194 lines
20 KiB
XML
194 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.7.105423" ID-Pensoft-Pub="2535-0730-2-253" ID-Pensoft-UUID="5A634470919E5CDBB7C3E87212C43D06" ID-ZooBank="C62F010297A64762B0EC1B98EB917868" ModsDocID="2535-0730-7-2-253" checkinTime="1690667349210" checkinUser="pensoft" docAuthor="Wagner, Gerhard" docDate="2023" docId="F79D030AEC73567BA48B19FBC1ADBDE2" docLanguage="en" docName="EvolutSyst 7(2): 253-256" docOrigin="Evolutionary Systematics 7 (2)" docPubDate="2023-07-28" docSource="http://dx.doi.org/10.3897/evolsyst.7.105423" docTitle="Thraustocolus (Prothraustocola) clypealis subsp. minor Wagner 2023, ssp. nov." docType="treatment" docUuid="3F8D8A70-EF4C-4A61-84F5-863A6C8DE2D1" docUuidSource="ZooBank" docVersion="1" id="5A634470919E5CDBB7C3E87212C43D06" lastPageNumber="253" masterDocId="5A634470919E5CDBB7C3E87212C43D06" masterDocTitle="A new species and subspecies of the darkling beetle genus Thraustocolus Kraatz, 1866 from Oman (Coleoptera, Tenebrionidae)" masterLastPageNumber="256" masterPageNumber="253" pageNumber="253" updateTime="1690667349210" updateUser="pensoft">
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<mods:title id="73B06E422FD631053F6E30BF89701E66">A new species and subspecies of the darkling beetle genus Thraustocolus Kraatz, 1866 from Oman (Coleoptera, Tenebrionidae)</mods:title>
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<mods:name id="C934534981B00299DD37F06AF685C45A" type="personal">
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<mods:roleTerm id="0DEC67D915879D55B864C423F554B6A0">Author</mods:roleTerm>
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<mods:namePart id="52AF5DCAF2110D4F60906D94B68D158E">Wagner, Gerhard</mods:namePart>
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<mods:affiliation id="70B7419D0AA8ADC16954EA73003B0312">Bilenbarg 27 B, 22397 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="B83C8188124612B33D2DC341B4C424DD" type="email">caeciliawagner@gmx.de</mods:nameIdentifier>
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<mods:title id="EBDE4FD9E9D8FA9AF5ECB18710F10D62">Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part id="AB56BB63D735B76D1CF8F6F012C67A6B">
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<mods:date id="77F38FBC42187E0F5D069BAA4047C320">2023</mods:date>
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<mods:number id="BB5D87EBD338FBD1C7E6566D93DFE8D7">2023-07-28</mods:number>
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<mods:number id="571C370C84FF29AFD6AEE4A5E003E344">7</mods:number>
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<mods:number id="23A6F48CD3B6B1541B7495FE7BE33F3A">2</mods:number>
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<mods:start id="53A5111874EFD16669831A125AEBA0C3">253</mods:start>
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<mods:end id="AF95ABEE0B24351E9D0E04D077A6D17F">256</mods:end>
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<mods:url id="472205DA540039144FE466FF7683D4CE">http://dx.doi.org/10.3897/evolsyst.7.105423</mods:url>
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<mods:classification id="86C49E66D8BD006E59BE762F2A7C2D06">journal article</mods:classification>
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<mods:identifier id="788FE3C8F217B5B159B234C5AB63AEEC" type="DOI">http://dx.doi.org/10.3897/evolsyst.7.105423</mods:identifier>
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<subSection id="FE0751CA82C0D8FE4EB09CDE4DA8ECC7" pageId="0" pageNumber="253" type="results">
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<treatment id="F79D030AEC73567BA48B19FBC1ADBDE2" LSID="urn:lsid:zoobank.org:act:3F8D8A70-EF4C-4A61-84F5-863A6C8DE2D1" httpUri="http://treatment.plazi.org/id/F79D030AEC73567BA48B19FBC1ADBDE2" lastPageNumber="253" pageId="0" pageNumber="253">
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<subSubSection id="95CA6D36BAED076EDC1CE4584789831C" pageId="0" pageNumber="253" type="nomenclature">
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<paragraph id="214871B5A90BBD368CD5110C4FEAA332" pageId="0" pageNumber="253">
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<taxonomicName id="37B115EC58DFB0A1F7F8323B93794721" LSID="https://zoobank.org/3F8D8A70-EF4C-4A61-84F5-863A6C8DE2D1" authority="Wagner, 2023" authorityName="Wagner" authorityYear="2023" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus (Prothraustocola) clypealis subsp. minor" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="subSpecies" species="clypealis" status=" ssp. nov." subGenus="Prothraustocola" subSpecies="minor">Thraustocolus (Prothraustocola) clypealis minor</taxonomicName>
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<taxonomicNameLabel id="28DFCE106E026C3DC6BC4050E6E8D654" pageId="0" pageNumber="253">ssp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="2FC0E3A4A64F24C2E67C8CC11F624447" pageId="0" pageNumber="253" type="description">
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<paragraph id="46AD6289299FF9D0D65B261EBC0AA52A" pageId="0" pageNumber="253">
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<figureCitation id="D5576AAED29E437B1ABA8C730DB1590D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thraustcolus clypealis sp. nov. Thraustcolus clypealis minor ssp. nov. dorsal view." httpUri="https://binary.pensoft.net/fig/885480" pageId="0" pageNumber="253">Fig. 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8B45EE991A877571A52A1440F89286BD" pageId="0" pageNumber="253" type="materials_examined">
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<paragraph id="F96EB4FB4BAEED303FBF7E9123396B40" pageId="0" pageNumber="253">Material examined.</paragraph>
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<paragraph id="E20C457C8656348C349249F3C07FEE91" pageId="0" pageNumber="253">
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<materialsCitation id="FE9CC9C3C43A7D87C671605FE1875921" collectingDate="2016-03-08" collectorName="C. & G. Wagner, Zoological Museum, Museum der Natur, Hamburg" country="Oman" latitude="24.333334" location="Fossil Valley" longLatPrecision="1252" longitude="55.9" municipality="Mahdah" specimenCount="♂" typeStatus="Holotype">
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<emphasis id="4FB5CE576F2A3C3188D810C087193CB4" bold="true" pageId="0" pageNumber="253">
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<emphasis id="692F90D06A01A7CC32F73ED4B3BA7CED" italics="true" pageId="0" pageNumber="253">
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<typeStatus id="24A66D11831C90CCE6DAD37117A7356A">Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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<specimenCount id="C8C328DDF95D410677652604FA8D8E4D">♂</specimenCount>
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:
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<collectingCountry id="DFD3E9B1101EB52C6AEF6EFA5EE19288" name="Oman">Oman</collectingCountry>
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,
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<location id="B6C9BE74235F546EE3D507014A75831B" LSID="urn:lsid:plazi:treatment:F79D030AEC73567BA48B19FBC1ADBDE2:B6C9BE74235F546EE3D507014A75831B" country="Oman" latitude="24.333334" longLatPrecision="1252" longitude="55.9" municipality="Mahdah" name="Fossil Valley">Fossil Valley</location>
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,
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<quantity id="C4F5DFAF1C589A4A88A826A9B6D946AE" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
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southwest of
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<collectingMunicipality id="5EDA183409E40490D46AD691345CD02F">Mahdah</collectingMunicipality>
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,
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<geoCoordinate id="4434D282640426BE25E94704C5AC6D5A" degrees="24" direction="north" minutes="20" orientation="latitude" precision="925" value="24.333334">24°20'N</geoCoordinate>
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,
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<geoCoordinate id="6C1A330DF665AEF53CDC32247888DDFE" degrees="55" direction="east" minutes="54" orientation="longitude" precision="925" value="55.9">55°54'E</geoCoordinate>
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,
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<collectingDate id="7D54475635CCF7115E111D6802452C07" value="2016-03-08">08.03.2016</collectingDate>
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, leg.
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<collectorName id="A0B9358DA7520B09B390F3A26EDEBA99">C. & G. Wagner</collectorName>
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, currently kept by the author, later in
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<collectorName id="7BC7127045FD9D3B4A7A64CCBF99D817">Zoological Museum</collectorName>
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,
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<collectorName id="FDCEA82EF42A8A44898C23E13BC33948">Museum der Natur</collectorName>
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,
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<collectorName id="C8039CE07D0082EA9E2FF06B3D3FED5E">Hamburg</collectorName>
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(ZMH)
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</materialsCitation>
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.
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<materialsCitation id="0FC5E19BCF66B45E6B6F49D22669BD0B" collectingDate="2016-03-08" collectorName="C. & G. Wagner, Zoological Museum, Museum der Natur, Hamburg" country="Oman" latitude="24.333334" location="Fossil Valley" longLatPrecision="1252" longitude="55.9" municipality="Mahdah" specimenCount="2" specimenCount-male="2" typeStatus="Paratypes">
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<emphasis id="DA5B81C60BBA9129228C25B956672111" bold="true" italics="true" pageId="0" pageNumber="253">
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<typeStatus id="EE8D2A387C07E8726F1DADCD043C7457">Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount id="4958F27D93F35B28CE81D0C9E4D2B60F" type="male">2♂</specimenCount>
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:
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<collectingCountry id="C0AB39BAB57E98DAA8C5E792AAD83ABD" name="Oman">Oman</collectingCountry>
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,
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<location id="FFD684E10C913DB9178D0C720D57B621" LSID="urn:lsid:plazi:treatment:F79D030AEC73567BA48B19FBC1ADBDE2:FFD684E10C913DB9178D0C720D57B621" country="Oman" latitude="24.333334" longLatPrecision="1252" longitude="55.9" municipality="Mahdah" name="Fossil Valley">Fossil Valley</location>
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,
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<quantity id="E293E14A2389374E12FAF51400BFEFB6" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
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southwest of
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<collectingMunicipality id="83BCCD2B15ED876DECB62AD3A86E0297">Mahdah</collectingMunicipality>
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,
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<geoCoordinate id="A70030A708F7C3056762E11BBD23F914" degrees="24" direction="north" minutes="20" orientation="latitude" precision="925" value="24.333334">24°20'N</geoCoordinate>
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,
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<geoCoordinate id="23A3FA6E9648E2C6CB91B447AA48FAC6" degrees="55" direction="east" minutes="54" orientation="longitude" precision="925" value="55.9">55°54'E</geoCoordinate>
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,
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<collectingDate id="73EEF27935CA185DB8FEEB3F0B5F3880" value="2016-03-08">08.03.2016</collectingDate>
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, leg.
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<collectorName id="0018DFA4F9A8FCA4CAC365DE2ABE48FF">C. & G. Wagner</collectorName>
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, currently kept by the author, later in
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<collectorName id="711C0260A489D0A528DDECF7CD5D3E4E">Zoological Museum</collectorName>
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,
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<collectorName id="7C6D1F81A004B92DEC10E75E4E331F60">Museum der Natur</collectorName>
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,
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<collectorName id="3792ACD8E066D10CEC82CD2AC5A300CC">Hamburg</collectorName>
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(ZMH)
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</materialsCitation>
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.
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</paragraph>
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<caption id="00E6EDF68B9F7A78227B75CB730E516B" httpUri="https://binary.pensoft.net/fig/885480" pageId="0" pageNumber="253" start="Figure 2" startId="F2">
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<paragraph id="91800F2763D92BC3D4D0493F83CA995A" pageId="0" pageNumber="253">
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<emphasis id="147A0968C66486F4BC6C9FE516E1229D" bold="true" pageId="0" pageNumber="253">Figure 2.</emphasis>
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<taxonomicName id="DD7B4577AE06A77D17F7988A1619822C" genus="Thraustcolus" lsidName="Thraustcolus clypealis" pageId="0" pageNumber="253" rank="species" species="clypealis">Thraustcolus clypealis</taxonomicName>
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sp. nov.
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<taxonomicName id="84190A5011E57E1C9E12A0B7101539AB" genus="Thraustcolus" lsidName="Thraustcolus minor" pageId="0" pageNumber="253" rank="species" species="minor">Thraustcolus clypealis minor</taxonomicName>
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ssp. nov. dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="9A09D233B8B6E9790DFE6696F41781D0" pageId="0" pageNumber="253" type="etymology">
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<paragraph id="D74EF9DBCB42453BBD373D113D0BC95D" pageId="0" pageNumber="253">Etymology.</paragraph>
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<paragraph id="7A8435E9BF6508272CCA69986955858A" pageId="0" pageNumber="253">
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The subspecies is named after the Latin
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<normalizedToken id="9F63629697F4D5018E7AE794EC7829AB" originalValue="“minor“">"minor"</normalizedToken>
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, because it differs essentially from the forma typica in its smaller size.
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</paragraph>
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</subSubSection>
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<subSubSection id="03BF17266FE8CBC9902FDAD46208FA20" pageId="0" pageNumber="253" type="description">
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<paragraph id="7537A9BE888B62BC7326A99C932211CB" pageId="0" pageNumber="253">Description.</paragraph>
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<paragraph id="FB8CE23A648A174CD0371E21008E4B55" pageId="0" pageNumber="253">
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Similar to
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<taxonomicName id="D525D2F1B9673002D62179A727E97074" authorityName="Wagner" authorityYear="2023" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus (Prothraustocola) clypealis" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="clypealis" subGenus="Prothraustocola">Thraustocolus (Prothraustocola) clypealis</taxonomicName>
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sp. nov., but noticeably smaller: length 6.6-7.6 mm, width 2.3-2.8 mm; sides of pronotum slightly less rounded, fore legs a little longer and more slender, brown-black.
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</paragraph>
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</subSubSection>
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<subSubSection id="BD2156AA7FAB8B88965B9D5224B3A40D" pageId="0" pageNumber="253" type="differential diagnosis">
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<paragraph id="DE168784BD22F5244DE8A4CC5E9200CA" pageId="0" pageNumber="253">Differential diagnosis.</paragraph>
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<paragraph id="1C87009B6F9DD5D2F0D71057C709991D" pageId="0" pageNumber="253">
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There are currently 6 species of the genus
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<taxonomicName id="240C3A8E63CE85A1CFE75B13E0863F43" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="genus">
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<emphasis id="E6669E542563C53D048BCA55E0CAF068" italics="true" pageId="0" pageNumber="253">Thraustocolus</emphasis>
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</taxonomicName>
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known from the Arabian Peninsula and one from southern Iran which are differentiated below from the new species and its subspecies. For this delimitation numerous characteristics are used which the new species and subspecies have in common. Distinguishing the new species and its subspecies from the previously known species of
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<taxonomicName id="D9B14F6C4F725EF93C1610366B28F71F" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="genus">
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<emphasis id="9EA6BC1319E37A8626235E952DA0D20A" italics="true" pageId="0" pageNumber="253">Thraustocolus</emphasis>
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</taxonomicName>
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from Oman, Iran and Saudi Arabia, can be done based on the following characters:
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<taxonomicName id="08B130551DD6A658E61D417C4EA56261" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus rugosus" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="rugosus">Thraustocolus (s. str.) rugosus</taxonomicName>
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Kaszab, 1979 from Oman, Saudi Arabia and the United Arab Emirates: Head and pronotum evenly punctured, sides of pronotum wrinkled punctured. Elytra sparse and finely rasp-like granulate. 7-7.8 mm.
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</paragraph>
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<paragraph id="C07D17CB2435B449CA6F41BA1626FDD0" pageId="0" pageNumber="253">
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<taxonomicName id="76381707375B93986CE7106A3BA522B5" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus arabicus" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="arabicus">Thraustocolus (s. str.) arabicus</taxonomicName>
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Kaszab, 1979 from Saudi Arabia: Head and pronotum evenly roughly punctured, pronotum punctured on the sides rasp-like. Elytra granulate. 6-10 mm. In contrast to these two species, the new species and its subspecies have an evenly roundly punctured pronotum, finer than head. Elytra not grained.
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</paragraph>
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<paragraph id="CCE8927A8ECE73A50C77C6847BD7A00D" pageId="0" pageNumber="253">
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<taxonomicName id="29B7DE6888AD44C07231B538EBB96BE1" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus catomoides" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="catomoides">Thraustocolus (s. str.) catomoides</taxonomicName>
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Kaszab, 1981 from Saudi Arabia: 3rd antennal segment 1.9 times as long as the 2nd, disc of the pronotum very sparsely punctured, the base dense, laterally vanishing and rasp-like densely granular-punctate. 4.3 mm. In the new species and subspecies the 3rd antennal segment is 2.8 times longer than the 2nd. Pronotum even with round punctures.
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</paragraph>
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<paragraph id="FD549A8B47FABCF7FB65E3373290EC08" pageId="0" pageNumber="253">
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<taxonomicName id="E801A0677DE1F007975E2F76AEE712D7" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus propheta" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="propheta">Thraustocolus (s. str.) propheta</taxonomicName>
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Koch, 1941 from Saudi Arabia: eyes large, completely detached, strong from the contours of genae, only slightly protruding from those of the temples, significantly projecting outwards above the level of genae. 3rd antennal segment only almost twice as long as the 2nd segment. 5-6 mm. In the new species and subspecies the eyes are flat and the 3rd antennal segment is 2.8 times as long as the 2rd.
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</paragraph>
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<paragraph id="FB80BFEC4E49C529B60632BA7367DBC0" pageId="0" pageNumber="253">
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<taxonomicName id="DFDD5BED5F08FA2241E121E4AF812C3C" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus (Prothraustocola) pitcheri" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="pitcheri" subGenus="Prothraustocola">Thraustocolus (Prothraustocola) pitcheri</taxonomicName>
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Johnson, 1989 from Saudi Arabia: eyes bulbous, protruding, the basal margin of the pronotum is briefly interrupted in the middle in ♂. 8.5 mm. In the new species and subspecies the eyes are flat and the base of the pronotum is clearly marginated throughout.
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</paragraph>
|
||
<paragraph id="41328CAFF75F0ABD696BB9E38A725DC9" pageId="0" pageNumber="253">
|
||
<taxonomicName id="07AC602A6D453E14CCC1EB043ACA17D6" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus (Prothraustocola) hormozganus" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="hormozganus" subGenus="Prothraustocola">Thraustocolus (Prothraustocola) hormozganus</taxonomicName>
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Grimm & Merkl, 2018 from Iran: punctures on pronotum rougher than on head, many punctures distinctly elongate. Prosternal apophysis between procoxae does not protrude backwards. Basal part of aedeagus is significantly shorter than the apical. 7.1-10.5 mm. In the new species and subspecies the pronotum is evenly provided with round punctures. Prosternal apophysis protrudes behind procoxae as a tip. Basal and apical sections of the aedeagus are about the same length.
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||
</paragraph>
|
||
<paragraph id="E72D2F2E4BC8C1F978E5AE01087A82E2" pageId="0" pageNumber="253">
|
||
<taxonomicName id="15D35014678BA91AD3D6B61B3942BB23" class="Insecta" family="Tenebrionidae" genus="Thraustocolus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraustocolus (P.) belutschistanica" order="Coleoptera" pageId="0" pageNumber="253" phylum="Arthropoda" rank="species" species="belutschistanica" subGenus="P.">Thraustocolus (P.) belutschistanica</taxonomicName>
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Kaszab, 1957 from Iran, Baluchistan: head dorsally finely and very sparsely punctured; pronotum almost cylindrical, lateral margin anteriorly completely extinguished. Prosternal apophysis not protruding beyond procoxae. 8.5 mm. In the new species and subspecies the head is dorsaly densely and clearly punctured. Pronotum clearly rounded laterally and completely marginated. Prosternal apophysis protrudes as a tip behind procoxae.
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||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
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</document> |