281 lines
31 KiB
XML
281 lines
31 KiB
XML
<document id="214A37A62918AC51ACD886961BAC2481" ID-CLB-Dataset="27556" ID-DOI="10.11646/zootaxa.4555.1.6" ID-GBIF-Dataset="6de65293-38cf-4b21-9a93-dacb22d1300d" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624177" ID-ZooBank="02458E32-85F2-4624-911A-2DB5FFCED3D1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554272039380" checkinUser="plazi" docAuthor="Purkayastha, Jayaditya & David, Patrick" docDate="2019" docId="03D5FA59FFBB4F455F84FF78FF26FE21" docLanguage="en" docName="zootaxa.4555.1.6.pdf" docOrigin="Zootaxa 4555 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hebius lacrima Purkayastha & David 2019, spec. nov." docType="treatment" docVersion="8" lastPageNumber="83" masterDocId="FFEC8221FFB94F415F13FFEFFFF4FFF1" masterDocTitle="A new species of the snake genus Hebius Thompson from Northeast India (Squamata: Natricidae)" masterLastPageNumber="90" masterPageNumber="79" pageNumber="81" updateTime="1698716362599" updateUser="ExternalLinkService">
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<mods:mods id="84868B1C88844FC28A8F7A013010DA56" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="70BE480AEA78054E9929680218D325C3">
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<mods:title id="61E7541A2A0570AAF7A83FD1F145EE7E">A new species of the snake genus Hebius Thompson from Northeast India (Squamata: Natricidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="23358210133C3FE574F3F7ACC7C134DA" type="personal">
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<mods:roleTerm id="5AAACB01E3A48C24BA0FA531B54D0A7F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EA7E98E51993A81874B90655D4BC7754">Purkayastha, Jayaditya</mods:namePart>
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</mods:name>
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<mods:name id="AF55FDAFBFD1E51471A586A3DF3CB5B9" type="personal">
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<mods:role id="A3B909184832AD760AE2720FDEA8FF47">
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<mods:roleTerm id="5445FD962FCC72604705DD1720924849">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="20C2EB31D1AAA173C8F5514E8B8FC442">David, Patrick</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="D4F121EC2395DE7D4EBB0B4D8F33BBE2">text</mods:typeOfResource>
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<mods:titleInfo id="90B2B990A853CA54F6D6CF84C7DEC91D">
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<mods:title id="55CAF0AD06465F152B831763943D066B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="593FA8DA7B8FC77DD563D16A6573A930">
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<mods:date id="EB3CB0E9B85C72AF9FAC941AAAB87585">2019</mods:date>
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<mods:detail id="D9126252298781EA4A17D5372093FB72" type="pubDate">
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<mods:number id="0574EE6E0B13D678ABBABA848814AEBB">2019-02-12</mods:number>
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</mods:detail>
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<mods:detail id="A6C7BB50A3900228283624E95495479D" type="volume">
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<mods:number id="1BCD697067CA3D3EE43BBF4B4CF68CA2">4555</mods:number>
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</mods:detail>
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<mods:detail id="2A6F5D767B963118DDA900A0E6CCD5CE" type="issue">
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<mods:number id="8788F3D421A65AE48AEEBEE8D30E91B5">1</mods:number>
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<mods:start id="C56F43E699106348C90BAC9D3E4487DD">79</mods:start>
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<mods:classification id="9DE31E6D6CB8492743372B0F60A85C48">journal article</mods:classification>
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<mods:identifier id="89CFE7F5D69710F2FF206D3C6B9681DF" type="CLB-Dataset">27556</mods:identifier>
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<mods:identifier id="B86B7930453EC60DDC35FF854E45D078" type="DOI">10.11646/zootaxa.4555.1.6</mods:identifier>
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<mods:identifier id="BE5D0F37674F66019124B6475883F151" type="GBIF-Dataset">6de65293-38cf-4b21-9a93-dacb22d1300d</mods:identifier>
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<mods:identifier id="5FA676D2085E7441BB3F10C8BDBFD9A0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6FC328F38362B638D9DFE72ABB9FDBFF" type="Zenodo-Dep">2624177</mods:identifier>
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<mods:identifier id="7AC05D1680BF83BAEB2322AA12C7C73D" type="ZooBank">02458E32-85F2-4624-911A-2DB5FFCED3D1</mods:identifier>
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</mods:mods>
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<treatment id="03D5FA59FFBB4F455F84FF78FF26FE21" ID-DOI="http://doi.org/10.5281/zenodo.5927633" ID-GBIF-Taxon="154533216" ID-Zenodo-Dep="5927633" LSID="urn:lsid:plazi:treatment:03D5FA59FFBB4F455F84FF78FF26FE21" httpUri="http://treatment.plazi.org/id/03D5FA59FFBB4F455F84FF78FF26FE21" lastPageId="4" lastPageNumber="83" pageId="2" pageNumber="81">
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<subSubSection id="C36618C4FFBB4F435F84FF78FE24FF43" box="[151,464,151,178]" pageId="2" pageNumber="81" type="nomenclature">
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<paragraph id="8BC34B4FFFBB4F435F84FF78FE24FF43" blockId="2.[151,464,151,211]" box="[151,464,151,178]" pageId="2" pageNumber="81">
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<heading id="D08BFC23FFBB4F435F84FF78FE24FF43" bold="true" box="[151,464,151,178]" fontSize="11" level="1" pageId="2" pageNumber="81" reason="1">
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<emphasis id="B908975DFFBB4F435F84FF78FE24FF43" bold="true" box="[151,464,151,178]" pageId="2" pageNumber="81">
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<taxonomicName id="4C7C30CCFFBB4F435F84FF78FEA6FF40" authority="Purkayastha & David, 2019" authorityName="Purkayastha & David" authorityYear="2019" box="[151,338,151,177]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="lacrima" status="sp. nov.">
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<emphasis id="B908975DFFBB4F435F84FF78FEA6FF40" bold="true" box="[151,338,151,177]" italics="true" pageId="2" pageNumber="81">Hebius lacrima</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23B2A26FFBB4F435E4AFF77FE24FF43" box="[345,464,152,178]" pageId="2" pageNumber="81" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36618C4FFBB4F435F84FF55FEDCFF22" box="[151,296,186,211]" pageId="2" pageNumber="81" type="description">
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<paragraph id="8BC34B4FFFBB4F435F84FF55FEDCFF22" blockId="2.[151,464,151,211]" box="[151,296,186,211]" pageId="2" pageNumber="81">
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||
(
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<figureCitation id="134757CAFFBB4F435F8CFF55FF18FF22" box="[159,236,186,211]" captionStart="FIGURE 1" captionStartId="3.[151,250,1126,1148]" captionTargetBox="[150,1436,193,1104]" captionTargetId="figure@3.[150,1436,193,1104]" captionTargetPageId="3" captionText="FIGURE 1. Preserved holotype of Hebius lacrima spec. nov., VR/ERS/ZSI/610, adult male. A: general view. B: close-up view of the dorsal pattern. C: left side of head. D: right side of head." httpUri="https://zenodo.org/record/2624179/files/figure.png" pageId="2" pageNumber="81">Figs. 1</figureCitation>
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&
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<figureCitation id="134757CAFFBB4F435E03FF54FEEBFF22" box="[272,287,187,211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1448,1470]" captionTargetBox="[150,1436,519,1426]" captionTargetId="figure@4.[150,1436,519,1426]" captionTargetPageId="4" captionText="FIGURE 2. Preserved holotype of Hebius lacrima spec. nov., VR/ERS/ZSI/610, adult male. A: Dorsal view of head. B: Ventral view of head. C: Ventral view." httpUri="https://zenodo.org/record/2624181/files/figure.png" pageId="2" pageNumber="81">2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36618C4FFBB4F435F84FEECFCB2FE92" pageId="2" pageNumber="81" type="materials_examined">
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<paragraph id="8BC34B4FFFBB4F435F84FEECFCB2FE92" blockId="2.[151,1436,258,1275]" pageId="2" pageNumber="81">
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<materialsCitation id="3B144112FFBB4F435F84FEECFCB2FE92" ID-GBIF-Occurrence="2234218747" collectingDate="2010-08-07" country="India" county="Jayaditya Purkayastha" latitude="27.980558" location="Basar" longLatPrecision="1" longitude="94.6885" municipality="State" pageId="2" pageNumber="81" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
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<emphasis id="B908975DFFBB4F435F84FEECFEF0FEEA" bold="true" box="[151,260,259,283]" pageId="2" pageNumber="81">
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<typeStatus id="54C7F5EDFFBB4F435F84FEECFEF0FEEA" box="[151,260,259,283]" pageId="2" pageNumber="81" type="holotype">Holotype</typeStatus>
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</emphasis>
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. VR/ERS/ZSI/610, adult male, from
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<location id="8EA31D94FFBB4F435DA2FEECFD05FEEB" LSID="urn:lsid:plazi:treatment:03D5FA59FFBB4F455F84FF78FF26FE21:8EA31D94FFBB4F435DA2FEECFD05FEEB" box="[689,753,259,282]" country="India" county="Jayaditya Purkayastha" latitude="27.980558" longLatPrecision="1" longitude="94.6885" municipality="State" name="Basar" pageId="2" pageNumber="81" stateProvince="Arunachal Pradesh">Basar</location>
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||
(
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<geoCoordinate id="EE482D88FFBB4F435C12FEEDFC68FEEA" box="[769,924,258,283]" direction="north" orientation="latitude" pageId="2" pageNumber="81" precision="1" value="27.980558">27.980559°N</geoCoordinate>
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,
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||
<geoCoordinate id="EE482D88FFBB4F435CB9FEEDFBB6FEEA" box="[938,1090,258,283]" direction="east" orientation="longitude" pageId="2" pageNumber="81" precision="1" value="94.6885">94.688496°E</geoCoordinate>
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),
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<location id="8EA31D94FFBB4F435B44FEECFAC3FEEA" LSID="urn:lsid:plazi:treatment:03D5FA59FFBB4F455F84FF78FF26FE21:8EA31D94FFBB4F435B44FEECFAC3FEEA" box="[1111,1335,258,283]" country="India" county="Jayaditya Purkayastha" latitude="27.980558" longLatPrecision="1" longitude="94.6885" municipality="State" name="West Siang District" pageId="2" pageNumber="81" stateProvince="Arunachal Pradesh">West Siang District</location>
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,
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<collectingMunicipality id="6BA7D135FFBB4F435A57FEECFA88FEEA" box="[1348,1404,259,283]" pageId="2" pageNumber="81">State</collectingMunicipality>
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of
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<collectingRegion id="49B885ADFFBB4F435F84FEC9FE9BFECE" box="[151,367,294,319]" country="India" name="Arunachal Pradesh" pageId="2" pageNumber="81">Arunachal Pradesh</collectingRegion>
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,
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<collectingCountry id="F36B0BDFFFBB4F435E6FFEC9FE4CFECE" box="[380,440,294,319]" name="India" pageId="2" pageNumber="81">India</collectingCountry>
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, at ca. 600 metres a.s.l. Collected by a villager, preserved in concentrated formalin and subsequently given to
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<collectingCounty id="62A233C3FFBB4F435E81FEA5FD7BFE92" box="[402,655,330,355]" pageId="2" pageNumber="81">Jayaditya Purkayastha</collectingCounty>
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,
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<date id="FFC26D8FFFBB4F435D89FEA4FCB7FE92" box="[666,835,331,355]" pageId="2" pageNumber="81" value="2010-08-07">
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<collectingDate id="EF869467FFBB4F435D89FEA4FCB7FE92" box="[666,835,331,355]" pageId="2" pageNumber="81" value="2010-08-07">7 August 2010</collectingDate>
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</date>
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||
.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36618C4FFBB4F435FD4FE81FA60FD0C" pageId="2" pageNumber="81" type="diagnosis">
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<paragraph id="8BC34B4FFFBB4F435FD4FE81FA60FD0C" blockId="2.[151,1436,258,1275]" pageId="2" pageNumber="81">
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<emphasis id="B908975DFFBB4F435FD4FE81FECEFE76" bold="true" box="[199,314,366,391]" pageId="2" pageNumber="81">Diagnosis</emphasis>
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. A species of the genus
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<taxonomicName id="4C7C30CCFFBB4F435D43FE81FD6BFE77" box="[592,671,366,390]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="81" phylum="Chordata" rank="genus">
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<emphasis id="B908975DFFBB4F435D43FE81FD6BFE77" box="[592,671,366,390]" italics="true" pageId="2" pageNumber="81">Hebius</emphasis>
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</taxonomicName>
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characterized by combination of the following characters: (1) body elongate; (2) tail amounting for 30.1 % of the total length; (3) 24 gradually enlarged maxillary teeth, followed, with a diastema, by 3 distinctly enlarged posterior teeth; (4) nostrils lateral; (5) internasals broad anteriorly; (6) 2 preoculars; (7) 1 anterior temporal, (8) 19–19–17 dorsal scale rows, distinctly keeled except scales of the 1
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<superScript id="7C09E607FFBB4F435A5FFE36FAA2FE16" attach="left" box="[1356,1366,473,487]" fontSize="6" pageId="2" pageNumber="81">st</superScript>
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rows, smooth but distinctly enlarged; (9) 147 ventrals, not keeled (+ 2 preventrals); (10) anal divided; (11) 89 subcaudals; (12) a white stripe extending on the middle of supralabials from the edge of rostral and 1
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<superScript id="7C09E607FFBB4F435B65FDCBFB74FDC3" attach="left" box="[1142,1152,548,562]" fontSize="6" pageId="2" pageNumber="81">st</superScript>
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SL to the anterior half of the 6
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<superScript id="7C09E607FFBB4F435FDDFDA4FF2EFDA8" attach="left" box="[206,218,587,601]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
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SL just below the eye; a second stripe, slightly higher on the labials and temporal regions than the first one, extends from the 7
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<superScript id="7C09E607FFBB4F435E7FFD9DFE8CFD71" attach="left" box="[364,376,626,640]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
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SL to the 9
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<superScript id="7C09E607FFBB4F435D13FD9DFDF8FD71" attach="left" box="[512,524,626,640]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
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SL then beyond the angle of jaw to the nape, producing a short V-like chevron; (13) a dark area on the posterior half of 6
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<superScript id="7C09E607FFBB4F435D7CFD76FD8FFD56" attach="left" box="[623,635,665,679]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
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SL, separating the two white stripes; (14) dorsum dark greyish-brown variegated with blackish-brown blotches, without stripe or aligned dorsolateral dots; (15) venter ivory with a large, elongate blotch parallel to the body axis near the tips of each ventral, forming a discontinuous ventrolateral stripe.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36618C4FFBB4F455FD4FCE7FE22FF24" lastPageId="4" lastPageNumber="83" pageId="2" pageNumber="81" type="description">
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<paragraph id="8BC34B4FFFBB4F435FD4FCE7FE5CFC40" blockId="2.[151,1436,258,1275]" pageId="2" pageNumber="81">
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<emphasis id="B908975DFFBB4F435FD4FCE7FE14FCD0" bold="true" box="[199,480,776,801]" pageId="2" pageNumber="81">
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Description of
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<typeStatus id="54C7F5EDFFBB4F435E6AFCE6FE14FCD0" box="[377,480,777,801]" pageId="2" pageNumber="81" type="holotype">holotype</typeStatus>
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</emphasis>
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. Body moderately stout; head average (3.6% of SVL), distinct from the neck; snout long, flattened, 34.7% of HL, 1.7 times as long as horizontal diameter of the eye, blunt from above, subrectangular in profile; nostril lateral, crescentic, piercing in the middle of divided nasal in its lower half; eye moderately sized, diameter 1.5 times greater than distance between its inferior margin and edge of upper lip; pupil round; tail cylindrical and tapering.
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</paragraph>
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<paragraph id="8BC34B4FFFBB4F435FD4FC52FB75FC24" blockId="2.[151,1436,258,1275]" box="[199,1153,956,981]" pageId="2" pageNumber="81">
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<emphasis id="B908975DFFBB4F435FD4FC52FF00FC25" box="[199,244,957,980]" italics="true" pageId="2" pageNumber="81">Size</emphasis>
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. SVL:
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<quantity id="4C84E6AAFFBB4F435E56FC52FE51FC25" box="[325,421,957,981]" metricMagnitude="-1" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="81" unit="mm" value="340.0">340 mm</quantity>
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; TaL:
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<quantity id="4C84E6AAFFBB4F435EFEFC52FDB9FC25" box="[493,589,957,981]" metricMagnitude="-1" metricUnit="m" metricValue="1.47" pageId="2" pageNumber="81" unit="mm" value="147.0">147 mm</quantity>
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; TL:
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<quantity id="4C84E6AAFFBB4F435D9AFC52FD1DFC25" box="[649,745,957,981]" metricMagnitude="-1" metricUnit="m" metricValue="4.87" pageId="2" pageNumber="81" unit="mm" value="487.0">487 mm</quantity>
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; HL:
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<quantity id="4C84E6AAFFBB4F435C3BFC52FC7BFC25" box="[808,911,957,981]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="2" pageNumber="81" unit="mm" value="12.4">12.4 mm</quantity>
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; ratio TaL/TL: 0.301.
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</paragraph>
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<paragraph id="8BC34B4FFFBB4F435FD4FC0EFDA4FBEC" blockId="2.[151,1436,258,1275]" pageId="2" pageNumber="81">
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<emphasis id="B908975DFFBB4F435FD4FC0EFEC5FC09" box="[199,305,993,1016]" italics="true" pageId="2" pageNumber="81">Dentition</emphasis>
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. A total of 27 maxillary teeth: 24 gradually enlarged + 3 distinctly enlarged posterior teeth, separated from anterior teeth by a short diastema.
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</paragraph>
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<paragraph id="8BC34B4FFFBB4F435FD4FBC6FD9EFB7D" blockId="2.[151,1436,258,1275]" pageId="2" pageNumber="81">
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<emphasis id="B908975DFFBB4F435FD4FBC6FECBFBB1" box="[199,319,1065,1088]" italics="true" pageId="2" pageNumber="81">Hemipenis</emphasis>
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. The organ was examined in situ. As it was in very bad condition and brittle because of the formalin used to preserve the specimen, we could observe only its general features. Hemipenis short, reaching the 5
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<superScript id="7C09E607FFBB4F435A83FBA3FA68FBAB" attach="none" box="[1424,1436,1100,1114]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
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subcaudal, simple and seemingly smooth.
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</paragraph>
|
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<paragraph id="8BC34B4FFFBB4F435FD4FB78FB45FB26" blockId="2.[151,1436,258,1275]" pageId="2" pageNumber="81">
|
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<emphasis id="B908975DFFBB4F435FD4FB78FE9AFB5E" box="[199,366,1175,1199]" italics="true" pageId="2" pageNumber="81">Body scalation</emphasis>
|
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. DSR: 19
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<emphasis id="B908975DFFBB4F435ECFFB77FE1EFB41" bold="true" box="[476,490,1176,1200]" pageId="2" pageNumber="81">–</emphasis>
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19
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<emphasis id="B908975DFFBB4F435D16FB77FDE7FB41" bold="true" box="[517,531,1176,1200]" pageId="2" pageNumber="81">–</emphasis>
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17, not notched at their posterior extremity, distinctly keeled with a narrow, sharp keel, more keeled on posterior half of body except scales of 1
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<superScript id="7C09E607FFBB4F435C5BFB55FCA6FB39" attach="left" box="[840,850,1210,1224]" fontSize="6" pageId="2" pageNumber="81">st</superScript>
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DSR, all smooth and enlarged.
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</paragraph>
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<paragraph id="8BC34B4FFFBB4F435FD4FB0DFE08FB0A" blockId="2.[151,1436,258,1275]" box="[199,508,1250,1275]" pageId="2" pageNumber="81">Dorsal scale row reduction:</paragraph>
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<paragraph id="8BC34B4FFFBB4F435FD4FAC1FE46FA67" blockId="2.[151,535,1321,1431]" pageId="2" pageNumber="81">
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4+5
|
||
<emphasis id="B908975DFFBB4F435FE9FAC6FEEDFAB9" bold="true" box="[250,281,1321,1352]" italics="true" pageId="2" pageNumber="81">Ą</emphasis>
|
||
4 (93) (left) 19 ———————————17 4+5
|
||
<emphasis id="B908975DFFBB4F435FE9FA97FEEDFA66" bold="true" box="[250,281,1400,1431]" italics="true" pageId="2" pageNumber="81">Ą</emphasis>
|
||
4 (97) (right)
|
||
</paragraph>
|
||
<paragraph id="8BC34B4FFFBB4F435FD4FA27FC91FA11" blockId="2.[151,1437,1479,2016]" box="[199,869,1479,1504]" pageId="2" pageNumber="81">
|
||
147
|
||
<collectingRegion id="49B885ADFFBB4F435FEBFA27FEC6FA2E" box="[248,306,1480,1503]" country="Austria" name="Wien" pageId="2" pageNumber="81">VEN</collectingRegion>
|
||
(+ 2 preventrals); 89 SC, all paired. Anal divided.
|
||
</paragraph>
|
||
<paragraph id="8BC34B4FFFBB4F435FD4FA04FF3BF9BD" blockId="2.[151,1437,1479,2016]" pageId="2" pageNumber="81">
|
||
Length of the portion of tail with 6 dorsal scale rows (expressed in number of subcaudals; see
|
||
<bibRefCitation id="EFED36BEFFBB4F435A33FA04FE95F9D9" author="Malnate, E. V. & Underwood, G." pageId="2" pageNumber="81" pagination="59 - 201" refId="ref6386" refString="Malnate, E. V. & Underwood, G. (1988) Australasian natricine snakes of the genus Tropidonophis. Proceedings of the Academy of Natural Sciences of Philadelphia, 140 (1), 59 - 201." type="journal article" year="1988">Malnate & Underwood 1988</bibRefCitation>
|
||
): 31; length of the portion of tail with 4 scale rows: 24. Ratio Length 6 rows / Length 4 rows: 1.29.
|
||
</paragraph>
|
||
<paragraph id="8BC34B4FFFBB4F435FD4F9B8FE97F811" blockId="2.[151,1437,1479,2016]" pageId="2" pageNumber="81">
|
||
<emphasis id="B908975DFFBB4F435FD4F9B8FE85F99E" box="[199,369,1623,1647]" italics="true" pageId="2" pageNumber="81">Head scalation</emphasis>
|
||
. Rostral hexagonal, wider than high; nasals subrectangular, longer than high, divided below the nostril, with crescentic, laterally opening nostril in its middle; internasals subtriangular and barely narrowing anteriorly, 1.1 times as long as wide and about 0.95 times as wide anteriorly than posteriorly; prefrontals rectangular, almost subequal, in contact with loreal; frontal hexagonal, large, 1.45 times as long as wide, with apex directed posteriorly, 1.8 times longer than suture between prefrontals; parietals twice as long as wide, in contact along a length 1.13 times as great as frontal length; 1/1 loreal, small, rectangular, elongate horizontally, 0.9 time as high as long, in broad contact with nasal; 2/2 preoculars; 3/3 postoculars, upper one much larger than the two lower ones; 9/10 supralabials, 1
|
||
<superScript id="7C09E607FFBB4F435EA7F8BDFE4AF891" attach="none" box="[436,446,1874,1888]" fontSize="6" pageId="2" pageNumber="81">st</superScript>
|
||
–3
|
||
<superScript id="7C09E607FFBB4F435EC8F8BDFE1CF891" attach="left" box="[475,488,1874,1888]" fontSize="6" pageId="2" pageNumber="81">rd</superScript>
|
||
in contact with nasal, 3
|
||
<superScript id="7C09E607FFBB4F435DE0F8BDFCF4F891" attach="none" box="[755,768,1874,1888]" fontSize="6" pageId="2" pageNumber="81">rd</superScript>
|
||
–4
|
||
<superScript id="7C09E607FFBB4F435C0FF8BDFCDCF891" attach="left" box="[796,808,1874,1888]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
|
||
in contact with loreal, 4
|
||
<superScript id="7C09E607FFBB4F435B29F8BDFBB2F891" attach="none" box="[1082,1094,1874,1888]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
|
||
–6
|
||
<superScript id="7C09E607FFBB4F435B71F8BDFB9AF891" attach="left" box="[1122,1134,1874,1888]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
|
||
entering orbit, 8
|
||
<superScript id="7C09E607FFBB4F435A39F8BDFAC2F891" attach="none" box="[1322,1334,1874,1888]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
|
||
–9
|
||
<superScript id="7C09E607FFBB4F435A42F8BDFAA9F891" attach="left" box="[1361,1373,1874,1888]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
|
||
/ 7
|
||
<superScript id="7C09E607FFBB4F435A91F8BDFA7AF891" attach="none" box="[1410,1422,1874,1888]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
|
||
– 8
|
||
<superScript id="7C09E607FFBB4F435FB6F896FF45F876" attach="left" box="[165,177,1913,1927]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
|
||
largest; temporals: 1+1 / 1+1, anterior one large and elongate; 9/10 infralabials, first pair in contact behind the mental, 1
|
||
<superScript id="7C09E607FFBB4F435FECF84FFEFDF85F" attach="none" box="[255,265,1952,1966]" fontSize="6" pageId="2" pageNumber="81">st</superScript>
|
||
–4
|
||
<superScript id="7C09E607FFBB4F435E36F84FFEC5F85F" attach="left" box="[293,305,1952,1966]" fontSize="6" pageId="2" pageNumber="81">th</superScript>
|
||
IL in contact with anterior chin shields; posterior chin shields longer than anterior ones, followed by one pair of gulars.
|
||
</paragraph>
|
||
<caption id="DF031BC7FFBA4F425F84FB89FCE7FB6B" httpUri="https://zenodo.org/record/2624179/files/figure.png" pageId="3" pageNumber="82" startId="3.[151,250,1126,1148]" targetBox="[150,1436,193,1104]" targetPageId="3">
|
||
<paragraph id="8BC34B4FFFBA4F425F84FB89FCE7FB6B" blockId="3.[151,1436,1126,1178]" pageId="3" pageNumber="82">
|
||
<emphasis id="B908975DFFBA4F425F84FB89FEE6FB8D" bold="true" box="[151,274,1126,1148]" pageId="3" pageNumber="82">FIGURE 1.</emphasis>
|
||
Preserved holotype of
|
||
<taxonomicName id="4C7C30CCFFBA4F425EE8FB89FD6CFB8D" authority="Purkayastha & David, 2019" authorityName="Purkayastha & David" authorityYear="2019" box="[507,664,1126,1148]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="82" phylum="Chordata" rank="species" species="lacrima" status="sp. nov.">
|
||
<emphasis id="B908975DFFBA4F425EE8FB89FD6CFB8D" box="[507,664,1126,1148]" italics="true" pageId="3" pageNumber="82">Hebius lacrima</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B908975DFFBA4F425D8EFB89FCF0FB8D" bold="true" box="[669,772,1126,1148]" pageId="3" pageNumber="82">
|
||
<taxonomicNameLabel id="A23B2A26FFBA4F425D8EFB89FCF0FB8D" box="[669,772,1126,1148]" pageId="3" pageNumber="82" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, VR/ERS/ZSI/610, adult male. A: general view. B: close-up view of the dorsal pattern. C: left side of head. D: right side of head.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC34B4FFFBA4F425FD4FB2DFE05FA66" blockId="3.[151,1437,1218,2024]" pageId="3" pageNumber="82">
|
||
<emphasis id="B908975DFFBA4F425FD4FB2DFE49FB2B" box="[199,445,1218,1242]" italics="true" pageId="3" pageNumber="82">Coloration in alcohol</emphasis>
|
||
. Body dark greyish-brown above, with scales irregularly marbled with blackish-brown and cream spots; on each side, three rows of irregular, faint dark blotches, not vertically aligned, usually on 3
|
||
<superScript id="7C09E607FFBA4F425A75FB0AFA87FB02" attach="none" box="[1382,1395,1253,1267]" fontSize="6" pageId="3" pageNumber="82">rd</superScript>
|
||
, 5
|
||
<superScript id="7C09E607FFBA4F425A83FB0AFA68FB02" attach="none" box="[1424,1436,1253,1267]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
and 9
|
||
<superScript id="7C09E607FFBA4F425FC5FAE3FF16FAEB" attach="left" box="[214,226,1292,1306]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
DSR; on each side of the anterior part of the body, a short dorsolateral row made of a total of six cream, elongate spots along a length equal to about 2.5 head lengths behind the nape region; scales of the first twelve scales of the 1
|
||
<superScript id="7C09E607FFBA4F425E29FAB9FEB0FA95" attach="left" box="[314,324,1366,1380]" fontSize="6" pageId="3" pageNumber="82">st</superScript>
|
||
DSR ivory, producing a short, conspicuous, ventrolateral stripe; the ivory area of scales of the 1
|
||
<superScript id="7C09E607FFBA4F425A81FAB9FA68FA95" attach="none" box="[1426,1436,1366,1380]" fontSize="6" pageId="3" pageNumber="82">st</superScript>
|
||
DSR decreases quickly in size.
|
||
</paragraph>
|
||
<paragraph id="8BC34B4FFFBA4F425FD4FA4DFC3EF8A0" blockId="3.[151,1437,1218,2024]" pageId="3" pageNumber="82">
|
||
Head dark brown above and on its sides, somewhat paler on the temporal region, with irregular paler brown vermiculation and some scattered beige dots; a short cream sagittal line just on the occipital region behind parietal suture; rostral brown; a broad but irregular, pure white stripe extends from the posterior edge of the rostral to the anterior half of 6
|
||
<superScript id="7C09E607FFBA4F425E70F9E2FE9BF9EA" attach="left" box="[355,367,1549,1563]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
SL across the lower part of nasal, the lower edge of loreal and the central part of 1
|
||
<superScript id="7C09E607FFBA4F425A79F9E2FA80F9EA" attach="right" box="[1386,1396,1549,1563]" fontSize="6" pageId="3" pageNumber="82">st</superScript>
|
||
–5
|
||
<superScript id="7C09E607FFBA4F425A83F9E2FA68F9EA" attach="none" box="[1424,1436,1549,1563]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
supralabials, covering about one third of the height of these supralabials with their upper and lower parts dark brown; posterior half of 6
|
||
<superScript id="7C09E607FFBA4F425EA9F9B7FE32F997" attach="left" box="[442,454,1624,1638]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
SL, just below the eye, dark brown, looking like a “tear” interrupting the white lateral stripe, the limit between the anterior white stripe and the dark area being curved towards the snout; a second pure white stripe, slightly higher on the head side than the anterior one, extends from the upper half of 7
|
||
<superScript id="7C09E607FFBA4F425BE2F94CFB09F940" attach="left" box="[1265,1277,1699,1713]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
SL across the central part of 8
|
||
<superScript id="7C09E607FFBA4F425E59F925FEA2F929" attach="left" box="[330,342,1738,1752]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
and 9
|
||
<superScript id="7C09E607FFBA4F425E88F925FE53F929" attach="left" box="[411,423,1738,1752]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
SL up to the corner of the mouth then farther on the nape, thick along a length of three or four scales then narrow, directed obliquely upwards, the branch of each side producing a conspicuous V-like chevron on the neck; eye black. The chin and throat are ivory; anterior chin shields marbled with irregular brown areas; mental and infralabials heavily marked with blackish-brown spots.
|
||
</paragraph>
|
||
<paragraph id="8BC34B4FFFBA4F455FD4F8B2FE22FF24" blockId="3.[151,1437,1218,2024]" lastBlockId="4.[151,1436,151,464]" lastPageId="4" lastPageNumber="83" pageId="3" pageNumber="82">
|
||
The venter is ivory; at the base of the triangular tip of each ventral a conspicuous, elongate blackish-brown blotch forming a discontinuous dark ventrolateral stripe that extends along the whole of the body; the ivory tip of each ventral contrasts with both the inner dark brown blotch of each ventral and the dark color of scales of the 1
|
||
<superScript id="7C09E607FFBA4F425A81F84BFA68F843" attach="none" box="[1426,1436,1956,1970]" fontSize="6" pageId="3" pageNumber="82">st</superScript>
|
||
DSR posterior to the 12
|
||
<superScript id="7C09E607FFBA4F425EB7F824FE44F828" attach="left" box="[420,432,1995,2009]" fontSize="6" pageId="3" pageNumber="82">th</superScript>
|
||
ventrals, producing a pale ventrolateral stripe. Under surface of tail ivory, with the outer part and tip of each subcaudal blackish-brown, producing a dark ventral stripe not separated from the dark color of the dorsal surface of the tail.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36618C4FFBD4F455FD4FF0FFF26FE21" pageId="4" pageNumber="83" type="etymology">
|
||
<paragraph id="8BC34B4FFFBD4F455FD4FF0FFA68FEB1" blockId="4.[151,1436,151,464]" pageId="4" pageNumber="83">
|
||
<emphasis id="B908975DFFBD4F455FD4FF0FFEB2FF09" bold="true" box="[199,326,224,248]" pageId="4" pageNumber="83">Etymology</emphasis>
|
||
. The species nomen derives from the Latin noun
|
||
<taxonomicName id="4C7C30CCFFBD4F455C7DFF0FFC31FF09" authorityName="Purkayastha & David" authorityYear="2019" box="[878,965,224,248]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="83" phylum="Chordata" rank="species" species="lacrima">
|
||
<emphasis id="B908975DFFBD4F455C7DFF0FFC31FF09" box="[878,965,224,248]" italics="true" pageId="4" pageNumber="83">lacrima</emphasis>
|
||
</taxonomicName>
|
||
(-
|
||
<emphasis id="B908975DFFBD4F455CCCFF0EFC0DFF09" box="[991,1017,225,248]" italics="true" pageId="4" pageNumber="83">ae</emphasis>
|
||
), meaning “a tear”, a reference to the dark area under the eye looking like a black tear which interrupts the white supralabial stripe. This species nomen is a noun in apposition and not an adjective.We suggest the following common names: Crying Keelback (English).
|
||
</paragraph>
|
||
<paragraph id="8BC34B4FFFBD4F455FD4FEA4FF26FE21" blockId="4.[151,1436,151,464]" pageId="4" pageNumber="83">
|
||
<emphasis id="B908975DFFBD4F455FD4FEA4FE73FE95" bold="true" box="[199,391,331,356]" pageId="4" pageNumber="83">Ecological notes</emphasis>
|
||
. The single known specimen was obtained from a rice field alongside a hill slope (
|
||
<figureCitation id="134757CAFFBD4F455A21FEA3FA81FE95" box="[1330,1397,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1750,1772]" captionTargetBox="[151,1435,1006,1728]" captionTargetId="figure@5.[151,1435,813,1729]" captionTargetPageId="5" captionText="FIGURE 3. Habitat of the holotype of Hebius lacrima spec. nov., West Siang District, State of Arunachal Pradesh, India, at ca. 600 metres a.s.l." httpUri="https://zenodo.org/record/2624185/files/figure.png" pageId="4" pageNumber="83">Fig. 3</figureCitation>
|
||
) in the outskirt of the city of Basar (
|
||
<figureCitation id="134757CAFFBD4F455D11FE80FDB3FE79" box="[514,583,367,392]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1476]" captionTargetBox="[178,1352,572,1377]" captionTargetId="figure@7.[150,1436,558,1432]" captionTargetPageId="7" captionText="FIGURE 4. A map of the state of Arunachal Pradesh showing the locality (Basar) of Hebius lacrima spec. nov. (solid red circle); Inset: a map of India pointing to (solid red circle) Arunachal Pradesh." httpUri="https://zenodo.org/record/2624187/files/figure.png" pageId="4" pageNumber="83">Fig. 4</figureCitation>
|
||
), so in a heavily disturbed area. A small stream was flowing adjacent to this field. An indigenous agricultural practice called Jhum (shifting) cultivation was done in the hills adjacent to the rice field.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |