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<document id="41A0C18529A4FD2FF36E458292B29D05" ID-DOI="10.11646/zootaxa.3697.1.1" ID-GBIF-Dataset="b2711d98-f97a-4771-bb6b-ff472a2f5a80" ID-ISSN="1175-5326" ID-Zenodo-Dep="284383" ID-ZooBank="492264BB-E919-447D-9D67-C226DE21A0CE" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460660988741" checkinUser="plazi" docAuthor="Reemer, Menno" docDate="2013" docId="03B4BF12FFE8FF82FF60FA9747D89311" docLanguage="en" docName="zt03697p088.pdf" docOrigin="Zootaxa 3697 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ceratophya scolopus Shannon 1927" docType="treatment" docVersion="7" lastPageNumber="23" masterDocId="FF8DC76AFFFDFF94FFF7FFAB43079029" masterDocTitle="Taxonomic exploration of Neotropical Microdontinae (Diptera: Syrphidae) mimicking stingless bees" masterLastPageNumber="88" masterPageNumber="1" pageNumber="22" updateTime="1698347572540" updateUser="plazi">
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<mods:title id="BE7B9C5CB67C621703B88FBCEAF90136">Taxonomic exploration of Neotropical Microdontinae (Diptera: Syrphidae) mimicking stingless bees</mods:title>
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<mods:namePart id="82D6C8E6F64C8CD5CE93E0B3C3BB9F27">Reemer, Menno</mods:namePart>
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<mods:date id="2FB14AD54415FE213732171570CDDC1B">2013</mods:date>
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<mods:number id="2E19DD57A4CCAA4A6705CDD9622D4ECA">3697</mods:number>
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<mods:identifier id="4A63678B1400205E797E3F91C3BAD685" type="DOI">10.11646/zootaxa.3697.1.1</mods:identifier>
<mods:identifier id="7CBF00FA59CAEBCD2CE2F2FD905C1C7E" type="GBIF-Dataset">b2711d98-f97a-4771-bb6b-ff472a2f5a80</mods:identifier>
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<treatment id="03B4BF12FFE8FF82FF60FA9747D89311" ID-DOI="http://doi.org/10.5281/zenodo.5624873" ID-GBIF-Taxon="127665195" ID-Zenodo-Dep="5624873" LSID="urn:lsid:plazi:treatment:03B4BF12FFE8FF82FF60FA9747D89311" httpUri="http://treatment.plazi.org/id/03B4BF12FFE8FF82FF60FA9747D89311" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="C3075D8FFFE8FF81FF60FA97403495F2" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BA20E04FFE8FF81FF60FA974162957F" blockId="21.[151,613,1340,1399]" box="[151,613,1340,1366]" pageId="21" pageNumber="22">
<heading id="D0EAB968FFE8FF81FF60FA974162957F" bold="true" box="[151,613,1340,1366]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="4C1D7587FFE8FF81FF60FA974162957F" ID-CoL="8BD53" authority="Shannon, 1927" authorityName="Shannon" authorityYear="1927" box="[151,613,1340,1366]" class="Insecta" family="Syrphidae" genus="Ceratophya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scolopus">
<emphasis id="B969D216FFE8FF81FF60FA974162957F" bold="true" box="[151,613,1340,1366]" pageId="21" pageNumber="22">
<emphasis id="B969D216FFE8FF81FF60FA974295957F" bold="true" box="[151,402,1340,1366]" italics="true" pageId="21" pageNumber="22">Ceratophya scolopus</emphasis>
(Shannon, 1927)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA20E04FFE8FF81FF60FAF54242955E" blockId="21.[151,613,1340,1399]" box="[151,325,1374,1399]" pageId="21" pageNumber="22">
<figureCitation id="13261281FFE8FF81FF60FAF54210955E" box="[151,279,1374,1399]" captionStart="FIGURES 42 – 54. 42 – 47" captionStartId="20.[151,264,1901,1923]" captionTargetBox="[180,1409,259,1853]" captionTargetId="figure@20.[151,1436,231,1880]" captionTargetPageId="20" captionText="FIGURES 42 – 54. 42 – 47. Ceratophya panamensis. 42. male holotype (above) &amp; female paratype (photo: American Museum of Natural History); 43. female, head frontal; 44. female, head lateral; 45. male, wing; 46. male, scutellum; 47. male, hind tarsus. 48 – 54. Ceratophya scolopus male (holotype). 48. habitus dorsal; 49. habitus lateral; 50. abdomen posterodorsal; 51. head frontal; 52. wing; 53. scutellum; 54. apex of hind tibia with basitarsus, lateral." httpUri="https://zenodo.org/record/284388/files/figure.png" pageId="21" pageNumber="22">Figs 48–54</figureCitation>
,
<figureCitation id="13261281FFE8FF81FED5FAF54246955E" box="[290,321,1374,1399]" captionStart="FIGURES 55 – 62. 55 – 57" captionStartId="23.[151,264,1880,1902]" captionTargetBox="[182,1407,217,1828]" captionTargetId="figure@23.[151,1436,193,1858]" captionTargetPageId="23" captionText="FIGURES 55 – 62. 55 – 57. Ceratophya, male genitalia. 55. C. notata (holotype); 56. C. panamensis (holotype); 57. C. scolopus (holotype). 58 – 62. Hypselosyrphus amazonicus male (Peru, coll. RMNH). 58. habitus dorsal; 59. habitus lateral; 60. head frontal; 61. head lateral; 62. wing." httpUri="https://zenodo.org/record/284389/files/figure.png" pageId="21" pageNumber="22">57</figureCitation>
.
</paragraph>
<paragraph id="8BA20E04FFE8FF81FF60FA0D403495F2" blockId="21.[151,864,1446,1499]" pageId="21" pageNumber="22">
<taxonomicName id="4C1D7587FFE8FF81FF60FA0D41279595" authority="Shannon, 1927: 20" authorityName="Shannon" authorityPageNumber="20" authorityYear="1927" box="[151,544,1446,1468]" class="Insecta" family="Syrphidae" genus="Microdon" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scolopus">
<emphasis id="B969D216FFE8FF81FF60FA0D425E9595" box="[151,345,1446,1468]" italics="true" pageId="21" pageNumber="22">Microdon scolopus</emphasis>
Shannon, 1927: 20
</taxonomicName>
.
<typeStatus id="54A6B0A6FFE8FF81FDDDFA0C415B9595" box="[554,604,1447,1468]" pageId="21" pageNumber="22">Type</typeStatus>
locality: Amazon region.
<taxonomicName id="4C1D7587FFE8FF81FF60FA6E412295F2" authority="Shannon" authorityName="Shannon" box="[151,549,1477,1499]" class="Insecta" family="Syrphidae" genus="Microdon" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scolopus" subGenus="Ubristes">
<emphasis id="B969D216FFE8FF81FF60FA6E43FD95F2" box="[151,250,1477,1499]" italics="true" pageId="21" pageNumber="22">Microdon</emphasis>
(
<emphasis id="B969D216FFE8FF81FEFFFA6E425995F2" box="[264,350,1477,1499]" italics="true" pageId="21" pageNumber="22">Ubristes</emphasis>
)
<emphasis id="B969D216FFE8FF81FE9AFA6E42C195F2" box="[365,454,1477,1499]" italics="true" pageId="21" pageNumber="22">scolopus</emphasis>
Shannon
</taxonomicName>
in Thompson
<emphasis id="B969D216FFE8FF81FD40FA6D41E395F2" box="[695,740,1477,1499]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. (1976)
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFE8FF81FF60F9AC4127964E" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BA20E04FFE8FF81FF60F9AC401E966D" blockId="21.[151,1437,1543,2037]" pageId="21" pageNumber="22">
<emphasis id="B969D216FFE8FF81FF60F9AC42B59609" bold="true" box="[151,434,1543,1568]" pageId="21" pageNumber="22">
Studied
<typeStatus id="54A6B0A6FFE8FF81FF0FF9AC422C9609" box="[248,299,1543,1568]" pageId="21" pageNumber="22">type</typeStatus>
specimens.
</emphasis>
<typeStatus id="54A6B0A6FFE8FF81FE4FF9A341489609" box="[440,591,1544,1568]" pageId="21" pageNumber="22" type="holotype">HOLOTYPE</typeStatus>
.
<collectingCountry id="F30A4E94FFE8FF81FDAEF9AC41919609" box="[601,662,1543,1568]" name="Malta" pageId="21" pageNumber="22">Male</collectingCountry>
. Label 1 (small, round, red-bordered): &quot;
<typeStatus id="54A6B0A6FFE8FF81FBBAF9AC47B99609" box="[1101,1214,1543,1568]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
&quot;; label 2: &quot;Amazon. 66 53&quot;; label 3: &quot;
<taxonomicName id="4C1D7587FFE8FF81FEA2F987416D966D" authority="Snn." authorityName="Snn." box="[341,618,1580,1605]" class="Insecta" family="Syrphidae" genus="Microdon" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="scolopus">
<emphasis id="B969D216FFE8FF81FEA2F987412B966C" box="[341,556,1580,1605]" italics="true" pageId="21" pageNumber="22">Microdon scolopus</emphasis>
Snn.
</taxonomicName>
&quot;. Coll. BMNH.
</paragraph>
<paragraph id="8BA20E04FFE8FF81FF30F9E44127964E" blockId="21.[151,1437,1543,2037]" box="[199,544,1615,1640]" pageId="21" pageNumber="22">
<emphasis id="B969D216FFE8FF81FF30F9E4424B9641" bold="true" box="[199,332,1615,1640]" pageId="21" pageNumber="22">Adult male</emphasis>
. Body size:
<quantity id="4CE5A3E1FFE8FF81FE2FF9FB411B964E" box="[472,540,1616,1639]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="22" unit="mm" value="7.0">7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFE8FF82FF30F9DF42A591DC" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="8BA20E04FFE8FF81FF30F9DF428F9769" blockId="21.[151,1437,1543,2037]" pageId="21" pageNumber="22">
<emphasis id="B969D216FFE8FF81FF30F9DF420996A4" bold="true" box="[199,270,1652,1677]" pageId="21" pageNumber="22">Head.</emphasis>
Face occupying about 1/3 of the head width in frontal view; parallel-sided; laterally weakly depressed; pale yellow, with blackish median vitta from antennal fossa to oral margin, where it is at its narrowest; entirely short yellow pilose. Face in profile straight, produced downward at anterior oral margin. Gena black. Frons and vertex black, short pilose. Occiput black. Eyes bare. Antennal fossa about as wide as high. Antenna brown, except scape yellowish basally; antennal ratio approximately 5:1:8; arista slender, slightly longer than half the length of the basoflagellomere.
</paragraph>
<paragraph id="8BA20E04FFE8FF81FF30F8E7402E97DC" blockId="21.[151,1437,1543,2037]" pageId="21" pageNumber="22">
<emphasis id="B969D216FFE8FF81FF30F8E74221974C" bold="true" box="[199,294,1868,1893]" pageId="21" pageNumber="22">Thorax.</emphasis>
Scutum black; short pilose. Postpronotum and postalar callus brown (perhaps yellow in better preserved specimens); short pilose. Scutellum yellow, with basal 1/3 and apical 1/4 blackish; apicomedially sulcate. Anterior and posterior part of anepisternum not differentiated; short pale pilose except on ventral 1/3. Anepimeron entirely white pilose. Katepisternum white pilose dorsally (ventral part not visible in
<typeStatus id="54A6B0A6FFE8FF81FB1DF812461D97F8" box="[1258,1306,1977,2001]" pageId="21" pageNumber="22">type</typeStatus>
specimen). Calypter greyish brown. Halters missing in
<typeStatus id="54A6B0A6FFE8FF81FD77F87641B797DC" box="[640,688,2013,2037]" pageId="21" pageNumber="22">type</typeStatus>
specimen.
</paragraph>
<paragraph id="8BA20E04FFEBFF82FF30FF3C42C190DE" blockId="22.[151,1437,151,824]" pageId="22" pageNumber="23">
Wing: hyaline; microtrichose, except bare on costal cells, on basal 1/3 of cell r1, on cell br except for traces of microtrichia around vena
<taxonomicName id="4C1D7587FFEBFF82FE4FFF1742F890FC" box="[440,511,188,213]" class="Insecta" family="Syrphidae" genus="Stipomorpha" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spuria">spuria</taxonomicName>
, on cell bm, posterobasal 1/3 of cell r4+5, on basal 1/6 of cells DM and CuA1, on anterobasal 1/2 of cell cup.
</paragraph>
<paragraph id="8BA20E04FFEBFF82FF30FEAE41A9914C" blockId="22.[151,1437,151,824]" pageId="22" pageNumber="23">
Legs: Blackish, with tarsi of pro- and mesolegs yellow.
<collectingCountry id="F30A4E94FFEBFF82FCC9FEAF40719134" box="[830,886,260,285]" name="India" pageId="22" pageNumber="23">Hind</collectingCountry>
leg with first tarsomere black, second tarsomere yellow (other tarsomeres missing in
<typeStatus id="54A6B0A6FFEBFF82FDB7FE8341779169" box="[576,624,296,320]" pageId="22" pageNumber="23">type</typeStatus>
specimen). First tarsomeres of all tarsi with strong basoventral tooth. Coxae and trochanters brownish, with pale pile.
</paragraph>
<paragraph id="8BA20E04FFEBFF82FF30FEC443E091F9" blockId="22.[151,1437,151,824]" pageId="22" pageNumber="23">
<emphasis id="B969D216FFEBFF82FF30FEC4424491A1" bold="true" box="[199,323,367,392]" pageId="22" pageNumber="23">Abdomen.</emphasis>
Blackish brown, with yellow posterior margin of tergite 4; mostly whitish pilose. In profile with tergite 4 almost perpendicular to tergite 2. Sternites 1-3 brown, sternite 4 yellow; short pale pilose. Genitalia as in fig. 57.
</paragraph>
<paragraph id="8BA20E04FFEBFF82FF30FE7742A591DC" blockId="22.[151,1437,151,824]" box="[199,418,476,501]" pageId="22" pageNumber="23">
<emphasis id="B969D216FFEBFF82FF30FE77422291DC" bold="true" box="[199,293,476,501]" pageId="22" pageNumber="23">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFEBFF82FF30FE5443E2933D" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BA20E04FFEBFF82FF30FE5441D89249" blockId="22.[151,1437,151,824]" pageId="22" pageNumber="23">
<emphasis id="B969D216FFEBFF82FF30FE5442469231" bold="true" box="[199,321,511,536]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
This species is unique among
<taxonomicName id="4C1D7587FFEBFF82FD6AFE5440269231" box="[669,801,511,536]" class="Insecta" family="Syrphidae" genus="Ceratophya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B969D216FFEBFF82FD6AFE5440269231" box="[669,801,511,536]" italics="true" pageId="22" pageNumber="23">Ceratophya</emphasis>
</taxonomicName>
-species in the presence of a strong basoventral tooth on the first tarsomeres of all tarsi. Furthermore, it's the only known
<taxonomicName id="4C1D7587FFEBFF82FC94FD8F40E09214" box="[867,999,548,573]" class="Insecta" family="Syrphidae" genus="Ceratophya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B969D216FFEBFF82FC94FD8F40E09214" box="[867,999,548,573]" italics="true" pageId="22" pageNumber="23">Ceratophya</emphasis>
</taxonomicName>
-species with an almost black abdomen (except for the yellow posterior margin of tergite 4).
</paragraph>
<paragraph id="8BA20E04FFEBFF82FF30FDC743E2933D" blockId="22.[151,1437,151,824]" pageId="22" pageNumber="23">
<emphasis id="B969D216FFEBFF82FF30FDC7421592AC" bold="true" box="[199,274,620,645]" pageId="22" pageNumber="23">Notes.</emphasis>
When Shannon (1927) described this species, he mentioned the similarity in general appearance to the stingless-bee mimics which he described in the subgenus
<taxonomicName id="4C1D7587FFEBFF82FC74FD2440E39281" box="[899,996,655,680]" class="Insecta" family="Syrphidae" genus="Ubristes" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B969D216FFEBFF82FC74FD2440E39281" box="[899,996,655,680]" italics="true" pageId="22" pageNumber="23">Ubristes</emphasis>
</taxonomicName>
. Nonetheless, he did not place it in
<taxonomicName id="4C1D7587FFEBFF82FF60FD1F43FF92E4" box="[151,248,692,717]" class="Insecta" family="Syrphidae" genus="Ubristes" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B969D216FFEBFF82FF60FD1F43FF92E4" box="[151,248,692,717]" italics="true" pageId="22" pageNumber="23">Ubristes</emphasis>
</taxonomicName>
, as Thompson
<emphasis id="B969D216FFEBFF82FE5AFD1E42E692E4" box="[429,481,692,717]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. (1976) did. Examination of the
<typeStatus id="54A6B0A6FFEBFF82FC92FD1E409192E4" box="[869,918,693,717]" pageId="22" pageNumber="23">type</typeStatus>
revealed that it has all the characters of the genus
<taxonomicName id="4C1D7587FFEBFF82FF13FD7C426C92D9" box="[228,363,727,752]" class="Insecta" family="Syrphidae" genus="Ceratophya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B969D216FFEBFF82FF13FD7C426C92D9" box="[228,363,727,752]" italics="true" pageId="22" pageNumber="23">Ceratophya</emphasis>
</taxonomicName>
. The specimen is a little dirty and greasy, so colours and pilosity are not always easy to assess.
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFEBFF82FF30FCB447D89311" box="[199,1247,799,824]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BA20E04FFEBFF82FF30FCB447D89311" blockId="22.[151,1437,151,824]" box="[199,1247,799,824]" pageId="22" pageNumber="23">
<emphasis id="B969D216FFEBFF82FF30FCB442679311" bold="true" box="[199,352,799,824]" pageId="22" pageNumber="23">Distribution.</emphasis>
Only known from the
<typeStatus id="54A6B0A6FFEBFF82FD97FCB441C59311" box="[608,706,799,824]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
from the Amazon region, probably from
<collectingCountry id="F30A4E94FFEBFF82FB64FCB447DC9311" box="[1171,1243,799,824]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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