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<document id="0FE5F35803920BC32FFFE8B06DB67F01" ID-CLB-Dataset="46207" ID-DOI="10.11646/zootaxa.3779.4.4" ID-GBIF-Dataset="c325c7c0-f7c1-4e82-8fcd-cb3eb2c4da3c" ID-ISSN="1175-5326" ID-Zenodo-Dep="229347" ID-ZooBank="A5762089-AADF-41B6-B9E9-70C0371C30C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460397980861" checkinUser="plazi" docAuthor="Jaschhof, Mathias" docDate="2014" docId="038D879309662002FF2AFE24FBF1F9C8" docLanguage="en" docName="zt03779p469.pdf" docOrigin="Zootaxa 3779 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Feltomyina polymera Alexander 1936" docType="treatment" docVersion="9" lastPageNumber="467" masterDocId="FFB4FFEB09622006FFBDFF93FFC0FFF8" masterDocTitle="A revision of the types of Neotropical Porricondylinae (Diptera: Cecidomyiidae)" masterLastPageNumber="469" masterPageNumber="463" pageNumber="467" updateTime="1698298894689" updateUser="ExternalLinkService">
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<mods:title id="4706FCB1E6723330B3488D0303CAACAE">A revision of the types of Neotropical Porricondylinae (Diptera: Cecidomyiidae)</mods:title>
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<mods:namePart id="5A5306EA84F031CED2DFB2290D180264">Jaschhof, Mathias</mods:namePart>
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<mods:date id="6294930CE10F6A4604A970EA58C5594A">2014</mods:date>
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<treatment id="038D879309662002FF2AFE24FBF1F9C8" ID-DOI="http://doi.org/10.5281/zenodo.6142924" ID-GBIF-Taxon="119418676" ID-Zenodo-Dep="6142924" LSID="urn:lsid:plazi:treatment:038D879309662002FF2AFE24FBF1F9C8" httpUri="http://treatment.plazi.org/id/038D879309662002FF2AFE24FBF1F9C8" lastPageNumber="467" pageId="4" pageNumber="467">
<subSubSection id="C33E650E09662002FF2AFE24FBBCFDC2" pageId="4" pageNumber="467" type="nomenclature">
<paragraph id="8B9B368509662002FF2AFE24FDA7FE28" blockId="4.[151,615,439,501]" box="[151,615,439,464]" pageId="4" pageNumber="467">
<heading id="D0D381E909662002FF2AFE24FDA7FE28" bold="true" box="[151,615,439,464]" fontSize="10" level="2" pageId="4" pageNumber="467" reason="2">
<taxonomicName id="4C244D0609662002FF2AFE24FDA7FE28" ID-CoL="3DY49" authority="Alexander, 1936" authorityName="Alexander" authorityYear="1936" box="[151,615,439,464]" class="Insecta" family="Cecidomyiidae" genus="Feltomyina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="polymera">
<emphasis id="B950EA9709662002FF2AFE24FDA7FE28" bold="true" box="[151,615,439,464]" pageId="4" pageNumber="467">
<emphasis id="B950EA9709662002FF2AFE24FE4EFE28" bold="true" box="[151,398,439,464]" italics="true" pageId="4" pageNumber="467">Feltomyina polymera</emphasis>
(
<bibRefCitation id="EFB54B7409662002FE26FE24FD9EFE28" author="Alexander" box="[411,606,439,464]" pageId="4" pageNumber="467" refString="Alexander, C. P. (1936) The maximum number of antennal segments in the order Diptera, with the description of a new genus of Cecidomyiidae. Bulletin of the Brooklyn Entomological Society, 31, 12 - 14." type="journal article" year="1936">Alexander, 1936</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9B368509662002FF2AFE4FFF38FE0C" blockId="4.[151,615,439,501]" box="[151,248,476,501]" pageId="4" pageNumber="467">
(
<figureCitation id="131F2A0009662002FF22FE4FFF31FE0C" box="[159,241,476,501]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1396,1419]" captionTargetBox="[341,1220,324,1319]" captionTargetId="figure@5.[264,1323,265,1394]" captionTargetPageId="5" captionText="FIGURES 7 11. Morphology of male Holoneurus occidentalis (7 9, 11) and female Feltomyina polymera (10). 7: genitalia, ventral. 8: gonostylus, dorsal. 9: fourth flagellomere, lateral. 10: ditto. 11: parameres and copulatory organ, ventral. Scale 0.05 mm. agd = accessory gland duct, p = paramere." httpUri="https://zenodo.org/record/229349/files/figure.png" pageId="4" pageNumber="467">Fig. 10</figureCitation>
)
</paragraph>
<paragraph id="8B9B368509662002FF2AFDB7FBBCFDC2" blockId="4.[151,1148,548,570]" box="[151,1148,548,570]" pageId="4" pageNumber="467">
<bibRefCitation id="EFB54B7409662002FF2AFDB7FEFDFDC2" author="Alexander" box="[151,317,548,570]" pageId="4" pageNumber="467" refString="Alexander, C. P. (1936) The maximum number of antennal segments in the order Diptera, with the description of a new genus of Cecidomyiidae. Bulletin of the Brooklyn Entomological Society, 31, 12 - 14." type="journal article" year="1936">Alexander 1936</bibRefCitation>
: 12, originally described as
<taxonomicName id="4C244D0609662002FDE5FDB7FD7EFDC2" box="[600,702,548,570]" class="Insecta" family="Cecidomyiidae" genus="Feltomyia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9709662002FDE5FDB7FD7EFDC2" box="[600,702,548,570]" italics="true" pageId="4" pageNumber="467">Feltomyia</emphasis>
</taxonomicName>
, a preoccupied name (see
<bibRefCitation id="EFB54B7409662002FC76FDB7FBB0FDC2" author="Alexander" box="[971,1136,548,570]" pageId="4" pageNumber="467" refString="Alexander, C. P. (1937) Change of name in Diptera. Bulletin of the Brooklyn Entomological Society, 32, 60." type="journal article" year="1937">Alexander 1937</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E650E09662002FF2AFDF5FAF8FD87" box="[151,1336,614,639]" pageId="4" pageNumber="467" type="materials_examined">
<paragraph id="8B9B368509662002FF2AFDF5FAF8FD87" blockId="4.[151,1436,614,1584]" box="[151,1336,614,639]" pageId="4" pageNumber="467">
<typeStatus id="549F882709662002FF2AFDF5FEC4FD87" box="[151,260,614,639]" pageId="4" pageNumber="467" type="holotype">
<emphasis id="B950EA9709662002FF2AFDF5FEC4FD87" bold="true" box="[151,260,614,639]" pageId="4" pageNumber="467">Holotype</emphasis>
</typeStatus>
. Female,
<collectingCountry id="F333761509662002FECFFDF4FE0EFD86" box="[370,462,615,638]" name="Panama" pageId="4" pageNumber="467">Panama</collectingCountry>
, Chiriquí, Potrerillos, at about
<quantity id="4CDC9B6009662002FC9AFDF4FCB0FD86" box="[807,880,615,639]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="467" unit="m" value="900.0">900 m</quantity>
,
<date id="FF9A104509662002FCC1FDF5FBC6FD87" box="[892,1030,614,639]" pageId="4" pageNumber="467" value="1935-05-06">6 May 1935</date>
, J.W. MacSwain (
<collectionCode id="ED35AE4009662002FB72FDF4FAEBFD87" box="[1231,1323,615,639]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="467">NMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E650E09662002FF7AFD19FC65FD13" pageId="4" pageNumber="467" type="discussion">
<paragraph id="8B9B368509662002FF7AFD19FC65FD13" blockId="4.[151,1436,614,1584]" pageId="4" pageNumber="467">
<emphasis id="B950EA9709662002FF7AFD19FE87FD5B" box="[199,327,650,675]" italics="true" pageId="4" pageNumber="467">Remarks. T</emphasis>
he previously dry specimen is now mounted on a microscope slide in
<collectingCountry id="F333761509662002FBDEFD19FB77FD5B" box="[1123,1207,650,675]" name="Canada" pageId="4" pageNumber="467">Canada</collectingCountry>
balsam. It is almost complete; one antenna, one wing, and a foreleg are missing. The remaining antenna, wing, and legs from one side were separated from the body and mounted under separate coverslips.
</paragraph>
</subSubSection>
<subSubSection id="C33E650E09662002FF7AFD65FBF1F9C8" pageId="4" pageNumber="467" type="description">
<paragraph id="8B9B368509662002FF7AFD65FB24FAA0" blockId="4.[151,1436,614,1584]" pageId="4" pageNumber="467">
<emphasis id="B950EA9709662002FF7AFD65FE3FFCF7" bold="true" box="[199,511,758,783]" pageId="4" pageNumber="467">Female description. Head.</emphasis>
Eye bridge dorsally 1011 ommatidia long. Antenna shorter than body. Scape and pedicel much darker than flagellum; scape broader and twice length of pedicel, both setose. Around 60 flagellomeres (63 according to
<bibRefCitation id="EFB54B7409662002FE44FCADFD0DFCAF" author="Alexander" box="[505,717,830,855]" pageId="4" pageNumber="467" refString="Alexander, C. P. (1936) The maximum number of antennal segments in the order Diptera, with the description of a new genus of Cecidomyiidae. Bulletin of the Brooklyn Entomological Society, 31, 12 - 14." type="journal article" year="1936">Alexander (1936))</bibRefCitation>
, in the one antenna retained, flagellomeres beyond the 50th are shriveled and apparently partly fused with each other; necks short, with microtrichia; nodes subglobular, 1 whorl of setae basally, usually 13 additional setae ventromedially, no sensory hairs, circumfila consisting of two rings interconnected by two longitudinal threads (
<figureCitation id="131F2A0009662002FD2EFC39FD2AFC3B" box="[659,746,938,963]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1396,1419]" captionTargetBox="[341,1220,324,1319]" captionTargetId="figure@5.[264,1323,265,1394]" captionTargetPageId="5" captionText="FIGURES 7 11. Morphology of male Holoneurus occidentalis (7 9, 11) and female Feltomyina polymera (10). 7: genitalia, ventral. 8: gonostylus, dorsal. 9: fourth flagellomere, lateral. 10: ditto. 11: parameres and copulatory organ, ventral. Scale 0.05 mm. agd = accessory gland duct, p = paramere." httpUri="https://zenodo.org/record/229349/files/figure.png" pageId="4" pageNumber="467">Fig. 10</figureCitation>
). Palpus vestigial, 2 short, largely fused segments.
<emphasis id="B950EA9709662002FA81FC39FA5CFC3B" bold="true" box="[1340,1436,938,963]" pageId="4" pageNumber="467">Thorax.</emphasis>
Number of setae on anepisternum, 11; preepisternum 2, 10; anepimeron, 8; metepisternum, 3.
<emphasis id="B950EA9709662002FB0DFC5DFB2FFC1F" bold="true" box="[1200,1263,974,999]" pageId="4" pageNumber="467">Legs.</emphasis>
Conspicuously bicolored due to alternating sections bearing either light or dark setae. Foreleg: femur completely dark, tibia subbasally and subapically dark, second tarsomere basally dark. Midleg: femur dorsally dark on distal three fifths, otherwise as foreleg. Hindleg: tibia dark except for base and apex. Basitarsi of all legs with strong spine. Claws very slightly bent, 1 large tooth basally. Empodia rudimentary.
<emphasis id="B950EA9709662002FCCEFBCDFC7BFB8F" bold="true" box="[883,955,1118,1143]" pageId="4" pageNumber="467">Wing.</emphasis>
Length 2.8 × width. Sc distinct, long; R1 long; R5 joining C beyond apex of wing; Rs somewhat longer than in other
<taxonomicName id="4C244D0609662002FC42FB16FBBDFB66" box="[1023,1149,1157,1182]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
, strongly oblique, in line with R5; r-m+m-cu moderately curved; M absent; CuA two-branched, CuA1 weak, separate from CuA2, CuA2 apically strongly curved.
<emphasis id="B950EA9709662002FE0AFB40FD96FB14" bold="true" box="[439,598,1235,1260]" pageId="4" pageNumber="467">Preabdomen.</emphasis>
Tergites IV and sternites IIVIII entire, with large, randomly distributed setae, tergites VIVIII with fewer, smaller setae and dense transverse rows of microtrichia. Pleural membrane setose.
<emphasis id="B950EA9709662002FF50FA88FEB7FACC" bold="true" box="[237,375,1307,1332]" pageId="4" pageNumber="467">Terminalia.</emphasis>
Ovipositor as in
<taxonomicName id="4C244D0609662002FDFBFA8FFD6EFACB" box="[582,686,1308,1331]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9709662002FDFBFA8FFD6EFACB" box="[582,686,1308,1331]" italics="true" pageId="4" pageNumber="467">Asynapta</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB54B7409662002FD7CFA88FBB9FACC" author="Jaschhof" box="[705,1145,1307,1332]" pageId="4" pageNumber="467" refString="Jaschhof, M. &amp; Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement, 20, 1 - 392." type="journal article" year="2013">Jaschhof &amp; Jaschhof 2013: fig. 153A</bibRefCitation>
), basicercus with 1 large bristle apicoventrally, disticercus with 1 large bristle both subapicolaterally and subapicoventrally.
</paragraph>
<paragraph id="8B9B368509662002FF7AFAF0FBF1F9C8" blockId="4.[151,1436,614,1584]" pageId="4" pageNumber="467">
<emphasis id="B950EA9709662002FF7AFAF0FEB0FA84" bold="true" box="[199,368,1379,1404]" pageId="4" pageNumber="467">Classification.</emphasis>
The presence of more than 14 flagellomeres and the kind of circumfila leave no doubt that
<taxonomicName id="4C244D0609662002FF2AFA14FE47FA58" box="[151,391,1415,1440]" class="Insecta" family="Cecidomyiidae" genus="Feltomyina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="polymera">
<emphasis id="B950EA9709662002FF2AFA14FE47FA58" box="[151,391,1415,1440]" italics="true" pageId="4" pageNumber="467">Feltomyina polymera</emphasis>
</taxonomicName>
is properly placed in the tribe
<taxonomicName id="4C244D0609662002FD61FA14FC97FA58" box="[732,855,1415,1440]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
(see
<bibRefCitation id="EFB54B7409662002FC2FFA14FBDEFA67" author="Gagne" box="[914,1054,1415,1439]" pageId="4" pageNumber="467" refString="Gagne, R. J. (1981) Cecidomyiidae. In: McAlpine, J. F., Peterson, B. V., Shewell, G. E., Teskey, H. J., Vockeroth, J. R. &amp; Wood, D. M. (Coordinators), Manual of Nearctic Diptera. Vol. 1. Research Branch, Agriculture Canada, Ontario, Monograph No. 27, pp. 257 - 292." type="book chapter" year="1981">Gagné 1981</bibRefCitation>
). The classification of this species in a genus of its own is justified. The number of female flagellomeres is considerably larger in
<taxonomicName id="4C244D0609662002FB68FA38FA95FA3C" box="[1237,1365,1451,1476]" class="Insecta" family="Cecidomyiidae" genus="Feltomyina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9709662002FB68FA38FA95FA3C" box="[1237,1365,1451,1476]" italics="true" pageId="4" pageNumber="467">Feltomyina</emphasis>
</taxonomicName>
(60+) than in any other
<taxonomicName id="4C244D0609662002FEE3FA5CFE19FA10" box="[350,473,1487,1512]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
(40+). Other derived character states found in
<taxonomicName id="4C244D0609662002FC51FA5CFBABFA10" box="[1004,1131,1487,1512]" class="Insecta" family="Cecidomyiidae" genus="Feltomyina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9709662002FC51FA5CFBABFA10" box="[1004,1131,1487,1512]" italics="true" pageId="4" pageNumber="467">Feltomyina</emphasis>
</taxonomicName>
are the absence of sensory hairs on flagellomeres and the possession of bicolored legs. Also, it is the only genus of
<taxonomicName id="4C244D0609662002FB57FA60FAA5F9F4" box="[1258,1381,1523,1548]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
and
<taxonomicName id="4C244D0609662002FF2AF984FE8AF9C8" box="[151,330,1559,1584]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
known to have setae on both preepisternum 2 and metepisternum.
</paragraph>
</subSubSection>
</treatment>
</document>