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<document id="EA77406CCFF2DF3CFF3843C8115DFBFC" ID-DOI="10.59893/bjc.23(2).004" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203251" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657628258" checkinUser="felipe" docAuthor="Minkina, Łukasz" docDate="2023" docId="038D87AAEE00FFA2A1601491FF30F96E" docLanguage="en" docName="BalticJColeopterol.23.2.205-208.pdf" docOrigin="Baltic Journal of Coleopterology (Basel, Switzerland) 23 (2)" docSource="http://dx.doi.org/10.3389/fpsyt.2024.1436142" docStyle="DocumentStyle:514B3A4252435B0AFD496C900C6D7572.1:BalticJColeop.2023-.journal_article" docStyleId="514B3A4252435B0AFD496C900C6D7572" docStyleName="BalticJColeop.2023-.journal_article" docStyleVersion="1" docTitle="Airapus madangensis Minkina, 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="208" masterDocId="FFB4FFD2EE01FFA1A1121637FFAAFFD0" masterDocTitle="A new species of the genusAirapus Stebnicka &amp; Howden, 1996 (Coleoptera: Scarabaeidae: Aphodiinae) from Papua New Guinea" masterLastPageNumber="208" masterPageNumber="205" pageNumber="206" updateTime="1722697181054" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="59300BD79FC4A8EB3068C4F253BE067D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="644CCF6F0A313C4F49FA549F9309E01F">A new species of the genusAirapus Stebnicka &amp; Howden, 1996 (Coleoptera: Scarabaeidae: Aphodiinae) from Papua New Guinea</mods:title>
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<mods:namePart id="66FFABC10918719E52E1FD792A2CE035">Minkina, Łukasz</mods:namePart>
</mods:name>
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<mods:title id="8103040BED2FA20E5D8FDF98AFD5F049">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="CFEF0763A1700CA77B07B763A13BEAE0">
<mods:date id="1649D7002EEE78599DB101F439D91900">2023</mods:date>
<mods:detail id="32C4346447DF57919B9D6459E994A60E" type="series">
<mods:title id="39DC3FF5144DAF307A1E1BD12C4C9F9E">Basel, Switzerland</mods:title>
</mods:detail>
<mods:detail id="FE38D336C2C1831DC46963A92B4D77D2" type="volume">
<mods:number id="F53E89BA953B057BBC09AD5EAC1A6152">23</mods:number>
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<mods:number id="79ECE08FE2BE68BF58188F99D04D85C9">2</mods:number>
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<mods:url id="C3C00D8BD22F783475BBA6542ECDEFFE">http://dx.doi.org/10.3389/fpsyt.2024.1436142</mods:url>
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<mods:classification id="9C3A30348714FF0D83D6B364E3769CC3">journal article</mods:classification>
<mods:identifier id="405BA2B02E7DBF2E9EDCD4AE43A8CC9C" type="DOI">10.59893/bjc.23(2).004</mods:identifier>
<mods:identifier id="3B980089ECC946AE4C3A4BF2F2D9A022" type="ISSN">1407-8619</mods:identifier>
<mods:identifier id="57C771EFC7247A929338B3D095F15475" type="Zenodo-Dep">13203251</mods:identifier>
</mods:mods>
<treatment id="038D87AAEE00FFA2A1601491FF30F96E" LSID="urn:lsid:plazi:treatment:038D87AAEE00FFA2A1601491FF30F96E" httpUri="http://treatment.plazi.org/id/038D87AAEE00FFA2A1601491FF30F96E" lastPageId="3" lastPageNumber="208" pageId="1" pageNumber="206">
<subSubSection id="C33E6537EE00FFA0A1601491FE11FD6D" box="[114,443,678,702]" pageId="1" pageNumber="206" type="nomenclature">
<paragraph id="8B9B36BCEE00FFA0A1601491FE11FD6D" blockId="1.[114,443,678,702]" box="[114,443,678,702]" pageId="1" pageNumber="206">
<heading id="D0D381D0EE00FFA0A1601491FE11FD6D" box="[114,443,678,702]" fontSize="10" level="2" pageId="1" pageNumber="206" reason="7">
<taxonomicName id="4C244D3FEE00FFA0A1601491FEF5FD6E" authorityName="Minkina" authorityYear="2023" box="[114,351,678,702]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="206" phylum="Arthropoda" rank="species" species="madangensis" status="sp. nov.">
<emphasis id="B950EAAEEE00FFA0A1601491FEF5FD6E" bold="true" box="[114,351,678,702]" italics="true" pageId="1" pageNumber="206">Airapus madangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357D5EE00FFA0A0741491FE11FD6D" box="[358,443,678,701]" pageId="1" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E6537EE00FFA0A16014D4FF76FD2A" box="[114,220,738,763]" pageId="1" pageNumber="206" type="description">
<paragraph id="8B9B36BCEE00FFA0A16014D4FF76FD2A" blockId="1.[114,220,738,763]" box="[114,220,738,763]" pageId="1" pageNumber="206">
(
<figureCitation id="131F2A39EE00FFA0A16B14D4FF78FD2A" box="[121,210,738,763]" captionStart-0="Figs 1-3" captionStart-1="Figs 4-5" captionStartId-0="1.[624,670,1480,1504]" captionStartId-1="2.[189,235,1258,1282]" captionTargetBox-0="[628,1108,848,1443]" captionTargetBox-1="[198,672,611,1204]" captionTargetId-0="figure-392@1.[624,1112,846,1443]" captionTargetId-1="figure-391@2.[189,677,600,1218]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs 1-3.Airapus madangensis sp. nov.,♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." captionText-1="Figs 4-5. Airapus madangensis sp. nov.,♀, holotype: 4- head; 5- epipharynx. Fig. 4-5: scale lines: Fig. 4: 1.0 mm; Fig. 5: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13203257" figureDoi-1="http://doi.org/10.5281/zenodo.13203259" httpUri-0="https://zenodo.org/record/13203257/files/figure.png" httpUri-1="https://zenodo.org/record/13203259/files/figure.png" pageId="1" pageNumber="206">Figs 1-5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E6537EE00FFA0A1601517FED9FB78" pageId="1" pageNumber="206" type="materials_examined">
<paragraph id="8B9B36BCEE00FFA0A1601517FE6CFC87" blockId="1.[114,593,800,855]" pageId="1" pageNumber="206">
<emphasis id="B950EAAEEE00FFA0A1601517FEA4FCE8" bold="true" box="[114,270,800,824]" pageId="1" pageNumber="206">Type locality.</emphasis>
<collectingCountry id="F333762CEE00FFA0A0071516FE42FCE8" box="[277,488,800,824]" name="Papua New Guinea" pageId="1" pageNumber="206">Papua New Guinea</collectingCountry>
,
<collectingRegion id="49E0F85EEE00FFA0A0E11517FF06FC86" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="206">Madang prov.</collectingRegion>
, Hansa Bay, Laing Island.
</paragraph>
<paragraph id="8B9B36BCEE00FFA0A3621685FC97FEB2" blockId="1.[624,1103,178,355]" pageId="1" pageNumber="206">coarse, with variable in size, rounded punctures; in the middle very dense, coarse, with somewhat variable in size, longitudinal punctures; anteriorly moderately dense, not so coarse, irregular in size, with rounded punctures.</paragraph>
<paragraph id="8B9B36BCEE00FFA0A36217BFFCF7FCFF" blockId="1.[624,1103,392,815]" pageId="1" pageNumber="206">
Epipharynx (
<figureCitation id="131F2A39EE00FFA0A3EF17BFFCEAFE70" box="[765,832,392,416]" captionStart="Figs 4-5" captionStartId="2.[189,235,1258,1282]" captionTargetBox="[198,672,611,1204]" captionTargetId="figure-391@2.[189,677,600,1218]" captionTargetPageId="2" captionText="Figs 4-5. Airapus madangensis sp. nov.,♀, holotype: 4- head; 5- epipharynx. Fig. 4-5: scale lines: Fig. 4: 1.0 mm; Fig. 5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13203259" httpUri="https://zenodo.org/record/13203259/files/figure.png" pageId="1" pageNumber="206">Fig. 5</figureCitation>
) transverse, with lateral sides broadly rounded, anterior margin of pedia concavely arcuate, corypha below of margin of pedia, indistinct, with two very short, thick celtes. Mesoepitorma shortened basally, somewhat pear-shaped. Acanthopariae with dense, very short and thin chaetae; acropariae with dense, short and thiner chaetae than on acanthopariae; chaetopariae with dense belt of thick, long chaetae; adelochaetae similar to chaetopariae, but less dense; prophobae very short and thin chaetae; chaetopediae without chaetae. Tormae long.
</paragraph>
<paragraph id="8B9B36BCEE00FFA0A160154BFEDEFBDF" blockId="1.[114,593,892,1192]" pageId="1" pageNumber="206">
<emphasis id="B950EAAEEE00FFA0A160154BFE85FC44" bold="true" box="[114,303,892,916]" pageId="1" pageNumber="206">Type material.</emphasis>
<materialsCitation id="3B4C3CE1EE00FFA0A058154BFEDEFBDF" collectingDate="1981-06-08" collectionCode="R, IRSN" collectorName="Coll. I. &amp; Sc. N. B. &amp; W. Laing Island &amp; J. Van Goethem" country="Papua New Guinea" county="Laing Island" location="Bay" municipality="Hansa" pageId="1" pageNumber="206" specimenCount="1" specimenCount-female="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="B950EAAEEE00FFA0A058154BFE12FC44" bold="true" box="[330,440,892,916]" pageId="1" pageNumber="206">
<typeStatus id="549F881EEE00FFA0A058154BFE12FC44" box="[330,440,892,916]" pageId="1" pageNumber="206" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D22FD35EE00FFA0A0C4154BFE40FC43" box="[470,490,892,915]" count="1" pageId="1" pageNumber="206" type="female">♀</specimenCount>
):
<collectorName id="26D1536AEE00FFA0A307154BFF2AFC63" pageId="1" pageNumber="206">Coll. I.</collectorName>
<collectionCode id="ED35AE79EE00FFA0A16D15ACFF3FFC63" box="[127,149,923,947]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="206">R</collectionCode>
.
<collectorName id="26D1536AEE00FFA0A18B15ACFF47FC63" box="[153,237,923,947]" pageId="1" pageNumber="206">Sc.N.B.</collectorName>
|
<collectingCountry id="F333762CEE00FFA0A01315ACFE7BFC63" box="[257,465,923,947]" name="Papua New Guinea" pageId="1" pageNumber="206">Papua New Guinea</collectingCountry>
|
<collectorName id="26D1536AEE00FFA0A0F615ACFF1CFC01" pageId="1" pageNumber="206">
W.
<collectingCounty id="62FA4E30EE00FFA0A31E15ACFF1CFC01" pageId="1" pageNumber="206">Laing Island</collectingCounty>
</collectorName>
,
<collectingMunicipality id="6BFFACC6EE00FFA0A1D4158DFEA3FC01" box="[198,265,954,977]" pageId="1" pageNumber="206">Hansa</collectingMunicipality>
|
<location id="8EFB6067EE00FFA0A035158DFEFEFC02" LSID="urn:lsid:plazi:treatment:038D87AAEE00FFA2A1601491FF30F96E:8EFB6067EE00FFA0A035158DFEFEFC02" box="[295,340,954,978]" country="Papua New Guinea" county="Laing Island" municipality="Hansa" name="Bay" pageId="1" pageNumber="206" stateProvince="Madang">Bay</location>
,
<collectingRegion id="49E0F85EEE00FFA0A077158EFDAAFC01" box="[357,512,953,977]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="206">Madang prov.</collectingRegion>
|
<location id="8EFB6067EE00FFA0A30C158DFDFBFC01" LSID="urn:lsid:plazi:treatment:038D87AAEE00FFA2A1601491FF30F96E:8EFB6067EE00FFA0A30C158DFDFBFC01" box="[542,593,954,977]" country="Papua New Guinea" county="Laing Island" municipality="Hansa" name="Wet" pageId="1" pageNumber="206" stateProvince="Madang">Wet</location>
leaf litter | I.G.: 26373 |
<date id="FF9A107CEE00FFA0A09B15EFFE44FC20" box="[393,494,984,1008]" pageId="1" pageNumber="206" value="1981-06-08">
<collectingDate id="EFDEE994EE00FFA0A09B15EFFE44FC20" box="[393,494,984,1008]" pageId="1" pageNumber="206" value="1981-06-08">8.vi.1981</collectingDate>
</date>
| leg.
<collectorName id="26D1536AEE00FFA0A32B15EEFEADFBDF" pageId="1" pageNumber="206">J. Van Goethem</collectorName>
|| (
<collectionCode id="ED35AE79EE00FFA0A03415C0FEC2FBDF" box="[294,360,1015,1039]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="206">IRSN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9B36BCEE00FFA0A1601222FED9FB78" blockId="1.[114,593,892,1192]" pageId="1" pageNumber="206">
<materialsCitation id="3B4C3CE1EE00FFA0A1601222FE27FBFE" box="[114,397,1045,1070]" collectionCode="R" collectorName="Coll. I." pageId="1" pageNumber="206" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B950EAAEEE00FFA0A1601222FF70FBFD" bold="true" box="[114,218,1045,1069]" pageId="1" pageNumber="206">
<typeStatus id="549F881EEE00FFA0A1601222FF70FBFD" box="[114,218,1045,1069]" pageId="1" pageNumber="206" type="paratype">Paratype</typeStatus>
</emphasis>
(
<specimenCount id="9D22FD35EE00FFA0A1E01221FEACFBFD" box="[242,262,1046,1069]" count="1" pageId="1" pageNumber="206" type="female">♀</specimenCount>
):
<collectorName id="26D1536AEE00FFA0A0371222FEDDFBFE" box="[293,375,1045,1070]" pageId="1" pageNumber="206">Coll. I.</collectorName>
<collectionCode id="ED35AE79EE00FFA0A0651221FE27FBFE" box="[375,397,1046,1070]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="206">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B4C3CE1EE00FFA0A0821221FEC5FB78" collectingDate="1978-05-09" collectionCode="ISEA" collectorName="Sc. N. B. &amp; J. Van Goethem" country="Papua New Guinea" location="Hills" pageId="1" pageNumber="206" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<collectorName id="26D1536AEE00FFA0A0821221FE4FFBFE" box="[400,485,1046,1070]" pageId="1" pageNumber="206">Sc.N.B.</collectorName>
|
<collectingCountry id="F333762CEE00FFA0A31B1221FEABFB9C" name="Papua New Guinea" pageId="1" pageNumber="206">Papua New Guinea</collectingCountry>
|
<location id="8EFB6067EE00FFA0A0341203FEF0FB9C" LSID="urn:lsid:plazi:treatment:038D87AAEE00FFA2A1601491FF30F96E:8EFB6067EE00FFA0A0341203FEF0FB9C" box="[294,346,1076,1100]" country="Papua New Guinea" name="Hills" pageId="1" pageNumber="206" stateProvince="Madang">Hills</location>
<locationDeviation id="9A292F9EEE00FFA0A0781203FD9FFB9C" box="[362,565,1076,1100]" location="Bogia" pageId="1" pageNumber="206">
<quantity id="4CDC9B59EE00FFA0A0781203FE03FB9C" box="[362,425,1076,1100]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="206" unit="km" value="4.0">4 km</quantity>
NW Bogia
</locationDeviation>
|
<collectingRegion id="49E0F85EEE00FFA0A1601264FEA1FBBA" box="[114,267,1107,1131]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="206">Madang prov.</collectingRegion>
, forest | gallery, wet leaf litter | near tree, I.G.: 25848 |
<date id="FF9A107CEE00FFA0A09C1246FE41FB59" box="[398,491,1137,1161]" pageId="1" pageNumber="206" value="1978-05-09">
<collectingDate id="EFDEE994EE00FFA0A09C1246FE41FB59" box="[398,491,1137,1161]" pageId="1" pageNumber="206" value="1978-05-09">9.v.1978</collectingDate>
</date>
| leg.
<collectorName id="26D1536AEE00FFA0A32B1245FEADFB78" pageId="1" pageNumber="206">J. Van Goethem</collectorName>
|| (
<collectionCode id="ED35AE79EE00FFA0A03412A7FECDFB78" box="[294,359,1168,1192]" pageId="1" pageNumber="206">ISEA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E6537EE00FFA3A16012FAFB8FFA01" lastPageId="2" lastPageNumber="207" pageId="1" pageNumber="206" type="description">
<paragraph id="8B9B36BCEE00FFA0A16012FAFD99FAF3" blockId="1.[114,593,1229,1315]" pageId="1" pageNumber="206">
<emphasis id="B950EAAEEE00FFA0A16012FAFEA9FB35" bold="true" box="[114,259,1229,1253]" pageId="1" pageNumber="206">Description.</emphasis>
Dorsum (
<figureCitation id="131F2A39EE00FFA0A07E12FAFE01FB36" box="[364,427,1229,1254]" captionStart="Figs 1-3" captionStartId="1.[624,670,1480,1504]" captionTargetBox="[628,1108,848,1443]" captionTargetId="figure-392@1.[624,1112,846,1443]" captionTargetPageId="1" captionText="Figs 1-3.Airapus madangensis sp. nov.,♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203257" httpUri="https://zenodo.org/record/13203257/files/figure.png" pageId="1" pageNumber="206">Fig. 1</figureCitation>
). Body length
<quantity id="4CDC9B59EE00FFA0A16012DBFF6CFAD3" box="[114,198,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="206" unit="mm" value="5.7">5.7 mm</quantity>
, elongate, shiny, blackish, sides and apex of elytra with distinct macrosetation.
</paragraph>
<paragraph id="8B9B36BCEE00FFA0A160137FFDFBF9A4" blockId="1.[114,593,1352,1652]" pageId="1" pageNumber="206">
Head (
<figureCitation id="131F2A39EE00FFA0A1AF137FFEA8FAB0" box="[189,258,1352,1376]" captionStart="Figs 4-5" captionStartId="2.[189,235,1258,1282]" captionTargetBox="[198,672,611,1204]" captionTargetId="figure-391@2.[189,677,600,1218]" captionTargetPageId="2" captionText="Figs 4-5. Airapus madangensis sp. nov.,♀, holotype: 4- head; 5- epipharynx. Fig. 4-5: scale lines: Fig. 4: 1.0 mm; Fig. 5: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13203259" httpUri="https://zenodo.org/record/13203259/files/figure.png" pageId="1" pageNumber="206">Fig. 4</figureCitation>
) relatively large, trapezoidal, convex, shiny, without microreticulation. Clypeus gently bordered, widely sinuate anteriorly, denticulate on sides, widely rounded laterally, not notched before genae, clypeal border without macrosetae. Genae obtuse, very distinctly exceeding eyes, with few very short, thin macrosetae. Frontal suture not noticeable, without gibbosities. Punctation simple: basally very dense, quite
</paragraph>
<caption id="DF5B6634EE00FFA0A36213FFFB9CF9CD" ID-DOI="http://doi.org/10.5281/zenodo.13203257" ID-Zenodo-Dep="13203257" httpUri="https://zenodo.org/record/13203257/files/figure.png" pageId="1" pageNumber="206" startId="1.[624,670,1480,1504]" targetBox="[628,1108,848,1443]" targetPageId="1" targetType="figure">
<paragraph id="8B9B36BCEE00FFA0A36213FFFB9CF9CD" blockId="1.[624,1103,1480,1565]" pageId="1" pageNumber="206">
Figs 1-3.
<taxonomicName id="4C244D3FEE00FFA0A3C413FFFC17FA30" authorityName="Minkina" authorityYear="2023" box="[726,957,1480,1504]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="206" phylum="Arthropoda" rank="species" species="madangensis" status="sp. nov.">
<emphasis id="B950EAAEEE00FFA0A3C413FFFC17FA30" box="[726,957,1480,1504]" italics="true" pageId="1" pageNumber="206">Airapus madangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357D5EE00FFA0A2DE13FEFB82FA30" box="[972,1064,1481,1504]" pageId="1" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
,♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B9B36BCEE00FFA3A1601091FD2CFE6E" blockId="1.[114,154,1702,1726]" lastBlockId="2.[189,669,178,447]" lastPageId="2" lastPageNumber="207" pageId="1" pageNumber="206">206 Pronotum transverse, approximately as wide as base of elytra, widest in the middle, convex, shiny, without microreticulation, with simple, quite regularly distributed punctation, punctures are coarser on disc, quite irregular in shape, mainly rounded. Base not distinctly bordered by a groove, anteriorly and on sides not bordered. Sides and base moderately long, quite thick macrosetae. Anterior angles widely rounded. Base before hind angles truncate.</paragraph>
<paragraph id="8B9B36BCEE03FFA3A1AF17D3FDB6FDEA" blockId="2.[189,668,484,570]" pageId="2" pageNumber="207">Scutellum small, triangular, with ogival sides, with few small punctures, moderately shiny, without microreticulation.</paragraph>
<caption id="DF5B6634EE03FFA3A1AF12DDFD20FAEE" ID-DOI="http://doi.org/10.5281/zenodo.13203259" ID-Zenodo-Dep="13203259" httpUri="https://zenodo.org/record/13203259/files/figure.png" pageId="2" pageNumber="207" startId="2.[189,235,1258,1282]" targetBox="[198,672,611,1204]" targetPageId="2" targetType="figure">
<paragraph id="8B9B36BCEE03FFA3A1AF12DDFD20FAEE" blockId="2.[189,668,1257,1343]" pageId="2" pageNumber="207">
Figs 4-5.
<taxonomicName id="4C244D3FEE03FFA3A03612DDFDA1FAD2" authorityName="Minkina" authorityYear="2023" box="[292,523,1258,1282]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="madangensis" status="sp. nov.">
<emphasis id="B950EAAEEE03FFA3A03612DDFDA1FAD2" box="[292,523,1258,1282]" italics="true" pageId="2" pageNumber="207">Airapus madangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357D5EE03FFA3A30B12DDFDDCFAD1" box="[537,630,1258,1281]" pageId="2" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
,♀, holotype: 4- head; 5- epipharynx. Fig. 4-5: scale lines: Fig. 4: 1.0 mm; Fig. 5: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B9B36BCEE03FFA3A1AF1353FBFDFE70" blockId="2.[189,668,1380,1650]" lastBlockId="2.[699,1179,178,416]" pageId="2" pageNumber="207">Elytra elongate, convex, parallel, shiny, with weak microreticulation (which is more distinct on sides and before apex), with short and thin macrosetae on sides and before apex; with small indistinct humeral denticles; with ten striae and ten intervals. Intervals shiny, convex, with quite regularly distributed, simple, coarse punctation; part of punctures nearby sides connected with punctures of striae and because of it striae seems indenting intervals margins here very distinctly; on sides because of that structure intervals seems to be somewhat tuberculate. Striae distinctly, quite densely punctate, with medium sized punctures; punctures distinctly indenting margins of intervals. All striae joined together before apex.</paragraph>
<paragraph id="8B9B36BCEE03FFA3A3A917F1FC1BFC18" blockId="2.[699,1179,453,968]" pageId="2" pageNumber="207">Legs. Femora shiny, without microreticulation, moderately coarsely and sparsely punctate, with punctures bearing very short macrosetae. Protibiae distinctly tridentate laterally, proximally not serrulate; dorsal side smooth, shiny, with few very fine punctures; apical spur long, moderately broad, gently downwardly and inwardly bent, with apex rounded. Meso- and metatibiae fimbriate apically with row of short spinules of equal length. Apex of metatibiae without accessory spine. Metatibiae superior apical spur very slightly shorter than basimetatarsomere, latter approximately as long as three of next metatarsomeres combined. Claws short, thin, distinctly arcuate.</paragraph>
<paragraph id="8B9B36BCEE03FFA3A3A915D9FCFDFBD5" blockId="2.[699,855,1006,1029]" box="[699,855,1006,1029]" pageId="2" pageNumber="207">Macropterous.</paragraph>
<paragraph id="8B9B36BCEE03FFA3A3A9121CFB8FFA01" blockId="2.[699,1178,1067,1489]" pageId="2" pageNumber="207">
Venter (
<figureCitation id="131F2A39EE03FFA3A20C121CFCC6FB93" box="[798,876,1067,1091]" captionStart="Figs 1-3" captionStartId="1.[624,670,1480,1504]" captionTargetBox="[628,1108,848,1443]" captionTargetId="figure-392@1.[624,1112,846,1443]" captionTargetPageId="1" captionText="Figs 1-3.Airapus madangensis sp. nov.,♀, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Figs 1-3: scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203257" httpUri="https://zenodo.org/record/13203257/files/figure.png" pageId="2" pageNumber="207">Fig. 2</figureCitation>
). Meso-metaventral plate shiny, weakly convex, with distinct, shallow, quite narrow longitudinal furrow in the middle; furrow with deeper part nearby base and nearby apex, additionaly with reversed-heart shaped concavity before apex; surface with moderately coarse, sparse punctures. Abdominal ventrites moderately shiny, anteriorly weakly fluted, with weak microreticulation, with sparse, moderately coarsely, irregularly spaced punctatures; all punctures bearing very short and quite thin macrosetae. Pygidium with similar structure to ventrites.
</paragraph>
</subSubSection>
<subSubSection id="C33E6537EE03FFA2A3A913C0FF30F96E" lastPageId="3" lastPageNumber="208" pageId="2" pageNumber="207" type="etymology">
<paragraph id="8B9B36BCEE03FFA3A3A913C0FBE3F99C" blockId="2.[699,1179,1527,1612]" pageId="2" pageNumber="207">
<emphasis id="B950EAAEEE03FFA3A3A913C0FCEEF9DF" bold="true" box="[699,836,1527,1551]" pageId="2" pageNumber="207">Etymology.</emphasis>
Toponymic; an adjective derived from the name of
<collectingRegion id="49E0F85EEE03FFA3A2C01022FB3CF9FD" box="[978,1174,1557,1581]" country="Papua New Guinea" name="Madang" pageId="2" pageNumber="207">Madang province</collectingRegion>
, where the new species was collected.
</paragraph>
<paragraph id="8B9B36BCEE03FFA3A57E1091FB3EF96E" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="207">207</paragraph>
<paragraph id="8B9B36BCEE02FFA2A1601686FDA7FF38" blockId="3.[114,593,177,232]" pageId="3" pageNumber="208">
<emphasis id="B950EAAEEE02FFA2A1601686FF5FFF19" bold="true" box="[114,245,177,201]" pageId="3" pageNumber="208">Variability.</emphasis>
Body length:
<quantity id="4CDC9B59EE02FFA2A0B81686FDE7FF19" box="[426,589,177,201]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="5.7" metricValueMin="5.6" pageId="3" pageNumber="208" unit="mm" value="5.65" valueMax="5.7" valueMin="5.6">5.6 – 5.7mm</quantity>
. Punctation of body is weakly variable.
</paragraph>
<paragraph id="8B9B36BCEE02FFA2A160173AFE67FEF5" blockId="3.[114,461,269,293]" box="[114,461,269,293]" pageId="3" pageNumber="208">
<emphasis id="B950EAAEEE02FFA2A160173AFEFCFEF5" bold="true" box="[114,342,269,293]" pageId="3" pageNumber="208">Sexual dimorphism.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B9B36BCEE02FFA2A160177CFEC7F9BB" blockId="3.[114,593,331,1643]" pageId="3" pageNumber="208">
<emphasis id="B950EAAEEE02FFA2A160177CFF72FEB3" bold="true" box="[114,216,331,355]" pageId="3" pageNumber="208">Affinity.</emphasis>
Newly described species at first glance, due to relatively regularly convex body, not very distinctly deplanate anterior angles of the pronotum and lack of typical deplanations on sides of the pronotum, as well as the relatively normally developed elytral intervals with macrosetae only on the sides and before apex, seems to be more similar to the genus
<taxonomicName id="4C244D3FEE02FFA2A0421477FDE7FD88" authority="Harold, 1867" authorityName="Harold" authorityYear="1867" box="[336,589,576,600]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAAEEE02FFA2A0421477FE07FD88" box="[336,429,576,600]" italics="true" pageId="3" pageNumber="208">Ataenius</emphasis>
Harold, 1867
</taxonomicName>
. Besides, large distance between mesocoxae, typical shape of meso-metaventral plate, denticulate clypeus and typical punctation of head definitely place newly described species in the genus
<taxonomicName id="4C244D3FEE02FFA2A07214EDFE8BFCC0" authority="Stebnicka &amp; Howden, 1996" authorityName="Stebnicka &amp; Howden" authorityYear="1996" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAAEEE02FFA2A07214EDFE1FFD22" box="[352,437,730,754]" italics="true" pageId="3" pageNumber="208">Airapus</emphasis>
Stebnicka &amp; Howden, 1996
</taxonomicName>
.
<taxonomicName id="4C244D3FEE02FFA2A03F14CFFDB6FCC0" authorityName="Minkina" authorityYear="2023" box="[301,540,760,784]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="madangensis" status="sp. nov.">
<emphasis id="B950EAAEEE02FFA2A03F14CFFDB6FCC0" box="[301,540,760,784]" italics="true" pageId="3" pageNumber="208">Airapus madangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357D5EE02FFA2A33E14CEFF0BFCFE" pageId="3" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from all other known species due to unique structure of elytral interals (quite smooth on the disc, slightly granulate on the sides; with characteristic punctation); additionaly it cannot be including to any group of species created by
<bibRefCitation id="EFB54B4DEE02FFA2A1FB15F8FE0BFC37" author="Stebnicka Z." box="[233,417,975,999]" pageId="3" pageNumber="208" pagination="833 - 895" refId="ref2021" refString="Stebnicka Z. 1998. The Aphodiinae of New Guinea, the Bismarck Archipelago and the Solomon Islands Invertebrate Taxonomy 12 (5): 833 - 895." type="journal article" year="1998">Stebnicka (1998)</bibRefCitation>
and due of it is easily distinguishable from other species from
<collectingCountry id="F333762CEE02FFA2A1A3123AFE20FBF4" box="[177,394,1036,1060]" name="Papua New Guinea" pageId="3" pageNumber="208">Papua New Guinea</collectingCountry>
. Additional body composites that allow it to be distinguished from other species are shape of meso-metaventral plate and shape of scutellum – both of which are distinctive and unique among different species of the genus. In any case, the two features mentioned need to be studied in the future in different manuscript, as they are not easy to determine (unique shapes of concavities before apex of meso-metaventral plates, unique clustering of scutellum punctation, etc.), and also – with this features it will probably be easier in the future to determine the relationship of affinities between species and possible new genera (as
<taxonomicName id="4C244D3FEE02FFA2A1BC13C0FEA9F9DF" authorityName="Stebnicka &amp; Howden" authorityYear="1996" box="[174,259,1527,1551]" class="Insecta" family="Scarabaeidae" genus="Airapus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAAEEE02FFA2A1BC13C0FEA9F9DF" box="[174,259,1527,1551]" italics="true" pageId="3" pageNumber="208">Airapus</emphasis>
</taxonomicName>
or even
<taxonomicName id="4C244D3FEE02FFA2A0B613C0FDABF9DF" authorityName="Harold" authorityYear="1867" box="[420,513,1527,1551]" class="Insecta" family="Scarabaeidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAAEEE02FFA2A0B613C0FDABF9DF" box="[420,513,1527,1551]" italics="true" pageId="3" pageNumber="208">Ataenius</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFB54B4DEE02FFA2A1601022FE84F9FE" author="Stebnicka Z." box="[114,302,1557,1582]" pageId="3" pageNumber="208" pagination="1 - 155" refId="ref2050" refString="Stebnicka Z. 2007. The genus Ataenius Harold, 1867 of New World. Iconography. Krakow: Institute of Systematics and Evolution of Animals Polish Academy of Sciences: 1 - 155." type="book chapter" year="2007">Stebnicka, 2007</bibRefCitation>
) probably paraphyletic, due to the presence of unique features between some species).
</paragraph>
<paragraph id="8B9B36BCEE02FFA2A1601091FF30F96E" blockId="3.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="3" pageNumber="208">208</paragraph>
</subSubSection>
</treatment>
</document>
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