C4FE86EE92" italics="true" pageId="2" pageNumber="56">Raphidopus indicus</emphasis>
</taxonomicName>
was described from Chennai (= Madras), eastern 
<collectingCountry id="F32BF9E59B4FFFEEFC8111A7FC29EEAD" box="[865,925,1735,1759]" name="India" pageId="2" pageNumber="56">India</collectingCountry>
, and the species has since been reported from Penang and along western Peninsular 
<collectingCountry id="F32BF9E59B4FFFEEFDA1118CFD1CEF76" box="[577,680,1772,1796]" name="Malaysia" pageId="2" pageNumber="56">Malaysia</collectingCountry>
(Ng &amp; 
<bibRefCitation id="EFADC4849B4FFFEEFD1E118CFC04EF76" author="Nakasone" box="[766,944,1772,1796]" pageId="2" pageNumber="56" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella Sankarankutty, 1961, with description of a new mangrove species, Raphidopus johnsoni from Singapore (Decapoda, Anomura). Crustaceana, 66 (1), 1 - 21." type="journal article" year="1994">Nakasone 1994</bibRefCitation>
: 7) and Phuket in western 
<collectingCountry id="F32BF9E59B4FFFEEFB39118CFA88EF76" box="[1241,1340,1772,1796]" name="Thailand" pageId="2" pageNumber="56">Thailand</collectingCountry>
(unpublished data). All these locations are in the Gulf of Bengal and Andaman Sea. 
<bibRefCitation id="EFADC4849B4FFFEEFC16106FFB02EF55" author="Haig" box="[1014,1206,1807,1831]" pageId="2" pageNumber="56" refString="Haig, J. (1981) Porcellanid crabs from the Indo-West Pacific, Part II. Steenstrupia, 7 (12), 269 - 291." type="journal article" year="1981">Haig (1981: 275)</bibRefCitation>
reported the species from Surabaya in eastern Java (
<collectingCountry id="F32BF9E59B4FFFEEFE1C1054FDD8EF3E" box="[508,620,1844,1868]" name="Indonesia" pageId="2" pageNumber="56">Indonesia</collectingCountry>
) but this record will need to be verified. 
<taxonomicName id="4C3CC2F69B4FFFEEFBDF1054FA99EF3E" box="[1087,1325,1844,1868]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B94865679B4FFFEEFBDF1054FA99EF3E" box="[1087,1325,1844,1868]" italics="true" pageId="2" pageNumber="56">Raphidopus johnsoni</emphasis>
</taxonomicName>
is known thus far only from 
<collectingCountry id="F32BF9E59B4FFFEEFE871037FE6DEF1D" box="[359,473,1879,1903]" name="Singapore" pageId="2" pageNumber="56">Singapore</collectingCountry>
(Ng &amp; 
<bibRefCitation id="EFADC4849B4FFFEEFDCE1037FD6BEF1D" author="Nakasone" box="[558,735,1879,1903]" pageId="2" pageNumber="56" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella Sankarankutty, 1961, with description of a new mangrove species, Raphidopus johnsoni from Singapore (Decapoda, Anomura). Crustaceana, 66 (1), 1 - 21." type="journal article" year="1994">Nakasone 1994</bibRefCitation>
: 9; Ng 
<emphasis id="B94865679B4FFFEEFCCE1039FCD2EF02" box="[814,870,1880,1904]" italics="true" pageId="2" pageNumber="56">et al.</emphasis>
2011: 27). Although both 
<taxonomicName id="4C3CC2F69B4FFFEEFB6D1039FAB8EF02" box="[1165,1292,1880,1904]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B94865679B4FFFEEFB6D1039FAB8EF02" box="[1165,1292,1880,1904]" italics="true" pageId="2" pageNumber="56">R. johnsoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3CC2F69B4FFFEEFAA21039FF0CEFE6" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B94865679B4FFFEEFAA21039FF0CEFE6" italics="true" pageId="2" pageNumber="56">R. ciliatus</emphasis>
</taxonomicName>
are present in 
<collectingCountry id="F32BF9E59B4FFFEEFEBF101CFE67EFE6" box="[351,467,1916,1940]" name="Singapore" pageId="2" pageNumber="56">Singapore</collectingCountry>
, the latter lives on subtidal soft muddy substrates whereas the first species is an intertidal mangrove species. All the specimens of 
<taxonomicName id="4C3CC2F69B4FFFEEFD7210C1FCBAEFCA" box="[658,782,1953,1976]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="56" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="B94865679B4FFFEEFD7210C1FCBAEFCA" box="[658,782,1953,1976]" italics="true" pageId="2" pageNumber="56">R. persicus</emphasis>
</taxonomicName>
obtained so far have been obtained from subtidal habitats.
</paragraph>
</subSubSection>
<subSubSection id="C326EAFE9B4FFFEEFF2910A3FB24EFAE" box="[201,1168,1987,2012]" pageId="2" pageNumber="56" type="distribution">
<paragraph id="8B83B9759B4FFFEEFF2910A3FB24EFAE" blockId="2.[151,1437,151,2012]" box="[201,1168,1987,2012]" pageId="2" pageNumber="56">
<emphasis id="B94865679B4FFFEEFF2910A3FED6EFAE" bold="true" box="[201,354,1987,2012]" pageId="2" pageNumber="56">Distribution.</emphasis>
The species is so far known only from the Persian Gulf, 
<quantity id="4CC414909B4FFFEEFC0010A4FBF0EFAE" box="[992,1092,1988,2012]" metricMagnitude="1" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="2" pageNumber="56" unit="m" value="18.0" valueMax="19.0" valueMin="17.0">17–19 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>