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<document id="69E3227BB674B91CFE26A5A09C266CBE" ID-DOI="10.5281/zenodo.5164350" ID-ISSN="1942-1354" ID-Zenodo-Dep="5164350" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628185324501" checkinUser="felipe" docAuthor="Gillett, Conrad P. D. T., Gillett, P. T., E. D. T, James &amp; Vaz-de-Mello, Fernando Z." docDate="2010" docId="B8512924EA7F8513FF2A78990E94CD57" docLanguage="en" docName="InsectaMundi.0118.1-19.pdf" docOrigin="Insecta Mundi 2010 (118)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Coprophanaeus (Coprophanaeus) acrisius" docType="treatment" docVersion="4" lastPageNumber="8" masterDocId="4468515CEA76851AFFEA785C0A38C90C" masterDocTitle="Diversity and distribution of the scarab beetle tribe Phanaeini in the northern states of the Brazilian Northeast (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="8" updateTime="1699035250239" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="0ED801D02BE6161ED5A9F20D1492A89A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9B75613D5D8235D8AD36983A2F6D259">
<mods:title id="3B1610344165D9AEB5665C95E47D2498">Diversity and distribution of the scarab beetle tribe Phanaeini in the northern states of the Brazilian Northeast (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="826462D1D601F77C17D486CAF2F4BBE3" type="personal">
<mods:role id="4F0C5DA97064194170452EBCD663EA65">
<mods:roleTerm id="26E19D04DF80B66AC4B14C31D0096C5D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C33C4C3ABAE2955944EF496930427146">Gillett, Conrad P. D. T.</mods:namePart>
</mods:name>
<mods:name id="3EA83A595AAFD09A91C3504D29ADCA27" type="personal">
<mods:role id="5F467BEBFACF777E7F42F141D8C70792">
<mods:roleTerm id="15A55DAE9F6684B44CC0485EAB04B1CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6AB5D9EF69FB489E25240EF312C8F202">Gillett, P. T.</mods:namePart>
</mods:name>
<mods:name id="FED3890CBF34CB7164232FF2F45EFA7E" type="personal">
<mods:role id="85DAFEA99FD9B08AF23B64080FCBA73F">
<mods:roleTerm id="0420E0E435DC7FD1A3DF3D28089BF383">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C950049FA25C3C8D2D0DB427477622A">E. D. T, James</mods:namePart>
</mods:name>
<mods:name id="B33022976E61AA8D71E40E1214A160E7" type="personal">
<mods:role id="366FE8FB94125A4B592747D6194714F3">
<mods:roleTerm id="5A7D1726FCD55CB17CD9906C24D924AE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DE74477F2E7CB0B72D1E7C0AB37E891E">Vaz-de-Mello, Fernando Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="42B822511AA416802BBBFEAEF1A4E571">text</mods:typeOfResource>
<mods:relatedItem id="E87FF010BDF6921B115BEFC898684F1B" type="host">
<mods:titleInfo id="21AFC4E0F05CDD8257A0CE863EC2D7D2">
<mods:title id="7316CB0BB668A0AA0929AC578582A2E0">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="B7617458CBBFE9E9311DB30C579E0BFA">
<mods:date id="DF17E89788237F4790F25637FAED77E5">2010</mods:date>
<mods:detail id="8E59263CC905EEC510B0DBA1128D1DBB" type="pubDate">
<mods:number id="46A9460302C966B61DD37750065F480D">2010-03-12</mods:number>
</mods:detail>
<mods:detail id="4A79BF2C9CAD3B9F3E8E501C36C3B85D" type="volume">
<mods:number id="73DEFB8A58DB5CB7C304808A29EA0248">2010</mods:number>
</mods:detail>
<mods:detail id="05330B70EDFD6EAE955E524F628583A5" type="issue">
<mods:number id="5557A61C06E8B574FC57D9A4613B8303">118</mods:number>
</mods:detail>
<mods:extent id="086BB5BE9DE24F8DEEF8FED66D32AC29" unit="page">
<mods:start id="70A10845435B35C255602B03324A7F2A">1</mods:start>
<mods:end id="5233F9F5FD162CE780DCF8B31E5B41E3">19</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3843F8273F37897859BCEF0D0921A46E">journal article</mods:classification>
<mods:identifier id="77EF604D829DFE52BD00614C68D81C6F" type="DOI">10.5281/zenodo.5164350</mods:identifier>
<mods:identifier id="0004D5E175A80D48831674D28A811589" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="0CFB29B0763EC6184D112A09629DB3CF" type="Zenodo-Dep">5164350</mods:identifier>
</mods:mods>
<treatment id="B8512924EA7F8513FF2A78990E94CD57" ID-DOI="http://doi.org/10.5281/zenodo.5167723" ID-Zenodo-Dep="5167723" LSID="urn:lsid:plazi:treatment:B8512924EA7F8513FF2A78990E94CD57" httpUri="http://treatment.plazi.org/id/B8512924EA7F8513FF2A78990E94CD57" lastPageNumber="8" pageId="9" pageNumber="8">
<subSubSection id="78E2CBB9EA7F8513FF2A789909F0C9D7" box="[192,968,195,220]" pageId="9" pageNumber="8" type="nomenclature">
<paragraph id="30479832EA7F8513FF2A789909F0C9D7" blockId="9.[192,968,195,251]" box="[192,968,195,220]" pageId="9" pageNumber="8">
<heading id="6B0F2F5EEA7F8513FF2A789909F0C9D7" bold="true" box="[192,968,195,220]" fontSize="10" level="2" pageId="9" pageNumber="8" reason="3">
<taxonomicName id="F7F8E3B1EA7F8513FF2A789909F0C9D7" ID-CoL="7ZMNX" authority="(Macleay, 1819)" baseAuthorityName="Macleay" baseAuthorityYear="1819" box="[192,968,195,220]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="acrisius" subGenus="Coprophanaeus">
<emphasis id="028C4420EA7F8513FF2A789909F0C9D7" bold="true" box="[192,968,195,220]" pageId="9" pageNumber="8">
<emphasis id="028C4420EA7F8513FF2A78990BAEC9D0" bold="true" box="[192,406,197,220]" italics="true" pageId="9" pageNumber="8">Coprophanaeus</emphasis>
(
<emphasis id="028C4420EA7F8513FE4E78990842C9D0" bold="true" box="[420,634,197,220]" italics="true" pageId="9" pageNumber="8">Coprophanaeus</emphasis>
)
<emphasis id="028C4420EA7F8513FD63789908CEC9D0" bold="true" box="[649,758,197,220]" italics="true" pageId="9" pageNumber="8">acrisius</emphasis>
(
<bibRefCitation id="5469E5C3EA7F8513FCE9789F0987C9D7" author="Macleay, W. S." box="[771,959,195,219]" pageId="9" pageNumber="8" refId="ref13883" refString="Macleay, W. S. 1819. Horae entomologicae: or essays on the annulose animals. Vol. 1, Pt. 1. Bagster; London. 160 p." type="book" year="1819">Macleay, 1819</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="78E2CBB9EA7F8513FF2A78BF0B78CB97" pageId="9" pageNumber="8" type="description">
<paragraph id="30479832EA7F8513FF2A78BF0B1EC9F7" blockId="9.[192,968,195,251]" box="[192,294,227,251]" pageId="9" pageNumber="8">
<figureCitation id="A8C384B7EA7F8513FF2A78BF0B1EC9F7" box="[192,294,227,251]" captionStart="Figure 7-13" captionStartId="8.[192,276,1091,1113]" captionTargetBox="[197,1441,202,1066]" captionTargetId="figure-282@8.[193,1444,193,1084]" captionTargetPageId="8" captionText="Figure 7-13. Habitus images. 7) Coprophanaeus cyanescens (Olsoufieff, 1924) male specimen from Maranguape, São Benedito, Sanctuario da Nossa Senhora de Penha (CE). 8) Coprophanaeus acrisius (Macleay, 1819) male specimen from Santa Quiteria de Maranhão, Fazenda Rodiador (MA). 9) Coprophanaeus dardanus (Macleay, 1819) male specimen from Baturité, Uirapuru (CE). 10) Coprophanaeus (Metallophanaeus) pertyi (Olsoufieff, 1924) male specimen from Maranguape, Tabatinga (CE). 11) Diabroctis mimas (Linnaeus, 1767) male specimen from Maranguape, Piraponga (CE). 12-13) Phanaeus melibaeus Blanchard, 1843 male specimen from Santa Quiteria de Maranhão, Fazenda Rodiador (MA). Photographs by M.P.T. Gillett." figureDoi="http://doi.org/10.5281/zenodo.5164356" httpUri="https://zenodo.org/record/5164356/files/figure.png" pageId="9" pageNumber="8">Figure 8</figureCitation>
</paragraph>
<paragraph id="30479832EA7F8513FF2A797F0840C857" blockId="9.[192,1441,290,539]" pageId="9" pageNumber="8">
<emphasis id="028C4420EA7F8513FF2A797F0BA4C837" bold="true" box="[192,412,291,315]" pageId="9" pageNumber="8">Collection data.</emphasis>
<materialsCitation id="8090926FEA7F8513FE49797E0E60C837" box="[419,1112,290,315]" collectingDate="2001-06-25" collectionCode="PICI, FVMC" location="Fortaleza" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Ceara">
<emphasis id="028C4420EA7F8513FE49797E0BC5C836" bold="true" box="[419,509,290,314]" pageId="9" pageNumber="8">
<collectingRegion id="F23C56D0EA7F8513FE49797E0BC0C836" box="[419,504,290,314]" country="Brazil" name="Ceara" pageId="9" pageNumber="8">Ceará</collectingRegion>
:
</emphasis>
<location id="3527CEE9EA7F8513FDEE797F0846C837" LSID="urn:lsid:plazi:treatment:B8512924EA7F8513FF2A78990E94CD57:3527CEE9EA7F8513FDEE797F0846C837" box="[516,638,291,315]" name="Fortaleza" pageId="9" pageNumber="8" stateProvince="Ceara">Fortaleza</location>
, campus do
<collectionCode id="56E900F7EA7F8513FCF9797F0969C837" box="[787,849,291,315]" pageId="9" pageNumber="8">PICI</collectionCode>
,
<date id="4446BEF2EA7F8513FCB6797F09D2C837" box="[860,1002,291,315]" pageId="9" pageNumber="8" value="2001-06-25">
<collectingDate id="5402471AEA7F8513FCB6797F09D2C837" box="[860,1002,291,315]" pageId="9" pageNumber="8" value="2001-06-25">25. vi. 2001</collectingDate>
</date>
(
<collectionCode id="56E900F7EA7F8513FC13797F0E68C837" box="[1017,1104,291,315]" pageId="9" pageNumber="8">FVMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8090926FEA7F8513FB89797F0849C857" collectingDate="2000-09-16" collectionCode="FVMC, NEW, STATE, RECORDS" location="Pindoretama" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Ceara">
<location id="3527CEE9EA7F8513FB89797F0F3FC837" LSID="urn:lsid:plazi:treatment:B8512924EA7F8513FF2A78990E94CD57:3527CEE9EA7F8513FB89797F0F3FC837" box="[1123,1287,291,315]" name="Pindoretama" pageId="9" pageNumber="8" stateProvince="Ceara">Pindoretama</location>
,
<date id="4446BEF2EA7F8513FAF9797F0F98C837" box="[1299,1440,291,315]" pageId="9" pageNumber="8" value="2000-09-16">
<collectingDate id="5402471AEA7F8513FAF9797F0F98C837" box="[1299,1440,291,315]" pageId="9" pageNumber="8" value="2000-09-16">16. ix. 2000</collectingDate>
</date>
(
<collectionCode id="56E900F7EA7F8513FF22791F0B27C857" box="[200,287,323,347]" pageId="9" pageNumber="8">FVMC</collectionCode>
).
<emphasis id="028C4420EA7F8513FEC5791F0849C857" bold="true" box="[303,625,323,347]" pageId="9" pageNumber="8">
<collectionCode id="56E900F7EA7F8513FEC5791F0B4BC857" box="[303,371,323,347]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="9" pageNumber="8">NEW</collectionCode>
<collectionCode id="56E900F7EA7F8513FE93791F0849C857" box="[377,625,323,347]" pageId="9" pageNumber="8">STATE RECORDS</collectionCode>
</emphasis>
</materialsCitation>
.
</paragraph>
<paragraph id="30479832EA7F8513FF1A793F0F58C8D7" blockId="9.[192,1441,290,539]" pageId="9" pageNumber="8">
<materialsCitation id="8090926FEA7F8513FF1A793F0EB9C877" box="[240,1153,354,379]" location="Fazenda Rodiador" longitude="-42.679447" municipality="Santa Quiteria" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Maranhao">
<emphasis id="028C4420EA7F8513FF1A793F0BBFC877" bold="true" box="[240,391,355,379]" pageId="9" pageNumber="8">
<collectingRegion id="F23C56D0EA7F8513FF1A793F0BB9C877" box="[240,385,355,379]" country="Brazil" name="Maranhao" pageId="9" pageNumber="8">Maranhão</collectingRegion>
:
</emphasis>
<collectingMunicipality id="D0230248EA7F8513FE67793F087FC877" box="[397,583,355,379]" pageId="9" pageNumber="8">Santa Quiteria</collectingMunicipality>
de
<collectingRegion id="F23C56D0EA7F8513FD98793F08CDC877" box="[626,757,355,379]" country="Brazil" name="Maranhao" pageId="9" pageNumber="8">Maranhão</collectingRegion>
,
<location id="3527CEE9EA7F8513FCEB793F09DFC877" LSID="urn:lsid:plazi:treatment:B8512924EA7F8513FF2A78990E94CD57:3527CEE9EA7F8513FCEB793F09DFC877" box="[769,999,355,379]" longitude="-42.679447" municipality="Santa Quiteria" name="Fazenda Rodiador" pageId="9" pageNumber="8" stateProvince="Maranhao">Fazenda Rodiador</location>
,
<geoCoordinate id="55CCFEF5EA7F8513FC18793F0EB9C877" box="[1010,1153,354,379]" degrees="42" direction="west" minutes="40" orientation="longitude" pageId="9" pageNumber="8" precision="15" seconds="46" value="-42.679447">
42
<superScript id="C78D357AEA7F8513FBFA793E0E20C87C" attach="none" box="[1040,1048,354,368]" fontSize="6" pageId="9" pageNumber="8">o</superScript>
40’46&quot;W
</geoCoordinate>
</materialsCitation>
;
<geoCoordinate id="55CCFEF5EA7F8513FB66793F0F2AC877" box="[1164,1298,354,379]" degrees="03" direction="south" minutes="24" orientation="latitude" pageId="9" pageNumber="8" precision="15" seconds="50" value="-3.413889">
03
<superScript id="C78D357AEA7F8513FB40793E0E8AC87C" attach="none" box="[1194,1202,354,368]" fontSize="6" pageId="9" pageNumber="8">o</superScript>
24’50&quot;S
</geoCoordinate>
,
<quantity id="F70035D7EA7F8513FAF4793F0F63C877" box="[1310,1371,355,379]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="9" pageNumber="8" unit="m" value="47.0">47 m</quantity>
palm forest, carrion traps,
<date id="4446BEF2EA7F8513FE2F79DF0869C897" box="[453,593,387,411]" pageId="9" pageNumber="8" value="2008-02-23">23. ii. 2008</date>
:
<quantity id="F70035D7EA7F8513FDB679DF08B3C897" box="[604,651,387,411]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="m" value="1.0">1 m</quantity>
;
<materialsCitation id="8090926FEA7F8513FD7279DF0F77C897" box="[664,1359,387,411]" collectingDate="2008-05-18" elevation="120" location="Fazenda Rodiador" longitude="-42.679447" municipality="Santa Quiteria" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Maranhao">
idem,
<quantity id="F70035D7EA7F8513FD0979DF0917C897" box="[739,815,387,411]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="8" unit="m" value="120.0">
<elevation id="BBD57F01EA7F8513FD0979DF0917C897" box="[739,815,387,411]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="8" unit="m" value="120.0">120 m</elevation>
</quantity>
chapada,
<collectingMethod id="E9B9E025EA7F8513FC4779DF0E69C897" box="[941,1105,387,411]" pageId="9" pageNumber="8">carrion traps</collectingMethod>
,
<date id="4446BEF2EA7F8513FBB779DF0EDEC897" box="[1117,1254,387,411]" pageId="9" pageNumber="8" value="2008-05-18">
<collectingDate id="5402471AEA7F8513FBB779DF0EDEC897" box="[1117,1254,387,411]" pageId="9" pageNumber="8" value="2008-05-18">18. v. 2008</collectingDate>
</date>
:
<quantity id="F70035D7EA7F8513FB1A79DF0F2FC897" box="[1264,1303,387,411]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="m" value="1.0">
<elevation id="BBD57F01EA7F8513FB1A79DF0F2FC897" box="[1264,1303,387,411]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="m" value="1.0">1m</elevation>
</quantity>
+ 1f
</materialsCitation>
;
<materialsCitation id="8090926FEA7F8513FAB179DF081EC8B7" collectingDate="2008-02-24" elevation="1" location="Fazenda Rodiador" longitude="-42.679447" municipality="Santa Quiteria" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Maranhao">
idem,
<collectingMethod id="E9B9E025EA7F8513FF2A79FF0B67C8B7" box="[192,351,419,443]" pageId="9" pageNumber="8">carrion traps</collectingMethod>
,
<date id="4446BEF2EA7F8513FE8379FF0BD6C8B7" box="[361,494,419,443]" pageId="9" pageNumber="8" value="2008-02-24">
<collectingDate id="5402471AEA7F8513FE8379FF0BD6C8B7" box="[361,494,419,443]" pageId="9" pageNumber="8" value="2008-02-24">24. ii. 2008</collectingDate>
</date>
:
<quantity id="F70035D7EA7F8513FE1279FF081EC8B7" box="[504,550,419,443]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="m" value="1.0">
<elevation id="BBD57F01EA7F8513FE1279FF081EC8B7" box="[504,550,419,443]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="m" value="1.0">1 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="8090926FEA7F8513FDDA79FF0926C8B7" box="[560,798,419,443]" collectingDate="2008-05-17" elevation="1" location="Fazenda Rodiador" longitude="-42.679447" municipality="Santa Quiteria" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Maranhao">
idem,
<date id="4446BEF2EA7F8513FD9379FF08C3C8B7" box="[633,763,419,443]" pageId="9" pageNumber="8" value="2008-05-17">
<collectingDate id="5402471AEA7F8513FD9379FF08C3C8B7" box="[633,763,419,443]" pageId="9" pageNumber="8" value="2008-05-17">17. v. 2008</collectingDate>
</date>
: 1f
</materialsCitation>
;
<materialsCitation id="8090926FEA7F8513FCC379FF0B93C8D7" collectingDate="2010-01-06" collectingDateMax="2010-01-07" collectingDateMin="2010-01-06" elevation="5" location="Fazenda Rodiador" longitude="-42.679447" municipality="Santa Quiteria" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Maranhao">
idem, open cerrado area, sandy soil,
<collectingMethod id="E9B9E025EA7F8513FB3279FF0F4DC8B7" box="[1240,1397,419,443]" pageId="9" pageNumber="8">carrion traps</collectingMethod>
<date id="4446BEF2EA7F8513FA9079FF0B7AC8D7" pageId="9" pageNumber="8" value="2010-01-06" valueMax="2010-01-07" valueMin="2010-01-06">
<collectingDate id="5402471AEA7F8513FA9079FF0B7AC8D7" pageId="9" pageNumber="8" value="2010-01-06" valueMax="2010-01-07" valueMin="2010-01-06">06- 07. i. 2010</collectingDate>
</date>
:
<quantity id="F70035D7EA7F8513FEA6799F0B4BC8D7" box="[332,371,451,475]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="8" unit="m" value="5.0">
<elevation id="BBD57F01EA7F8513FEA6799F0B4BC8D7" box="[332,371,451,475]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="8" unit="m" value="5.0">5m</elevation>
</quantity>
+ 2f
</materialsCitation>
;
<materialsCitation id="8090926FEA7F8513FE5C799F0889C8D7" box="[438,689,451,475]" collectingDate="2010-01-06" collectingDateMax="2010-01-07" collectingDateMin="2010-01-06" elevation="5" location="Fazenda Rodiador" longitude="-42.679447" municipality="Santa Quiteria" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Maranhao">
idem,
<collectingMethod id="E9B9E025EA7F8513FDE8799F08B4C8D7" box="[514,652,451,475]" pageId="9" pageNumber="8">dung traps</collectingMethod>
: 2f
</materialsCitation>
;
<materialsCitation id="8090926FEA7F8513FD56799F0F58C8D7" box="[700,1376,451,475]" collectingDate="2010-01-06" collectingDateMax="2010-01-07" collectingDateMin="2010-01-06" elevation="2" location="Fazenda Rodiador" longitude="-42.679447" municipality="Santa Quiteria" pageId="9" pageNumber="8" specimenCount="2" stateProvince="Maranhao">
idem, palm forest,
<collectingMethod id="E9B9E025EA7F8513FC48799F0E7AC8D7" box="[930,1090,451,475]" pageId="9" pageNumber="8">carrion traps</collectingMethod>
<date id="4446BEF2EA7F8513FBA3799F0ECBC8D7" box="[1097,1267,451,475]" pageId="9" pageNumber="8" value="2010-01-06" valueMax="2010-01-07" valueMin="2010-01-06">
<collectingDate id="5402471AEA7F8513FBA3799F0ECBC8D7" box="[1097,1267,451,475]" pageId="9" pageNumber="8" value="2010-01-06" valueMax="2010-01-07" valueMin="2010-01-06">06-07. i. 2010</collectingDate>
</date>
:
<quantity id="F70035D7EA7F8513FB17799F0F1BC8D7" box="[1277,1315,451,475]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="8" unit="m" value="2.0">
<elevation id="BBD57F01EA7F8513FB17799F0F1BC8D7" box="[1277,1315,451,475]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="8" unit="m" value="2.0">2m</elevation>
</quantity>
+
<specimenCount id="26FE53BBEA7F8513FAAB799F0F58C8D7" box="[1345,1376,451,475]" pageId="9" pageNumber="8" type="generic">2f.</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="30479832EA7F8513FF1A79BF0B7ECB17" blockId="9.[192,1441,290,539]" pageId="9" pageNumber="8">
<emphasis id="028C4420EA7F8513FF1A79BF0B9BC8F7" bold="true" box="[240,419,483,507]" pageId="9" pageNumber="8">
<collectingRegion id="F23C56D0EA7F8513FF1A79BF0BA6C8F7" box="[240,414,483,507]" country="Brazil" name="Pernambuco" pageId="9" pageNumber="8">Pernambuco</collectingRegion>
:
</emphasis>
Recife, campus of Universidade Federal de
<collectingRegion id="F23C56D0EA7F8513FC7479BF0E0FC8F7" box="[926,1079,483,507]" country="Brazil" name="Pernambuco" pageId="9" pageNumber="8">Pernambuco</collectingRegion>
, no date:
<quantity id="F70035D7EA7F8513FB4B79BF0EFFC8F7" box="[1185,1223,483,507]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="m" value="1.0">1m</quantity>
+
<specimenCount id="26FE53BBEA7F8513FB3379BF0ECDC8F7" box="[1241,1269,483,507]" pageId="9" pageNumber="8" type="generic">1f.</specimenCount>
<emphasis id="028C4420EA7F8513FB1379BF0B7ECB17" bold="true" pageId="9" pageNumber="8">NEW STATE RECORD.</emphasis>
</paragraph>
<paragraph id="30479832EA7F8513FF2A7A1F0B78CB97" blockId="9.[192,1440,579,667]" pageId="9" pageNumber="8">
<emphasis id="028C4420EA7F8513FF2A7A1F0B17CB57" bold="true" box="[192,303,579,603]" pageId="9" pageNumber="8">Results.</emphasis>
2008 data:
<specimenCount id="26FE53BBEA7F8513FE547A1F086ECB57" box="[446,598,579,603]" pageId="9" pageNumber="8" type="generic">5 specimens</specimenCount>
;
<specimenCount id="26FE53BBEA7F8513FD887A1F08F8CB57" box="[610,704,579,603]" pageId="9" pageNumber="8" type="male">3 males</specimenCount>
and
<specimenCount id="26FE53BBEA7F8513FD167A1F094DCB57" box="[764,885,579,603]" pageId="9" pageNumber="8" type="female">2 females</specimenCount>
. Male: Female sex ratio 1.50. Bait preference (2008 data): Carrion (5). Temporal distribution (all data): January (13), February (2), May (3), July, September
</paragraph>
</subSubSection>
<subSubSection id="78E2CBB9EA7F8513FF2A7A9F09C6CA17" pageId="9" pageNumber="8" type="distribution">
<paragraph id="30479832EA7F8513FF2A7A9F09C6CA17" blockId="9.[192,1440,707,795]" pageId="9" pageNumber="8">
<emphasis id="028C4420EA7F8513FF2A7A9F0B4ACBD7" bold="true" box="[192,370,707,731]" pageId="9" pageNumber="8">Distribution.</emphasis>
This species has been recorded from
<collectingCountry id="48EFD8A2EA7F8513FCC07A9F094DCBD7" box="[810,885,707,731]" name="Brazil" pageId="9" pageNumber="8">Brazil</collectingCountry>
(Pará),
<collectingCountry id="48EFD8A2EA7F8513FC397A9F0E11CBD7" box="[979,1065,707,731]" name="Bolivia" pageId="9" pageNumber="8">Bolivia</collectingCountry>
(
<collectingRegion id="F23C56D0EA7F8513FBDD7A9F0EF9CBD7" box="[1079,1217,707,731]" country="Bolivia" name="Santa Cruz" pageId="9" pageNumber="8">Santa Cruz</collectingRegion>
) and
<collectingCountry id="48EFD8A2EA7F8513FAE87A9F0F46CBD7" box="[1282,1406,707,731]" name="Argentina" pageId="9" pageNumber="8">Argentina</collectingCountry>
by Arnaud (2002). The species was first recorded from MA by
<bibRefCitation id="5469E5C3EA7F8513FC7E7ABF0EA6CBF7" author="Sampaio, D. V. &amp; C. Arzabe &amp; A. A. J. F. Castro &amp; F. Z. Vaz-de-Mello" box="[916,1182,739,763]" pageId="9" pageNumber="8" pagination="1 - 4" refId="ref14410" refString="Sampaio, D. V., C. Arzabe, A. A. J. F. Castro, and F. Z. Vaz-de-Mello. 2009. Consideracoes sobre a fauna de besouros rola-bosta (Coleoptera: Scarabaeidae: Scarabaeinae) em duas areas de cerrado litoraneo no Maranhao, Brasil. Anais do IX Congresso de Ecologia do Brasil, 13 a 17 de Setembro de 2009, Sao Lourenco - MG: 1 - 4." type="book chapter" year="2009">Sampaio et al. (2009)</bibRefCitation>
. It is associated with cerrado habitats in
<collectingCountry id="48EFD8A2EA7F8513FE467B5F0BC2CA17" box="[428,506,771,795]" name="Brazil" pageId="9" pageNumber="8">Brazil</collectingCountry>
, occurring in both forested and open areas.
</paragraph>
</subSubSection>
<subSubSection id="78E2CBB9EA7F8513FF2A7B1F0E94CD57" pageId="9" pageNumber="8" type="discussion">
<paragraph id="30479832EA7F8513FF2A7B1F0E94CD57" blockId="9.[192,1440,835,1116]" pageId="9" pageNumber="8">
<emphasis id="028C4420EA7F8513FF2A7B1F0B7CCA57" bold="true" box="[192,324,835,859]" pageId="9" pageNumber="8">Remarks.</emphasis>
<taxonomicName id="F7F8E3B1EA7F8513FEA07B1F0848CA57" box="[330,624,835,859]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="acrisius">
<emphasis id="028C4420EA7F8513FEA07B1F0848CA57" box="[330,624,835,859]" italics="true" pageId="9" pageNumber="8">Coprophanaeus acrisius</emphasis>
</taxonomicName>
was preferentially attracted to carrion traps but can be observed in smaller numbers attracted to dung traps (records from MA,
<date id="4446BEF2EA7F8513FC777B380E71CA77" box="[925,1097,867,892]" pageId="9" pageNumber="8" value="2010-01">January 2010</date>
) in cerrado-type habitat (locally called ‘chapada’), which corresponds to slightly elevated regions of open dry woodland with grass. It was also present in an area of dry forest thickets within palm forest. It appears to be an uncommon and patchily distributed species that is sympatric with the more abundant
<taxonomicName id="F7F8E3B1EA7F8513FBDE7B980EE5CAD7" baseAuthorityName="Olsoufieff" baseAuthorityYear="1924" box="[1076,1245,964,987]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cyanescens">
<emphasis id="028C4420EA7F8513FBDE7B980EE5CAD7" box="[1076,1245,964,987]" italics="true" pageId="9" pageNumber="8">C. cyanescens</emphasis>
</taxonomicName>
.
<bibRefCitation id="5469E5C3EA7F8513FB077B9F0B34CAF7" author="Sampaio, D. V. &amp; C. Arzabe &amp; A. A. J. F. Castro &amp; F. Z. Vaz-de-Mello" pageId="9" pageNumber="8" pagination="1 - 4" refId="ref14410" refString="Sampaio, D. V., C. Arzabe, A. A. J. F. Castro, and F. Z. Vaz-de-Mello. 2009. Consideracoes sobre a fauna de besouros rola-bosta (Coleoptera: Scarabaeidae: Scarabaeinae) em duas areas de cerrado litoraneo no Maranhao, Brasil. Anais do IX Congresso de Ecologia do Brasil, 13 a 17 de Setembro de 2009, Sao Lourenco - MG: 1 - 4." type="book chapter" year="2009">Sampaio et al. (2009)</bibRefCitation>
recorded a single individual of this species from an area of coastal cerrado habitat in MA. In MA, it has been recorded only on sandy soils. The specimens from MA match the
<typeStatus id="EF432690EA7F8513FBDE7C5F0EA7CD17" box="[1076,1183,1027,1051]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
, and other specimens housed in the BMNH, closely in color. The dorsal surface of the elytra and the pronotum (except for the lateral basal areas) are bright greenish, with copper-green colored elytral intervals.
</paragraph>
</subSubSection>
</treatment>
</document>
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