166 lines
18 KiB
XML
166 lines
18 KiB
XML
<document id="E5C9D0AE0289649C7D28D38C31149177" ID-DOI="10.11646/zootaxa.5284.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7923218" ID-ZooBank="2A865757-B6B4-48CD-A9AD-334F7E7B508B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683785948286" checkinUser="plazi" docAuthor="Anichtchenko, Alexander & Valainis, Uldis" docDate="2023" docId="03B46F27FFCBFF826DC3F9DCFDA693C6" docLanguage="en" docName="zootaxa.5284.2.1.pdf" docOrigin="Zootaxa 5284 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5284.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omophron (Omophron) schuelei Anichtchenko & Valainis 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="217" masterDocId="FF8D175FFFC4FF926D54FFFCFF8B900F" masterDocTitle="Revision of the subgenus Omophron (s. str.) Latreille, 1802 (Coleoptera: Carabidae Omophron) of the Afrotropical region" masterLastPageNumber="246" masterPageNumber="201" pageNumber="216" updateTime="1683786577624" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="65E9F0BE4D2D51BCE888E15745C73D83">Revision of the subgenus Omophron (s. str.) Latreille, 1802 (Coleoptera: Carabidae Omophron) of the Afrotropical region</mods:title>
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<mods:name id="4F4C391B01AB02382788793BD80AF79D" type="personal">
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<mods:role id="6B1E456333D003027DCDD9206965AD51">
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<mods:roleTerm id="F15E1DCE386F184AA5E87B57F41EC0F7">Author</mods:roleTerm>
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<mods:namePart id="13380FC07B4B1E2733B782C9535AF783">Anichtchenko, Alexander</mods:namePart>
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<mods:nameIdentifier id="94BC53F13E5C910F0BB1E9E46B934503" type="email">alexander.anishchenko@du.lv</mods:nameIdentifier>
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<mods:roleTerm id="B63F7631DC1AD65B21FD78220C342E22">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1BF1F8F6056D5C0B73C320571EEEE05E">Valainis, Uldis</mods:namePart>
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<mods:title id="855EA5F09E8EABBD43FC83835BB35468">Zootaxa</mods:title>
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<mods:part id="94BB9CAAA7B4E5FC34D11074DEF7C1A6">
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<mods:date id="BF6B03088E2077BB95D67418CB92FB92">2023</mods:date>
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<mods:number id="667F1D930AEDC03D6E7AB159EB4CE643">2023-05-11</mods:number>
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<mods:number id="B2D33E20966A23C428924E689414980D">5284</mods:number>
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<mods:url id="F5340600BFBC4059603507A637A6A64B">http://dx.doi.org/10.11646/zootaxa.5284.2.1</mods:url>
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<mods:classification id="72BA3DA2A9F159E693AB9D1BB3449A73">journal article</mods:classification>
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<mods:identifier id="171CD13C1935E0964DD0BB2D1C8AE432" type="DOI">10.11646/zootaxa.5284.2.1</mods:identifier>
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<mods:identifier id="F1A33C91E860AFE1C2E12C58F0A967D1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B46F27FFCBFF826DC3F9DCFDA693C6" ID-DOI="http://doi.org/10.5281/zenodo.7923248" ID-Zenodo-Dep="7923248" LSID="urn:lsid:plazi:treatment:03B46F27FFCBFF826DC3F9DCFDA693C6" httpUri="http://treatment.plazi.org/id/03B46F27FFCBFF826DC3F9DCFDA693C6" lastPageId="16" lastPageNumber="217" pageId="15" pageNumber="216">
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<subSubSection id="C3078DBAFFCBFF9D6DC3F9DCFDE59634" box="[151,622,1568,1595]" pageId="15" pageNumber="216" type="nomenclature">
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<paragraph id="8BA2DE31FFCBFF9D6DC3F9DCFDE59634" blockId="15.[151,622,1568,1595]" box="[151,622,1568,1595]" pageId="15" pageNumber="216">
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<heading id="D0EA695DFFCBFF9D6DC3F9DCFDE59634" bold="true" box="[151,622,1568,1595]" fontSize="11" level="1" pageId="15" pageNumber="216" reason="1">
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<emphasis id="B9690223FFCBFF9D6DC3F9DCFDE59634" bold="true" box="[151,622,1568,1595]" pageId="15" pageNumber="216">
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<taxonomicName id="4C1DA5B2FFCBFF9D6DC3F9DCFDAF9634" authority="Anichtchenko & Valainis, 2023" authorityName="Anichtchenko & Valainis" authorityYear="2023" box="[151,548,1568,1595]" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="schuelei" status="sp. n." subGenus="Omophron">
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<emphasis id="B9690223FFCBFF9D6DC3F9DCFDAF9634" bold="true" box="[151,548,1568,1595]" italics="true" pageId="15" pageNumber="216">Omophron (Omophron) schuelei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25ABF58FFCBFF9D6F78F9DDFDE59634" box="[556,622,1569,1595]" pageId="15" pageNumber="216" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3078DBAFFCBFF9D6DC3F994FCA796A8" pageId="15" pageNumber="216" type="materials_examined">
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<paragraph id="8BA2DE31FFCBFF9D6DC3F994FCA796A8" blockId="15.[151,1437,1640,2026]" pageId="15" pageNumber="216">
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<emphasis id="B9690223FFCBFF9D6DC3F994FE0E968D" bold="true" box="[151,389,1640,1666]" pageId="15" pageNumber="216">Material examined.</emphasis>
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<materialsCitation id="3B75D46CFFCBFF9D6CC4F995FCA796A8" collectingDate="2010-01-18" collectionCode="PSC" collectorName="Southern Kafue NP & NW Nanzhila Plains & P. Schule" country="Zambia" latitude="-16.209732" location="Southern Kafue" longLatPrecision="1" longitude="25.79454" municipality="Nanzhila Plains" pageId="15" pageNumber="216" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<emphasis id="B9690223FFCBFF9D6CC4F995FD8F968C" bold="true" box="[400,516,1641,1667]" pageId="15" pageNumber="216">
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<typeStatus id="54A66093FFCBFF9D6CC4F995FE74968C" box="[400,511,1641,1667]" pageId="15" pageNumber="216">Holotype</typeStatus>
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,
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</emphasis>
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<specimenCount id="9D1B15B8FFCBFF9D6F44F995FDAE968C" box="[528,549,1641,1667]" pageId="15" pageNumber="216" type="female">♀</specimenCount>
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:
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<collectingCountry id="F30A9EA1FFCBFF9D6F6DF994FD1A968D" box="[569,657,1640,1666]" name="Zambia" pageId="15" pageNumber="216">Zambia</collectingCountry>
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,
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<collectorName id="26E8BBE7FFCBFF9D6FF5F994FB0A968D" box="[673,1153,1640,1667]" pageId="15" pageNumber="216">
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<location id="8EC288EAFFCBFF9D6FF5F994FCDD968D" LSID="urn:lsid:plazi:treatment:03B46F27FFCBFF826DC3F9DCFDA693C6:8EC288EAFFCBFF9D6FF5F994FCDD968D" box="[673,854,1640,1666]" country="Zambia" latitude="-16.209732" longLatPrecision="1" longitude="25.79454" municipality="Nanzhila Plains" name="Southern Kafue" pageId="15" pageNumber="216">Southern Kafue</location>
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NP NW
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<collectingMunicipality id="6BC6444BFFCBFF9D6E9EF994FB0A968D" box="[970,1153,1640,1666]" pageId="15" pageNumber="216">Nanzhila Plains</collectingMunicipality>
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</collectorName>
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,
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<geoCoordinate id="EE29B8F6FFCBFF9D69C5F994FA87968C" box="[1169,1292,1640,1667]" degrees="16" direction="south" minutes="12" orientation="latitude" pageId="15" pageNumber="216" precision="15" seconds="35" value="-16.209723">16°12’35S</geoCoordinate>
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,
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<geoCoordinate id="EE29B8F6FFCBFF9D6848F994FA13968C" box="[1308,1432,1640,1667]" degrees="25" direction="east" minutes="47" orientation="longitude" pageId="15" pageNumber="216" precision="15" seconds="40" value="25.794443">25°47’40E</geoCoordinate>
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,
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<date id="FFA3F8F1FFCBFF9D6DC3F971FE9396A8" box="[151,280,1677,1703]" pageId="15" pageNumber="216" value="2010-01-18">
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<collectingDate id="EFE70119FFCBFF9D6DC3F971FE9396A8" box="[151,280,1677,1703]" pageId="15" pageNumber="216" value="2010-01-18">18.01.2010</collectingDate>
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</date>
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,
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<collectorName id="26E8BBE7FFCBFF9D6C77F971FE0196A9" box="[291,394,1676,1703]" pageId="15" pageNumber="216">P. Schüle</collectorName>
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leg. [
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<geoCoordinate id="EE29B8F6FFCBFF9D6C9CF970FDC696A9" box="[456,589,1676,1702]" degrees="16.209732" direction="south" orientation="latitude" pageId="15" pageNumber="216" precision="1" value="-16.209732">-16.209732</geoCoordinate>
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,
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<geoCoordinate id="EE29B8F6FFCBFF9D6F0DF970FD5896A8" box="[601,723,1676,1703]" degrees="25.794541" direction="east" orientation="longitude" pageId="15" pageNumber="216" precision="1" value="25.79454">25.794541</geoCoordinate>
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] (
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<collectionCode id="ED0C46F4FFCBFF9D6FBCF971FCAB96A8" box="[744,800,1677,1703]" pageId="15" pageNumber="216">PSC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3078DBAFFCBFF9D6D93F94CFB2396E1" pageId="15" pageNumber="216" type="diagnosis">
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<paragraph id="8BA2DE31FFCBFF9D6D93F94CFB2396E1" blockId="15.[151,1437,1640,2026]" pageId="15" pageNumber="216">
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<emphasis id="B9690223FFCBFF9D6D93F94CFE4496C5" bold="true" box="[199,463,1712,1738]" pageId="15" pageNumber="216">Differential diagnosis.</emphasis>
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A very unusual species, easily distinguished from all known species by the following combination of characters: size large, pronotum strongly oblate, with widely explanate sides.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3078DBAFFCBFF826D93F904FE7F9352" lastPageId="16" lastPageNumber="217" pageId="15" pageNumber="216" type="description">
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<paragraph id="8BA2DE31FFCBFF9D6D93F904FC4D971C" blockId="15.[151,1437,1640,2026]" box="[199,966,1784,1811]" pageId="15" pageNumber="216">
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<emphasis id="B9690223FFCBFF9D6D93F904FED2971D" bold="true" box="[199,345,1784,1810]" pageId="15" pageNumber="216">Description.</emphasis>
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Body length
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<quantity id="4CE573D4FFCBFF9D6CA4F904FDC2971C" box="[496,585,1784,1811]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="15" pageNumber="216" unit="mm" value="6.4">6.4 mm</quantity>
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; width
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<quantity id="4CE573D4FFCBFF9D6FCFF905FD7F971C" box="[667,756,1785,1811]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="216" unit="mm" value="4.2">4.2 mm</quantity>
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. Habitus (
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<figureCitation id="1326C2B4FFCBFF9D6E31F904FC31971C" box="[869,954,1784,1811]" captionStart="FIGURES 21–23" captionStartId="14.[152,270,1376,1400]" captionTargetBox="[156,1435,526,1353]" captionTargetId="figure-169@14.[152,1436,525,1353]" captionTargetPageId="14" captionText="FIGURES 21–23. Habitus of Omophron “minutum”—species group, dorsal view: 21—O. (O.) muellerae sp. n.; 22—O. (O.) schuelei sp. n. (Holotype); 23—O. (O) mimeticum sp. n. (Holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7923246" httpUri="https://zenodo.org/record/7923246/files/figure.png" pageId="15" pageNumber="216">Fig. 22</figureCitation>
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).
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</paragraph>
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<paragraph id="8BA2DE31FFCBFF9D6D93F8E0FD1E97AD" blockId="15.[151,1437,1640,2026]" pageId="15" pageNumber="216">Color: head uniformly dark brown, almost black, except narrow brown margin of clypeus. Labrum pearl color. Mandibles dark brown. Pronotum deep brown, almost black, with wide, reddish lateral margins. Elytra deep brown, almost black, with narrow reddish brown lateral pattern, lateral interval of pearl color. Legs, antennae and mouth parts light reddish brown. Venter dark brown.</paragraph>
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<paragraph id="8BA2DE31FFCBFF826D93F850FA0890F6" blockId="15.[151,1437,1640,2026]" lastBlockId="16.[151,1437,151,969]" lastPageId="16" lastPageNumber="217" pageId="15" pageNumber="216">
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Head relatively flat, evenly, densely and coarsely punctate, moderately wrinkled near the eyes, with a deep impression between frons and clypeus. Subocular ridge well developed, curved upwards and slightly rounded. Clypeus moderately convex, smooth, bisetose. Labrum slightly emarginate, with moderately rounded sides and six setae. Antennal scape unisetose apically, antennomeres 3 and 4 smooth, with a few longer setae around the apex, 5– 11 pubescent. Gula smooth, gena coarsely and densely punctate, strongly wrinkled near the eyes. HW:
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<quantity id="4CE573D4FFD4FF82684CFF23FAF490F6" box="[1304,1407,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="16" pageNumber="217" unit="mm" value="2.18">2.18 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BA2DE31FFD4FF826D93FEFFFCDA91FA" blockId="16.[151,1437,151,969]" pageId="16" pageNumber="217">
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The shape of the pronotum is peculiar and easy distinguishable from that of other African
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<taxonomicName id="4C1DA5B2FFD4FF826990FEFFFA169112" box="[1220,1437,259,285]" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B9690223FFD4FF826990FEFFFAB79112" box="[1220,1340,259,285]" italics="true" pageId="16" pageNumber="217">Omophron</emphasis>
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species.
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</taxonomicName>
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Pronotum (PL:
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<quantity id="4CE573D4FFD4FF826C11FEDBFE20914E" box="[325,427,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.86" pageId="16" pageNumber="217" unit="mm" value="1.86">1.86 mm</quantity>
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, PW:
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<quantity id="4CE573D4FFD4FF826CBAFEDBFDDD914E" box="[494,598,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.64" pageId="16" pageNumber="217" unit="mm" value="3.64">3.64 mm</quantity>
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.) strongly oblate on disc, base bisinuate on each side sides evenly rounded, with widely explanate border. Pale margin of pronotum wide, extended from base to anterior margin, becoming obsolete at the edge Anterior angles strongly acute. Disc of pronotum with wide transverse impression in anterior half, and with two wide impressions on the base; coarsely and densely punctate, sides smooth. Median impression absent Prosternum coarsely but sparsely punctate, proepisternum coarsely and more densely punctate, proepipleura almost smooth, with only 1–2 small punctures in anterior part.
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</paragraph>
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<paragraph id="8BA2DE31FFD4FF826D93FE03FE5C928A" blockId="16.[151,1437,151,969]" pageId="16" pageNumber="217">
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Elytra (EL:
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<quantity id="4CE573D4FFD4FF826C1FFE03FE3B9216" box="[331,432,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="4.37" pageId="16" pageNumber="217" unit="mm" value="4.37">4.37 mm</quantity>
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, EW:
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<quantity id="4CE573D4FFD4FF826CA6FE03FDD29216" box="[498,601,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="4.23" pageId="16" pageNumber="217" unit="mm" value="4.23">4.23 mm</quantity>
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.) moderately convex, flattened on disc and expressly oblate near pronotum, ovate, moderately rounded laterally, strongly rounded at apex. Border of elytra widened near shoulders and narrower towards apex. Elytra with 15 striae. Striae deep, densely but finely punctate. Intervals weakly convex on the disc, moderately convex on sides.
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</paragraph>
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<paragraph id="8BA2DE31FFD4FF826D93FD73FD419336" blockId="16.[151,1437,151,969]" pageId="16" pageNumber="217">
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Venter (
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<figureCitation id="1326C2B4FFD4FF826C77FD73FEF092A6" box="[291,379,654,681]" captionStart="FIGURES 24–26" captionStartId="15.[152,270,1426,1450]" captionTargetBox="[153,1434,744,1401]" captionTargetId="figure-229@15.[151,1435,741,1403]" captionTargetPageId="15" captionText="FIGURES 24–26. Habitus of Omophron “minutum”—species group, ventral view: 24—O. (O.) muellerae sp. n.; 25—O. (O.) schuelei sp. n. (Holotype); 26—O. (O) mimeticum sp. n. (Holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7923250" httpUri="https://zenodo.org/record/7923250/files/figure.png" pageId="16" pageNumber="217">Fig. 25</figureCitation>
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). Mesepisternum punctate. Metasternum coarsely but sparsely punctate, metepisternum and metepimeron coarsely but unevenly punctate, slightly wrinkled near epipleura. Elytral epipleura almost smooth, with only 1–2 small punctures. Ventrites I–III coarsely but sparsely punctate, ventrites IV–V with 2–3 large punctures on sides and a row of small punctures along transversal sulcus; last ventrite VI with deep and very large punctures. Metacoxa bisetose, with 2–4 additional punctures.
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</paragraph>
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<paragraph id="8BA2DE31FFD4FF826D93FCBFFE7F9352" blockId="16.[151,1437,151,969]" box="[199,500,835,861]" pageId="16" pageNumber="217">Aedeagus. Male unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C3078DBAFFD4FF826D93FC9BFE1593AA" pageId="16" pageNumber="217" type="etymology">
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<paragraph id="8BA2DE31FFD4FF826D93FC9BFE1593AA" blockId="16.[151,1437,151,969]" pageId="16" pageNumber="217">
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<emphasis id="B9690223FFD4FF826D93FC9BFEC5938E" bold="true" box="[199,334,871,897]" pageId="16" pageNumber="217">Etymology.</emphasis>
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New species named after our friend and colleague Peter Schüle (Herrenberg,
|
||
<collectingCountry id="F30A9EA1FFD4FF8269BCFC9BFADE938E" box="[1256,1365,871,897]" name="Germany" pageId="16" pageNumber="217">Germany</collectingCountry>
|
||
), who collected the specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3078DBAFFD4FF826D93FC52FDA693C6" box="[199,557,942,969]" pageId="16" pageNumber="217" type="distribution">
|
||
<paragraph id="8BA2DE31FFD4FF826D93FC52FDA693C6" blockId="16.[151,1437,151,969]" box="[199,557,942,969]" pageId="16" pageNumber="217">
|
||
<emphasis id="B9690223FFD4FF826D93FC52FEEA93C7" bold="true" box="[199,353,942,968]" pageId="16" pageNumber="217">Distribution.</emphasis>
|
||
<collectingCountry id="F30A9EA1FFD4FF826C33FC53FE3693C6" box="[359,445,943,969]" name="Zambia" pageId="16" pageNumber="217">Zambia</collectingCountry>
|
||
(Map. 2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |