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<document id="D0380D4C4B369E2E27A5C2AA893B37A3" ID-CLB-Dataset="32700" ID-DOI="10.11646/zootaxa.4290.3.7" ID-GBIF-Dataset="48f25ad8-5841-4eab-a32a-f03853045968" ID-ISSN="1175-5326" ID-Zenodo-Dep="829273" ID-ZooBank="66A83CED-6200-4758-A679-39B5A0656B38" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500111116054" checkinUser="plazi" docAuthor="Márquez, Roberto, Mejía-Vargas, Daniel, Palacios-Rodríguez, Pablo, Ramírez-Castañeda, Valeria &amp; Amézquita, Adolfo" docDate="2017" docId="038D87893575D921FF767896FEB4F98B" docLanguage="en" docName="zootaxa.4290.3.7.pdf" docOrigin="Zootaxa 4290 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andinobates victimatus Márquez, Mejía-Vargas, Palacios-Rodríguez, Ramírez-Castañeda &amp; Amézquita, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="543" masterDocId="FFB4FFF13576D92DFFE17D54FFEEFF80" masterDocTitle="A new species of Andinobates (Anura: Dendrobatidae) from the Urabá region of Colombia" masterLastPageNumber="546" masterPageNumber="531" pageNumber="534" updateTime="1698465473380" updateUser="ExternalLinkService">
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<mods:title id="74D6C228C21DBCD7A6EBDEEDA42D1A01">A new species of Andinobates (Anura: Dendrobatidae) from the Urabá region of Colombia</mods:title>
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<mods:namePart id="52CAE6F1978E916ADD2F5A331EB2155A">Ramírez-Castañeda, Valeria</mods:namePart>
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<paragraph id="8B9B369F3575D92EFF767896FE04FA7E" blockId="3.[151,530,1474,1534]" pageId="3" pageNumber="534">
<taxonomicName id="4C244D1C3575D92EFF767896FE5CFA5C" ID-CoL="66N9G" box="[151,434,1474,1500]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="534" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3575D92EFF767896FE5CFA5C" bold="true" box="[151,434,1474,1500]" italics="true" pageId="3" pageNumber="534">Andinobates victimatus</emphasis>
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<emphasis id="B950EA8D3575D92EFE587897FDFCFA5C" bold="true" box="[441,530,1475,1500]" pageId="3" pageNumber="534">sp. nov.</emphasis>
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<figureCitation id="131F2A1A3575D92EFF7678B1FEEEFA7E" box="[151,256,1509,1534]" captionStart="FIGURE 2" captionStartId="4.[151,250,1806,1828]" captionTargetBox="[193,1386,458,1743]" captionTargetId="figure@4.[151,1436,409,1785]" captionTargetPageId="4" captionText="FIGURE 2. Maximum likelihood mtDNA genealogy inferred from three concatenated loci (16 S, COI, Cytb). Numbers on internodes correspond to nodal support in the form of Bayesian posterior probability / ML bootstrap support. Support for withinspecies nodes is not shown (unless mentioned in the text) for better visualization. Branch colors follow the legend in Figure 1. Photo of A. tolimensis by Ricardo Medina-Rengifo, all others by Roberto Márquez." httpUri="https://zenodo.org/record/829277/files/figure.png" pageId="3" pageNumber="534">Figures 2</figureCitation>
and 
<figureCitation id="131F2A1A3575D92EFED778B2FEAAFA7E" box="[310,324,1510,1534]" captionStart="FIGURE 4" captionStartId="6.[151,250,1682,1704]" captionTargetBox="[187,1409,225,1623]" captionTargetId="figure@6.[151,1436,193,1661]" captionTargetPageId="6" captionText="FIGURE 4. Andinobates victimatus sp. nov. (A – K) and A. fulguritus (L – S). A – D: Lateral and dorsal views of an unsexed adult from the type locality. E – F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H – K: Juveniles of A. victimatus sp. nov. exhibiting intermediate stages of coloration development at ~ 1.5 months (H – I), ~ 2.5 months (J), and ~ 3 months (K). L – S: Unsexed A. fulguritus adults from Playa de Oro, Chocó (L – M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q – S), Photos in A – D and H – K by Roberto Márquez, E – F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L – S by Daniel Mejía- Vargas." httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="3" pageNumber="534">4</figureCitation>
A–K. 
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.
</paragraph>
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<paragraph id="8B9B369F3575D92EFF767B79FB83F9EA" blockId="3.[151,1437,1581,2038]" pageId="3" pageNumber="534">
<emphasis id="B950EA8D3575D92EFF767B79FEE5F9C6" bold="true" box="[151,267,1581,1606]" pageId="3" pageNumber="534">
<typeStatus id="549F883D3575D92EFF767B79FEE8F9C6" box="[151,262,1581,1606]" pageId="3" pageNumber="534" type="holotype">Holotype</typeStatus>
.
</emphasis>
An adult female (ANDES-A 3686) collected by Daniel Mejía-Vargas in Chichiridó, Dabeiba, 
<collectingRegion id="49E0F87D3575D92EFACD7B79FA72F9C6" box="[1324,1436,1581,1606]" country="Colombia" name="Antioquia" pageId="3" pageNumber="534">Antioquia</collectingRegion>
(7.02°, -76.38°), located in the Llorona Canyon, at 280 meters above sea level (m.a.s.l.).
</paragraph>
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<emphasis id="B950EA8D3575D92EFF267B21FEA8F90E" bold="true" box="[199,326,1653,1678]" pageId="3" pageNumber="534">
<typeStatus id="549F883D3575D92EFF267B21FEAFF90E" box="[199,321,1653,1678]" pageId="3" pageNumber="534" type="paratype">Paratypes</typeStatus>
.
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Five adults (
<specimenCount id="9D22FD163575D92EFE047B22FDAEF90E" box="[485,576,1653,1678]" pageId="3" pageNumber="534" type="male">4 males</specimenCount>
and 
<specimenCount id="9D22FD163575D92EFD617B22FD07F90E" box="[640,745,1653,1678]" pageId="3" pageNumber="534" type="female">1 female</specimenCount>
; ANDES-A 3684, 3685, 3687–3689), collected at the 
<typeStatus id="549F883D3575D92EFA8D7B22FA72F90E" box="[1388,1436,1654,1678]" pageId="3" pageNumber="534">type</typeStatus>
locality, along with the 
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.
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<subSubSection id="C33E65143575D92EFF267BE9FDACF8C2" pageId="3" pageNumber="534" type="etymology">
<paragraph id="8B9B369F3575D92EFF267BE9FDACF8C2" blockId="3.[151,1437,1581,2038]" pageId="3" pageNumber="534">
<emphasis id="B950EA8D3575D92EFF267BE9FEA0F956" bold="true" box="[199,334,1725,1750]" pageId="3" pageNumber="534">Etymology.</emphasis>
From the Latin 
<taxonomicName id="4C244D1C3575D92EFDF27BEAFD68F955" box="[531,646,1726,1749]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="534" phylum="Chordata" rank="species" species="victimatus">
<emphasis id="B950EA8D3575D92EFDF27BEAFD68F955" box="[531,646,1726,1749]" italics="true" pageId="3" pageNumber="534">victimatus</emphasis>
</taxonomicName>
(the victimized); noun in apposition. The Urabá region, where this species occurs, has historically been flailed by Colombia’s longstanding armed conflicts, perhaps more than most other regions of the country, leaving a virtually countless trail of innocent victims. We name this species in honor and remembrance of all these victims.
</paragraph>
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<paragraph id="8B9B369F3575D929FF267A19FE3BFEE5" blockId="3.[151,1437,1581,2038]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="535" pageId="3" pageNumber="534">
<emphasis id="B950EA8D3575D92EFF267A19FE1FF8E6" bold="true" box="[199,497,1869,1894]" pageId="3" pageNumber="534">Definition and diagnosis.</emphasis>
<taxonomicName id="4C244D1C3575D92EFE197A19FD11F8E5" box="[504,767,1869,1894]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="534" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3575D92EFE197A19FD11F8E5" box="[504,767,1869,1894]" italics="true" pageId="3" pageNumber="534">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63575D92EFCE67A19FC8FF8E6" box="[775,865,1869,1894]" pageId="3" pageNumber="534" rank="species">
<emphasis id="B950EA8D3575D92EFCE67A19FC8FF8E6" bold="true" box="[775,865,1869,1894]" pageId="3" pageNumber="534">sp. nov.</emphasis>
</taxonomicNameLabel>
is a minute (SVL mean = 14.23 ± SD = 
<quantity id="4CDC9B7A3575D92EFACE7A19FA79F8E5" box="[1327,1431,1869,1894]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="534" unit="mm" value="0.67">0.67 mm</quantity>
; 
<tableCitation id="C6A603243575D92EFF767A25FF02F80A" box="[151,236,1905,1930]" captionStart="TABLE 1" captionStartId="7.[151,239,260,283]" captionText="TABLE 1. Measurements of the type series of A. victimatus sp. nov. in mm. See methods section for explanation of abbreviations." pageId="3" pageNumber="534">Table 1</tableCitation>
) dendrobatid frog, characterized by having homogeneous scarlet red dorsal coloration, which extends throughout the limbs. Fingers and toe tips are light metallic gray to metallic gray. The venter and flanks are also scarlet red, with patches degrading into vermilion and finally black, forming irregular blotches (
<figureCitation id="131F2A1A3575D92EFB007AEDFAC8F852" box="[1249,1318,1977,2002]" captionStart="FIGURE 4" captionStartId="6.[151,250,1682,1704]" captionTargetBox="[187,1409,225,1623]" captionTargetId="figure@6.[151,1436,193,1661]" captionTargetPageId="6" captionText="FIGURE 4. Andinobates victimatus sp. nov. (A – K) and A. fulguritus (L – S). A – D: Lateral and dorsal views of an unsexed adult from the type locality. E – F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H – K: Juveniles of A. victimatus sp. nov. exhibiting intermediate stages of coloration development at ~ 1.5 months (H – I), ~ 2.5 months (J), and ~ 3 months (K). L – S: Unsexed A. fulguritus adults from Playa de Oro, Chocó (L – M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q – S), Photos in A – D and H – K by Roberto Márquez, E – F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L – S by Daniel Mejía- Vargas." httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="3" pageNumber="534">Fig. 4</figureCitation>
A–F). The skin is slightly granular, with granularity increasing towards the urostyle and hind limbs. This species is assignable to 
<taxonomicName id="4C244D1C3572D929FF597DC3FEAAFF30" box="[184,324,151,176]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="535" phylum="Chordata" rank="genus">
<emphasis id="B950EA8D3572D929FF597DC3FEAAFF30" box="[184,324,151,176]" italics="true" pageId="4" pageNumber="535">Andinobates</emphasis>
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based on the following traits: small adult size (SVL &lt;
<quantity id="4CDC9B7A3572D929FC387DCCFBDEFF30" box="[985,1072,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="535" unit="mm" value="20.0">20 mm</quantity>
), bright dorsal coloration, limb reticulation absent, first finger distinctively shorter than second, median lingual process absent, and maxillary and premaxillary teeth absent. Molecular phylogenetic analyses support this placement. Within 
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<emphasis id="B950EA8D3572D929FB7F7D8BFAC4FF78" box="[1182,1322,223,248]" italics="true" pageId="4" pageNumber="535">Andinobates</emphasis>
</taxonomicName>
, we place 
<taxonomicName id="4C244D1C3572D929FF767C50FEC0FE9C" box="[151,302,260,284]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="535" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3572D929FF767C50FEC0FE9C" box="[151,302,260,284]" italics="true" pageId="4" pageNumber="535">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63572D929FED87C51FE61FE9C" box="[313,399,261,284]" pageId="4" pageNumber="535" rank="species">sp. nov</taxonomicNameLabel>
in the 
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<emphasis id="B950EA8D3572D929FE0B7C50FD95FE9D" box="[490,635,260,285]" italics="true" pageId="4" pageNumber="535">A. fulguritus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EFB54B6E3572D929FD3A7C50FBA9FE9C" author="Brown" box="[731,1095,260,285]" pageId="4" pageNumber="535" refString="Brown, J. L., Twomey, E., Amezquita, A., Souza, M. B. de, Caldwell, J. - L. P., Lotters, S., von May, R., Melo-Sampaio, P. R., Mejia-Vargas, D., Perez-Pena, P., Pepper, M., Poelman, E. H., Sanchez-Rodriguez, M. &amp; Summers, K. (2011) A taxonomic revision of the Neotropical poison frog genus Ranitomeya (Amphibia: Dendrobatidae). Zootaxa, 3083, 1 - 120." type="journal article" year="2011">
Brown &amp; Twomey 
<emphasis id="B950EA8D3572D929FC257C51FBEFFE9D" box="[964,1025,260,285]" italics="true" pageId="4" pageNumber="535">et al.</emphasis>
2011
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), also based on phylogenetic analyses, juvenile color pattern (
<figureCitation id="131F2A1A3572D929FDE97C73FDA2FEC0" box="[520,588,295,320]" captionStart="FIGURE 4" captionStartId="6.[151,250,1682,1704]" captionTargetBox="[187,1409,225,1623]" captionTargetId="figure@6.[151,1436,193,1661]" captionTargetPageId="6" captionText="FIGURE 4. Andinobates victimatus sp. nov. (A – K) and A. fulguritus (L – S). A – D: Lateral and dorsal views of an unsexed adult from the type locality. E – F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H – K: Juveniles of A. victimatus sp. nov. exhibiting intermediate stages of coloration development at ~ 1.5 months (H – I), ~ 2.5 months (J), and ~ 3 months (K). L – S: Unsexed A. fulguritus adults from Playa de Oro, Chocó (L – M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q – S), Photos in A – D and H – K by Roberto Márquez, E – F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L – S by Daniel Mejía- Vargas." httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="4" pageNumber="535">Fig. 4</figureCitation>
H–K; see Coloration in life), and calls of the “short buzz” 
<typeStatus id="549F883D3572D929FB117C7CFACEFEC0" box="[1264,1312,296,320]" pageId="4" pageNumber="535">type</typeStatus>
(
<bibRefCitation id="EFB54B6E3572D929FAD17C7CFE96FEE4" author="Brown" pageId="4" pageNumber="535" refString="Brown, J. L., Twomey, E., Amezquita, A., Souza, M. B. de, Caldwell, J. - L. P., Lotters, S., von May, R., Melo-Sampaio, P. R., Mejia-Vargas, D., Perez-Pena, P., Pepper, M., Poelman, E. H., Sanchez-Rodriguez, M. &amp; Summers, K. (2011) A taxonomic revision of the Neotropical poison frog genus Ranitomeya (Amphibia: Dendrobatidae). Zootaxa, 3083, 1 - 120." type="journal article" year="2011">
Brown &amp; Twomey 
<emphasis id="B950EA8D3572D929FF1E7C19FEDFFEE5" box="[255,305,332,357]" italics="true" pageId="4" pageNumber="535">et al</emphasis>
. 2011
</bibRefCitation>
; 
<figureCitation id="131F2A1A3572D929FE657C18FE29FEE5" box="[388,455,332,357]" captionStart="FIGURE 5" captionStartId="9.[151,250,1650,1672]" captionTargetBox="[187,1401,447,1629]" captionTargetId="figure@9.[151,1436,409,1629]" captionTargetPageId="9" captionText="FIGURE 5. Oscillogram (below, in relative units) and spectrogram (above) of one spontaneous advertisement call of Andinobates victimatus sp. nov. The male was captured at the type locality and recorded in captivity. The distribution of sound energy is represented by the power spectrum (overlaid white line, amplitude in X axis is relative). The temperature of the substrate around the calling male was 19.4 ° C." httpUri="https://zenodo.org/record/829283/files/figure.png" pageId="4" pageNumber="535">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF5B66173572D929FF767A5AFC0DF800" httpUri="https://zenodo.org/record/829277/files/figure.png" pageId="4" pageNumber="535" targetBox="[193,1386,458,1743]" targetPageId="4">
<paragraph id="8B9B369F3572D929FF767A5AFC0DF800" blockId="4.[151,1436,1806,1920]" pageId="4" pageNumber="535">
<emphasis id="B950EA8D3572D929FF767A5AFEFBF8A4" bold="true" box="[151,277,1806,1828]" pageId="4" pageNumber="535">FIGURE 2.</emphasis>
Maximum likelihood mtDNA genealogy inferred from three concatenated loci (16S, COI, Cytb). Numbers on internodes correspond to nodal support in the form of Bayesian posterior probability/ML bootstrap support. Support for withinspecies nodes is not shown (unless mentioned in the text) for better visualization. Branch colors follow the legend in Figure 1. Photo of 
<taxonomicName id="4C244D1C3572D929FF157A3FFE99F800" box="[244,375,1899,1920]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="535" phylum="Chordata" rank="species" species="tolimensis">
<emphasis id="B950EA8D3572D929FF157A3FFE99F800" box="[244,375,1899,1920]" italics="true" pageId="4" pageNumber="535">A. tolimensis</emphasis>
</taxonomicName>
by Ricardo Medina-Rengifo, all others by Roberto Márquez.
</paragraph>
</caption>
<caption id="DF5B66173573D928FF7679F2FD36FB79" httpUri="https://zenodo.org/record/829279/files/figure.png" pageId="5" pageNumber="536" targetBox="[176,1415,217,1138]" targetPageId="5">
<paragraph id="8B9B369F3573D928FF7679F2FD36FB79" blockId="5.[151,1436,1190,1273]" pageId="5" pageNumber="536">
<emphasis id="B950EA8D3573D928FF7679F2FEF6FB3B" bold="true" box="[151,280,1190,1212]" pageId="5" pageNumber="536">FIGURE 3.</emphasis>
Mean pairwise intra and interspecific Kimura two-parameter (K2P) distances between the Cytb sequences analyzed in this study. Since only one sequence per species was used for the 
<taxonomicName id="4C244D1C3573D928FC7C7991FBF7FB5A" box="[925,1049,1221,1242]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="bombetes">
<emphasis id="B950EA8D3573D928FC7C7991FBF7FB5A" box="[925,1049,1221,1242]" italics="true" pageId="5" pageNumber="536">A. bombetes</emphasis>
</taxonomicName>
group, we obtained intraspecific Cytb distances for these species from Amézquita 
<emphasis id="B950EA8D3573D928FDB579B0FD6FFB78" box="[596,641,1251,1273]" italics="true" pageId="5" pageNumber="536">et al</emphasis>
. (2013).
</paragraph>
</caption>
<paragraph id="8B9B369F3573D928FF267874FB92F849" blockId="5.[151,1436,1312,1994]" pageId="5" pageNumber="536">
The new species can be easily distinguished from most other known members of 
<taxonomicName id="4C244D1C3573D928FBB07874FB33FAB9" box="[1105,1245,1312,1337]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="genus">
<emphasis id="B950EA8D3573D928FBB07874FB33FAB9" box="[1105,1245,1312,1337]" italics="true" pageId="5" pageNumber="536">Andinobates</emphasis>
</taxonomicName>
by its distinctive solid red dorsal coloration (
<figureCitation id="131F2A1A3573D928FE2D7810FDFCFADD" box="[460,530,1348,1373]" captionStart="FIGURE 4" captionStartId="6.[151,250,1682,1704]" captionTargetBox="[187,1409,225,1623]" captionTargetId="figure@6.[151,1436,193,1661]" captionTargetPageId="6" captionText="FIGURE 4. Andinobates victimatus sp. nov. (A – K) and A. fulguritus (L – S). A – D: Lateral and dorsal views of an unsexed adult from the type locality. E – F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H – K: Juveniles of A. victimatus sp. nov. exhibiting intermediate stages of coloration development at ~ 1.5 months (H – I), ~ 2.5 months (J), and ~ 3 months (K). L – S: Unsexed A. fulguritus adults from Playa de Oro, Chocó (L – M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q – S), Photos in A – D and H – K by Roberto Márquez, E – F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L – S by Daniel Mejía- Vargas." httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="5" pageNumber="536">Fig. 4</figureCitation>
). The only exception is the recently described 
<taxonomicName id="4C244D1C3573D928FBFC7811FA76FADD" authority="Batista et al. 2014" authorityName="Batista et al." authorityYear="2014" box="[1053,1432,1348,1373]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="geminisae">
<emphasis id="B950EA8D3573D928FBFC7811FB40FADC" box="[1053,1198,1349,1372]" italics="true" pageId="5" pageNumber="536">A. geminisae</emphasis>
(
<bibRefCitation id="EFB54B6E3573D928FB5C7810FA7EFADD" author="Batista" box="[1213,1424,1348,1373]" pageId="5" pageNumber="536" refString="Batista, A., Jaramillo, C. A., Ponce, M. &amp; Crawford, A. J. (2014) A new species of Andinobates (Amphibia: Anura: Dendrobatidae) from west central Panama. Zootaxa, 3866 (3), 333 - 352." type="journal article" year="2014">
Batista 
<emphasis id="B950EA8D3573D928FAF57812FAA3FADD" box="[1300,1357,1348,1373]" italics="true" pageId="5" pageNumber="536">et al.</emphasis>
2014
</bibRefCitation>
)
</taxonomicName>
, which has uniform orange dorsal coloration, but lacks light gray/tan pigmentation on finger and toe tips, has uniform ventral coloration, and produces calls of the “tonal buzz” 
<typeStatus id="549F883D3573D928FC9C78D9FC5EFA25" box="[893,944,1421,1445]" pageId="5" pageNumber="536">type</typeStatus>
, with distinctive clicks at the beginning and end of the call, as opposed to the “short buzz” calls of 
<taxonomicName id="4C244D1C3573D928FCE578E5FC79FA49" box="[772,919,1457,1481]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="victimatus">
<emphasis id="B950EA8D3573D928FCE578E5FC79FA49" box="[772,919,1457,1481]" italics="true" pageId="5" pageNumber="536">A. victimatus</emphasis>
</taxonomicName>
. Moreover, calls of 
<taxonomicName id="4C244D1C3573D928FB9D78E5FAE1FA49" box="[1148,1295,1457,1481]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3573D928FB9D78E5FAE1FA49" box="[1148,1295,1457,1481]" italics="true" pageId="5" pageNumber="536">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63573D928FAF678E4FA9CFA49" box="[1303,1394,1456,1481]" pageId="5" pageNumber="536" rank="species">
<emphasis id="B950EA8D3573D928FAF678E4FA9CFA49" bold="true" box="[1303,1394,1456,1481]" pageId="5" pageNumber="536">sp. nov.</emphasis>
</taxonomicNameLabel>
are shorter in duration (0.42– 
<geoCoordinate id="EE1050583573D928FE5B7880FE10FA6C" box="[442,510,1492,1517]" direction="south" orientation="latitude" pageId="5" pageNumber="536" precision="555" value="-0.58">0.58 s</geoCoordinate>
vs. 1.52– 
<geoCoordinate id="EE1050583573D928FD8E7880FD4BFA6D" box="[623,677,1492,1517]" direction="south" orientation="latitude" pageId="5" pageNumber="536" precision="5555" value="-1.6">1.6 s</geoCoordinate>
) and higher in peak frequency (5.65–5.86 kHz vs. 4.40–4.50 kHz; 
<figureCitation id="131F2A1A3573D928FF7678ACFF35F991" box="[151,219,1528,1553]" captionStart="FIGURE 6" captionStartId="10.[151,250,998,1020]" captionTargetBox="[154,1436,510,955]" captionTargetId="figure@10.[151,1436,510,959]" captionTargetPageId="10" captionText="FIGURE 6. Comparisons of SVL and call traits between lowland species of Andinobates. Rectangles indicate the full range of values observed or reported in the literature. When available, means are represented as diamonds, with bars denoting 95 % confidence intervals (assuming normality; estimated as 1.96 times the standard error of the mean). SVL data were obtained from Batista et al. (2014), Jungfer et al. (2000), and Silverstone (1975), and bioacoustic data from Batista et al. (2014) and Brown &amp; Twomey et al. (2011). A. vic = A. victimatus, A. alt = A. altobueyensis, A. ful = A. fulguritus, A. vir = A. viridis, A. min = A. minutus, A. cla = A. claudiae, A. gem = A. geminisae." httpUri="https://zenodo.org/record/829285/files/figure.png" pageId="5" pageNumber="536">Fig. 6</figureCitation>
) than those of 
<taxonomicName id="4C244D1C3573D928FE6778ADFDF7F991" box="[390,537,1529,1553]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="geminisae">
<emphasis id="B950EA8D3573D928FE6778ADFDF7F991" box="[390,537,1529,1553]" italics="true" pageId="5" pageNumber="536">A. geminisae</emphasis>
</taxonomicName>
. Finally, 
<taxonomicName id="4C244D1C3573D928FD6578ADFCF9F991" box="[644,791,1529,1553]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="geminisae">
<emphasis id="B950EA8D3573D928FD6578ADFCF9F991" box="[644,791,1529,1553]" italics="true" pageId="5" pageNumber="536">A. geminisae</emphasis>
</taxonomicName>
is, on average, smaller than 
<taxonomicName id="4C244D1C3573D928FBBE78ADFB1CF991" box="[1119,1266,1529,1553]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3573D928FBBE78ADFB1CF991" box="[1119,1266,1529,1553]" italics="true" pageId="5" pageNumber="536">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63573D928FB1A78ACFAB8F991" box="[1275,1366,1528,1553]" pageId="5" pageNumber="536" rank="species">
<emphasis id="B950EA8D3573D928FB1A78ACFAB8F991" bold="true" box="[1275,1366,1528,1553]" pageId="5" pageNumber="536">sp. nov.</emphasis>
</taxonomicNameLabel>
(SVL 12.68 ± 
<quantity id="4CDC9B7A3573D928FF147B49FEB4F9B4" box="[245,346,1565,1589]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="5" pageNumber="536" unit="mm" value="0.87">0.87 mm</quantity>
vs. 14.23 ± 
<quantity id="4CDC9B7A3573D928FE087B48FDBFF9B4" box="[489,593,1564,1589]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="5" pageNumber="536" unit="mm" value="0.67">0.67 mm</quantity>
; 
<emphasis id="B950EA8D3573D928FDBC7B4AFD9AF9B9" box="[605,628,1566,1593]" italics="true" pageId="5" pageNumber="536">t15</emphasis>
= 3.78, p = 0.002, N = 6 
<emphasis id="B950EA8D3573D928FC7A7B49FC0AF9B4" box="[923,996,1565,1588]" italics="true" pageId="5" pageNumber="536">
<taxonomicName id="4C244D1C3573D928FC7A7B49FC31F9B4" box="[923,991,1565,1588]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="vic">A. vic</taxonomicName>
,
</emphasis>
11 
<taxonomicName id="4C244D1C3573D928FBEF7B49FBB3F9B5" box="[1038,1117,1565,1589]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="gem">
<emphasis id="B950EA8D3573D928FBEF7B49FBB3F9B5" box="[1038,1117,1565,1589]" italics="true" pageId="5" pageNumber="536">A. gem</emphasis>
</taxonomicName>
; 
<figureCitation id="131F2A1A3573D928FB8C7B48FB5CF9B5" box="[1133,1202,1564,1589]" captionStart="FIGURE 6" captionStartId="10.[151,250,998,1020]" captionTargetBox="[154,1436,510,955]" captionTargetId="figure@10.[151,1436,510,959]" captionTargetPageId="10" captionText="FIGURE 6. Comparisons of SVL and call traits between lowland species of Andinobates. Rectangles indicate the full range of values observed or reported in the literature. When available, means are represented as diamonds, with bars denoting 95 % confidence intervals (assuming normality; estimated as 1.96 times the standard error of the mean). SVL data were obtained from Batista et al. (2014), Jungfer et al. (2000), and Silverstone (1975), and bioacoustic data from Batista et al. (2014) and Brown &amp; Twomey et al. (2011). A. vic = A. victimatus, A. alt = A. altobueyensis, A. ful = A. fulguritus, A. vir = A. viridis, A. min = A. minutus, A. cla = A. claudiae, A. gem = A. geminisae." httpUri="https://zenodo.org/record/829285/files/figure.png" pageId="5" pageNumber="536">Fig. 6</figureCitation>
). However, the SVL ranges of both species overlap slightly (
<taxonomicName id="4C244D1C3573D928FDBB7B10FD02F9DB" box="[602,748,1604,1627]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="geminisae">
<emphasis id="B950EA8D3573D928FDBB7B10FD02F9DB" box="[602,748,1604,1627]" italics="true" pageId="5" pageNumber="536">A. geminisae</emphasis>
</taxonomicName>
: 
<quantity id="4CDC9B7A3573D928FD1A7B17FC55F9DB" box="[763,955,1603,1628]" metricMagnitude="-2" metricUnit="m" metricValue="1.263" metricValueMax="1.363" metricValueMin="1.163" pageId="5" pageNumber="536" unit="mm" value="12.63" valueMax="13.63" valueMin="11.63">11.63–13.63 mm</quantity>
, 
<taxonomicName id="4C244D1C3573D928FC297B10FBB5F9DB" box="[968,1115,1604,1627]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3573D928FC297B10FBB5F9DB" box="[968,1115,1604,1627]" italics="true" pageId="5" pageNumber="536">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63573D928FB837B17FB53F9DC" box="[1122,1213,1603,1628]" pageId="5" pageNumber="536" rank="species">
<emphasis id="B950EA8D3573D928FB837B17FB53F9DC" bold="true" box="[1122,1213,1603,1628]" pageId="5" pageNumber="536">sp. nov.</emphasis>
</taxonomicNameLabel>
: 
<quantity id="4CDC9B7A3573D928FB2D7B10FA7FF9DC" box="[1228,1425,1604,1628]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.498" metricValueMin="1.322" pageId="5" pageNumber="536" unit="mm" value="14.1" valueMax="14.98" valueMin="13.22">13.22–14.98 mm</quantity>
). Some individuals of 
<taxonomicName id="4C244D1C3573D928FE9F7B3CFDDFF900" box="[382,561,1639,1664]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="B950EA8D3573D928FE9F7B3CFDDFF900" box="[382,561,1639,1664]" italics="true" pageId="5" pageNumber="536">A. opisthomelas</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C244D1C3573D928FDDF7B3CFD31F900" box="[574,735,1639,1664]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="B950EA8D3573D928FDDF7B3CFD31F900" box="[574,735,1639,1664]" italics="true" pageId="5" pageNumber="536">A. virolinensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C244D1C3573D928FCFA7B3CFC32F900" box="[795,988,1639,1664]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="cassidyhornae">
<emphasis id="B950EA8D3573D928FCFA7B3CFC32F900" box="[795,988,1639,1664]" italics="true" pageId="5" pageNumber="536">A. cassidyhornae</emphasis>
</taxonomicName>
can have nearly homogenous red dorsal coloration (
<bibRefCitation id="EFB54B6E3573D928FEF97BDFFDF7F924" author="Amezquita" box="[280,537,1675,1700]" pageId="5" pageNumber="536" refString="Amezquita, A., Marquez, R., Medina, R., Mejia-Vargas, D., Kahn, T. R., Suarez, G. &amp; Mazariegos, L. (2013) A new species of Andean poison frog, Andinobates (Anura: Dendrobatidae), from the northwestern Andes of Colombia. Zootaxa, 3620 (1), 163 - 178." type="journal article" year="2013">
Amézquita 
<emphasis id="B950EA8D3573D928FE7C7BD8FE38F924" box="[413,470,1675,1700]" italics="true" pageId="5" pageNumber="536">et al.</emphasis>
2013
</bibRefCitation>
; 
<bibRefCitation id="EFB54B6E3573D928FDC77BDFFC0AF924" author="Ruiz-Carranza" box="[550,996,1675,1700]" pageId="5" pageNumber="536" refString="Ruiz-Carranza, P. &amp; Ramirez-Pinilla, M. P. (1992) Una nueva especie de Minyobates (Anura: Dendrobatidae) de Colombia. Lozania (Acta Zoologica Colombiana), 1992, 1 - 16." type="journal article" year="1992">Ruiz-Carranza &amp; Ramírez-Pinilla 1992</bibRefCitation>
; 
<bibRefCitation id="EFB54B6E3573D928FC107BDFFB5DF924" author="Silverstone" box="[1009,1203,1675,1700]" pageId="5" pageNumber="536" refString="Silverstone, P. A. (1975) A revision of the poison-arrow frogs of the genus Dendrobates Wagler. Natural History Museum of Los Angeles County, Science Bulletin, 21, 1 - 55." type="journal article" year="1975">Silverstone 1975</bibRefCitation>
). However, they can be distinguished from 
<taxonomicName id="4C244D1C3573D928FE7D7BE4FDDCF947" box="[412,562,1712,1735]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3573D928FE7D7BE4FDDCF947" box="[412,562,1712,1735]" italics="true" pageId="5" pageNumber="536">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63573D928FDDD7BFBFD77F948" box="[572,665,1711,1736]" pageId="5" pageNumber="536" rank="species">
<emphasis id="B950EA8D3573D928FDDD7BFBFD77F948" bold="true" box="[572,665,1711,1736]" pageId="5" pageNumber="536">sp. nov.</emphasis>
</taxonomicNameLabel>
by their ventral pattern which consists of red blotches on a black background in 
<taxonomicName id="4C244D1C3573D928FEB17B80FDF7F96C" box="[336,537,1747,1772]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="cassidyhornae">
<emphasis id="B950EA8D3573D928FEB17B80FDF7F96C" box="[336,537,1747,1772]" italics="true" pageId="5" pageNumber="536">A. cassidyhornae</emphasis>
</taxonomicName>
, light gray spots/marbling in 
<taxonomicName id="4C244D1C3573D928FC6E7B80FBD7F96C" box="[911,1081,1747,1772]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="B950EA8D3573D928FC6E7B80FBD7F96C" box="[911,1081,1747,1772]" italics="true" pageId="5" pageNumber="536">A. virolinensis</emphasis>
</taxonomicName>
and some populations of 
<taxonomicName id="4C244D1C3573D928FA657B80FEC5F890" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="B950EA8D3573D928FA657B80FEC5F890" italics="true" pageId="5" pageNumber="536">A. opisthomelas</emphasis>
</taxonomicName>
, and dark brown with irregular suffusions of red from the flanks in other populations of 
<taxonomicName id="4C244D1C3573D928FA657BACFEC5F8B4" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="B950EA8D3573D928FA657BACFEC5F8B4" italics="true" pageId="5" pageNumber="536">A. opisthomelas</emphasis>
</taxonomicName>
. Moreover, the three species are, on average, larger than 
<taxonomicName id="4C244D1C3573D928FC5C7A48FBBFF8B3" box="[957,1105,1820,1843]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="victimatus">
<emphasis id="B950EA8D3573D928FC5C7A48FBBFF8B3" box="[957,1105,1820,1843]" italics="true" pageId="5" pageNumber="536">A. victimatus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C244D1C3573D928FB857A48FAC8F8B4" box="[1124,1318,1819,1844]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="cassidyhornae">
<emphasis id="B950EA8D3573D928FB857A48FAC8F8B4" box="[1124,1318,1819,1844]" italics="true" pageId="5" pageNumber="536">A. cassidyhornae</emphasis>
</taxonomicName>
: 19.031 ± 
<quantity id="4CDC9B7A3573D928FF767A14FF13F8D7" box="[151,253,1856,1880]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="536" unit="mm" value="0.31">0.31 mm</quantity>
, 
<emphasis id="B950EA8D3573D928FEE87A14FEF1F8DC" box="[265,287,1856,1884]" italics="true" pageId="5" pageNumber="536">t11</emphasis>
= 17.04, p &lt;0.0001, N = 6 
<emphasis id="B950EA8D3573D928FDB67A14FD71F8D7" box="[599,671,1856,1879]" italics="true" pageId="5" pageNumber="536">
<taxonomicName id="4C244D1C3573D928FDB67A14FD74F8D7" box="[599,666,1856,1879]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="vic">A. vic</taxonomicName>
,
</emphasis>
7 
<taxonomicName id="4C244D1C3573D928FD5A7A14FCEEF8D7" box="[699,768,1856,1879]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="cas">
<emphasis id="B950EA8D3573D928FD5A7A14FCEEF8D7" box="[699,768,1856,1879]" italics="true" pageId="5" pageNumber="536">A. cas</emphasis>
</taxonomicName>
; 
<taxonomicName id="4C244D1C3573D928FCEE7A14FC5EF8D8" box="[783,944,1855,1880]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="B950EA8D3573D928FCEE7A14FC5EF8D8" box="[783,944,1855,1880]" italics="true" pageId="5" pageNumber="536">A. virolinensis</emphasis>
</taxonomicName>
: 16.86 ± 
<quantity id="4CDC9B7A3573D928FBFD7A14FB6DF8D7" box="[1052,1155,1856,1880]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="5" pageNumber="536" unit="mm" value="0.91">0.91 mm</quantity>
, 
<emphasis id="B950EA8D3573D928FB6F7A14FB4BF8DC" box="[1166,1189,1856,1884]" italics="true" pageId="5" pageNumber="536">t58</emphasis>
= 6.85, p &lt;0.0001, N = 6 
<emphasis id="B950EA8D3573D928FF277A33FEE1F8FE" box="[198,271,1895,1918]" italics="true" pageId="5" pageNumber="536">
<taxonomicName id="4C244D1C3573D928FF277A33FEE5F8FE" box="[198,267,1895,1918]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="vic">A. vic</taxonomicName>
,
</emphasis>
54 
<taxonomicName id="4C244D1C3573D928FEDC7A33FE93F8FE" box="[317,381,1895,1918]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="vir">
<emphasis id="B950EA8D3573D928FEDC7A33FE93F8FE" box="[317,381,1895,1918]" italics="true" pageId="5" pageNumber="536">A. vir</emphasis>
</taxonomicName>
; 
<taxonomicName id="4C244D1C3573D928FE6F7A33FDADF8FF" box="[398,579,1894,1919]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="B950EA8D3573D928FE6F7A33FDADF8FF" box="[398,579,1894,1919]" italics="true" pageId="5" pageNumber="536">A. opisthomelas</emphasis>
</taxonomicName>
: 16.80± 
<quantity id="4CDC9B7A3573D928FD437A32FCE4F8FE" box="[674,778,1894,1919]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="5" pageNumber="536" unit="mm" value="1.24">1.24 mm</quantity>
, 
<emphasis id="B950EA8D3573D928FCF67A33FCC0F803" box="[791,814,1895,1923]" italics="true" pageId="5" pageNumber="536">t53</emphasis>
= 5.14, p &lt;0.0001, N = 6 
<emphasis id="B950EA8D3573D928FB837A33FB45F8FE" box="[1122,1195,1895,1918]" italics="true" pageId="5" pageNumber="536">
<taxonomicName id="4C244D1C3573D928FB837A33FB48F8FE" box="[1122,1190,1895,1918]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="vic">A. vic</taxonomicName>
,
</emphasis>
49 
<taxonomicName id="4C244D1C3573D928FB397A33FAF3F8FE" box="[1240,1309,1895,1918]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="opi">
<emphasis id="B950EA8D3573D928FB397A33FAF3F8FE" box="[1240,1309,1895,1918]" italics="true" pageId="5" pageNumber="536">A. opi</emphasis>
</taxonomicName>
; data from original descriptions and 
<bibRefCitation id="EFB54B6E3573D928FE557AD8FD9AF825" author="Silverstone" box="[436,628,1932,1957]" pageId="5" pageNumber="536" refString="Silverstone, P. A. (1975) A revision of the poison-arrow frogs of the genus Dendrobates Wagler. Natural History Museum of Los Angeles County, Science Bulletin, 21, 1 - 55." type="journal article" year="1975">Silverstone 1975</bibRefCitation>
), although, again, with slight range overlap with 
<taxonomicName id="4C244D1C3573D928FB7C7AD9FAD0F825" box="[1181,1342,1932,1957]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="B950EA8D3573D928FB7C7AD9FAD0F825" box="[1181,1342,1932,1957]" italics="true" pageId="5" pageNumber="536">A. virolinensis</emphasis>
</taxonomicName>
(
<quantity id="4CDC9B7A3573D928FAAC7AD9FEE7F849" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.498" metricValueMin="1.322" pageId="5" pageNumber="536" unit="mm" value="14.1" valueMax="14.98" valueMin="13.22">13.22– 14.98 mm</quantity>
vs. 
<quantity id="4CDC9B7A3573D928FED77AE5FE35F849" box="[310,475,1969,1994]" metricMagnitude="-2" metricUnit="m" metricValue="1.675" metricValueMax="1.89" metricValueMin="1.46" pageId="5" pageNumber="536" unit="mm" value="16.75" valueMax="18.9" valueMin="14.6">14.6–18.9 mm</quantity>
) and 
<taxonomicName id="4C244D1C3573D928FDF97AE6FD25F84A" box="[536,715,1969,1994]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="536" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="B950EA8D3573D928FDF97AE6FD25F84A" box="[536,715,1969,1994]" italics="true" pageId="5" pageNumber="536">A. opisthomelas</emphasis>
</taxonomicName>
(
<quantity id="4CDC9B7A3573D928FD3B7AE5FC75F849" box="[730,923,1969,1993]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.498" metricValueMin="1.322" pageId="5" pageNumber="536" unit="mm" value="14.1" valueMax="14.98" valueMin="13.22">13.22–14.98 mm</quantity>
vs. 
<quantity id="4CDC9B7A3573D928FC297AE5FB81F849" box="[968,1135,1969,1994]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.95" metricValueMin="1.45" pageId="5" pageNumber="536" unit="mm" value="17.0" valueMax="19.5" valueMin="14.5">14.5–19.5 mm</quantity>
).
</paragraph>
<caption id="DF5B66173570D92BFF767BC6FF0DF8E0" httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="6" pageNumber="537" targetBox="[187,1409,225,1623]" targetPageId="6">
<paragraph id="8B9B369F3570D92BFF767BC6FF0DF8E0" blockId="6.[151,1436,1682,1888]" pageId="6" pageNumber="537">
<emphasis id="B950EA8D3570D92BFF767BC6FEFCF927" bold="true" box="[151,274,1682,1704]" pageId="6" pageNumber="537">FIGURE 4.</emphasis>
<taxonomicName id="4C244D1C3570D92BFEF97BC7FDEAF928" box="[280,516,1683,1704]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3570D92BFEF97BC7FDEAF928" box="[280,516,1683,1704]" italics="true" pageId="6" pageNumber="537">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63570D92BFDEB7BC7FDB4F928" box="[522,602,1683,1704]" pageId="6" pageNumber="537" rank="species">
<emphasis id="B950EA8D3570D92BFDEB7BC7FDB4F928" bold="true" box="[522,602,1683,1704]" pageId="6" pageNumber="537">sp. nov.</emphasis>
</taxonomicNameLabel>
(A–K) and 
<taxonomicName id="4C244D1C3570D92BFD337BC7FCBFF928" box="[722,849,1683,1704]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D3570D92BFD337BC7FCBFF928" box="[722,849,1683,1704]" italics="true" pageId="6" pageNumber="537">A. fulguritus</emphasis>
</taxonomicName>
(L–S). A–D: Lateral and dorsal views of an unsexed adult from the type locality. E–F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H–K: Juveniles of 
<taxonomicName id="4C244D1C3570D92BFE227B84FDA7F965" box="[451,585,1744,1765]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3570D92BFE227B84FDA7F965" box="[451,585,1744,1765]" italics="true" pageId="6" pageNumber="537">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63570D92BFDB37B84FD4AF965" box="[594,676,1744,1765]" pageId="6" pageNumber="537" rank="species">
<emphasis id="B950EA8D3570D92BFDB37B84FD4AF965" bold="true" box="[594,676,1744,1765]" pageId="6" pageNumber="537">sp. nov.</emphasis>
</taxonomicNameLabel>
exhibiting intermediate stages of coloration development at ~1.5 months (H–I), ~2.5 months (J), and ~3 months (K). L–S: Unsexed 
<taxonomicName id="4C244D1C3570D92BFD0D7BBBFC82F883" box="[748,876,1774,1796]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D3570D92BFD0D7BBBFC82F883" box="[748,876,1774,1796]" italics="true" pageId="6" pageNumber="537">A. fulguritus</emphasis>
</taxonomicName>
adults from Playa de Oro, Chocó (L–M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q–S), Photos in A–D and H–K by Roberto Márquez, E–F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L–S by Daniel Mejía- Vargas.
</paragraph>
</caption>
<paragraph id="8B9B369F3570D92AFF267ADCFD12FF54" blockId="6.[151,1436,1927,2024]" lastBlockId="7.[151,1436,151,213]" lastPageId="7" lastPageNumber="538" pageId="6" pageNumber="537">
In addition to coloration, 
<taxonomicName id="4C244D1C3570D92BFDE67ADCFD4CF820" box="[519,674,1928,1952]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3570D92BFDE67ADCFD4CF820" box="[519,674,1928,1952]" italics="true" pageId="6" pageNumber="537">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63570D92BFD537AD3FCFBF820" box="[690,789,1927,1952]" pageId="6" pageNumber="537" rank="species">
<emphasis id="B950EA8D3570D92BFD537AD3FCFBF820" bold="true" box="[690,789,1927,1952]" pageId="6" pageNumber="537">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from other lowland species of 
<taxonomicName id="4C244D1C3570D92BFF767AFFFECDF844" box="[151,291,1963,1988]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="genus">
<emphasis id="B950EA8D3570D92BFF767AFFFECDF844" box="[151,291,1963,1988]" italics="true" pageId="6" pageNumber="537">Andinobates</emphasis>
</taxonomicName>
(i.e. 
<taxonomicName id="4C244D1C3570D92BFEBD7AF8FE3AF844" box="[348,468,1964,1988]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B950EA8D3570D92BFEBD7AF8FE3AF844" box="[348,468,1964,1988]" italics="true" pageId="6" pageNumber="537">A. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C244D1C3570D92BFDEB7AF8FD78F844" box="[522,662,1963,1988]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D3570D92BFDEB7AF8FD78F844" box="[522,662,1963,1988]" italics="true" pageId="6" pageNumber="537">A. fulguritus</emphasis>
</taxonomicName>
groups) in the following ways (
<figureCitation id="131F2A1A3570D92BFC1D7AFFFBAFF844" box="[1020,1089,1963,1988]" captionStart="FIGURE 6" captionStartId="10.[151,250,998,1020]" captionTargetBox="[154,1436,510,955]" captionTargetId="figure@10.[151,1436,510,959]" captionTargetPageId="10" captionText="FIGURE 6. Comparisons of SVL and call traits between lowland species of Andinobates. Rectangles indicate the full range of values observed or reported in the literature. When available, means are represented as diamonds, with bars denoting 95 % confidence intervals (assuming normality; estimated as 1.96 times the standard error of the mean). SVL data were obtained from Batista et al. (2014), Jungfer et al. (2000), and Silverstone (1975), and bioacoustic data from Batista et al. (2014) and Brown &amp; Twomey et al. (2011). A. vic = A. victimatus, A. alt = A. altobueyensis, A. ful = A. fulguritus, A. vir = A. viridis, A. min = A. minutus, A. cla = A. claudiae, A. gem = A. geminisae." httpUri="https://zenodo.org/record/829285/files/figure.png" pageId="6" pageNumber="537">Fig. 6</figureCitation>
): The calls of 
<taxonomicName id="4C244D1C3570D92BFB007AF8FA9DF844" box="[1249,1395,1964,1988]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3570D92BFB007AF8FA9DF844" box="[1249,1395,1964,1988]" italics="true" pageId="6" pageNumber="537">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63570D92BFA9B7AFFFF26F868" pageId="6" pageNumber="537" rank="species">
<emphasis id="B950EA8D3570D92BFA9B7AFFFF26F868" bold="true" pageId="6" pageNumber="537">sp. nov.</emphasis>
</taxonomicNameLabel>
are shorter than those of 
<taxonomicName id="4C244D1C3570D92BFE127A84FD9BF868" box="[499,629,1999,2024]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="537" phylum="Chordata" rank="species" species="claudiae">
<emphasis id="B950EA8D3570D92BFE127A84FD9BF868" box="[499,629,1999,2024]" italics="true" pageId="6" pageNumber="537">A. claudiae</emphasis>
</taxonomicName>
(0.42– 
<geoCoordinate id="EE1050583570D92BFD267A9BFCE2F867" box="[711,780,1999,2024]" direction="south" orientation="latitude" pageId="6" pageNumber="537" precision="555" value="-0.58">0.58 s</geoCoordinate>
vs. 0.85– 
<geoCoordinate id="EE1050583570D92BFC617A9BFC2AF868" box="[896,964,1999,2024]" direction="south" orientation="latitude" pageId="6" pageNumber="537" precision="555" value="-1.03">1.03 s</geoCoordinate>
), and have a higher peak frequency than those of 
<taxonomicName id="4C244D1C3571D92AFF1D7DCCFE64FF30" box="[252,394,151,176]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="538" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D3571D92AFF1D7DCCFE64FF30" box="[252,394,151,176]" italics="true" pageId="7" pageNumber="538">A. fulguritus</emphasis>
</taxonomicName>
(5.65–5.86 kHz vs. 4.83–5.16 kHz); 
<taxonomicName id="4C244D1C3571D92AFCDB7DCCFC21FF30" box="[826,975,152,176]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="538" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3571D92AFCDB7DCCFC21FF30" box="[826,975,152,176]" italics="true" pageId="7" pageNumber="538">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63571D92AFC397DC3FBDAFF30" box="[984,1076,151,176]" pageId="7" pageNumber="538" rank="species">
<emphasis id="B950EA8D3571D92AFC397DC3FBDAFF30" bold="true" box="[984,1076,151,176]" pageId="7" pageNumber="538">sp. nov.</emphasis>
</taxonomicNameLabel>
is smaller in body size than 
<taxonomicName id="4C244D1C3571D92AFA657DCCFEC1FF55" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="538" phylum="Chordata" rank="species" species="altobueyensis">
<emphasis id="B950EA8D3571D92AFA657DCCFEC1FF55" italics="true" pageId="7" pageNumber="538">A. altobueyensis</emphasis>
</taxonomicName>
(
<quantity id="4CDC9B7A3571D92AFEDF7DE8FE11FF54" box="[318,511,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.498" metricValueMin="1.322" pageId="7" pageNumber="538" unit="mm" value="14.1" valueMax="14.98" valueMin="13.22">13.22–14.98 mm</quantity>
vs. 16.30–17.00 mm).
</paragraph>
</subSubSection>
<caption id="DF5B66173571D92AFF767C50FEC0FEBB" pageId="7" pageNumber="538">
<paragraph id="8B9B369F3571D92AFF767C50FEC0FEBB" blockId="7.[151,1436,260,315]" pageId="7" pageNumber="538">
<emphasis id="B950EA8D3571D92AFF767C50FEE2FE9B" bold="true" box="[151,268,260,283]" pageId="7" pageNumber="538">TABLE 1.</emphasis>
Measurements of the type series of 
<taxonomicName id="4C244D1C3571D92AFD417C50FCC3FE9B" box="[672,813,260,283]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="538" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D3571D92AFD417C50FCC3FE9B" box="[672,813,260,283]" italics="true" pageId="7" pageNumber="538">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F63571D92AFCD67C50FC61FE9B" box="[823,911,260,283]" pageId="7" pageNumber="538" rank="species">
<emphasis id="B950EA8D3571D92AFCD67C50FC61FE9B" bold="true" box="[823,911,260,283]" pageId="7" pageNumber="538">sp. nov.</emphasis>
</taxonomicNameLabel>
in mm. See methods section for explanation of abbreviations.
</paragraph>
</caption>
<paragraph id="8B9B369F3571D92AFF7E7C02FEE9FB54" pageId="7" pageNumber="538">
<table id="F924C43F357126D2FF767C02FA97FB54" box="[151,1401,342,1236]" gridcols="10" gridrows="8" pageId="7" pageNumber="538">
<tr id="351434DD357126D2FF767C02FA97FE34" box="[151,1401,342,436]" gridrow="0" pageId="7" pageNumber="538">
<th id="76C55DA1357126D2FF767C02FEE0FE34" box="[151,270,342,436]" gridcol="0" gridrow="0" pageId="7" pageNumber="538">Dimension SVL</th>
<th id="76C55DA1357126D2FEC77C02FE7BFE34" box="[294,405,342,436]" gridcol="1" gridrow="0" pageId="7" pageNumber="538">ANDES-A 3684 ♂ 14.42</th>
<th id="76C55DA1357126D2FE527C02FDCDFE34" box="[435,547,342,436]" gridcol="2" gridrow="0" pageId="7" pageNumber="538">ANDES-A 3685 ♀ 14.5</th>
<th id="76C55DA1357126D2FDA97C02FD59FE34" box="[584,695,342,436]" gridcol="3" gridrow="0" pageId="7" pageNumber="538">ANDES-A 3686* ♀ 13.22</th>
<th id="76C55DA1357126D2FD3C7C02FCA2FE34" box="[733,844,342,436]" gridcol="4" gridrow="0" pageId="7" pageNumber="538">ANDES-A 3687 ♂ 13.62</th>
<th id="76C55DA1357126D2FC937C02FC0FFE34" box="[882,993,342,436]" gridcol="5" gridrow="0" pageId="7" pageNumber="538">ANDES-A 3688 ♂ 14.67</th>
<th id="76C55DA1357126D2FBE67C02FB98FE34" box="[1031,1142,342,436]" gridcol="6" gridrow="0" pageId="7" pageNumber="538">ANDES-A 3689 ♂ 14.98</th>
<th id="76C55DA1357126D2FB727C02FB23FE34" box="[1171,1229,342,436]" gridcol="7" gridrow="0" pageId="7" pageNumber="538">Mean 14.23</th>
<th id="76C55DA1357126D2FB177C02FACCFE34" box="[1270,1314,342,436]" gridcol="8" gridrow="0" pageId="7" pageNumber="538">SE 0.27</th>
<th id="76C55DA1357126D2FAAC7C02FA97FE34" box="[1357,1401,342,436]" gridcol="9" gridrow="0" pageId="7" pageNumber="538">SD 0.67</th>
</tr>
<tr id="351434DD357126D2FF767C9CFA97FDB0" box="[151,1401,456,560]" gridrow="1" pageId="7" pageNumber="538">
<th id="76C55DA1357126D2FF767C9CFEE0FDB0" box="[151,270,456,560]" gridcol="0" gridrow="1" pageId="7" pageNumber="538">TL GBW HW</th>
<td id="76C55DA1357126D2FEC77C9CFE7BFDB0" box="[294,405,456,560]" gridcol="1" gridrow="1" pageId="7" pageNumber="538">6.17 4.6 4.74</td>
<td id="76C55DA1357126D2FE527C9CFDCDFDB0" box="[435,547,456,560]" gridcol="2" gridrow="1" pageId="7" pageNumber="538">6.31 4.16 4.31</td>
<td id="76C55DA1357126D2FDA97C9CFD59FDB0" box="[584,695,456,560]" gridcol="3" gridrow="1" pageId="7" pageNumber="538">6.32 3.89 3.90</td>
<td id="76C55DA1357126D2FD3C7C9CFCA2FDB0" box="[733,844,456,560]" gridcol="4" gridrow="1" pageId="7" pageNumber="538">6.57 3.93 4.64</td>
<td id="76C55DA1357126D2FC937C9CFC0FFDB0" box="[882,993,456,560]" gridcol="5" gridrow="1" pageId="7" pageNumber="538">6.35 3.78 4.54</td>
<td id="76C55DA1357126D2FBE67C9CFB98FDB0" box="[1031,1142,456,560]" gridcol="6" gridrow="1" pageId="7" pageNumber="538">6.50 4.64 4.60</td>
<td id="76C55DA1357126D2FB727C9CFB23FDB0" box="[1171,1229,456,560]" gridcol="7" gridrow="1" pageId="7" pageNumber="538">6.37 4.17 4.46</td>
<td id="76C55DA1357126D2FB177C9CFACCFDB0" box="[1270,1314,456,560]" gridcol="8" gridrow="1" pageId="7" pageNumber="538">0.06 0.15 0.13</td>
<td id="76C55DA1357126D2FAAC7C9CFA97FDB0" box="[1357,1401,456,560]" gridcol="9" gridrow="1" pageId="7" pageNumber="538">0.14 0.37 0.31</td>
</tr>
<tr id="351434DD357126D2FF767F10FA97FD2C" box="[151,1401,580,684]" gridrow="2" pageId="7" pageNumber="538">
<th id="76C55DA1357126D2FF767F10FEE0FD2C" box="[151,270,580,684]" gridcol="0" gridrow="2" pageId="7" pageNumber="538">IOD HL TSCN</th>
<td id="76C55DA1357126D2FEC77F10FE7BFD2C" box="[294,405,580,684]" gridcol="1" gridrow="2" pageId="7" pageNumber="538">3.92 3.42 0.69</td>
<td id="76C55DA1357126D2FE527F10FDCDFD2C" box="[435,547,580,684]" gridcol="2" gridrow="2" pageId="7" pageNumber="538">4.01 3.57 0.80</td>
<td id="76C55DA1357126D2FDA97F10FD59FD2C" box="[584,695,580,684]" gridcol="3" gridrow="2" pageId="7" pageNumber="538">3.58 3.31 0.76</td>
<td id="76C55DA1357126D2FD3C7F10FCA2FD2C" box="[733,844,580,684]" gridcol="4" gridrow="2" pageId="7" pageNumber="538">4.04 3.45 0.83</td>
<td id="76C55DA1357126D2FC937F10FC0FFD2C" box="[882,993,580,684]" gridcol="5" gridrow="2" pageId="7" pageNumber="538">3.44 3.12 0.79</td>
<td id="76C55DA1357126D2FBE67F10FB98FD2C" box="[1031,1142,580,684]" gridcol="6" gridrow="2" pageId="7" pageNumber="538">3.80 3.61 0.84</td>
<td id="76C55DA1357126D2FB727F10FB23FD2C" box="[1171,1229,580,684]" gridcol="7" gridrow="2" pageId="7" pageNumber="538">3.80 3.41 0.78</td>
<td id="76C55DA1357126D2FB177F10FACCFD2C" box="[1270,1314,580,684]" gridcol="8" gridrow="2" pageId="7" pageNumber="538">0.10 0.07 0.02</td>
<td id="76C55DA1357126D2FAAC7F10FA97FD2C" box="[1357,1401,580,684]" gridcol="9" gridrow="2" pageId="7" pageNumber="538">0.24 0.18 0.05</td>
</tr>
<tr id="351434DD357126D2FF767F94FA97FCA8" box="[151,1401,704,808]" gridrow="3" pageId="7" pageNumber="538">
<th id="76C55DA1357126D2FF767F94FEE0FCA8" box="[151,270,704,808]" gridcol="0" gridrow="3" pageId="7" pageNumber="538">NED IND EL</th>
<td id="76C55DA1357126D2FEC77F94FE7BFCA8" box="[294,405,704,808]" gridcol="1" gridrow="3" pageId="7" pageNumber="538">1.02 1.85 1.70</td>
<td id="76C55DA1357126D2FE527F94FDCDFCA8" box="[435,547,704,808]" gridcol="2" gridrow="3" pageId="7" pageNumber="538">1.14 1.97 1.87</td>
<td id="76C55DA1357126D2FDA97F94FD59FCA8" box="[584,695,704,808]" gridcol="3" gridrow="3" pageId="7" pageNumber="538">0.94 1.81 1.74</td>
<td id="76C55DA1357126D2FD3C7F94FCA2FCA8" box="[733,844,704,808]" gridcol="4" gridrow="3" pageId="7" pageNumber="538">1.18 1.94 1.79</td>
<td id="76C55DA1357126D2FC937F94FC0FFCA8" box="[882,993,704,808]" gridcol="5" gridrow="3" pageId="7" pageNumber="538">0.97 1.79 1.80</td>
<td id="76C55DA1357126D2FBE67F94FB98FCA8" box="[1031,1142,704,808]" gridcol="6" gridrow="3" pageId="7" pageNumber="538">1.43 1.56 1.95</td>
<td id="76C55DA1357126D2FB727F94FB23FCA8" box="[1171,1229,704,808]" gridcol="7" gridrow="3" pageId="7" pageNumber="538">1.11 1.82 1.81</td>
<td id="76C55DA1357126D2FB177F94FACCFCA8" box="[1270,1314,704,808]" gridcol="8" gridrow="3" pageId="7" pageNumber="538">0.07 0.06 0.04</td>
<td id="76C55DA1357126D2FAAC7F94FA97FCA8" box="[1357,1401,704,808]" gridcol="9" gridrow="3" pageId="7" pageNumber="538">0.18 0.15 0.09</td>
</tr>
<tr id="351434DD357126D2FF767E68FA97FC24" box="[151,1401,828,932]" gridrow="4" pageId="7" pageNumber="538">
<th id="76C55DA1357126D2FF767E68FEE0FC24" box="[151,270,828,932]" gridcol="0" gridrow="4" pageId="7" pageNumber="538">HDT MTD HaL</th>
<td id="76C55DA1357126D2FEC77E68FE7BFC24" box="[294,405,828,932]" gridcol="1" gridrow="4" pageId="7" pageNumber="538">0.69 0.30 3.68</td>
<td id="76C55DA1357126D2FE527E68FDCDFC24" box="[435,547,828,932]" gridcol="2" gridrow="4" pageId="7" pageNumber="538">0.91 0.33 3.74</td>
<td id="76C55DA1357126D2FDA97E68FD59FC24" box="[584,695,828,932]" gridcol="3" gridrow="4" pageId="7" pageNumber="538">0.72 0.32 3.52</td>
<td id="76C55DA1357126D2FD3C7E68FCA2FC24" box="[733,844,828,932]" gridcol="4" gridrow="4" pageId="7" pageNumber="538">0.64 0.33 3.61</td>
<td id="76C55DA1357126D2FC937E68FC0FFC24" box="[882,993,828,932]" gridcol="5" gridrow="4" pageId="7" pageNumber="538">0.64 0.34 3.76</td>
<td id="76C55DA1357126D2FBE67E68FB98FC24" box="[1031,1142,828,932]" gridcol="6" gridrow="4" pageId="7" pageNumber="538">0.97 0.72 3.72</td>
<td id="76C55DA1357126D2FB727E68FB23FC24" box="[1171,1229,828,932]" gridcol="7" gridrow="4" pageId="7" pageNumber="538">0.76 0.39 3.67</td>
<td id="76C55DA1357126D2FB177E68FACCFC24" box="[1270,1314,828,932]" gridcol="8" gridrow="4" pageId="7" pageNumber="538">0.06 0.07 0.04</td>
<td id="76C55DA1357126D2FAAC7E68FA97FC24" box="[1357,1401,828,932]" gridcol="9" gridrow="4" pageId="7" pageNumber="538">0.14 0.16 0.09</td>
</tr>
<tr id="351434DD357126D2FF767EECFA97FBA0" box="[151,1401,952,1056]" gridrow="5" pageId="7" pageNumber="538">
<th id="76C55DA1357126D2FF767EECFEE0FBA0" box="[151,270,952,1056]" gridcol="0" gridrow="5" pageId="7" pageNumber="538">W3FD W3F W3TD</th>
<td id="76C55DA1357126D2FEC77EECFE7BFBA0" box="[294,405,952,1056]" gridcol="1" gridrow="5" pageId="7" pageNumber="538">0.42 0.22 0.32</td>
<td id="76C55DA1357126D2FE527EECFDCDFBA0" box="[435,547,952,1056]" gridcol="2" gridrow="5" pageId="7" pageNumber="538">0.6 0.27 0.4</td>
<td id="76C55DA1357126D2FDA97EECFD59FBA0" box="[584,695,952,1056]" gridcol="3" gridrow="5" pageId="7" pageNumber="538">0.51 0.27 0.46</td>
<td id="76C55DA1357126D2FD3C7EECFCA2FBA0" box="[733,844,952,1056]" gridcol="4" gridrow="5" pageId="7" pageNumber="538">0.55 0.3 0.34</td>
<td id="76C55DA1357126D2FC937EECFC0FFBA0" box="[882,993,952,1056]" gridcol="5" gridrow="5" pageId="7" pageNumber="538">0.63 0.32 0.38</td>
<td id="76C55DA1357126D2FBE67EECFB98FBA0" box="[1031,1142,952,1056]" gridcol="6" gridrow="5" pageId="7" pageNumber="538">0.66 0.49 0.40</td>
<td id="76C55DA1357126D2FB727EECFB23FBA0" box="[1171,1229,952,1056]" gridcol="7" gridrow="5" pageId="7" pageNumber="538">0.56 0.31 0.38</td>
<td id="76C55DA1357126D2FB177EECFACCFBA0" box="[1270,1314,952,1056]" gridcol="8" gridrow="5" pageId="7" pageNumber="538">0.04 0.04 0.02</td>
<td id="76C55DA1357126D2FAAC7EECFA97FBA0" box="[1357,1401,952,1056]" gridcol="9" gridrow="5" pageId="7" pageNumber="538">0.09 0.09 0.05</td>
</tr>
<tr id="351434DD357126D2FF767960FA97FB1C" box="[151,1401,1076,1180]" gridrow="6" pageId="7" pageNumber="538">
<th id="76C55DA1357126D2FF767960FEE0FB1C" box="[151,270,1076,1180]" gridcol="0" gridrow="6" pageId="7" pageNumber="538">W3T W4TD W4T</th>
<td id="76C55DA1357126D2FEC77960FE7BFB1C" box="[294,405,1076,1180]" gridcol="1" gridrow="6" pageId="7" pageNumber="538">0.23 0.39 0.26</td>
<td id="76C55DA1357126D2FE527960FDCDFB1C" box="[435,547,1076,1180]" gridcol="2" gridrow="6" pageId="7" pageNumber="538">0.28 0.49 0.25</td>
<td id="76C55DA1357126D2FDA97960FD59FB1C" box="[584,695,1076,1180]" gridcol="3" gridrow="6" pageId="7" pageNumber="538">0.26 0.52 0.34</td>
<td id="76C55DA1357126D2FD3C7960FCA2FB1C" box="[733,844,1076,1180]" gridcol="4" gridrow="6" pageId="7" pageNumber="538">0.21 0.41 0.32</td>
<td id="76C55DA1357126D2FC937960FC0FFB1C" box="[882,993,1076,1180]" gridcol="5" gridrow="6" pageId="7" pageNumber="538">0.27 0.45 0.30</td>
<td id="76C55DA1357126D2FBE67960FB98FB1C" box="[1031,1142,1076,1180]" gridcol="6" gridrow="6" pageId="7" pageNumber="538">0.38 0.40 0.40</td>
<td id="76C55DA1357126D2FB727960FB23FB1C" box="[1171,1229,1076,1180]" gridcol="7" gridrow="6" pageId="7" pageNumber="538">0.27 0.44 0.31</td>
<td id="76C55DA1357126D2FB177960FACCFB1C" box="[1270,1314,1076,1180]" gridcol="8" gridrow="6" pageId="7" pageNumber="538">0.02 0.02 0.02</td>
<td id="76C55DA1357126D2FAAC7960FA97FB1C" box="[1357,1401,1076,1180]" gridcol="9" gridrow="6" pageId="7" pageNumber="538">0.06 0.05 0.06</td>
</tr>
<tr id="351434DD357126D2FF7679E9FA97FB54" box="[151,1401,1213,1236]" gridrow="7" pageId="7" pageNumber="538" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76C55DA1357126D2FF7679E9FEE0FB54" box="[151,270,1213,1236]" gridcol="0" gridrow="7" pageId="7" pageNumber="538">*Holotype</th>
</tr>
</table>
</paragraph>
<subSubSection id="C33E65143571D925FF2679A9FE0BFA40" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538" type="description">
<paragraph id="8B9B369F3571D92AFF2679A9FB4EF9FE" blockId="7.[151,1436,1277,2022]" pageId="7" pageNumber="538">
<emphasis id="B950EA8D3571D92AFF2679A9FDF4FA96" bold="true" box="[199,538,1277,1302]" pageId="7" pageNumber="538">
Description of the 
<typeStatus id="549F883D3571D92AFE4D79A9FDFBFA96" box="[428,533,1277,1302]" pageId="7" pageNumber="538" type="holotype">holotype</typeStatus>
.
</emphasis>
An adult female, 
<quantity id="4CDC9B7A3571D92AFD0E79AAFC8AFA95" box="[751,868,1278,1302]" metricMagnitude="-2" metricUnit="m" metricValue="1.322" pageId="7" pageNumber="538" unit="mm" value="13.22">13.22 mm</quantity>
in snout-vent length. Snout dorsally subtruncate, laterally rounded and slightly protrusive; 
<emphasis id="B950EA8D3571D92AFD857875FCCEFABA" box="[612,800,1313,1338]" italics="true" pageId="7" pageNumber="538">canthus rostralis</emphasis>
rounded; loreal region vertical and slightly concave. The head is wider than it is long, and the same width as the body (HW/GBW = 1.002); head width 29.5% of SVL; head length 25.0% of SVL; nares 1.24 times closer to tip of snout than to eyes; pupil rounded and horizontally elliptical; eye length 52.6% of head length. Tympanic ring visible, oval shaped, of moderate size (HDT 41.4% of EL); posterior and dorsal margins hidden under skin. Maxillary and premaxillary teeth absent; median lingual process absent; tongue about two times longer than wide; anterior margin of tongue not indented; posterior third of tongue not attached to floor of mouth. Hands of moderate size (HaL 26.6% of SVL); adpressed finger lengths are I&lt;II&lt;IV&lt;III; adpressed toe lengths are I&lt;II&lt;V&lt;III&lt;IV; discs of fingers II, III, and IV moderately expanded; toe IV disc weakly expanded; globular, rounded subarticular tubercles; outer palmar tubercle larger than inner palmar tubercle; outer plantar tubercle smaller than inner plantar tubercle; no foot or hand webbing.
</paragraph>
<paragraph id="8B9B369F3571D92AFF267BDEFB65F81E" blockId="7.[151,1436,1277,2022]" pageId="7" pageNumber="538">In life, the dorsum was homogeneous scarlet red, with faintly visible dorsolateral stripes; flanks were mostly uniform vermilion, with a few small, dark specks towards the ventral area. The venter was also vermilion with lesssaturated blotches transitioning from vermilion to black towards the chest and throat. Fingers and toes were light metallic gray, and palms and soles were black. The skin was slightly granular, especially in the posterior venter, cloaca, and dorsal surface of hind limbs. In ethanol (70%), the dorsum and dorsal surfaces of limbs are metallic gray; hands, feet, and articulations are tan to light brown. The venter and flanks are also metallic gray, with some scattered dark brown specks; a large dark brown blotch is present on the throat. The skin is mostly smooth, except for the ventral surface of the hind limbs and cloaca, where weak granularity is still visible.</paragraph>
<paragraph id="8B9B369F3571D925FF267AFDFF08FE08" blockId="7.[151,1436,1277,2022]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="539" pageId="7" pageNumber="538">
<emphasis id="B950EA8D3571D92AFF267AFDFD85F842" bold="true" box="[199,619,1961,1986]" pageId="7" pageNumber="538">
Coloration in life (
<figureCitation id="131F2A1A3571D92AFE487AFDFDF7F842" box="[425,537,1961,1986]" captionStart="FIGURE 4" captionStartId="6.[151,250,1682,1704]" captionTargetBox="[187,1409,225,1623]" captionTargetId="figure@6.[151,1436,193,1661]" captionTargetPageId="6" captionText="FIGURE 4. Andinobates victimatus sp. nov. (A – K) and A. fulguritus (L – S). A – D: Lateral and dorsal views of an unsexed adult from the type locality. E – F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H – K: Juveniles of A. victimatus sp. nov. exhibiting intermediate stages of coloration development at ~ 1.5 months (H – I), ~ 2.5 months (J), and ~ 3 months (K). L – S: Unsexed A. fulguritus adults from Playa de Oro, Chocó (L – M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q – S), Photos in A – D and H – K by Roberto Márquez, E – F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L – S by Daniel Mejía- Vargas." httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="7" pageNumber="538">Figs. 4, 4</figureCitation>
A–K).
</emphasis>
The dorsum and dorsal surfaces of limbs of all individuals in the 
<typeStatus id="549F883D3571D92AFA8D7AFEFA72F842" box="[1388,1436,1962,1986]" pageId="7" pageNumber="538">type</typeStatus>
series are homogenous scarlet red. In some collected and uncollected individuals, a faint, complete dorsolateral stripe (
<typeStatus id="549F883D357ED925FF057DCCFEF8FF30" box="[228,278,152,176]" pageId="8" pageNumber="539">type</typeStatus>
A in 
<bibRefCitation id="EFB54B6E357ED925FEB57DCCFDFDFF30" author="Grant" box="[340,531,151,176]" pageId="8" pageNumber="539" refString="Grant, T., Frost, D. R., Caldwell, J. P., Gagliardo, R., Haddad, C. F. B., Kok, P. J. R., Means, D. B., Noonan, B. P., Schargel, W. E. &amp; Wheeler, W. C. (2006) Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, 299, 1 - 262. https: // doi. org / 10.1206 / 0003 - 0090 (2006) 299 [1: PSODFA] 2.0. CO; 2" type="journal article" year="2006">
Grant 
<emphasis id="B950EA8D357ED925FE7A7DCDFE3DFF30" box="[411,467,151,176]" italics="true" pageId="8" pageNumber="539">et al.</emphasis>
2006
</bibRefCitation>
) of a more saturated shade of red is visible (
<figureCitation id="131F2A1A357ED925FBE07DC3FBD3FF30" box="[1025,1085,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1682,1704]" captionTargetBox="[187,1409,225,1623]" captionTargetId="figure@6.[151,1436,193,1661]" captionTargetPageId="6" captionText="FIGURE 4. Andinobates victimatus sp. nov. (A – K) and A. fulguritus (L – S). A – D: Lateral and dorsal views of an unsexed adult from the type locality. E – F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H – K: Juveniles of A. victimatus sp. nov. exhibiting intermediate stages of coloration development at ~ 1.5 months (H – I), ~ 2.5 months (J), and ~ 3 months (K). L – S: Unsexed A. fulguritus adults from Playa de Oro, Chocó (L – M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q – S), Photos in A – D and H – K by Roberto Márquez, E – F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L – S by Daniel Mejía- Vargas." httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="8" pageNumber="539">Fig 4</figureCitation>
A-D). The scarlet red becomes less saturated in the flanks and venter, transitioning between scarlet-red, vermilion, and black in gradually changing blotches, especially in the throat, chest, and the dorsal surface of the thighs (forming a seat patch; 
<figureCitation id="131F2A1A357ED925FA807D8BFA72FF78" box="[1377,1436,223,248]" captionStart="FIGURE 4" captionStartId="6.[151,250,1682,1704]" captionTargetBox="[187,1409,225,1623]" captionTargetId="figure@6.[151,1436,193,1661]" captionTargetPageId="6" captionText="FIGURE 4. Andinobates victimatus sp. nov. (A – K) and A. fulguritus (L – S). A – D: Lateral and dorsal views of an unsexed adult from the type locality. E – F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H – K: Juveniles of A. victimatus sp. nov. exhibiting intermediate stages of coloration development at ~ 1.5 months (H – I), ~ 2.5 months (J), and ~ 3 months (K). L – S: Unsexed A. fulguritus adults from Playa de Oro, Chocó (L – M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q – S), Photos in A – D and H – K by Roberto Márquez, E – F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L – S by Daniel Mejía- Vargas." httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="8" pageNumber="539">Fig 4</figureCitation>
E–F). The pupil and iris are both black, and nearly indistinguishable. The nares are surrounded by thin black margins, and the anterior edge of the tympanic ring is also black, with the first quarter of the tympanum degrading from black to vermilion. Fingers and toes are light metallic gray to metallic gray, and palms and soles black to dark brown.
</paragraph>
<paragraph id="8B9B369F357ED925FF267CC0FB86FD04" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="539">
Juveniles reared in the laboratory (
<figureCitation id="131F2A1A357ED925FD8A7CC0FD5AFE2C" box="[619,692,404,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1682,1704]" captionTargetBox="[187,1409,225,1623]" captionTargetId="figure@6.[151,1436,193,1661]" captionTargetPageId="6" captionText="FIGURE 4. Andinobates victimatus sp. nov. (A – K) and A. fulguritus (L – S). A – D: Lateral and dorsal views of an unsexed adult from the type locality. E – F: Ventral views of unsexed adults at the type locality. G: Palm and sole of the hand and foot of the holotype. H – K: Juveniles of A. victimatus sp. nov. exhibiting intermediate stages of coloration development at ~ 1.5 months (H – I), ~ 2.5 months (J), and ~ 3 months (K). L – S: Unsexed A. fulguritus adults from Playa de Oro, Chocó (L – M, type locality), Santa Cecilia, Risaralda (N), Nuquí, Chocó (O), Calima, Valle del Cauca (P), Buenaventura, Valle del Cauca (Q – S), Photos in A – D and H – K by Roberto Márquez, E – F by Jan Van der Meulen, G by Pablo Palacios-Rodríguez, and L – S by Daniel Mejía- Vargas." httpUri="https://zenodo.org/record/829281/files/figure.png" pageId="8" pageNumber="539">Fig. 4</figureCitation>
H–K) developed tan/orange dorsolateral stripes as metamorphs (around stage 40), and later acquired a faint incomplete dorsal median stripe that extended from the tip of the snout to the posterior-dorsal region, and a labial stripe extending from just below the snout to the insertion of the arm, both reminiscent of the color pattern of some 
<taxonomicName id="4C244D1C357ED925FD407F54FCC0FD98" box="[673,814,511,536]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357ED925FD407F54FCC0FD98" box="[673,814,511,536]" italics="true" pageId="8" pageNumber="539">A. fulguritus</emphasis>
</taxonomicName>
. As time went by, the stripes became a more saturated red, and the dark brown background gave way to homogenous scarlet red pigmentation. Adult coloration was attained around 3.5 months after leaving the water. These individuals were reared at a lower temperature (~18–22 °C) than their natural habitat (~22–28 °C), so this process is probably quicker in nature.
</paragraph>
<paragraph id="8B9B369F357ED925FF267FDBFB9AFC95" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="539">
<emphasis id="B950EA8D357ED925FF267FDBFD2FFD28" bold="true" box="[199,705,655,680]" pageId="8" pageNumber="539">Coloration in preservative (70% ethanol).</emphasis>
In preservative, the red color turns metallic gray. Fingers, toes, and tubercles become light tan/brown color, and the pupil and cornea become light gray to white. The heterogeneity in ventral coloration becomes more defined in preservative, as the black blotches become dark brown, or stay black, while the scarlet red/vermilion background becomes metallic gray.
</paragraph>
<paragraph id="8B9B369F357ED925FF267E4BFBBDFC48" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="539">
<emphasis id="B950EA8D357ED925FF267E4BFE0AFCB8" bold="true" box="[199,484,799,824]" pageId="8" pageNumber="539">
Vocalization (
<figureCitation id="131F2A1A357ED925FE887E4BFE39FCB8" box="[361,471,799,824]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="9.[151,250,1650,1672]" captionStartId-1="10.[151,250,998,1020]" captionTargetBox-0="[187,1401,447,1629]" captionTargetBox-1="[154,1436,510,955]" captionTargetId-0="figure@9.[151,1436,409,1629]" captionTargetId-1="figure@10.[151,1436,510,959]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 5. Oscillogram (below, in relative units) and spectrogram (above) of one spontaneous advertisement call of Andinobates victimatus sp. nov. The male was captured at the type locality and recorded in captivity. The distribution of sound energy is represented by the power spectrum (overlaid white line, amplitude in X axis is relative). The temperature of the substrate around the calling male was 19.4 ° C." captionText-1="FIGURE 6. Comparisons of SVL and call traits between lowland species of Andinobates. Rectangles indicate the full range of values observed or reported in the literature. When available, means are represented as diamonds, with bars denoting 95 % confidence intervals (assuming normality; estimated as 1.96 times the standard error of the mean). SVL data were obtained from Batista et al. (2014), Jungfer et al. (2000), and Silverstone (1975), and bioacoustic data from Batista et al. (2014) and Brown &amp; Twomey et al. (2011). A. vic = A. victimatus, A. alt = A. altobueyensis, A. ful = A. fulguritus, A. vir = A. viridis, A. min = A. minutus, A. cla = A. claudiae, A. gem = A. geminisae." httpUri-0="https://zenodo.org/record/829283/files/figure.png" httpUri-1="https://zenodo.org/record/829285/files/figure.png" pageId="8" pageNumber="539">Figs. 5–6</figureCitation>
).
</emphasis>
Similar to other members of the 
<taxonomicName id="4C244D1C357ED925FCB87E74FC0BFCB8" box="[857,997,799,824]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357ED925FCB87E74FC0BFCB8" box="[857,997,799,824]" italics="true" pageId="8" pageNumber="539">A. fulguritus</emphasis>
</taxonomicName>
group, 
<taxonomicName id="4C244D1C357ED925FBDD7E74FB21FCB8" box="[1084,1231,800,824]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357ED925FBDD7E74FB21FCB8" box="[1084,1231,800,824]" italics="true" pageId="8" pageNumber="539">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357ED925FB377E4BFADEFCB8" box="[1238,1328,799,824]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B950EA8D357ED925FB377E4BFADEFCB8" bold="true" box="[1238,1328,799,824]" pageId="8" pageNumber="539">sp. nov.</emphasis>
</taxonomicNameLabel>
produces calls of the “short buzz” 
<typeStatus id="549F883D357ED925FE4F7E11FE0FFCDD" box="[430,481,837,861]" pageId="8" pageNumber="539">type</typeStatus>
, consisting of a series of pulses modulated in amplitude but not frequency (
<bibRefCitation id="EFB54B6E357ED925FAD07E10FE98FC00" author="Brown" pageId="8" pageNumber="539" refString="Brown, J. L., Twomey, E., Amezquita, A., Souza, M. B. de, Caldwell, J. - L. P., Lotters, S., von May, R., Melo-Sampaio, P. R., Mejia-Vargas, D., Perez-Pena, P., Pepper, M., Poelman, E. H., Sanchez-Rodriguez, M. &amp; Summers, K. (2011) A taxonomic revision of the Neotropical poison frog genus Ranitomeya (Amphibia: Dendrobatidae). Zootaxa, 3083, 1 - 120." type="journal article" year="2011">
Brown &amp; Twomey 
<emphasis id="B950EA8D357ED925FF1E7E3DFED9FC00" box="[255,311,871,896]" italics="true" pageId="8" pageNumber="539">et al.</emphasis>
2011
</bibRefCitation>
; 
<bibRefCitation id="EFB54B6E357ED925FE637E3CFD8BFC00" author="Myers" box="[386,613,871,896]" pageId="8" pageNumber="539" refString="Myers, C. W. &amp; Daly, J. W. (1976) Preliminary evaluation of skin toxins and vocalizations in taxonomic and evolutionary studies of poison-dart frogs (Dendrobatidae). Bulletin of the American Museum of Natural History, 157, 173 - 262." type="journal article" year="1976">Myers &amp; Daly 1976</bibRefCitation>
). Calls range from 0.42– 
<geoCoordinate id="EE105058357ED925FC987E33FC54FCFF" box="[889,954,871,896]" direction="south" orientation="latitude" pageId="8" pageNumber="539" precision="555" value="-0.58">0.58 s</geoCoordinate>
in length (mean = 0.49 ± SD = 
<geoCoordinate id="EE105058357ED925FAFB7E33FAB0FC00" box="[1306,1374,871,896]" direction="south" orientation="latitude" pageId="8" pageNumber="539" precision="555" value="-0.06">0.06 s</geoCoordinate>
), and contain 99–190 pulses per call (142 ± 42.6). The peak frequency ranges between 5.65–5.86 kHz (5.75 ± 0.08 kHz), and the frequency interquartile bandwidth between 0.76–1.03 kHz (0.88 ± 0.11 kHz).
</paragraph>
<paragraph id="8B9B369F357ED925FF267E80FBDFFBFD" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="539">
<emphasis id="B950EA8D357ED925FF267E80FE6AFC6D" bold="true" box="[199,388,980,1005]" pageId="8" pageNumber="539">Natural history.</emphasis>
<taxonomicName id="4C244D1C357ED925FE6A7E80FDF3FC6C" box="[395,541,980,1004]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357ED925FE6A7E80FDF3FC6C" box="[395,541,980,1004]" italics="true" pageId="8" pageNumber="539">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357ED925FDC57E80FD93FC6D" box="[548,637,980,1005]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B950EA8D357ED925FDC57E80FD93FC6D" bold="true" box="[548,637,980,1005]" pageId="8" pageNumber="539">sp. nov.</emphasis>
</taxonomicNameLabel>
is a terrestrial species, which inhabits the understory of lowland forests in the Urabá region of Northwestern 
<collectingCountry id="F333760F357ED925FDD47EA3FD4DFB90" box="[565,675,1015,1040]" name="Colombia" pageId="8" pageNumber="539">Colombia</collectingCountry>
(
<figureCitation id="131F2A1A357ED925FD537EA3FD18FB90" box="[690,758,1015,1040]" captionStart="FIGURE 7" captionStartId="10.[151,250,1871,1893]" captionTargetBox="[162,1428,1254,1839]" captionTargetId="figure@10.[151,1436,1245,1849]" captionTargetPageId="10" captionText="FIGURE 7. Type locality of A. victimatus sp. nov. in Dabeiba, Antioquia, Colombia. A: Río Sucio at the Llorona Canyon. B: Stream at Chichiridó C: The forests of the Llorona Canyon. D: Guzmania musaica bromeliad, used by A. victimatus sp. nov. to deposit larvae. E: Understory habitat of A. victimatus sp. nov. All photos by Jan Van der Meulen." httpUri="https://zenodo.org/record/829287/files/figure.png" pageId="8" pageNumber="539">Fig. 7</figureCitation>
). Males call to defend territories and attract mates, and were typically observed doing so from the leaf litter, or perched on low branches (&lt;
<quantity id="4CDC9B7A357ED925FBFE7948FBA5FBB4" box="[1055,1099,1052,1076]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="539" unit="m" value="1.0">1 m</quantity>
above the ground) and fallen logs. A few males were also heard calling from bromeliads up to ~ 
<quantity id="4CDC9B7A357ED925FC477914FC3DFBD7" box="[934,979,1088,1112]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="539" unit="m" value="3.0">3 m</quantity>
above the ground. Call activity peaked between 7:30 and 9:30 AM, with occasional calling throughout the rest of the day.
</paragraph>
<paragraph id="8B9B369F357ED925FF2679DDFE0BFA40" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="539">
Egg clutches are presumably laid within male territories, and tadpoles are transported by males, one at a time, to bromeliad tanks. Tadpoles were observed in bromeliads up to 
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above the ground (although we did not survey plants above this height). All of the observed tadpoles were in tanks of 
<taxonomicName id="4C244D1C357ED925FC247984FB70FB68" box="[965,1182,1232,1256]" class="Liliopsida" family="Bromeliaceae" genus="Guzmania" kingdom="Plantae" order="Poales" pageId="8" pageNumber="539" phylum="Tracheophyta" rank="species" species="musaica">
<emphasis id="B950EA8D357ED925FC247984FB70FB68" box="[965,1182,1232,1256]" italics="true" pageId="8" pageNumber="539">Guzmania musaica</emphasis>
</taxonomicName>
bromeliads (
<figureCitation id="131F2A1A357ED925FAD5799BFA96FB68" box="[1332,1400,1231,1256]" captionStart="FIGURE 7" captionStartId="10.[151,250,1871,1893]" captionTargetBox="[162,1428,1254,1839]" captionTargetId="figure@10.[151,1436,1245,1849]" captionTargetPageId="10" captionText="FIGURE 7. Type locality of A. victimatus sp. nov. in Dabeiba, Antioquia, Colombia. A: Río Sucio at the Llorona Canyon. B: Stream at Chichiridó C: The forests of the Llorona Canyon. D: Guzmania musaica bromeliad, used by A. victimatus sp. nov. to deposit larvae. E: Understory habitat of A. victimatus sp. nov. All photos by Jan Van der Meulen." httpUri="https://zenodo.org/record/829287/files/figure.png" pageId="8" pageNumber="539">Fig. 7</figureCitation>
D), which leads us to speculate that 
<taxonomicName id="4C244D1C357ED925FE1D79A0FD60FA8C" box="[508,654,1268,1292]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357ED925FE1D79A0FD60FA8C" box="[508,654,1268,1292]" italics="true" pageId="8" pageNumber="539">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357ED925FD7579A0FD03FA8D" box="[660,749,1268,1293]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B950EA8D357ED925FD7579A0FD03FA8D" bold="true" box="[660,749,1268,1293]" pageId="8" pageNumber="539">sp. nov.</emphasis>
</taxonomicNameLabel>
prefers this bromeliad over other similarly-sized species (e.g. other 
<taxonomicName id="4C244D1C357ED925FF38784CFEA2FAAF" box="[217,332,1304,1327]" class="Liliopsida" family="Bromeliaceae" genus="Guzmania" kingdom="Plantae" order="Poales" pageId="8" pageNumber="539" phylum="Tracheophyta" rank="genus">
<emphasis id="B950EA8D357ED925FF38784CFEA2FAAF" box="[217,332,1304,1327]" italics="true" pageId="8" pageNumber="539">Guzmania</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C244D1C357ED925FE947843FE0BFAB0" box="[373,485,1303,1328]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="539" phylum="Tracheophyta" rank="genus">
<emphasis id="B950EA8D357ED925FE947843FE0BFAB0" box="[373,485,1303,1328]" italics="true" pageId="8" pageNumber="539">Tillandsia</emphasis>
</taxonomicName>
), for yet unknown reasons. At the 
<typeStatus id="549F883D357ED925FC97784DFC48FAB1" box="[886,934,1305,1329]" pageId="8" pageNumber="539">type</typeStatus>
locality, 
<taxonomicName id="4C244D1C357ED925FBF0784CFB4BFAB0" box="[1041,1189,1304,1328]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357ED925FBF0784CFB4BFAB0" box="[1041,1189,1304,1328]" italics="true" pageId="8" pageNumber="539">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357ED925FB4F7843FAE7FAB0" box="[1198,1289,1303,1328]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B950EA8D357ED925FB4F7843FAE7FAB0" bold="true" box="[1198,1289,1303,1328]" pageId="8" pageNumber="539">sp. nov.</emphasis>
</taxonomicNameLabel>
is sympatric with 
<taxonomicName id="4C244D1C357ED925FF2E7868FE59FAD5" box="[207,439,1340,1365]" class="Amphibia" family="Dendrobatidae" genus="Oophaga" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="histrionica">
<emphasis id="B950EA8D357ED925FF2E7868FE59FAD5" box="[207,439,1340,1365]" italics="true" pageId="8" pageNumber="539">Oophaga histrionica</emphasis>
</taxonomicName>
(
<taxonomicName id="4C244D1C357ED925FE247868FDFFFAD4" box="[453,529,1340,1364]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="order">Anura</taxonomicName>
: 
<taxonomicName id="4C244D1C357ED925FDFA7868FD2BFAD5" box="[539,709,1340,1365]" class="Amphibia" family="Dendrobatidae" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="family">Dendrobatidae</taxonomicName>
), which deposits larvae in similar phytotelmata. However, we did not find any 
<taxonomicName id="4C244D1C357ED925FEC77834FE2CFAF8" box="[294,450,1375,1400]" class="Amphibia" family="Dendrobatidae" genus="Oophaga" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="histrionica">
<emphasis id="B950EA8D357ED925FEC77834FE2CFAF8" box="[294,450,1375,1400]" italics="true" pageId="8" pageNumber="539">O. histrionica</emphasis>
</taxonomicName>
tadpoles in 
<taxonomicName id="4C244D1C357ED925FDAA7834FD29FAF8" box="[587,711,1376,1400]" class="Liliopsida" family="Bromeliaceae" genus="Guzmania" kingdom="Plantae" order="Poales" pageId="8" pageNumber="539" phylum="Tracheophyta" rank="species" species="musaica">
<emphasis id="B950EA8D357ED925FDAA7834FD29FAF8" box="[587,711,1376,1400]" italics="true" pageId="8" pageNumber="539">G. musaica</emphasis>
</taxonomicName>
tanks; only in smaller bromeliad species. Whether this indicates some instance of resource partitioning, competition, or another kind of ecological interaction between the two species remains to be studied.
</paragraph>
</subSubSection>
<subSubSection id="C33E6514357ED926FF267898F9F0FDD5" lastPageId="11" lastPageNumber="542" pageId="8" pageNumber="539" type="distribution">
<paragraph id="8B9B369F357ED925FF267898FEBAF814" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="539">
<emphasis id="B950EA8D357ED925FF267898FD66FA65" bold="true" box="[199,648,1484,1509]" pageId="8" pageNumber="539">Distribution and conservation status.</emphasis>
This species occurs in the Urabá region of Northwestern 
<collectingCountry id="F333760F357ED925FAC77898FA79FA65" box="[1318,1431,1484,1509]" name="Colombia" pageId="8" pageNumber="539">Colombia</collectingCountry>
, inhabiting tropical rain forests at ~ 
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200– 
<specimenCount id="9D22FD16357ED925FDB878BBFD47F988" box="[601,681,1519,1544]" pageId="8" pageNumber="539" type="generic">600 m</specimenCount>
</quantity>
.a.s.l. (
<figureCitation id="131F2A1A357ED925FD0C78BBFCD2F988" box="[749,828,1519,1544]" captionStart="FIGURE 1" captionStartId="2.[151,250,1421,1443]" captionTargetBox="[188,1429,219,1382]" captionTargetId="figure@2.[151,1436,195,1397]" captionTargetPageId="2" captionText="FIGURE 1. Reported (triangles), tentative (squares), and sampled (circles) localities of species in the Andinobates fulguritus group. Sampled localities are those from which individuals and / or tissue samples were obtained for this study, or sequences were available in GenBank. Reported localities were either obtained from the Global Biodiversity Information Facility (www. gbif. org; accessed on Oct. 19 th, 2016), or approximated from Silverstone (1975) and Batista et al. (2014). Tentative localities derive from personal communications (Carlos Bran-Castrillón) or museum specimen databases (see text for details)." httpUri="https://zenodo.org/record/829275/files/figure.png" pageId="8" pageNumber="539">Figs. 1</figureCitation>
, 
<figureCitation id="131F2A1A357ED925FCAA78A4FCB4F988" box="[843,858,1520,1544]" captionStart="FIGURE 7" captionStartId="10.[151,250,1871,1893]" captionTargetBox="[162,1428,1254,1839]" captionTargetId="figure@10.[151,1436,1245,1849]" captionTargetPageId="10" captionText="FIGURE 7. Type locality of A. victimatus sp. nov. in Dabeiba, Antioquia, Colombia. A: Río Sucio at the Llorona Canyon. B: Stream at Chichiridó C: The forests of the Llorona Canyon. D: Guzmania musaica bromeliad, used by A. victimatus sp. nov. to deposit larvae. E: Understory habitat of A. victimatus sp. nov. All photos by Jan Van der Meulen." httpUri="https://zenodo.org/record/829287/files/figure.png" pageId="8" pageNumber="539">7</figureCitation>
) in the foothills of the 
<collectingRegion id="49E0F87D357ED925FB8578A5FB51F988" box="[1124,1215,1521,1544]" country="American Samoa" name="Western" pageId="8" pageNumber="539">Western</collectingRegion>
Andes, west of the Paramillo Massif, on the Atrato River drainage. The 
<typeStatus id="549F883D357ED925FD087B41FCF7F9AD" box="[745,793,1557,1581]" pageId="8" pageNumber="539">type</typeStatus>
locality is in the Llorona Canyon in Dabeiba, 
<collectingRegion id="49E0F87D357ED925FAC47B40FA76F9AD" box="[1317,1432,1556,1581]" country="Colombia" name="Antioquia" pageId="8" pageNumber="539">Antioquia</collectingRegion>
, 
<collectingCountry id="F333760F357ED925FF767B63FEE6F9D0" box="[151,264,1591,1616]" name="Colombia" pageId="8" pageNumber="539">Colombia</collectingCountry>
, just southwest of the Massif. We have become aware of reports of similar frogs from other biologists and conservationists working in the Urabá region (Carlos Bran-Castrillón, pers. com.), which would extend the new species’ distribution north into the municipalities of Mutatá and Carepa, almost to the Gulf of Urabá, on the Caribbean coast (
<figureCitation id="131F2A1A357ED925FE877BF0FE43F93C" box="[358,429,1700,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1421,1443]" captionTargetBox="[188,1429,219,1382]" captionTargetId="figure@2.[151,1436,195,1397]" captionTargetPageId="2" captionText="FIGURE 1. Reported (triangles), tentative (squares), and sampled (circles) localities of species in the Andinobates fulguritus group. Sampled localities are those from which individuals and / or tissue samples were obtained for this study, or sequences were available in GenBank. Reported localities were either obtained from the Global Biodiversity Information Facility (www. gbif. org; accessed on Oct. 19 th, 2016), or approximated from Silverstone (1975) and Batista et al. (2014). Tentative localities derive from personal communications (Carlos Bran-Castrillón) or museum specimen databases (see text for details)." httpUri="https://zenodo.org/record/829275/files/figure.png" pageId="8" pageNumber="539">Fig. 1</figureCitation>
). In addition, two specimens at the Natural History Museum of the Universidad de 
<collectingRegion id="49E0F87D357ED925FF767B93FEE9F960" box="[151,263,1735,1760]" country="Colombia" name="Antioquia" pageId="8" pageNumber="539">Antioquia</collectingRegion>
(MHUA-A 3382 and 3864), currently assigned to 
<taxonomicName id="4C244D1C357ED925FC977B9CFBC1F960" box="[886,1071,1735,1760]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="B950EA8D357ED925FC977B9CFBC1F960" box="[886,1071,1735,1760]" italics="true" pageId="8" pageNumber="539">A. opisthomelas</emphasis>
</taxonomicName>
, are from the municipality of Tierralta, 
<collectingRegion id="49E0F87D357ED925FEE87BB8FE82F885" box="[265,364,1772,1797]" country="Colombia" name="Cordoba" pageId="8" pageNumber="539">Córdoba</collectingRegion>
, at ~ 
<specimenCount id="9D22FD16357ED925FE4B7BB8FDEFF885" box="[426,513,1772,1797]" pageId="8" pageNumber="539" type="generic">
<quantity id="4CDC9B7A357ED925FE4B7BB8FE13F885" box="[426,509,1772,1797]" metricMagnitude="2" metricUnit="m" metricValue="5.9" pageId="8" pageNumber="539" unit="m" value="590.0">590 m</quantity>
.
</specimenCount>
a.s.l (
<figureCitation id="131F2A1A357ED925FDDE7BB8FD6BF884" box="[575,645,1772,1797]" captionStart="FIGURE 1" captionStartId="2.[151,250,1421,1443]" captionTargetBox="[188,1429,219,1382]" captionTargetId="figure@2.[151,1436,195,1397]" captionTargetPageId="2" captionText="FIGURE 1. Reported (triangles), tentative (squares), and sampled (circles) localities of species in the Andinobates fulguritus group. Sampled localities are those from which individuals and / or tissue samples were obtained for this study, or sequences were available in GenBank. Reported localities were either obtained from the Global Biodiversity Information Facility (www. gbif. org; accessed on Oct. 19 th, 2016), or approximated from Silverstone (1975) and Batista et al. (2014). Tentative localities derive from personal communications (Carlos Bran-Castrillón) or museum specimen databases (see text for details)." httpUri="https://zenodo.org/record/829275/files/figure.png" pageId="8" pageNumber="539">Fig. 1</figureCitation>
). It is possible that these two specimens actually correspond to 
<taxonomicName id="4C244D1C357ED925FA657BB9FEE4F8A8" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357ED925FA657BB9FEE4F8A8" italics="true" pageId="8" pageNumber="539">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357ED925FEF37A5BFE83F8A8" box="[274,365,1807,1832]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B950EA8D357ED925FEF37A5BFE83F8A8" bold="true" box="[274,365,1807,1832]" pageId="8" pageNumber="539">sp. nov.</emphasis>
</taxonomicNameLabel>
, since 
<taxonomicName id="4C244D1C357ED925FE5C7A44FD9FF8A8" box="[445,625,1807,1832]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="B950EA8D357ED925FE5C7A44FD9FF8A8" box="[445,625,1807,1832]" italics="true" pageId="8" pageNumber="539">A. opisthomelas</emphasis>
</taxonomicName>
does not normally occur below 
<quantity id="4CDC9B7A357ED925FC017A44FBD5F8A8" box="[992,1083,1807,1832]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="539" unit="m" value="1100.0">1100 m</quantity>
.a.s.l (
<bibRefCitation id="EFB54B6E357ED925FB987A5BFAACF8A8" author="Silverstone" box="[1145,1346,1807,1832]" pageId="8" pageNumber="539" refString="Silverstone, P. A. (1975) A revision of the poison-arrow frogs of the genus Dendrobates Wagler. Natural History Museum of Los Angeles County, Science Bulletin, 21, 1 - 55." type="journal article" year="1975">Silverstone, 1975</bibRefCitation>
), which would mean that this species is distributed around the northern end of the Paramillo Massif, into the Sinú River drainage. As we have not examined specimens, alive or preserved, from any of these localities, we consider them tentative reports.
</paragraph>
<paragraph id="8B9B369F357ED924FF2F7AF4FE4DFEE5" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1437,151,357]" lastPageId="9" lastPageNumber="540" pageId="8" pageNumber="539">
<taxonomicName id="4C244D1C357ED925FF2F7AF4FE3AF838" box="[206,468,1952,1977]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="539" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357ED925FF2F7AF4FE3AF838" box="[206,468,1952,1977]" italics="true" pageId="8" pageNumber="539">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357ED925FE3A7AF4FDDBF839" box="[475,565,1952,1977]" pageId="8" pageNumber="539" rank="species">
<emphasis id="B950EA8D357ED925FE3A7AF4FDDBF839" bold="true" box="[475,565,1952,1977]" pageId="8" pageNumber="539">sp. nov.</emphasis>
</taxonomicNameLabel>
has, to the best of our knowledge, not been reported south or west of the 
<typeStatus id="549F883D357ED925FA8D7AF5FA72F839" box="[1388,1436,1953,1977]" pageId="8" pageNumber="539">type</typeStatus>
locality. However, given the continuous rain forests and gentle topography of the Northeastern 
<collectingRegion id="49E0F87D357ED925FB307A90FAF3F85D" box="[1233,1309,1988,2013]" country="Panama" name="Embera-Wounaan" pageId="8" pageNumber="539">Chocó</collectingRegion>
, it is likely that this species does extend into these areas. Nonetheless, Silverstone did not find it when collecting 
<taxonomicName id="4C244D1C357FD924FAEE7DCCFA72FF30" box="[1295,1436,151,176]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="540" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357FD924FAEE7DCCFA72FF30" box="[1295,1436,151,176]" italics="true" pageId="9" pageNumber="540">A. fulguritus</emphasis>
</taxonomicName>
in the Arquía river, ~ 
<quantity id="4CDC9B7A357FD924FE657DE8FE22FF55" box="[388,460,188,213]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="540" unit="km" value="85.0">85 km</quantity>
south of the 
<typeStatus id="549F883D357FD924FDBE7DE9FD61FF55" box="[607,655,189,213]" pageId="9" pageNumber="540">type</typeStatus>
locality, in the 1960s, and it has not been reported in the vicinities of Quibdó (~ 
<quantity id="4CDC9B7A357FD924FEEE7DB4FE82FF78" box="[271,364,223,248]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="540" unit="km" value="130.0">130 km</quantity>
south) or the San Juan River drainage (&gt; 
<quantity id="4CDC9B7A357FD924FC8B7DB4FC29FF78" box="[874,967,223,248]" metricMagnitude="5" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="540" unit="km" value="180.0">180 km</quantity>
south), both well sampled in terms of Dendrobatid frogs. Therefore, we suspect that this species’ range does not extend too far south of the 
<typeStatus id="549F883D357FD924FAEA7C51FAD5FE9D" box="[1291,1339,261,285]" pageId="9" pageNumber="540">type</typeStatus>
locality. Further exploration and inspection of museum specimens are needed to get a better grasp of this species’ geographic distribution.
</paragraph>
<caption id="DF5B6617357FD924FF767B26FD82F964" httpUri="https://zenodo.org/record/829283/files/figure.png" pageId="9" pageNumber="540" targetBox="[187,1401,447,1629]" targetPageId="9">
<paragraph id="8B9B369F357FD924FF767B26FD82F964" blockId="9.[151,1436,1650,1764]" pageId="9" pageNumber="540">
<emphasis id="B950EA8D357FD924FF767B26FEF7F908" bold="true" box="[151,281,1650,1672]" pageId="9" pageNumber="540">FIGURE 5.</emphasis>
Oscillogram (below, in relative units) and spectrogram (above) of one spontaneous advertisement call of 
<taxonomicName id="4C244D1C357FD924FF767BC5FE6AF927" box="[151,388,1681,1703]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="540" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357FD924FF767BC5FE6AF927" box="[151,388,1681,1703]" italics="true" pageId="9" pageNumber="540">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357FD924FE6B7BC5FE35F926" box="[394,475,1681,1702]" pageId="9" pageNumber="540" rank="species">
<emphasis id="B950EA8D357FD924FE6B7BC5FE35F926" bold="true" box="[394,475,1681,1702]" pageId="9" pageNumber="540">sp. nov.</emphasis>
</taxonomicNameLabel>
The male was captured at the type locality and recorded in captivity. The distribution of sound energy is represented by the power spectrum (overlaid white line, amplitude in X axis is relative). The temperature of the substrate around the calling male was 19.4 °C.
</paragraph>
</caption>
<paragraph id="8B9B369F357FD927FF267A46FC85FE2F" blockId="9.[151,1437,1809,2016]" lastBlockId="10.[151,1436,151,432]" lastPageId="10" lastPageNumber="541" pageId="9" pageNumber="540">
In spite of the current uncertainty regarding the distribution of 
<taxonomicName id="4C244D1C357FD924FC7C7A46FBDCF8A9" box="[925,1074,1810,1833]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="540" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357FD924FC7C7A46FBDCF8A9" box="[925,1074,1810,1833]" italics="true" pageId="9" pageNumber="540">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357FD924FBDA7A45FB79F8AA" box="[1083,1175,1809,1834]" pageId="9" pageNumber="540" rank="species">
<emphasis id="B950EA8D357FD924FBDA7A45FB79F8AA" bold="true" box="[1083,1175,1809,1834]" pageId="9" pageNumber="540">sp. nov.</emphasis>
</taxonomicNameLabel>
, it seems clear that its extent of occurrence is less than 
<quantity id="4CDC9B7A357FD924FDC07A63FD61F8D0" box="[545,655,1847,1872]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="540" unit="km" value="5000.0">5,000 km</quantity>
2. It is currently known from one locality with certainty, plus five tentative additional ones, and the forests in this species’ range have been, and will probably continue to be under strong deforestation pressures due to mining (both legal and illegal), cattle farming, and large-scale agriculture (e.g. banana and plantain plantations), which reduce the amount and quality of available habitat, and increase its fragmentation. Therefore, we tentatively designate 
<taxonomicName id="4C244D1C357FD924FD027A9CFC96F860" box="[739,888,1992,2016]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="540" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357FD924FD027A9CFC96F860" box="[739,888,1992,2016]" italics="true" pageId="9" pageNumber="540">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357FD924FC627A93FC31F860" box="[899,991,1991,2016]" pageId="9" pageNumber="540" rank="species">
<emphasis id="B950EA8D357FD924FC627A93FC31F860" bold="true" box="[899,991,1991,2016]" pageId="9" pageNumber="540">sp. nov.</emphasis>
</taxonomicNameLabel>
as Endangered (EN: B1a, biii; IUCN, 2001). However, it is possible that the range of 
<taxonomicName id="4C244D1C357CD927FD587DCFFCA3FF33" box="[697,845,155,179]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357CD927FD587DCFFCA3FF33" box="[697,845,155,179]" italics="true" pageId="10" pageNumber="541">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357CD927FCB77DCEFC5FFF33" box="[854,945,154,179]" pageId="10" pageNumber="541" rank="species">
<emphasis id="B950EA8D357CD927FCB77DCEFC5FFF33" bold="true" box="[854,945,154,179]" pageId="10" pageNumber="541">sp. nov.</emphasis>
</taxonomicNameLabel>
does exceed 
<quantity id="4CDC9B7A357CD927FBAD7DCEFB56FF33" box="[1100,1208,154,179]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="541" unit="km" value="5000.0">5,000 km</quantity>
2, in which case this species should be listed as Vulnerable (VU; B1a, biii). Most of the range of 
<taxonomicName id="4C244D1C357CD927FBE37DEBFB78FF57" box="[1026,1174,191,215]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357CD927FBE37DEBFB78FF57" box="[1026,1174,191,215]" italics="true" pageId="10" pageNumber="541">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357CD927FB7F7DEAFB14FF57" box="[1182,1274,190,215]" pageId="10" pageNumber="541" rank="species">
<emphasis id="B950EA8D357CD927FB7F7DEAFB14FF57" bold="true" box="[1182,1274,190,215]" pageId="10" pageNumber="541">sp. nov.</emphasis>
</taxonomicNameLabel>
surrounds the Nudo de Paramillo National Natural Park, a 460,000 ha nature reserve, managed by the Colombian government, which is closed to the public. However, we are unaware of any reports of 
<taxonomicName id="4C244D1C357CD927FC0C7C53FB6CFE9F" box="[1005,1154,263,287]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357CD927FC0C7C53FB6CFE9F" box="[1005,1154,263,287]" italics="true" pageId="10" pageNumber="541">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357CD927FB6A7C5CFB30FE9F" box="[1163,1246,264,287]" pageId="10" pageNumber="541" rank="species">sp. nov</taxonomicNameLabel>
within the park, which encompasses most of the Paramillo Massif above 
<specimenCount id="9D22FD16357CD927FCD47C7FFC65FEC3" box="[821,907,298,323]" pageId="10" pageNumber="541" type="generic">
<quantity id="4CDC9B7A357CD927FCD47C7FFC66FEC3" box="[821,904,298,323]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="541" unit="m" value="700.0">700 m</quantity>
.
</specimenCount>
a.s.l. This national park, nonetheless, protects ~250,000 ha in the 
<quantity id="4CDC9B7A357CD927FE657C1BFDF3FEE7" box="[388,541,334,359]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="10" pageNumber="541" unit="m" value="850.0" valueMax="1000.0" valueMin="700.0">700–1000 m</quantity>
.a.s.l elevation belt, where 
<taxonomicName id="4C244D1C357CD927FCB67C1BFC01FEE7" box="[855,1007,335,359]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="victimatus">
<emphasis id="B950EA8D357CD927FCB67C1BFC01FEE7" box="[855,1007,335,359]" italics="true" pageId="10" pageNumber="541">A. victimatus</emphasis>
</taxonomicName>
could exist at the higher end of its elevational range. Considerably more research on this species’ distribution, habitat requirements, and population ecology is needed to confidently assign it to a category of threat.
</paragraph>
<caption id="DF5B6617357CD927FF767EB2FD0AFB16" httpUri="https://zenodo.org/record/829285/files/figure.png" pageId="10" pageNumber="541" targetBox="[154,1436,510,955]" targetPageId="10">
<paragraph id="8B9B369F357CD927FF767EB2FD0AFB16" blockId="10.[151,1436,998,1174]" pageId="10" pageNumber="541">
<emphasis id="B950EA8D357CD927FF767EB2FEFCFC7C" bold="true" box="[151,274,998,1020]" pageId="10" pageNumber="541">FIGURE 6.</emphasis>
Comparisons of SVL and call traits between lowland species of 
<taxonomicName id="4C244D1C357CD927FC437EB3FBCEFC7C" box="[930,1056,999,1020]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="genus">
<emphasis id="B950EA8D357CD927FC437EB3FBCEFC7C" box="[930,1056,999,1020]" italics="true" pageId="10" pageNumber="541">Andinobates</emphasis>
</taxonomicName>
. Rectangles indicate the full range of values observed or reported in the literature. When available, means are represented as diamonds, with bars denoting 95% confidence intervals (assuming normality; estimated as 1.96 times the standard error of the mean). SVL data were obtained from Batista 
<emphasis id="B950EA8D357CD927FEFE7917FEA0FBD8" box="[287,334,1091,1112]" italics="true" pageId="10" pageNumber="541">et al</emphasis>
. (2014), Jungfer 
<emphasis id="B950EA8D357CD927FDE27917FDD6FBD8" box="[515,568,1091,1112]" italics="true" pageId="10" pageNumber="541">et al.</emphasis>
(2000), and Silverstone (1975), and bioacoustic data from Batista 
<emphasis id="B950EA8D357CD927FB0F7917FAF3FBD8" box="[1262,1309,1091,1112]" italics="true" pageId="10" pageNumber="541">et al</emphasis>
. (2014) and Brown &amp; Twomey 
<emphasis id="B950EA8D357CD927FEB87936FE6BFBF6" box="[345,389,1121,1143]" italics="true" pageId="10" pageNumber="541">et al</emphasis>
. (2011). 
<taxonomicName id="4C244D1C357CD927FE017936FDF6FBF7" box="[480,536,1122,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="vic">
<emphasis id="B950EA8D357CD927FE017936FDF6FBF7" box="[480,536,1122,1143]" italics="true" pageId="10" pageNumber="541">A. vic</emphasis>
</taxonomicName>
= 
<emphasis id="B950EA8D357CD927FDD27936FD17FBF6" box="[563,761,1121,1143]" italics="true" pageId="10" pageNumber="541">
<taxonomicName id="4C244D1C357CD927FDD27936FD54FBF7" box="[563,698,1122,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="victimatus">A. victimatus</taxonomicName>
, 
<taxonomicName id="4C244D1C357CD927FD227936FD17FBF6" box="[707,761,1121,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="alt">A. alt</taxonomicName>
</emphasis>
= 
<emphasis id="B950EA8D357CD927FCF57936FC12FBF6" box="[788,1020,1121,1143]" italics="true" pageId="10" pageNumber="541">
<taxonomicName id="4C244D1C357CD927FCF57936FC52FBF6" box="[788,956,1121,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="altobueyensis">A. altobueyensis</taxonomicName>
, 
<taxonomicName id="4C244D1C357CD927FC247936FC12FBF6" box="[965,1020,1121,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="ful">A. ful</taxonomicName>
</emphasis>
= 
<emphasis id="B950EA8D357CD927FBF47936FB39FBF7" box="[1045,1239,1121,1143]" italics="true" pageId="10" pageNumber="541">
<taxonomicName id="4C244D1C357CD927FBF47936FB78FBF6" box="[1045,1174,1121,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="fulguritus">A. fulguritus</taxonomicName>
, 
<taxonomicName id="4C244D1C357CD927FB417936FB39FBF7" box="[1184,1239,1122,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="vir">A. vir</taxonomicName>
</emphasis>
= 
<taxonomicName id="4C244D1C357CD927FB137936FAA0FBF6" box="[1266,1358,1121,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="viridis">
<emphasis id="B950EA8D357CD927FB137936FAA0FBF6" box="[1266,1358,1121,1143]" italics="true" pageId="10" pageNumber="541">A. viridis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C244D1C357CD927FABB7936FA72FBF7" box="[1370,1436,1122,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="min">
<emphasis id="B950EA8D357CD927FABB7936FA72FBF7" box="[1370,1436,1122,1143]" italics="true" pageId="10" pageNumber="541">A. min</emphasis>
</taxonomicName>
= 
<taxonomicName id="4C244D1C357CD927FF4A79D4FEF9FB16" box="[171,279,1152,1174]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B950EA8D357CD927FF4A79D4FEF9FB16" box="[171,279,1152,1174]" italics="true" pageId="10" pageNumber="541">A. minutus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C244D1C357CD927FEC579D4FEB1FB15" box="[292,351,1152,1173]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="cla">
<emphasis id="B950EA8D357CD927FEC579D4FEB1FB15" box="[292,351,1152,1173]" italics="true" pageId="10" pageNumber="541">A. cla</emphasis>
</taxonomicName>
= 
<taxonomicName id="4C244D1C357CD927FE9B79D4FE03FB15" box="[378,493,1152,1173]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="claudiae">
<emphasis id="B950EA8D357CD927FE9B79D4FE03FB15" box="[378,493,1152,1173]" italics="true" pageId="10" pageNumber="541">A. claudiae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C244D1C357CD927FE1B79D4FDAEFB16" box="[506,576,1152,1174]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="gem">
<emphasis id="B950EA8D357CD927FE1B79D4FDAEFB16" box="[506,576,1152,1174]" italics="true" pageId="10" pageNumber="541">A. gem</emphasis>
</taxonomicName>
= 
<taxonomicName id="4C244D1C357CD927FDBD79D4FD30FB16" box="[604,734,1152,1174]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="geminisae">
<emphasis id="B950EA8D357CD927FDBD79D4FD30FB16" box="[604,734,1152,1174]" italics="true" pageId="10" pageNumber="541">A. geminisae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF5B6617357CD927FF767A1BFB81F822" httpUri="https://zenodo.org/record/829287/files/figure.png" pageId="10" pageNumber="541" targetBox="[162,1428,1254,1839]" targetPageId="10">
<paragraph id="8B9B369F357CD927FF767A1BFB81F822" blockId="10.[151,1436,1871,1954]" pageId="10" pageNumber="541">
<emphasis id="B950EA8D357CD927FF767A1BFEFDF8E4" bold="true" box="[151,275,1871,1893]" pageId="10" pageNumber="541">FIGURE 7.</emphasis>
Type locality of 
<taxonomicName id="4C244D1C357CD927FE237A1BFDA9F8E5" box="[450,583,1871,1893]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357CD927FE237A1BFDA9F8E5" box="[450,583,1871,1893]" italics="true" pageId="10" pageNumber="541">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357CD927FDAC7A1BFD70F8E4" box="[589,670,1871,1892]" pageId="10" pageNumber="541" rank="species">
<emphasis id="B950EA8D357CD927FDAC7A1BFD70F8E4" bold="true" box="[589,670,1871,1892]" pageId="10" pageNumber="541">sp. nov.</emphasis>
</taxonomicNameLabel>
in Dabeiba, Antioquia, Colombia. A: Río Sucio at the Llorona Canyon. B: Stream at Chichiridó C: The forests of the Llorona Canyon. D: 
<taxonomicName id="4C244D1C357CD927FCF77A3AFC37F803" box="[790,985,1902,1923]" class="Liliopsida" family="Bromeliaceae" genus="Guzmania" kingdom="Plantae" order="Poales" pageId="10" pageNumber="541" phylum="Tracheophyta" rank="species" species="musaica">
<emphasis id="B950EA8D357CD927FCF77A3AFC37F803" box="[790,985,1902,1923]" italics="true" pageId="10" pageNumber="541">Guzmania musaica</emphasis>
</taxonomicName>
bromeliad, used by 
<taxonomicName id="4C244D1C357CD927FB467A3AFAC5F803" box="[1191,1323,1902,1923]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357CD927FB467A3AFAC5F803" box="[1191,1323,1902,1923]" italics="true" pageId="10" pageNumber="541">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357CD927FAD07A3AFA6FF803" box="[1329,1409,1902,1923]" pageId="10" pageNumber="541" rank="species">
<emphasis id="B950EA8D357CD927FAD07A3AFA6FF803" bold="true" box="[1329,1409,1902,1923]" pageId="10" pageNumber="541">sp. nov.</emphasis>
</taxonomicNameLabel>
to deposit larvae. E: Understory habitat of 
<taxonomicName id="4C244D1C357CD927FDCD7AD8FD5FF822" box="[556,689,1932,1954]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="541" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357CD927FDCD7AD8FD5FF822" box="[556,689,1932,1954]" italics="true" pageId="10" pageNumber="541">A. victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357CD927FD567AD8FCE9F821" box="[695,775,1932,1953]" pageId="10" pageNumber="541" rank="species">
<emphasis id="B950EA8D357CD927FD567AD8FCE9F821" bold="true" box="[695,775,1932,1953]" pageId="10" pageNumber="541">sp. nov.</emphasis>
</taxonomicNameLabel>
All photos by Jan Van der Meulen.
</paragraph>
</caption>
<caption id="DF5B6617357DD926FF9C7F41FA8FFDAA" box="[125,1377,533,554]" pageId="11" pageNumber="542">
<paragraph id="8B9B369F357DD926FF9C7F41FA8FFDAA" blockId="11.[125,1788,533,644]" box="[125,1377,533,554]" pageId="11" pageNumber="542">
<emphasis id="B950EA8D357DD926FF9C7F41FF08FDAA" bold="true" box="[125,230,533,554]" pageId="11" pageNumber="542">TABLE 2.</emphasis>
Specimen information for individuals used in molecular and phylogenetic analyses. A dash indicates no data available.
</paragraph>
</caption>
<paragraph id="8B9B369F357DD926FF9C7F14F9F0FDD5" blockId="11.[125,1788,533,644]" box="[125,1566,576,597]" pageId="11" pageNumber="542">
<emphasis id="B950EA8D357DD926FF9C7F14F9F0FDD5" bold="true" box="[125,1566,576,597]" pageId="11" pageNumber="542">Voucher Species Locality Study GenBank Accession</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33E6514357DD921FAB27F3BFEB4F98B" lastPageId="12" lastPageNumber="543" pageId="11" pageNumber="542" type="description">
<paragraph id="8B9B369F357DD926FAB27F3BF887F909" blockId="11.[125,1788,533,644]" lastBlockId="11.[1595,1897,1651,1673]" pageId="11" pageNumber="542">
<emphasis id="B950EA8D357DD926FAB27F3BF912FD04" bold="true" box="[1363,1788,623,644]" pageId="11" pageNumber="542">16S COI Cytb</emphasis>
<emphasis id="B950EA8D357DD926F9DA7B27F887F909" box="[1595,1897,1651,1673]" italics="true" pageId="11" pageNumber="542">……continued on the next page</emphasis>
</paragraph>
<paragraph id="8B9B369F357DD926FF9C7FC9F8DAF9DB" pageId="11" pageNumber="542">
<table id="F924C43F357D26D2FF9C7FC8F887F9DB" box="[125,1897,668,1627]" gridcols="7" gridrows="18" pageId="11" pageNumber="542">
<tr id="351434DD357D26D2FF9C7FC8F887FD69" box="[125,1897,668,745]" gridrow="0" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7FC8FEC6FD69" box="[125,296,668,745]" gridcol="0" gridrow="0" pageId="11" pageNumber="542">JDL 24489 GECOH 1321</th>
<th id="76C55DA1357D26D2FEAB7FC8FD90FD69" box="[330,638,668,745]" gridcol="1" gridrow="0" pageId="11" pageNumber="542">
<emphasis id="B950EA8D357DD926FEAB7FC9FDC8FD69" italics="true" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7FC9FDBAFD32" box="[330,596,668,690]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="ventrimaculata">Ranitomeya ventrimaculata</taxonomicName>
<taxonomicName id="4C244D1C357DD926FEAB7F87FDC8FD69" box="[330,550,723,745]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">Andinobates fulguritus</taxonomicName>
</emphasis>
</th>
<th id="76C55DA1357D26D2FD237FC8FBB5FD69" box="[706,1115,668,745]" gridcol="2" gridrow="0" pageId="11" pageNumber="542">
Km 11, Vía Leticia 
<emphasis id="B950EA8D357DD926FC607FC9FC63FD32" box="[897,909,669,690]" italics="true" pageId="11" pageNumber="542">–</emphasis>
Tarapacá. Leticia, Amazonas, Colombia Playa de Oro, Tadó, Chocó, Colombia
</th>
<th id="76C55DA1357D26D2FB9A7FC8FAC3FD69" box="[1147,1325,668,745]" gridcol="3" gridrow="0" pageId="11" pageNumber="542">Grant et al. 2006 This study</th>
<th id="76C55DA1357D26D2FAB27FC8FA50FD69" box="[1363,1470,668,745]" gridcol="4" gridrow="0" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27FC9FA53FD32" box="[1363,1469,669,690]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ502266" pageId="11" pageNumber="542">DQ502266</accessionNumber>
<accessionNumber id="9477AB7C357DD926FAB27F80FA53FD69" box="[1363,1469,724,745]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885030" pageId="11" pageNumber="542">KY885030</accessionNumber>
</th>
<th id="76C55DA1357D26D2F9ED7FC8F993FD69" box="[1548,1661,668,745]" gridcol="5" gridrow="0" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7FC9F999FD32" box="[1548,1655,669,690]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ502928" pageId="11" pageNumber="542">DQ502928</accessionNumber>
<accessionNumber id="9477AB7C357DD926F9ED7F80F999FD69" box="[1548,1655,724,745]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885053" pageId="11" pageNumber="542">KY885053</accessionNumber>
</th>
<th id="76C55DA1357D26D2F9287FC8F887FD69" box="[1737,1897,668,745]" gridcol="6" gridrow="0" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287FC9F8DAFD32" box="[1737,1844,669,690]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ502697" pageId="11" pageNumber="542">DQ502697</accessionNumber>
<accessionNumber id="9477AB7C357DD926F9287F80F8DDFD69" box="[1737,1843,724,745]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885076" pageId="11" pageNumber="542">KY885076</accessionNumber>
</th>
</tr>
<tr id="351434DD357D26D2FF9C7E55F887FC97" box="[125,1897,769,791]" gridrow="1" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7E55FEC6FC97" box="[125,296,769,791]" gridcol="0" gridrow="1" pageId="11" pageNumber="542">GECOH 1612</th>
<td id="76C55DA1357D26D2FEAB7E55FD90FC97" box="[330,638,769,791]" gridcol="1" gridrow="1" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7E55FDC8FC97" box="[330,550,769,791]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB7E55FDC8FC97" box="[330,550,769,791]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD237E55FBB5FC97" box="[706,1115,769,791]" gridcol="2" gridrow="1" pageId="11" pageNumber="542">Playa de Oro, Tadó, Chocó, Colombia</td>
<td id="76C55DA1357D26D2FB9A7E55FAC3FC97" box="[1147,1325,769,791]" gridcol="3" gridrow="1" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB27E55FA50FC97" box="[1363,1470,769,791]" gridcol="4" gridrow="1" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27E56FA50FC97" box="[1363,1470,770,791]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885031" pageId="11" pageNumber="542">KY885031</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED7E55F993FC97" box="[1548,1661,769,791]" gridcol="5" gridrow="1" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7E56F999FC97" box="[1548,1655,770,791]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885055" pageId="11" pageNumber="542">KY885055</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9287E55F887FC97" box="[1737,1897,769,791]" gridcol="6" gridrow="1" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287E56F8DAFC97" box="[1737,1844,770,791]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885078" pageId="11" pageNumber="542">KY885078</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C7E7AF887FCC4" box="[125,1897,814,836]" gridrow="2" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7E7AFEC6FCC4" box="[125,296,814,836]" gridcol="0" gridrow="2" pageId="11" pageNumber="542">GECOH 1613</th>
<td id="76C55DA1357D26D2FEAB7E7AFD90FCC4" box="[330,638,814,836]" gridcol="1" gridrow="2" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7E7AFDC8FCC4" box="[330,550,814,836]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB7E7AFDC8FCC4" box="[330,550,814,836]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD237E7AFBB5FCC4" box="[706,1115,814,836]" gridcol="2" gridrow="2" pageId="11" pageNumber="542">Playa de Oro, Tadó, Chocó, Colombia</td>
<td id="76C55DA1357D26D2FB9A7E7AFAC3FCC4" box="[1147,1325,814,836]" gridcol="3" gridrow="2" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB27E7AFA50FCC4" box="[1363,1470,814,836]" gridcol="4" gridrow="2" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27E7BFA53FCC4" box="[1363,1469,815,836]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885032" pageId="11" pageNumber="542">KY885032</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED7E7AF993FCC4" box="[1548,1661,814,836]" gridcol="5" gridrow="2" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7E7BF998FCC4" box="[1548,1654,815,836]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885056" pageId="11" pageNumber="542">KY885056</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9287E7AF887FCC4" box="[1737,1897,814,836]" gridcol="6" gridrow="2" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287E7BF8DAFCC4" box="[1737,1844,815,836]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885079" pageId="11" pageNumber="542">KY885079</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C7E0FF887FC29" box="[125,1897,859,937]" gridrow="3" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7E0FFEC6FC29" box="[125,296,859,937]" gridcol="0" gridrow="3" pageId="11" pageNumber="542">MHNUC 340 GECOH 1320</th>
<td id="76C55DA1357D26D2FEAB7E0FFD90FC29" box="[330,638,859,937]" gridcol="1" gridrow="3" pageId="11" pageNumber="542">
<emphasis id="B950EA8D357DD926FEAB7E0FFDC8FC29" italics="true" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7E0FFDC8FCF1" box="[330,550,859,881]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">Andinobates fulguritus</taxonomicName>
<taxonomicName id="4C244D1C357DD926FEAB7EC7FDC8FC29" box="[330,550,915,937]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">Andinobates fulguritus</taxonomicName>
</emphasis>
</td>
<td id="76C55DA1357D26D2FD237E0FFBB5FC29" box="[706,1115,859,937]" gridcol="2" gridrow="3" pageId="11" pageNumber="542">Sierra Mecana (260 m). Bahía Solano, Chocó, Colombia Playa de Oro, Tadó, Chocó, Colombia</td>
<td id="76C55DA1357D26D2FB9A7E0FFAC3FC29" box="[1147,1325,859,937]" gridcol="3" gridrow="3" pageId="11" pageNumber="542">Grant et al. 2006 This study</td>
<td id="76C55DA1357D26D2FAB27E0FFA50FC29" box="[1363,1470,859,937]" gridcol="4" gridrow="3" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27E08FA53FCF1" box="[1363,1469,860,881]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ502106" pageId="11" pageNumber="542">DQ502106</accessionNumber>
-
</td>
<td id="76C55DA1357D26D2F9ED7E0FF993FC29" box="[1548,1661,859,937]" gridcol="5" gridrow="3" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7E08F999FCF1" box="[1548,1655,860,881]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ502817" pageId="11" pageNumber="542">DQ502817</accessionNumber>
<accessionNumber id="9477AB7C357DD926F9ED7EC0F998FC29" box="[1548,1654,916,937]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885052" pageId="11" pageNumber="542">KY885052</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9287E0FF887FC29" box="[1737,1897,859,937]" gridcol="6" gridrow="3" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287E08F8DAFCF1" box="[1737,1844,860,881]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ502538" pageId="11" pageNumber="542">DQ502538</accessionNumber>
<accessionNumber id="9477AB7C357DD926F9287EC0F8DAFC29" box="[1737,1844,916,937]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885075" pageId="11" pageNumber="542">KY885075</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C7E94F887FC56" box="[125,1897,960,982]" gridrow="4" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7E94FEC6FC56" box="[125,296,960,982]" gridcol="0" gridrow="4" pageId="11" pageNumber="542">GECOH 860</th>
<td id="76C55DA1357D26D2FEAB7E94FD90FC56" box="[330,638,960,982]" gridcol="1" gridrow="4" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7E94FDC8FC56" box="[330,550,960,982]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB7E94FDC8FC56" box="[330,550,960,982]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD237E94FBB5FC56" box="[706,1115,960,982]" gridcol="2" gridrow="4" pageId="11" pageNumber="542">Arusí, Nuquí, Chocó, Colombia</td>
<td id="76C55DA1357D26D2FB9A7E94FAC3FC56" box="[1147,1325,960,982]" gridcol="3" gridrow="4" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB27E94FA50FC56" box="[1363,1470,960,982]" gridcol="4" gridrow="4" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27E95FA53FC56" box="[1363,1469,961,982]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885048" pageId="11" pageNumber="542">KY885048</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED7E94F993FC56" box="[1548,1661,960,982]" gridcol="5" gridrow="4" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7E95F998FC56" box="[1548,1654,961,982]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885072" pageId="11" pageNumber="542">KY885072</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9287E94F887FC56" box="[1737,1897,960,982]" gridcol="6" gridrow="4" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287E95F8DAFC56" box="[1737,1844,961,982]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885095" pageId="11" pageNumber="542">KY885095</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C7EB9F887FB83" box="[125,1897,1005,1027]" gridrow="5" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7EB9FEC6FB83" box="[125,296,1005,1027]" gridcol="0" gridrow="5" pageId="11" pageNumber="542">GECOH 861</th>
<td id="76C55DA1357D26D2FEAB7EB9FD90FB83" box="[330,638,1005,1027]" gridcol="1" gridrow="5" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7EB9FDC8FB83" box="[330,550,1005,1027]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB7EB9FDC8FB83" box="[330,550,1005,1027]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD237EB9FBB5FB83" box="[706,1115,1005,1027]" gridcol="2" gridrow="5" pageId="11" pageNumber="542">Arusí, Nuquí, Chocó, Colombia</td>
<td id="76C55DA1357D26D2FB9A7EB9FAC3FB83" box="[1147,1325,1005,1027]" gridcol="3" gridrow="5" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB27EB9FA50FB83" box="[1363,1470,1005,1027]" gridcol="4" gridrow="5" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27EBAFA53FB83" box="[1363,1469,1006,1027]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885049" pageId="11" pageNumber="542">KY885049</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED7EB9F993FB83" box="[1548,1661,1005,1027]" gridcol="5" gridrow="5" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7EBAF999FB83" box="[1548,1655,1006,1027]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885073" pageId="11" pageNumber="542">KY885073</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9287EB9F887FB83" box="[1737,1897,1005,1027]" gridcol="6" gridrow="5" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287EBAF8DDFB83" box="[1737,1843,1006,1027]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885096" pageId="11" pageNumber="542">KY885096</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C794FF887FBB1" box="[125,1897,1051,1073]" gridrow="6" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C794FFEC6FBB1" box="[125,296,1051,1073]" gridcol="0" gridrow="6" pageId="11" pageNumber="542">GECOH 862</th>
<td id="76C55DA1357D26D2FEAB794FFD90FBB1" box="[330,638,1051,1073]" gridcol="1" gridrow="6" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB794FFDC8FBB1" box="[330,550,1051,1073]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB794FFDC8FBB1" box="[330,550,1051,1073]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD23794FFBB5FBB1" box="[706,1115,1051,1073]" gridcol="2" gridrow="6" pageId="11" pageNumber="542">Arusí, Nuquí, Chocó, Colombia</td>
<td id="76C55DA1357D26D2FB9A794FFAC3FBB1" box="[1147,1325,1051,1073]" gridcol="3" gridrow="6" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB2794FFA50FBB1" box="[1363,1470,1051,1073]" gridcol="4" gridrow="6" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27948FA53FBB1" box="[1363,1469,1052,1073]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885050" pageId="11" pageNumber="542">KY885050</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED794FF993FBB1" box="[1548,1661,1051,1073]" gridcol="5" gridrow="6" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7948F998FBB1" box="[1548,1654,1052,1073]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885074" pageId="11" pageNumber="542">KY885074</accessionNumber>
</td>
<td id="76C55DA1357D26D2F928794FF887FBB1" box="[1737,1897,1051,1073]" gridcol="6" gridrow="6" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287948F8DAFBB1" box="[1737,1844,1052,1073]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885097" pageId="11" pageNumber="542">KY885097</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C791CF887FBDE" box="[125,1897,1096,1118]" gridrow="7" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C791CFEC6FBDE" box="[125,296,1096,1118]" gridcol="0" gridrow="7" pageId="11" pageNumber="542">GECOH 1402</th>
<td id="76C55DA1357D26D2FEAB791CFD90FBDE" box="[330,638,1096,1118]" gridcol="1" gridrow="7" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB791CFDC8FBDE" box="[330,550,1096,1118]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB791CFDC8FBDE" box="[330,550,1096,1118]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD23791CFBB5FBDE" box="[706,1115,1096,1118]" gridcol="2" gridrow="7" pageId="11" pageNumber="542">Arusí, Nuquí, Chocó, Colombia</td>
<td id="76C55DA1357D26D2FB9A791CFAC3FBDE" box="[1147,1325,1096,1118]" gridcol="3" gridrow="7" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB2791CFA50FBDE" box="[1363,1470,1096,1118]" gridcol="4" gridrow="7" pageId="11" pageNumber="542">-</td>
<td id="76C55DA1357D26D2F9ED791CF993FBDE" box="[1548,1661,1096,1118]" gridcol="5" gridrow="7" pageId="11" pageNumber="542">-</td>
<td id="76C55DA1357D26D2F928791CF887FBDE" box="[1737,1897,1096,1118]" gridcol="6" gridrow="7" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F928791DF8DAFBDE" box="[1737,1844,1097,1118]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885077" pageId="11" pageNumber="542">KY885077</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C7921F887FB0B" box="[125,1897,1141,1163]" gridrow="8" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7921FEC6FB0B" box="[125,296,1141,1163]" gridcol="0" gridrow="8" pageId="11" pageNumber="542">GECOH 546</th>
<td id="76C55DA1357D26D2FEAB7921FD90FB0B" box="[330,638,1141,1163]" gridcol="1" gridrow="8" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7921FDC8FB0B" box="[330,550,1141,1163]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB7921FDC8FB0B" box="[330,550,1141,1163]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD237921FBB5FB0B" box="[706,1115,1141,1163]" gridcol="2" gridrow="8" pageId="11" pageNumber="542">Itaurí, Pueblo Rico, Risaralda, Colombia</td>
<td id="76C55DA1357D26D2FB9A7921FAC3FB0B" box="[1147,1325,1141,1163]" gridcol="3" gridrow="8" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB27921FA50FB0B" box="[1363,1470,1141,1163]" gridcol="4" gridrow="8" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27922FA53FB0B" box="[1363,1469,1142,1163]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885047" pageId="11" pageNumber="542">KY885047</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED7921F993FB0B" box="[1548,1661,1141,1163]" gridcol="5" gridrow="8" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7922F999FB0B" box="[1548,1655,1142,1163]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885071" pageId="11" pageNumber="542">KY885071</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9287921F887FB0B" box="[1737,1897,1141,1163]" gridcol="6" gridrow="8" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287922F8DDFB0B" box="[1737,1843,1142,1163]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885094" pageId="11" pageNumber="542">KY885094</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C79F7F887FB39" box="[125,1897,1187,1209]" gridrow="9" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C79F7FEC6FB39" box="[125,296,1187,1209]" gridcol="0" gridrow="9" pageId="11" pageNumber="542">GECOH 2093</th>
<td id="76C55DA1357D26D2FEAB79F7FD90FB39" box="[330,638,1187,1209]" gridcol="1" gridrow="9" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB79F7FDC8FB39" box="[330,550,1187,1209]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB79F7FDC8FB39" box="[330,550,1187,1209]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD2379F7FBB5FB39" box="[706,1115,1187,1209]" gridcol="2" gridrow="9" pageId="11" pageNumber="542">Itaurí, Pueblo Rico, Risaralda, Colombia</td>
<td id="76C55DA1357D26D2FB9A79F7FAC3FB39" box="[1147,1325,1187,1209]" gridcol="3" gridrow="9" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB279F7FA50FB39" box="[1363,1470,1187,1209]" gridcol="4" gridrow="9" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB279F0FA53FB39" box="[1363,1469,1188,1209]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885042" pageId="11" pageNumber="542">KY885042</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED79F7F993FB39" box="[1548,1661,1187,1209]" gridcol="5" gridrow="9" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED79F0F998FB39" box="[1548,1654,1188,1209]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885064" pageId="11" pageNumber="542">KY885064</accessionNumber>
</td>
<td id="76C55DA1357D26D2F92879F7F887FB39" box="[1737,1897,1187,1209]" gridcol="6" gridrow="9" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F92879F0F8DAFB39" box="[1737,1844,1188,1209]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885089" pageId="11" pageNumber="542">KY885089</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C7984F887FB66" box="[125,1897,1232,1254]" gridrow="10" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7984FEC6FB66" box="[125,296,1232,1254]" gridcol="0" gridrow="10" pageId="11" pageNumber="542">GCOH 2094</th>
<td id="76C55DA1357D26D2FEAB7984FD90FB66" box="[330,638,1232,1254]" gridcol="1" gridrow="10" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7984FDC8FB66" box="[330,550,1232,1254]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB7984FDC8FB66" box="[330,550,1232,1254]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD237984FBB5FB66" box="[706,1115,1232,1254]" gridcol="2" gridrow="10" pageId="11" pageNumber="542">Itaurí, Pueblo Rico, Risaralda, Colombia</td>
<td id="76C55DA1357D26D2FB9A7984FAC3FB66" box="[1147,1325,1232,1254]" gridcol="3" gridrow="10" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB27984FA50FB66" box="[1363,1470,1232,1254]" gridcol="4" gridrow="10" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27985FA50FB66" box="[1363,1470,1233,1254]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885043" pageId="11" pageNumber="542">KY885043</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED7984F993FB66" box="[1548,1661,1232,1254]" gridcol="5" gridrow="10" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9F37985F993FB66" box="[1554,1661,1233,1254]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885065" pageId="11" pageNumber="542">KY885065</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9287984F887FB66" box="[1737,1897,1232,1254]" gridcol="6" gridrow="10" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287985F8DDFB66" box="[1737,1843,1233,1254]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885090" pageId="11" pageNumber="542">KY885090</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C79A9F887FA93" box="[125,1897,1277,1299]" gridrow="11" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C79A9FEC6FA93" box="[125,296,1277,1299]" gridcol="0" gridrow="11" pageId="11" pageNumber="542">GECOH 545</th>
<td id="76C55DA1357D26D2FEAB79A9FD90FA93" box="[330,638,1277,1299]" gridcol="1" gridrow="11" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB79A9FDC8FA93" box="[330,550,1277,1299]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB79A9FDC8FA93" box="[330,550,1277,1299]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD2379A9FBB5FA93" box="[706,1115,1277,1299]" gridcol="2" gridrow="11" pageId="11" pageNumber="542">Itaurí, Pueblo Rico, Risaralda, Colombia</td>
<td id="76C55DA1357D26D2FB9A79A9FAC3FA93" box="[1147,1325,1277,1299]" gridcol="3" gridrow="11" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB279A9FA50FA93" box="[1363,1470,1277,1299]" gridcol="4" gridrow="11" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB279AAFA53FA93" box="[1363,1469,1278,1299]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885046" pageId="11" pageNumber="542">KY885046</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED79A9F993FA93" box="[1548,1661,1277,1299]" gridcol="5" gridrow="11" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED79AAF998FA93" box="[1548,1654,1278,1299]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885070" pageId="11" pageNumber="542">KY885070</accessionNumber>
</td>
<td id="76C55DA1357D26D2F92879A9F887FA93" box="[1737,1897,1277,1299]" gridcol="6" gridrow="11" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F92879AAF8DAFA93" box="[1737,1844,1278,1299]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885093" pageId="11" pageNumber="542">KY885093</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C787FF887FAC1" box="[125,1897,1323,1345]" gridrow="12" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C787FFEC6FAC1" box="[125,296,1323,1345]" gridcol="0" gridrow="12" pageId="11" pageNumber="542">-</th>
<td id="76C55DA1357D26D2FEAB787FFD90FAC1" box="[330,638,1323,1345]" gridcol="1" gridrow="12" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB787FFDC8FAC1" box="[330,550,1323,1345]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="fulguritus">
<emphasis id="B950EA8D357DD926FEAB787FFDC8FAC1" box="[330,550,1323,1345]" italics="true" pageId="11" pageNumber="542">Andinobates fulguritus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357D26D2FD23787FFBB5FAC1" box="[706,1115,1323,1345]" gridcol="2" gridrow="12" pageId="11" pageNumber="542">San Blas, Panamá</td>
<td id="76C55DA1357D26D2FB9A787FFAC3FAC1" box="[1147,1325,1323,1345]" gridcol="3" gridrow="12" pageId="11" pageNumber="542">Vences et al. 2000</td>
<td id="76C55DA1357D26D2FAB2787FFA50FAC1" box="[1363,1470,1323,1345]" gridcol="4" gridrow="12" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27878FA57FAC1" box="[1363,1465,1324,1345]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF124116" pageId="11" pageNumber="542">AF124116</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED787FF993FAC1" box="[1548,1661,1323,1345]" gridcol="5" gridrow="12" pageId="11" pageNumber="542">-</td>
<td id="76C55DA1357D26D2F928787FF887FAC1" box="[1737,1897,1323,1345]" gridcol="6" gridrow="12" pageId="11" pageNumber="542">-</td>
</tr>
<tr id="351434DD357D26D2FF9C780CF887FAEE" box="[125,1897,1368,1390]" gridrow="13" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C780CFEC6FAEE" box="[125,296,1368,1390]" gridcol="0" gridrow="13" pageId="11" pageNumber="542">ANDES-A 3683</th>
<td id="76C55DA1357D26D2FEAB780CFD90FAEE" box="[330,638,1368,1390]" gridcol="1" gridrow="13" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB780CFDC2FAEE" box="[330,556,1368,1390]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357DD926FEAB780CFDC2FAEE" box="[330,556,1368,1390]" italics="true" pageId="11" pageNumber="542">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357DD926FDD0780DFD90FAEE" box="[561,638,1369,1390]" pageId="11" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<td id="76C55DA1357D26D2FD23780CFBB5FAEE" box="[706,1115,1368,1390]" gridcol="2" gridrow="13" pageId="11" pageNumber="542">Chichiridó, Dabeiba, Antioquia, Colombia</td>
<td id="76C55DA1357D26D2FB9A780CFAC3FAEE" box="[1147,1325,1368,1390]" gridcol="3" gridrow="13" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB2780CFA50FAEE" box="[1363,1470,1368,1390]" gridcol="4" gridrow="13" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB2780DFA50FAEE" box="[1363,1470,1369,1390]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885033" pageId="11" pageNumber="542">KY885033</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED780CF993FAEE" box="[1548,1661,1368,1390]" gridcol="5" gridrow="13" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED780DF998FAEE" box="[1548,1654,1369,1390]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885057" pageId="11" pageNumber="542">KY885057</accessionNumber>
</td>
<td id="76C55DA1357D26D2F928780CF887FAEE" box="[1737,1897,1368,1390]" gridcol="6" gridrow="13" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F928780DF8DDFAEE" box="[1737,1843,1369,1390]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885080" pageId="11" pageNumber="542">KY885080</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C78D1F887FA1B" box="[125,1897,1413,1435]" gridrow="14" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C78D1FEC6FA1B" box="[125,296,1413,1435]" gridcol="0" gridrow="14" pageId="11" pageNumber="542">ANDES-A 3684*</th>
<td id="76C55DA1357D26D2FEAB78D1FD90FA1B" box="[330,638,1413,1435]" gridcol="1" gridrow="14" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB78D1FDC2FA1B" box="[330,556,1413,1435]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357DD926FEAB78D1FDC2FA1B" box="[330,556,1413,1435]" italics="true" pageId="11" pageNumber="542">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357DD926FDD078D2FD90FA1B" box="[561,638,1414,1435]" pageId="11" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<td id="76C55DA1357D26D2FD2378D1FBB5FA1B" box="[706,1115,1413,1435]" gridcol="2" gridrow="14" pageId="11" pageNumber="542">Chichiridó, Dabeiba, Antioquia, Colombia</td>
<td id="76C55DA1357D26D2FB9A78D1FAC3FA1B" box="[1147,1325,1413,1435]" gridcol="3" gridrow="14" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB278D1FA50FA1B" box="[1363,1470,1413,1435]" gridcol="4" gridrow="14" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB278D2FA53FA1B" box="[1363,1469,1414,1435]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885034" pageId="11" pageNumber="542">KY885034</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED78D1F993FA1B" box="[1548,1661,1413,1435]" gridcol="5" gridrow="14" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED78D2F998FA1B" box="[1548,1654,1414,1435]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885058" pageId="11" pageNumber="542">KY885058</accessionNumber>
</td>
<td id="76C55DA1357D26D2F92878D1F887FA1B" box="[1737,1897,1413,1435]" gridcol="6" gridrow="14" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F92878D2F8DAFA1B" box="[1737,1844,1414,1435]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885081" pageId="11" pageNumber="542">KY885081</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C78E7F887FA49" box="[125,1897,1459,1481]" gridrow="15" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C78E7FEC6FA49" box="[125,296,1459,1481]" gridcol="0" gridrow="15" pageId="11" pageNumber="542">ANDES-A 3685*</th>
<td id="76C55DA1357D26D2FEAB78E7FD90FA49" box="[330,638,1459,1481]" gridcol="1" gridrow="15" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB78E7FDC2FA49" box="[330,556,1459,1481]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357DD926FEAB78E7FDC2FA49" box="[330,556,1459,1481]" italics="true" pageId="11" pageNumber="542">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357DD926FDD078E0FD90FA49" box="[561,638,1460,1481]" pageId="11" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<td id="76C55DA1357D26D2FD2378E7FBB5FA49" box="[706,1115,1459,1481]" gridcol="2" gridrow="15" pageId="11" pageNumber="542">Chichiridó, Dabeiba, Antioquia, Colombia</td>
<td id="76C55DA1357D26D2FB9A78E7FAC3FA49" box="[1147,1325,1459,1481]" gridcol="3" gridrow="15" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB278E7FA50FA49" box="[1363,1470,1459,1481]" gridcol="4" gridrow="15" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB278E0FA53FA49" box="[1363,1469,1460,1481]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885035" pageId="11" pageNumber="542">KY885035</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED78E7F993FA49" box="[1548,1661,1459,1481]" gridcol="5" gridrow="15" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED78E0F998FA49" box="[1548,1654,1460,1481]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885059" pageId="11" pageNumber="542">KY885059</accessionNumber>
</td>
<td id="76C55DA1357D26D2F92878E7F887FA49" box="[1737,1897,1459,1481]" gridcol="6" gridrow="15" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F92878E0F8DDFA49" box="[1737,1843,1460,1481]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885082" pageId="11" pageNumber="542">KY885082</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C78B4F887F9AD" box="[125,1897,1504,1581]" gridrow="16" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C78B4FEC6F9AD" box="[125,296,1504,1581]" gridcol="0" gridrow="16" pageId="11" pageNumber="542">ANDES-A 3686** ANDES-A 3687*</th>
<td id="76C55DA1357D26D2FEAB78B4FD90F9AD" box="[330,638,1504,1581]" gridcol="1" gridrow="16" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB78B4FDC2FA76" box="[330,556,1504,1526]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357DD926FEAB78B4FDC2FA76" box="[330,556,1504,1526]" italics="true" pageId="11" pageNumber="542">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357DD926FDD078B5FD90FA76" box="[561,638,1505,1526]" pageId="11" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="4C244D1C357DD926FEAB7B43FDC2F9AD" box="[330,556,1559,1581]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357DD926FEAB7B43FDC2F9AD" box="[330,556,1559,1581]" italics="true" pageId="11" pageNumber="542">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357DD926FDD07B4CFD90F9AD" box="[561,638,1560,1581]" pageId="11" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<td id="76C55DA1357D26D2FD2378B4FBB5F9AD" box="[706,1115,1504,1581]" gridcol="2" gridrow="16" pageId="11" pageNumber="542">Chichiridó, Dabeiba, Antioquia, Colombia Chichiridó, Dabeiba, Antioquia, Colombia</td>
<td id="76C55DA1357D26D2FB9A78B4FAC3F9AD" box="[1147,1325,1504,1581]" gridcol="3" gridrow="16" pageId="11" pageNumber="542">This study This study</td>
<td id="76C55DA1357D26D2FAB278B4FA50F9AD" box="[1363,1470,1504,1581]" gridcol="4" gridrow="16" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB278B5FA53FA76" box="[1363,1469,1505,1526]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885036" pageId="11" pageNumber="542">KY885036</accessionNumber>
<accessionNumber id="9477AB7C357DD926FAB27B4CFA53F9AD" box="[1363,1469,1560,1581]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885037" pageId="11" pageNumber="542">KY885037</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED78B4F993F9AD" box="[1548,1661,1504,1581]" gridcol="5" gridrow="16" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED78B5F998FA76" box="[1548,1654,1505,1526]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885060" pageId="11" pageNumber="542">KY885060</accessionNumber>
<accessionNumber id="9477AB7C357DD926F9ED7B4CF999F9AD" box="[1548,1655,1560,1581]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885061" pageId="11" pageNumber="542">KY885061</accessionNumber>
</td>
<td id="76C55DA1357D26D2F92878B4F887F9AD" box="[1737,1897,1504,1581]" gridcol="6" gridrow="16" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F92878B5F8DAFA76" box="[1737,1844,1505,1526]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885083" pageId="11" pageNumber="542">KY885083</accessionNumber>
<accessionNumber id="9477AB7C357DD926F9287B4CF8DDF9AD" box="[1737,1843,1560,1581]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885084" pageId="11" pageNumber="542">KY885084</accessionNumber>
</td>
</tr>
<tr id="351434DD357D26D2FF9C7B11F887F9DB" box="[125,1897,1605,1627]" gridrow="17" pageId="11" pageNumber="542">
<th id="76C55DA1357D26D2FF9C7B11FEC6F9DB" box="[125,296,1605,1627]" gridcol="0" gridrow="17" pageId="11" pageNumber="542">ANDES-A 3688*</th>
<td id="76C55DA1357D26D2FEAB7B11FD90F9DB" box="[330,638,1605,1627]" gridcol="1" gridrow="17" pageId="11" pageNumber="542">
<taxonomicName id="4C244D1C357DD926FEAB7B11FDC2F9DB" box="[330,556,1605,1627]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="victimatus" status="sp. nov.">
<emphasis id="B950EA8D357DD926FEAB7B11FDC2F9DB" box="[330,556,1605,1627]" italics="true" pageId="11" pageNumber="542">Andinobates victimatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357F6357DD926FDD07B12FD90F9DB" box="[561,638,1606,1627]" pageId="11" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</td>
<td id="76C55DA1357D26D2FD237B11FBB5F9DB" box="[706,1115,1605,1627]" gridcol="2" gridrow="17" pageId="11" pageNumber="542">Chichiridó, Dabeiba, Antioquia, Colombia</td>
<td id="76C55DA1357D26D2FB9A7B11FAC3F9DB" box="[1147,1325,1605,1627]" gridcol="3" gridrow="17" pageId="11" pageNumber="542">This study</td>
<td id="76C55DA1357D26D2FAB27B11FA50F9DB" box="[1363,1470,1605,1627]" gridcol="4" gridrow="17" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926FAB27B12FA53F9DB" box="[1363,1469,1606,1627]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885038" pageId="11" pageNumber="542">KY885038</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9ED7B11F993F9DB" box="[1548,1661,1605,1627]" gridcol="5" gridrow="17" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9ED7B12F998F9DB" box="[1548,1654,1606,1627]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885062" pageId="11" pageNumber="542">KY885062</accessionNumber>
</td>
<td id="76C55DA1357D26D2F9287B11F887F9DB" box="[1737,1897,1605,1627]" gridcol="6" gridrow="17" pageId="11" pageNumber="542">
<accessionNumber id="9477AB7C357DD926F9287B12F8DAF9DB" box="[1737,1844,1606,1627]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885085" pageId="11" pageNumber="542">KY885085</accessionNumber>
</td>
</tr>
</table>
</paragraph>
<caption id="DF5B6617357AD921FF967FE0FEBAFD49" box="[119,340,692,713]" pageId="12" pageNumber="543">
<paragraph id="8B9B369F357AD921FF967FE0FEBAFD49" blockId="12.[119,1738,692,805]" box="[119,340,692,713]" pageId="12" pageNumber="543">
<emphasis id="B950EA8D357AD921FF967FE0FF30FD49" bold="true" box="[119,222,692,713]" pageId="12" pageNumber="543">TABLE 2.</emphasis>
(CƟntinueđ)
</paragraph>
</caption>
<paragraph id="8B9B369F357AD921FF967FB7FA1CFD78" blockId="12.[119,1738,692,805]" box="[119,1522,739,760]" pageId="12" pageNumber="543">
<emphasis id="B950EA8D357AD921FF967FB7FA1CFD78" bold="true" box="[119,1522,739,760]" pageId="12" pageNumber="543">Voucher Species Locality Study GenBank Accession</emphasis>
</paragraph>
<paragraph id="8B9B369F357AD921FACC7E44F917FB8C" blockId="12.[119,1738,692,805]" lastBlockId="12.[119,1793,1014,1493]" pageId="12" pageNumber="543">
<emphasis id="B950EA8D357AD921FACC7E44F924FCA5" bold="true" box="[1325,1738,784,805]" pageId="12" pageNumber="543">16S COI Cytb</emphasis>
GECOH 1216 
<taxonomicName id="4C244D1C357AD921FEC17EA2FDF8FB8B" box="[288,534,1014,1035]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="B950EA8D357AD921FEC17EA2FDF8FB8B" box="[288,534,1014,1035]" italics="true" pageId="12" pageNumber="543">Andinobates opisthomelas</emphasis>
</taxonomicName>
I la đel SƟl, Guatapé, AntiƟquia, Amézquita et al. 
<accessionNumber id="9477AB7C357AD921FACC7EA3FA63FB8C" box="[1325,1421,1015,1036]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ936634" pageId="12" pageNumber="543">JQ936634</accessionNumber>
<accessionNumber id="9477AB7C357AD921FA017EA3F9A7FB8C" box="[1504,1609,1015,1036]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885051" pageId="12" pageNumber="543">KY885051</accessionNumber>
<accessionNumber id="9477AB7C357AD921F9797EA3F917FB8C" box="[1688,1785,1015,1036]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ936620" pageId="12" pageNumber="543">JQ936620</accessionNumber>
</paragraph>
<paragraph id="8B9B369F357AD921FF967E69F8EEFC60" pageId="12" pageNumber="543">
<table id="F924C43F357A26D2FF967E68F8EFFC60" box="[119,1793,828,992]" gridcols="9" gridrows="3" pageId="12" pageNumber="543">
<tr id="351434DD357A26D2FF967E68F8EFFC07" box="[119,1793,828,903]" gridrow="0" pageId="12" pageNumber="543">
<th id="76C55DA1357A26D2FF967E68FF12FC07" box="[119,252,828,903]" gridcol="0" gridrow="0" pageId="12" pageNumber="543">ANDES-A 3689* GECOH 1751</th>
<th id="76C55DA1357A26D2FEC17E68FE14FC07" box="[288,506,828,903]" gridcol="1" gridrow="0" pageId="12" pageNumber="543">
<emphasis id="B950EA8D357AD921FEC17E68FE14FC07" italics="true" pageId="12" pageNumber="543">
<taxonomicName id="4C244D1C357AD921FEC17E68FE14FCD1" box="[288,506,828,849]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="victimatus">Andinobates victimatus</taxonomicName>
<taxonomicName id="4C244D1C357AD921FEC17E26FE14FC07" box="[288,506,882,903]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="victimatus">Andinobates victimatus</taxonomicName>
</emphasis>
</th>
<th id="76C55DA1357A26D2FDE87E68FDA5FC07" box="[521,587,828,903]" gridcol="2" gridrow="0" pageId="12" pageNumber="543">p. nƟv. p. nƟv.</th>
<th id="76C55DA1357A26D2FD4C7E68FBD5FC07" box="[685,1083,828,903]" gridcol="3" gridrow="0" pageId="12" pageNumber="543">Chichiriđó, Dabeiba, AntiƟquia, CƟlƟmbia Chichiriđó, Dabeiba, AntiƟquia, CƟlƟmbia</th>
<th id="76C55DA1357A26D2FBBB7E68FB95FC07" box="[1114,1147,828,903]" gridcol="4" gridrow="0" pageId="12" pageNumber="543">Thi Thi</th>
<th id="76C55DA1357A26D2FB737E68FB52FC07" box="[1170,1212,828,903]" gridcol="5" gridrow="0" pageId="12" pageNumber="543">tuđy tuđy</th>
<th id="76C55DA1357A26D2FACD7E68FA74FC07" box="[1324,1434,828,903]" gridcol="6" gridrow="0" pageId="12" pageNumber="543">
<accessionNumber id="9477AB7C357AD921FACD7E69FA7AFCD2" box="[1324,1428,829,850]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885044" pageId="12" pageNumber="543">KY885044</accessionNumber>
<accessionNumber id="9477AB7C357AD921FACD7E26FA7AFC07" box="[1324,1428,882,903]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885039" pageId="12" pageNumber="543">KY885039</accessionNumber>
</th>
<th id="76C55DA1357A26D2FA017E68F9A7FC07" box="[1504,1609,828,903]" gridcol="7" gridrow="0" pageId="12" pageNumber="543">
<accessionNumber id="9477AB7C357AD921FA017E69F9A7FCD2" box="[1504,1609,829,850]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885066" pageId="12" pageNumber="543">KY885066</accessionNumber>
-
</th>
<th id="76C55DA1357A26D2F9797E68F8EFFC07" box="[1688,1793,828,903]" gridcol="8" gridrow="0" pageId="12" pageNumber="543">
<accessionNumber id="9477AB7C357AD921F9797E69F8EEFCD2" box="[1688,1792,829,850]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885091" pageId="12" pageNumber="543">KY885091</accessionNumber>
<accessionNumber id="9477AB7C357AD921F9797E26F8EFFC07" box="[1688,1793,882,903]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885086" pageId="12" pageNumber="543">KY885086</accessionNumber>
</th>
</tr>
<tr id="351434DD357A26D2FF967ECAF8EFFC33" box="[119,1793,926,947]" gridrow="1" pageId="12" pageNumber="543">
<th id="76C55DA1357A26D2FF967ECAFF12FC33" box="[119,252,926,947]" gridcol="0" gridrow="1" pageId="12" pageNumber="543">GECOH 1756</th>
<td id="76C55DA1357A26D2FEC17ECAFE14FC33" box="[288,506,926,947]" gridcol="1" gridrow="1" pageId="12" pageNumber="543">
<taxonomicName id="4C244D1C357AD921FEC17ECAFE14FC33" box="[288,506,926,947]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="victimatus">
<emphasis id="B950EA8D357AD921FEC17ECAFE14FC33" box="[288,506,926,947]" italics="true" pageId="12" pageNumber="543">Andinobates victimatus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357A26D2FDE87ECAFDA5FC33" box="[521,587,926,947]" gridcol="2" gridrow="1" pageId="12" pageNumber="543">p. nƟv.</td>
<td id="76C55DA1357A26D2FD4C7ECAFBD5FC33" box="[685,1083,926,947]" gridcol="3" gridrow="1" pageId="12" pageNumber="543">Chichiriđó, Dabeiba, AntiƟquia, CƟlƟmbia</td>
<td id="76C55DA1357A26D2FBBB7ECAFB95FC33" box="[1114,1147,926,947]" gridcol="4" gridrow="1" pageId="12" pageNumber="543">Thi</td>
<td id="76C55DA1357A26D2FB737ECAFB52FC33" box="[1170,1212,926,947]" gridcol="5" gridrow="1" pageId="12" pageNumber="543">tuđy</td>
<td id="76C55DA1357A26D2FACD7ECAFA74FC33" box="[1324,1434,926,947]" gridcol="6" gridrow="1" pageId="12" pageNumber="543">
<accessionNumber id="9477AB7C357AD921FACD7ECAFA7AFC33" box="[1324,1428,926,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885040" pageId="12" pageNumber="543">KY885040</accessionNumber>
</td>
<td id="76C55DA1357A26D2FA017ECAF9A7FC33" box="[1504,1609,926,947]" gridcol="7" gridrow="1" pageId="12" pageNumber="543">
<accessionNumber id="9477AB7C357AD921FA017ECAF9A7FC33" box="[1504,1609,926,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885063" pageId="12" pageNumber="543">KY885063</accessionNumber>
</td>
<td id="76C55DA1357A26D2F9797ECAF8EFFC33" box="[1688,1793,926,947]" gridcol="8" gridrow="1" pageId="12" pageNumber="543">
<accessionNumber id="9477AB7C357AD921F9797ECAF8EEFC33" box="[1688,1792,926,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885087" pageId="12" pageNumber="543">KY885087</accessionNumber>
</td>
</tr>
<tr id="351434DD357A26D2FF967E9FF8EFFC60" box="[119,1793,971,992]" gridrow="2" pageId="12" pageNumber="543">
<th id="76C55DA1357A26D2FF967E9FFF12FC60" box="[119,252,971,992]" gridcol="0" gridrow="2" pageId="12" pageNumber="543">GECOH 1757</th>
<td id="76C55DA1357A26D2FEC17E9FFE14FC60" box="[288,506,971,992]" gridcol="1" gridrow="2" pageId="12" pageNumber="543">
<taxonomicName id="4C244D1C357AD921FEC17E9FFE14FC60" box="[288,506,971,992]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="victimatus">
<emphasis id="B950EA8D357AD921FEC17E9FFE14FC60" box="[288,506,971,992]" italics="true" pageId="12" pageNumber="543">Andinobates victimatus</emphasis>
</taxonomicName>
</td>
<td id="76C55DA1357A26D2FDE87E9FFDA5FC60" box="[521,587,971,992]" gridcol="2" gridrow="2" pageId="12" pageNumber="543">p. nƟv.</td>
<td id="76C55DA1357A26D2FD4C7E9FFBD5FC60" box="[685,1083,971,992]" gridcol="3" gridrow="2" pageId="12" pageNumber="543">Chichiriđó, Dabeiba, AntiƟquia, CƟlƟmbia</td>
<td id="76C55DA1357A26D2FBBB7E9FFB95FC60" box="[1114,1147,971,992]" gridcol="4" gridrow="2" pageId="12" pageNumber="543">Thi</td>
<td id="76C55DA1357A26D2FB737E9FFB52FC60" box="[1170,1212,971,992]" gridcol="5" gridrow="2" pageId="12" pageNumber="543">tuđy</td>
<td id="76C55DA1357A26D2FACD7E9FFA74FC60" box="[1324,1434,971,992]" gridcol="6" gridrow="2" pageId="12" pageNumber="543">
<accessionNumber id="9477AB7C357AD921FAD07E9FFA74FC60" box="[1329,1434,971,992]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885041" pageId="12" pageNumber="543">KY885041</accessionNumber>
</td>
<td id="76C55DA1357A26D2FA017E9FF9A7FC60" box="[1504,1609,971,992]" gridcol="7" gridrow="2" pageId="12" pageNumber="543">-</td>
<td id="76C55DA1357A26D2F9797E9FF8EFFC60" box="[1688,1793,971,992]" gridcol="8" gridrow="2" pageId="12" pageNumber="543">
<accessionNumber id="9477AB7C357AD921F9797E9FF8EEFC60" box="[1688,1792,971,992]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885088" pageId="12" pageNumber="543">KY885088</accessionNumber>
</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9B369F357AD921FD4C7946FB67FBA7" blockId="12.[119,1793,1014,1493]" box="[685,1161,1042,1063]" pageId="12" pageNumber="543">CƟlƟmbia 2013</paragraph>
<paragraph id="8B9B369F357AD921FF967978F8EFFBC1" blockId="12.[119,1793,1014,1493]" box="[119,1793,1068,1090]" pageId="12" pageNumber="543">
GECOH 297 
<taxonomicName id="4C244D1C357AD921FEC17978FD1CFBC2" authority="Bosque" authorityName="Bosque" box="[288,754,1068,1090]" class="Amphibia" family="Dendrobatidae" genus="Andnobates" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="bombetes">
<emphasis id="B950EA8D357AD921FEC17978FE02FBC1" box="[288,492,1068,1089]" italics="true" pageId="12" pageNumber="543">Andnobates bombetes</emphasis>
Bosque
</taxonomicName>
YƟtƟcƟ Natural Re erve, YƟtƟcƟ, Thi tuđy 
<accessionNumber id="9477AB7C357AD921FACD7978FA7AFBC1" box="[1324,1428,1068,1089]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885045" pageId="12" pageNumber="543">KY885045</accessionNumber>
<accessionNumber id="9477AB7C357AD921FA017978F9A6FBC1" box="[1504,1608,1068,1089]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885067" pageId="12" pageNumber="543">KY885067</accessionNumber>
<accessionNumber id="9477AB7C357AD921F9797978F8EFFBC1" box="[1688,1793,1068,1089]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885092" pageId="12" pageNumber="543">KY885092</accessionNumber>
</paragraph>
<paragraph id="8B9B369F357AD921FD4C7913FC47FBDC" blockId="12.[119,1793,1014,1493]" box="[685,937,1095,1116]" pageId="12" pageNumber="543">
<collectingRegion id="49E0F87D357AD921FD4C7913FCAAFBDC" box="[685,836,1095,1116]" country="Colombia" name="Valle del Cauca" pageId="12" pageNumber="543">Valle đel Cauca</collectingRegion>
, CƟlƟmbia
</paragraph>
<paragraph id="8B9B369F357AD921FF967936F917FBF7" blockId="12.[119,1793,1014,1493]" box="[119,1785,1122,1143]" pageId="12" pageNumber="543">
<materialsCitation id="3B4C3CC2357AD921FF967936F917FBF7" ID-GBIF-Occurrence="1572326090" box="[119,1785,1122,1143]" collectionCode="GECOH" location="Santander" pageId="12" pageNumber="543" specimenCode="GECOH 305" specimenCount="1" stateProvince="Santander">
<specimenCode id="DB829EE4357AD921FF967936FF1EFBF7" box="[119,240,1122,1143]" collectionCode="GECOH" pageId="12" pageNumber="543">
<collectionCode id="ED35AE5A357AD921FF967936FF29FBF7" box="[119,199,1122,1143]" pageId="12" pageNumber="543">GECOH</collectionCode>
305
</specimenCode>
<taxonomicName id="4C244D1C357AD921FEC17936FDE9FBF7" box="[288,519,1122,1143]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="B950EA8D357AD921FEC17936FDE9FBF7" box="[288,519,1122,1143]" italics="true" pageId="12" pageNumber="543">Andinobates virolinensis</emphasis>
</taxonomicName>
CƟ tilla đe fara, VirƟlín, 
<collectingRegion id="49E0F87D357AD921FC777936FC18FBF7" box="[918,1014,1122,1143]" country="Colombia" name="Santander" pageId="12" pageNumber="543">Santanđer</collectingRegion>
, ƂrƟwn et al. 2011; 
<accessionNumber id="9477AB7C357AD921FACC7936FA63FBF7" box="[1325,1421,1122,1143]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN635873" pageId="12" pageNumber="543">JN635873</accessionNumber>
<accessionNumber id="9477AB7C357AD921FA017936F9A6FBF7" box="[1504,1608,1122,1143]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885068" pageId="12" pageNumber="543">KY885068</accessionNumber>
<accessionNumber id="9477AB7C357AD921F9797936F917FBF7" box="[1688,1785,1122,1143]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ936626" pageId="12" pageNumber="543">JQ936626</accessionNumber>
</materialsCitation>
</paragraph>
<paragraph id="8B9B369F357AD921FD4C7929FB1AFB12" blockId="12.[119,1793,1014,1493]" box="[685,1268,1149,1170]" pageId="12" pageNumber="543">
CƟlƟmbia 
<bibRefCitation id="EFB54B6E357AD921FBBB7929FB1AFB12" author="Amezquita" box="[1114,1268,1149,1170]" pageId="12" pageNumber="543" refString="Amezquita, A., Marquez, R., Medina, R., Mejia-Vargas, D., Kahn, T. R., Suarez, G. &amp; Mazariegos, L. (2013) A new species of Andean poison frog, Andinobates (Anura: Dendrobatidae), from the northwestern Andes of Colombia. Zootaxa, 3620 (1), 163 - 178." type="journal article" year="2013">Amézquita et al.</bibRefCitation>
</paragraph>
<paragraph id="8B9B369F357AD921FBBB79CCFB67FB2D" blockId="12.[119,1793,1014,1493]" box="[1114,1161,1176,1197]" pageId="12" pageNumber="543">2013</paragraph>
<paragraph id="8B9B369F357AD921FF9679E7F917FB48" blockId="12.[119,1793,1014,1493]" box="[119,1785,1203,1224]" pageId="12" pageNumber="543">
GECOH 474 
<taxonomicName id="4C244D1C357AD921FEC179E7FD0DFB48" authority="Falan" authorityName="Falan" box="[288,739,1203,1224]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="tolimensis">
<emphasis id="B950EA8D357AD921FEC179E7FE17FB48" box="[288,505,1203,1224]" italics="true" pageId="12" pageNumber="543">Andinobates tolimensis</emphasis>
Falan
</taxonomicName>
, TƟlima, CƟlƟmbia ƂrƟwn et al. 2011; 
<accessionNumber id="9477AB7C357AD921FACC79E7FA63FB48" box="[1325,1421,1203,1224]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN635857" pageId="12" pageNumber="543">JN635857</accessionNumber>
<accessionNumber id="9477AB7C357AD921FA0179E7F9A6FB48" box="[1504,1608,1203,1224]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885069" pageId="12" pageNumber="543">KY885069</accessionNumber>
<accessionNumber id="9477AB7C357AD921F97979E7F917FB48" box="[1688,1785,1203,1224]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ936632" pageId="12" pageNumber="543">JQ936632</accessionNumber>
</paragraph>
<bibRefCitation id="EFB54B6E357AD921FBBB799AFB67FB7E" author="Amezquita" pageId="12" pageNumber="543" refString="Amezquita, A., Marquez, R., Medina, R., Mejia-Vargas, D., Kahn, T. R., Suarez, G. &amp; Mazariegos, L. (2013) A new species of Andean poison frog, Andinobates (Anura: Dendrobatidae), from the northwestern Andes of Colombia. Zootaxa, 3620 (1), 163 - 178." type="journal article" year="2013">
<paragraph id="8B9B369F357AD921FBBB799AFB1AFB63" blockId="12.[119,1793,1014,1493]" box="[1114,1268,1230,1251]" pageId="12" pageNumber="543">Amézquita et al.</paragraph>
<paragraph id="8B9B369F357AD921FBBB79BDFB67FB7E" blockId="12.[119,1793,1014,1493]" box="[1114,1161,1257,1278]" pageId="12" pageNumber="543">2013</paragraph>
</bibRefCitation>
<paragraph id="8B9B369F357AD921FF967850F917FA99" blockId="12.[119,1793,1014,1493]" box="[119,1785,1283,1305]" pageId="12" pageNumber="543">
GECOH 1523 
<taxonomicName id="4C244D1C357AD921FEC17857FD04FA99" authority="La Me" authorityName="La Me" box="[288,746,1283,1305]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="cassidyhornae">
<emphasis id="B950EA8D357AD921FEC17857FDCFFA98" box="[288,545,1283,1304]" italics="true" pageId="12" pageNumber="543">Andinobates cassidyhornae</emphasis>
La Me
</taxonomicName>
enia, Jarđín, AntiƟquia, CƟlƟmbia Amézquita et al. 
<accessionNumber id="9477AB7C357AD921FACC7850FA63FA99" box="[1325,1421,1284,1305]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ936636" pageId="12" pageNumber="543">JQ936636</accessionNumber>
<accessionNumber id="9477AB7C357AD921FA017850F9A7FA99" box="[1504,1609,1284,1305]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY885054" pageId="12" pageNumber="543">KY885054</accessionNumber>
<accessionNumber id="9477AB7C357AD921F9797850F917FA99" box="[1688,1785,1284,1305]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ936622" pageId="12" pageNumber="543">JQ936622</accessionNumber>
</paragraph>
<paragraph id="8B9B369F357AD921FBBB784AFB67FAB3" blockId="12.[119,1793,1014,1493]" box="[1114,1161,1310,1331]" pageId="12" pageNumber="543">2013</paragraph>
<paragraph id="8B9B369F357AD921FF96786DFBDCFAE9" blockId="12.[119,1793,1014,1493]" pageId="12" pageNumber="543">
MVUP 2428 
<taxonomicName id="4C244D1C357AD921FEC1786DFE17FACE" box="[288,505,1337,1358]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="geminisae">
<emphasis id="B950EA8D357AD921FEC1786DFE17FACE" box="[288,505,1337,1358]" italics="true" pageId="12" pageNumber="543">Andinobates geminisae</emphasis>
</taxonomicName>
6 Km ea t Ɵf El Carmen (RíƟ Ƃelén), Ƃati ta et al. 2014 
<accessionNumber id="9477AB7C357AD921FACD786DFA76FACE" box="[1324,1432,1337,1358]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM212167" pageId="12" pageNumber="543">KM212167</accessionNumber>
<accessionNumber id="9477AB7C357AD921FA01786DF9A2FACE" box="[1504,1612,1337,1358]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM212166" pageId="12" pageNumber="543">KM212166</accessionNumber>
- Heađwater Ɵf RíƟ CanƟ, DƟnƟ Ɵ, CƟlón,
</paragraph>
<paragraph id="8B9B369F357AD921FD4C783BFD18FA04" blockId="12.[119,1793,1014,1493]" box="[685,758,1391,1412]" pageId="12" pageNumber="543">Panamá</paragraph>
<paragraph id="8B9B369F357AD921FF9678DEF8EEFA1F" blockId="12.[119,1793,1014,1493]" box="[119,1792,1417,1439]" pageId="12" pageNumber="543">
USNM-FS 
<taxonomicName id="4C244D1C357AD921FEC178DDFE05FA1E" box="[288,491,1417,1438]" class="Amphibia" family="Dendrobatidae" genus="Andinobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="claudiae">
<emphasis id="B950EA8D357AD921FEC178DDFE05FA1E" box="[288,491,1417,1438]" italics="true" pageId="12" pageNumber="543">Andinobates claudiae</emphasis>
</taxonomicName>
SƟuth enđ Ɵf I la PƟpa, 
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59980 SunwƟƟđ Channel, ƂƟca đel TƟrƟ, 
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