310 lines
34 KiB
XML
310 lines
34 KiB
XML
<document id="E364D8AAA6AD69D81132208DB950771F" ID-DOI="10.26879/756" ID-ISSN="1094-8074" ID-Zenodo-Dep="11062937" ID-ZooBank="5EE56527-5F98-4724-8120-18F7471BB497" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1714012330871" checkinUser="felipe" docAuthor="López-Torres, Sergi, Silcox, Mary T. & Holroyd, Patricia A." docDate="2018" docId="038D87899230486621E0FC5CFC94B1A2" docLanguage="en" docName="PalaeontolElectron.21.3.37A.pdf" docOrigin="Palaeontologia Electronica (37 A) 21 (3)" docSource="http://dx.doi.org/10.26879/756" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Walshina López-Torres & Silcox & Holroyd 2018, gen. nov." docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="FFB4FFF192354860222AFFE4F808B326" masterDocTitle="New omomyoids (Euprimates, Mammalia) from the late Uintan of southern California, USA, and the question of the extinction of the Paromomyidae (Plesiadapiformes, Primates)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="6" updateTime="1715625529881" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="9E12D79F9FB733617C4FC3CDD5322CEB">New omomyoids (Euprimates, Mammalia) from the late Uintan of southern California, USA, and the question of the extinction of the Paromomyidae (Plesiadapiformes, Primates)</mods:title>
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<mods:namePart id="F2DFD7E30DFFC7AEE08DD3238CF9FD36">López-Torres, Sergi</mods:namePart>
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<mods:namePart id="E7AD5D9D987301937FE6963115A60A4E">Silcox, Mary T.</mods:namePart>
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<mods:namePart id="DF14F57A0891420BD4EB3F874B49B92F">Holroyd, Patricia A.</mods:namePart>
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<mods:typeOfResource id="365661901ECD65B21285726B47755354">text</mods:typeOfResource>
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<mods:title id="3A543DA58992856D8523F3EF4792BE17">Palaeontologia Electronica</mods:title>
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<mods:part id="43217CAEFBFE42CA16D104AE2A9EEC0A">
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<mods:date id="F687F5650466AE72A6FCF286751E043A">2018</mods:date>
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<mods:title id="19EA8ECCF191CB07DB63D097C8E0F928">37 A</mods:title>
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<treatment id="038D87899230486621E0FC5CFC94B1A2" LSID="urn:lsid:plazi:treatment:038D87899230486621E0FC5CFC94B1A2" httpUri="http://treatment.plazi.org/id/038D87899230486621E0FC5CFC94B1A2" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<subSubSection id="C33E65149230486521E0FC5CFD17B0F4" box="[970,1311,952,978]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="8B9B369F9230486521E0FC5CFD17B0F4" blockId="5.[869,1411,760,1010]" box="[970,1311,952,978]" pageId="5" pageNumber="6">
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<heading id="D0D381F39230486521E0FC5CFD17B0F4" box="[970,1311,952,978]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="6" reason="2">
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Genus
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<taxonomicName id="4C244D1C92304865260AFC5CFCA3B0F7" authority="López-Torres & Silcox & Holroyd, 2018" authorityName="López-Torres & Silcox & Holroyd" authorityYear="2018" box="[1056,1195,952,977]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B950EA8D92304865260AFC5CFCA3B0F7" box="[1056,1195,952,977]" italics="true" pageId="5" pageNumber="6">WALSHINA</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26357F6923048652698FC5DFD17B0F4" box="[1202,1311,953,978]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E6514923048652181FC3DFC0AB6BB" pageId="5" pageNumber="6" type="description">
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<paragraph id="8B9B369F923048652181FC3DFD35B0D4" blockId="5.[869,1411,760,1010]" box="[939,1341,985,1010]" pageId="5" pageNumber="6">
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||
<figureCitation id="131F2A1A923048652181FC3DFC6AB0D4" box="[939,1122,985,1010]" captionStart="FIGURE 4" captionStartId="6.[196,287,856,879]" captionTargetBox="[241,760,175,815]" captionTargetId="figure-613@6.[201,788,162,832]" captionTargetPageId="6" captionText="FIGURE 4. Environmental SEM images of four teeth of Walshina esmaraldensis gen. et sp. nov. 1: left M3, SDSNH 76276; 2: right M1, SDSNH 76337; 3: right M2, SDSNH 76338; 4: right M3, SDSNH 72583. All teeth are in occlusal view. Arrow indicates the location of the fovea." figureDoi="http://doi.org/10.5281/zenodo.11062945" httpUri="https://zenodo.org/record/11062945/files/figure.png" pageId="5" pageNumber="6">Figures 4.1-4.4</figureCitation>
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,
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<figureCitation id="131F2A1A923048652647FC3DFCD9B0D4" box="[1133,1233,985,1010]" captionStart="FIGURE 5" captionStartId="7.[188,278,1450,1473]" captionTargetBox="[366,1296,178,1422]" captionTargetId="figure-75@7.[318,1311,168,1422]" captionTargetPageId="7" captionText="FIGURE 5. Photographs taken with digital camera (1, 4-11) and Micro-CT scan reconstructions generated using Avizo 7 (2, 3). Walshina shifrae comb. nov. (1, 4, 7, 10) – 1: right M1, CM 15797 (holotype; mirrored), in occlusal view; 4: left M2, CM 15103, in occlusal view; 7: left M, CM 21637, in occlusal view; 10: left M; CM 15726, in occlusal view. 2 3 Walshina esmaraldensis gen. et sp. nov. (2, 5) – 2: left M1, LACM 40198 (holotype), in occlusal view; 5: left M2, SDSNH 62850, in occlusal view. Walshina mcgrewi comb. nov. (3, 6, 8, 9, 11) – 3: left M1, CM 15635 (holotype), in occlusal view; 6: left M2, CM15794, in occlusal view; 8, 9, 11: left mandibular fragment with M, CM 29005, in occlusal 2 (8), buccal (9), and lingual (11) views." figureDoi="http://doi.org/10.5281/zenodo.11062947" httpUri="https://zenodo.org/record/11062947/files/figure.png" pageId="5" pageNumber="6">5.1-5.11</figureCitation>
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,
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<figureCitation id="131F2A1A9230486526F6FC3DFD35B0D4" box="[1244,1341,985,1010]" captionStart="FIGURE 6" captionStartId="8.[198,289,1210,1233]" captionTargetBox="[467,1188,193,1157]" captionTargetId="figure-245@8.[412,1221,164,1192]" captionTargetPageId="8" captionText="FIGURE 6. Micro-CT scan reconstructions of specimens of Walshina esmaraldensis gen. et sp. nov. generated using Avizo 7: 1: left M1, LACM 40198 (holotype), in occlusal view; 2: left M2, SDSNH 87336, in occlusal view; 3: lingual fragment of a left M2, SDSNH 87337, in occlusal view; 4: lingual fragment of a right M2, SDSNH 42268, in occlusal view; 5, 6, 9, 10: left M2, SDSNH 87332, in occlusal (5), buccal (6), mesial (9) and lingual (10) views; 7, 8, 11, 12: mesial fragment of a left M1, SDSNH 87331, in occlusal (7), buccal (8), mesial (11) and lingual (12) views; 13, 14, 17, 18: left M3, SDSNH 87334, in buccal (13), occlusal (14), lingual (17) and mesial (18) views; 15, 16, 19: distal fragment of a right M3, SDSNH 87335, in buccal (15), occlusal (16) and lingual (19) views." figureDoi="http://doi.org/10.5281/zenodo.11062949" httpUri="https://zenodo.org/record/11062949/files/figure.png" pageId="5" pageNumber="6">6.1-6.19</figureCitation>
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</paragraph>
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<paragraph id="8B9B369F923048652162FBE7FD86B731" blockId="5.[840,1422,1027,1047]" box="[840,1422,1027,1047]" pageId="5" pageNumber="6">
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<emphasis id="B950EA8D923048652162FBE7FD86B731" bold="true" box="[840,1422,1027,1047]" pageId="5" pageNumber="6">
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zoobank.org/
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<uuid id="FF820C4A9230486521FBFBE7FD86B731" box="[977,1422,1027,1047]" pageId="5" pageNumber="6">5F051939-CD72-4EBC-8BBC-3E6B637DC806</uuid>
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</emphasis>
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||
</paragraph>
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<paragraph id="8B9B369F923048652162FBCDFBEBB77B" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
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1968
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<taxonomicName id="4C244D1C923048652191FBCDFD05B766" authority="(Matthew, 1915)" baseAuthorityName="Matthew" baseAuthorityYear="1915" box="[955,1293,1065,1088]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
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<emphasis id="B950EA8D923048652191FBCDFC5DB766" box="[955,1109,1065,1088]" italics="true" pageId="5" pageNumber="6">Phenacolemur</emphasis>
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(Matthew, 1915)
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</taxonomicName>
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||
; Robinson, p. 324
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</paragraph>
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<paragraph id="8B9B369F923048652162FB8DFCC1B7BB" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
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1976
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<taxonomicName id="4C244D1C923048652191FB8DFD52B7A6" authority="(Matthew and Granger, 1921)" baseAuthorityName="Matthew and Granger" baseAuthorityYear="1921" box="[955,1370,1128,1152]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
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<emphasis id="B950EA8D923048652191FB8DFC1CB7A6" box="[955,1044,1129,1152]" italics="true" pageId="5" pageNumber="6">Ignacius</emphasis>
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(Matthew and Granger, 1921)
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</taxonomicName>
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||
; Bown and Rose, p. 112 (in part)
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</paragraph>
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<paragraph id="8B9B369F923048652162FB4DFC32B7FB" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
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1978
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<taxonomicName id="4C244D1C923048652191FB4DFD04B7E6" authority="(Matthew, 1915)" baseAuthorityName="Matthew" baseAuthorityYear="1915" box="[955,1292,1193,1216]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
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<emphasis id="B950EA8D923048652191FB4DFC5DB7E6" box="[955,1109,1193,1216]" italics="true" pageId="5" pageNumber="6">Phenacolemur</emphasis>
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(Matthew, 1915)
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</taxonomicName>
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; Krishtalka, p. 338, fig. 2-4
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</paragraph>
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<paragraph id="8B9B369F923048652162FB0DFBE7B63B" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
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1990
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<taxonomicName id="4C244D1C923048652191FB0DFD06B626" authority="(Matthew, 1915)" baseAuthorityName="Matthew" baseAuthorityYear="1915" box="[955,1294,1257,1280]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
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<emphasis id="B950EA8D923048652191FB0DFC5EB626" box="[955,1110,1257,1280]" italics="true" pageId="5" pageNumber="6">Phenacolemur</emphasis>
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||
(Matthew, 1915)
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</taxonomicName>
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; Mason, p. 2, fig. 2
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</paragraph>
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<paragraph id="8B9B369F923048652162FACDFC30B67B" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
|
||
1991a cf.
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<taxonomicName id="4C244D1C9230486521F7FACDFD38B666" authority="(Matthew, 1915)" baseAuthorityName="Matthew" baseAuthorityYear="1915" box="[989,1328,1321,1344]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
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||
<emphasis id="B950EA8D9230486521F7FACDFC7FB666" box="[989,1143,1321,1344]" italics="true" pageId="5" pageNumber="6">Phenacolemur</emphasis>
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||
(Matthew, 1915)
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||
</taxonomicName>
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||
; Walsh, p. 166, table 1
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||
</paragraph>
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||
<paragraph id="8B9B369F923048652162FA8DFC0AB6BB" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
|
||
1996
|
||
<taxonomicName id="4C244D1C923048652191FA8DFD03B6A6" authority="(Matthew, 1915)" baseAuthorityName="Matthew" baseAuthorityYear="1915" box="[955,1291,1385,1408]" class="Mammalia" family="Paromomyidae" genus="Phenacolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D923048652191FA8DFC5EB6A6" box="[955,1110,1385,1408]" italics="true" pageId="5" pageNumber="6">Phenacolemur</emphasis>
|
||
(Matthew, 1915)
|
||
</taxonomicName>
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||
; Walsh, p. 85, table 2
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C33E6514923048652162FA4DFC67B544" pageId="5" pageNumber="6" type="type_taxon">
|
||
<paragraph id="8B9B369F923048652162FA4DFB71B6C4" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
|
||
<emphasis id="B950EA8D923048652162FA4DFBFDB6E4" bold="true" box="[840,1013,1449,1474]" pageId="5" pageNumber="6">
|
||
<typeStatus id="549F883D923048652162FA4DFB8CB6E4" box="[840,900,1449,1474]" pageId="5" pageNumber="6">Type</typeStatus>
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||
species.
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||
</emphasis>
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||
<taxonomicName id="4C244D1C9230486521D6FA4DFD14B6E4" authority="López-Torres & Silcox & Holroyd, 2018" authorityName="López-Torres & Silcox & Holroyd" authorityYear="2018" box="[1020,1308,1449,1474]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="esmaraldensis" status="gen. et sp.">
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||
<emphasis id="B950EA8D9230486521D6FA4DFD14B6E4" box="[1020,1308,1449,1474]" italics="true" pageId="5" pageNumber="6">Walshina esmaraldensis</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A26357F692304865270EFA4DFDA9B6E4" box="[1316,1441,1449,1474]" pageId="5" pageNumber="6">gen. et sp.</taxonomicNameLabel>
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||
nov.
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</paragraph>
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||
<paragraph id="8B9B369F923048652162FA0DFC67B544" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
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||
<emphasis id="B950EA8D923048652162FA0DFC35B524" bold="true" box="[840,1085,1513,1538]" pageId="5" pageNumber="6">Included species.</emphasis>
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||
<typeStatus id="549F883D923048652671FA0DFC9DB524" box="[1115,1173,1513,1538]" pageId="5" pageNumber="6">Type</typeStatus>
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||
species,
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||
<taxonomicName id="4C244D1C923048652719FA0DFDA8B524" authorityName="López-Torres & Silcox & Holroyd" authorityYear="2018" box="[1331,1440,1513,1538]" pageId="5" pageNumber="6" status="comb. nov.">
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||
<emphasis id="B950EA8D923048652719FA0DFDA8B524" box="[1331,1440,1513,1538]" italics="true" pageId="5" pageNumber="6">Walshina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B950EA8D923048652162F9EDFBA3B504" box="[840,939,1545,1570]" italics="true" pageId="5" pageNumber="6">mcgrewi</emphasis>
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||
<taxonomicNameLabel id="A26357F6923048652195F9EDFC45B504" box="[959,1101,1545,1570]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(=
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||
<taxonomicName id="4C244D1C9230486526A4F9EDFD95B504" authority="Robinson" authorityName="Robinson" box="[1166,1437,1545,1570]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="mcgrewi">
|
||
<emphasis id="B950EA8D9230486526A4F9EDFD1CB504" box="[1166,1300,1545,1570]" italics="true" pageId="5" pageNumber="6">I. mcgrewi</emphasis>
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Robinson
|
||
</taxonomicName>
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, 1968), and
|
||
<taxonomicName id="4C244D1C9230486521F4F9CDFCA2B567" authorityName="López-Torres & Silcox & Holroyd" authorityYear="2018" baseAuthorityName="López-Torres & Silcox & Holroyd" baseAuthorityYear="2018" box="[990,1194,1576,1602]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="shifrae" status="comb. nov.">
|
||
<emphasis id="B950EA8D9230486521F4F9CDFCA2B567" box="[990,1194,1576,1602]" italics="true" pageId="5" pageNumber="6">Walshina shifrae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26357F6923048652693F9CDFD4AB564" box="[1209,1346,1577,1602]" pageId="5" pageNumber="6" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(=
|
||
<emphasis id="B950EA8D923048652752F9CDFDA8B564" box="[1400,1440,1577,1602]" italics="true" pageId="5" pageNumber="6">Ph.</emphasis>
|
||
<taxonomicName id="4C244D1C923048652162F9ACFC6CB544" authority="Krishtalka, 1978" authorityName="Krishtalka" authorityYear="1978" box="[840,1124,1608,1634]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="shifrae">
|
||
<emphasis id="B950EA8D923048652162F9ACFB90B547" box="[840,920,1608,1633]" italics="true" pageId="5" pageNumber="6">shifrae</emphasis>
|
||
Krishtalka, 1978
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6514923048652162F98DFCA7B584" pageId="5" pageNumber="6" type="distribution">
|
||
<paragraph id="8B9B369F923048652162F98DFCA7B584" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
|
||
<emphasis id="B950EA8D923048652162F98DFBEDB5A4" bold="true" box="[840,997,1641,1666]" pageId="5" pageNumber="6">Distribution.</emphasis>
|
||
Uintan and Duchesnean of Wyoming, and Uintan of
|
||
<collectingRegion id="49E0F87D92304865261CF96CFCA3B584" box="[1078,1195,1672,1698]" country="United States of America" name="California" pageId="5" pageNumber="6">California</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6514923048652162F94DFD83B427" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="8B9B369F923048652162F94DFD83B427" blockId="5.[840,1441,1065,1895]" pageId="5" pageNumber="6">
|
||
<emphasis id="B950EA8D923048652162F94DFBDEB5E4" bold="true" box="[840,982,1705,1730]" pageId="5" pageNumber="6">Etymology.</emphasis>
|
||
In memory of Stephen L. Walsh of the San Diego Museum of Natural History, in recognition of his work on the San Diego County faunas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6514923048662162F8EDFAF7B45A" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="diagnosis">
|
||
<paragraph id="8B9B369F923048662162F8EDFAF7B45A" blockId="5.[840,1441,1065,1895]" lastBlockId="6.[192,793,1092,1916]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
<emphasis id="B950EA8D923048652162F8EDFBC6B404" bold="true" box="[840,974,1801,1826]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
|
||
Paracristid of M
|
||
<subScript id="17A034DA9230486526B1F8F7FCAFB401" attach="left" box="[1179,1191,1811,1831]" fontSize="8" pageId="5" pageNumber="6">1</subScript>
|
||
relatively long as in
|
||
<taxonomicName id="4C244D1C923048652162F8CAFBC6B461" authorityName=", Gunnell" authorityYear="1995" box="[840,974,1838,1863]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D923048652162F8CAFBC6B461" box="[840,974,1838,1863]" italics="true" pageId="5" pageNumber="6">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C244D1C92304865263AF8C9FCB8B461" authorityName="Gunnell" authorityYear="1995" box="[1040,1200,1837,1863]" class="Mammalia" family="Omomyidae" genus="Sphacorhysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D92304865263AF8C9FCB8B461" box="[1040,1200,1837,1863]" italics="true" pageId="5" pageNumber="6">Sphacorhysis</emphasis>
|
||
</taxonomicName>
|
||
, but paraconid less clearly distinct from the paracristid. Differs from
|
||
<taxonomicName id="4C244D1C9233486622EAFBA0F968B778" authorityName="Gunnell" authorityYear="1995" box="[192,352,1092,1118]" class="Mammalia" family="Omomyidae" genus="Sphacorhysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486622EAFBA0F968B778" box="[192,352,1092,1118]" italics="true" pageId="6" pageNumber="7">Sphacorhysis</emphasis>
|
||
</taxonomicName>
|
||
(but not
|
||
<taxonomicName id="4C244D1C9233486623E6FBA0FA5AB77B" authorityName=", Gunnell" authorityYear="1995" box="[460,594,1092,1117]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486623E6FBA0FA5AB77B" box="[460,594,1092,1117]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
) in having lower molar talonid basins that are relatively deep with smooth enamel. Differs from
|
||
<taxonomicName id="4C244D1C92334866200DFB60FAA5B7BB" authorityName=", Gunnell" authorityYear="1995" box="[551,685,1156,1181]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D92334866200DFB60FAA5B7BB" box="[551,685,1156,1181]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
(but not
|
||
<taxonomicName id="4C244D1C9233486622EAFB40F968B798" authorityName="Gunnell" authorityYear="1995" box="[192,352,1188,1214]" class="Mammalia" family="Omomyidae" genus="Sphacorhysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486622EAFB40F968B798" box="[192,352,1188,1214]" italics="true" pageId="6" pageNumber="7">Sphacorhysis</emphasis>
|
||
</taxonomicName>
|
||
) in that the cristid obliqua of M
|
||
<subScript id="17A034DA9233486620FAFB4BFAD4B7E5" attach="left" box="[720,732,1199,1219]" fontSize="8" pageId="6" pageNumber="7">1</subScript>
|
||
contacts the postvallid distal to the protoconid rather than between the protoconid and metaconid. Unlike the other trogolemurins, M
|
||
<subScript id="17A034DA923348662058FAF0FA76B60E" attach="left" box="[626,638,1300,1320]" fontSize="8" pageId="6" pageNumber="7">1</subScript>
|
||
and M
|
||
<subScript id="17A034DA9233486620CEFAF0FAF8B60E" attach="left" box="[740,752,1300,1320]" fontSize="8" pageId="6" pageNumber="7">2</subScript>
|
||
of
|
||
<taxonomicName id="4C244D1C9233486622EAFACAF925B661" authority="López-Torres & Silcox & Holroyd, 2018" authorityName="López-Torres & Silcox & Holroyd" authorityYear="2018" box="[192,301,1326,1351]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B950EA8D9233486622EAFACAF925B661" box="[192,301,1326,1351]" italics="true" pageId="6" pageNumber="7">Walshina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26357F692334866231CFACAF9ADB661" box="[310,421,1326,1351]" pageId="6" pageNumber="7" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
have strong hypoconulids with distinct foveae located below and buccal to the hypoconulid. As in
|
||
<taxonomicName id="4C244D1C92334866238EFA8AFA4CB6AE" authorityName="Gunnell" authorityYear="1995" box="[420,580,1390,1416]" class="Mammalia" family="Omomyidae" genus="Sphacorhysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D92334866238EFA8AFA4CB6AE" box="[420,580,1390,1416]" italics="true" pageId="6" pageNumber="7">Sphacorhysis</emphasis>
|
||
</taxonomicName>
|
||
, the distal aspect of M
|
||
<subScript id="17A034DA9233486622DCFA7DF90AB68B" attach="left" box="[246,258,1433,1453]" fontSize="8" pageId="6" pageNumber="7">1</subScript>
|
||
and M
|
||
<subScript id="17A034DA92334866237DFA7DF96BB68B" attach="left" box="[343,355,1433,1453]" fontSize="8" pageId="6" pageNumber="7">2</subScript>
|
||
is convex, whereas in
|
||
<taxonomicName id="4C244D1C923348662051FA6AFB09B681" authorityName=", Gunnell" authorityYear="1995" box="[635,769,1422,1447]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D923348662051FA6AFB09B681" box="[635,769,1422,1447]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
it is concave. M
|
||
<subScript id="17A034DA92334866234FFA5AF979B6F4" attach="left" box="[357,369,1470,1490]" fontSize="8" pageId="6" pageNumber="7">3</subScript>
|
||
hypoconulid narrower than in other trogolemurins. M
|
||
<subScript id="17A034DA9233486623ACFA07F99AB6D1" attach="left" box="[390,402,1507,1527]" fontSize="8" pageId="6" pageNumber="7">3</subScript>
|
||
trigonid significantly taller than in
|
||
<taxonomicName id="4C244D1C9233486622EAFA19F968B531" authorityName="Gunnell" authorityYear="1995" box="[192,352,1533,1559]" class="Mammalia" family="Omomyidae" genus="Sphacorhysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486622EAFA19F968B531" box="[192,352,1533,1559]" italics="true" pageId="6" pageNumber="7">Sphacorhysis</emphasis>
|
||
</taxonomicName>
|
||
(but not
|
||
<taxonomicName id="4C244D1C9233486623FCFA19FA54B530" authorityName=", Gunnell" authorityYear="1995" box="[470,604,1533,1558]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486623FCFA19FA54B530" box="[470,604,1533,1558]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
). Like
|
||
<taxonomicName id="4C244D1C92334866209FFA19F90DB510" authorityName="Gunnell" authorityYear="1995" class="Mammalia" family="Omomyidae" genus="Sphacorhysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D92334866209FFA19F90DB510" italics="true" pageId="6" pageNumber="7">Sphacorhysis</emphasis>
|
||
</taxonomicName>
|
||
, lower molar entocristids form a rounded contour (i.e., forming a U-shaped entocristid) in lingual view, in contrast to the V-shaped entocristid in
|
||
<taxonomicName id="4C244D1C9233486622EAF999F94DB5B0" authorityName=", Gunnell" authorityYear="1995" box="[192,325,1661,1686]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486622EAF999F94DB5B0" box="[192,325,1661,1686]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
. Further differs from
|
||
<taxonomicName id="4C244D1C92334866207DF999FAD5B5B0" authorityName=", Gunnell" authorityYear="1995" box="[599,733,1661,1686]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D92334866207DF999FAD5B5B0" box="[599,733,1661,1686]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C244D1C9233486622EAF979F968B591" authorityName="Gunnell" authorityYear="1995" box="[192,352,1693,1719]" class="Mammalia" family="Omomyidae" genus="Sphacorhysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486622EAF979F968B591" box="[192,352,1693,1719]" italics="true" pageId="6" pageNumber="7">Sphacorhysis</emphasis>
|
||
</taxonomicName>
|
||
in having much weaker buccal cingulids. Notably stronger precingulum on M
|
||
<superScript id="7C519BD792334866209EF959FAC8B5F7" attach="left" box="[692,704,1725,1745]" fontSize="8" pageId="6" pageNumber="7">2</superScript>
|
||
than in
|
||
<taxonomicName id="4C244D1C9233486622EAF904F94DB5DF" authorityName=", Gunnell" authorityYear="1995" box="[192,325,1760,1785]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486622EAF904F94DB5DF" box="[192,325,1760,1785]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
. Protocone lingual expansion on the upper molars not as pronounced as in
|
||
<taxonomicName id="4C244D1C9233486620A6F8E4FB19B43F" authorityName=", Gunnell" authorityYear="1995" box="[652,785,1792,1817]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D9233486620A6F8E4FB19B43F" box="[652,785,1792,1817]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
. Compared to
|
||
<taxonomicName id="4C244D1C92334866235EF8C7F9F1B41A" authorityName=", Gunnell" authorityYear="1995" box="[372,505,1827,1852]" class="Mammalia" family="Omomyidae" genus="Trogolemur" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EA8D92334866235EF8C7F9F1B41A" box="[372,505,1827,1852]" italics="true" pageId="6" pageNumber="7">Trogolemur</emphasis>
|
||
</taxonomicName>
|
||
, mesial aspect of M
|
||
<superScript id="7C519BD7923348662126F8C4FB10B412" attach="left" box="[780,792,1824,1844]" fontSize="8" pageId="6" pageNumber="7">3</superScript>
|
||
straighter, and the lingual border of that tooth much shorter mesiodistally relative to its buccal border.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E6514923348662162FF42FC94B1A2" pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph id="8B9B369F923348662162FF42FC94B1A2" blockId="6.[840,1441,166,644]" pageId="6" pageNumber="7">
|
||
<emphasis id="B950EA8D923348662162FF42FBD5B399" bold="true" box="[840,989,166,191]" pageId="6" pageNumber="7">Discussion.</emphasis>
|
||
All trogolemurins share a distally expanded distolingual basin of the upper molars (particularly marked in
|
||
<taxonomicName id="4C244D1C923348662643FF02FCF2B3D9" authorityName="López-Torres & Silcox & Holroyd" authorityYear="2018" baseAuthorityName="López-Torres & Silcox & Holroyd" baseAuthorityYear="2018" box="[1129,1274,230,255]" class="Mammalia" family="Paromomyidae" genus="Ignacius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcgrewi" status="comb. nov.">
|
||
<emphasis id="B950EA8D923348662643FF02FCF2B3D9" box="[1129,1274,230,255]" italics="true" pageId="6" pageNumber="7">W. mcgrewi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26357F6923348662723FF02FD9EB3D9" box="[1289,1430,230,255]" pageId="6" pageNumber="7" rank="species">comb. nov.</taxonomicNameLabel>
|
||
), which is quite similar to that observed in paromomyid plesiadapiforms. This similarity is likely one reason why some members of
|
||
<taxonomicName id="4C244D1C923348662696FEA2FD21B279" authority="López-Torres & Silcox & Holroyd, 2018" authorityName="López-Torres & Silcox & Holroyd" authorityYear="2018" box="[1212,1321,326,351]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B950EA8D923348662696FEA2FD21B279" box="[1212,1321,326,351]" italics="true" pageId="6" pageNumber="7">Walshina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26357F6923348662719FEA2FDA9B279" box="[1331,1441,326,351]" pageId="6" pageNumber="7" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
have previously been considered paromomyids. However, in other ways the morphology of trogolemurins is inconsistent with that of paromomyids, including the presence of paraconids on M
|
||
<subScript id="17A034DA9233486627A7FE35FD91B2C3" attach="left" box="[1421,1433,465,485]" fontSize="8" pageId="6" pageNumber="7">3</subScript>
|
||
.
|
||
<taxonomicName id="4C244D1C923348662162FE0FFBBDB122" authority="López-Torres & Silcox & Holroyd, 2018" authorityName="López-Torres & Silcox & Holroyd" authorityYear="2018" box="[840,949,491,516]" class="Mammalia" family="Omomyidae" genus="Walshina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="6" pageNumber="7" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B950EA8D923348662162FE0FFBBDB122" box="[840,949,491,516]" italics="true" pageId="6" pageNumber="7">Walshina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26357F69233486621E9FE0FFC3DB122" box="[963,1077,491,516]" pageId="6" pageNumber="7" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
remains quite poorly known, with the only record being isolated upper and lower molars. One likely reason for this limited record is that the genus includes the smallest North American omomyoids (see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |