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<document id="9D8C1BE3A6D2F22E0CC76B6C8D2AAA43" ID-DOI="10.11646/zootaxa.5406.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10628032" ID-ZooBank="A1F77DF6-4F1B-4858-85FB-22620BCE0484" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1707284769071" checkinUser="plazi" docAuthor="Zuñiga, Ronald &amp; Hanson, Paul" docDate="2024" docId="052387912125FFBBFF4DFDA9FA22444D" docLanguage="en" docName="zootaxa.5406.3.3.pdf" docOrigin="Zootaxa 5406 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5406.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epirhyssa Zuñiga &amp; Hanson 2024" docType="key" docVersion="3" lastPageNumber="444" masterDocId="F91AFFE92124FFB8FFDAFFC2FFBF4605" masterDocTitle="Two new species of Epirhyssa (Hymenoptera: Ichneumonidae: Rhyssinae) from Costa Rica" masterLastPageNumber="450" masterPageNumber="441" pageNumber="442" updateTime="1707315348456" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="E37316755E28245008BE0D4FECF569C8">Two new species of Epirhyssa (Hymenoptera: Ichneumonidae: Rhyssinae) from Costa Rica</mods:title>
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<mods:name id="9AD049256E51EFA8EA977A26970A40CF" type="personal">
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<mods:namePart id="16E7C73E86415E574D73A4DB565052AD">Zuñiga, Ronald</mods:namePart>
<mods:affiliation id="85687ACF62CBC38DC0FFF6741F95E904">Guanacaste Dry Forest Conservation Fund, Sector Santa Rosa, Area de Conservación Guanacaste, Costa Rica.</mods:affiliation>
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<mods:namePart id="8817400628F56F4B93DBCD24A4040B69">Hanson, Paul</mods:namePart>
<mods:affiliation id="987C7607847D2A13398EC68953411460">Centro de Investigación en Biodiversidad y Ecología Tropical, Escuela de Biologia, Universidad de Costa Rica, San Pedro, 11501 - 2060, San José, Costa Rica.</mods:affiliation>
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<mods:title id="F87C835C3E0E4519E1CA73DD9B1AC3DD">Zootaxa</mods:title>
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<mods:date id="0F01C7A19F23D0ED6913F4F43C4D124C">2024</mods:date>
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<treatment id="052387912125FFBBFF4DFDA9FA22444D" LSID="urn:lsid:plazi:treatment:052387912125FFBBFF4DFDA9FA22444D" httpUri="http://treatment.plazi.org/id/052387912125FFBBFF4DFDA9FA22444D" lastPageId="3" lastPageNumber="444" pageId="1" pageNumber="442">
<subSubSection id="C590650C2125FFB9FF4DFDA9FCBF4480" box="[151,768,619,646]" pageId="1" pageNumber="442" type="nomenclature">
<paragraph id="8D3536872125FFB9FF4DFDA9FCBF4480" blockId="1.[151,768,619,646]" box="[151,768,619,646]" pageId="1" pageNumber="442">
<heading id="D67D81EB2125FFB9FF4DFDA9FCBF4480" bold="true" box="[151,768,619,646]" fontSize="11" level="1" pageId="1" pageNumber="442" reason="1">
<emphasis id="BFFEEA952125FFB9FF4DFDA9FCBF4480" bold="true" box="[151,768,619,646]" pageId="1" pageNumber="442">
Key to Costa Rican species of
<taxonomicName id="4A8A4D042125FFB9FDD4FDA9FD384483" ID-CoL="4DBX" ID-ENA="144467" authorityName="Zuñiga &amp; Hanson" authorityYear="2024" box="[526,647,619,646]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="BFFEEA952125FFB9FDD4FDA9FD384483" bold="true" box="[526,647,619,646]" italics="true" pageId="1" pageNumber="442">Epirhyssa</emphasis>
</taxonomicName>
(females)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C590650C2125FFB9FF4DFD71FB2744F4" pageId="1" pageNumber="442" type="discussion">
<paragraph id="8D3536872125FFB9FF4DFD71FB2744F4" blockId="1.[151,1437,691,754]" pageId="1" pageNumber="442">
Modified from
<bibRefCitation id="E91B4B762125FFB9FE98FD71FE6944C8" author="Gauld, I." box="[322,470,691,717]" pageId="1" pageNumber="442" pagination="1 - 589" refId="ref5115" refString="Gauld, I. (1991) The Ichneumonidae of Costa Rica, 1. Memoirs of the American Entomological Institute, 47, 1 - 589." type="journal article" year="1991">Gauld (1991)</bibRefCitation>
and
<bibRefCitation id="E91B4B762125FFB9FDD1FD71FCAD44C8" author="Gauld, I. D. &amp; Ward, S." box="[523,786,691,717]" pageId="1" pageNumber="442" pagination="447 - 453" refId="ref5252" refString="Gauld, I. D. &amp; Ward, S. (1997) Appendix 1. Subfamily Rhyssinae. Supplement to Volume 1. In: Gauld, I., The Ichneumonidae of Costa Rica, 2. Memoirs of the American Entomological Institute, 57, pp. 447 - 453." type="journal article" year="1997">Gauld and Ward (1997)</bibRefCitation>
; additional illustrations can be found in these publications.
<taxonomicName id="4A8A4D042125FFB9FF4DFD15FE0144F4" authority="Gauld" authorityName="Gauld" authorityYear="1991" box="[151,446,727,753]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="corralesi">
<emphasis id="BFFEEA952125FFB9FF4DFD15FECC44F4" box="[151,371,727,753]" italics="true" pageId="1" pageNumber="442">Epirhyssa corralesi</emphasis>
Gauld
</taxonomicName>
was synonymized with
<taxonomicName id="4A8A4D042125FFB9FD17FD1AFC3744F7" authority="Porter" authorityName="Porter" authorityYear="1978" box="[717,904,728,754]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="oaxaca">
<emphasis id="BFFEEA952125FFB9FD17FD1AFC8244F4" box="[717,829,728,753]" italics="true" pageId="1" pageNumber="442">E. oaxaca</emphasis>
Porter
</taxonomicName>
by
<bibRefCitation id="E91B4B762125FFB9FC68FD15FB2B44F4" author="Khalaim, A. I. &amp; Ruiz-Cancino, E. &amp; Coronado-Blanco, J. J." box="[946,1172,727,754]" pageId="1" pageNumber="442" pagination="305 - 313" refId="ref5472" refString="Khalaim, A. I., Ruiz-Cancino, E. &amp; Coronado-Blanco, J. J. (2021) Darwin wasps (Hymenoptera: Ichneumonidae) of Mexico: subfamily Rhyssinae. Russian Entomological Journal, 30 (3), 305 - 313. https: // doi. org / 10.15298 / rusentj. 30.3.08" type="journal article" year="2021">
Khalaim
<emphasis id="BFFEEA952125FFB9FBC0FD1AFBF444F4" box="[1050,1099,727,753]" italics="true" pageId="1" pageNumber="442">et al</emphasis>
. 2021
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C590650C2125FFBBFF4DFCDEFA22444D" lastPageId="3" lastPageNumber="444" pageId="1" pageNumber="442" type="key">
<keyStep id="367E2E222125FFB9FF4DFCDEFA23454A" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DFCDEFA234534" blockId="1.[151,1436,796,847]" box="[151,1436,796,818]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFCDEFA234534" box="[151,1436,796,818]" pageId="1" pageNumber="442">1. Fore wing without blackish mark at apex.................................................................. 2</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DFCF8FA23454A" blockId="1.[151,1436,796,847]" box="[151,1436,825,847]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFCF8FA23454A" box="[151,1436,825,847]" pageId="1" pageNumber="442">- Fore wing with a distinct blackish mark at apex, or at least weakly infumate...................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DFCB6FA2345A2" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DFCB6FA23458C" blockId="1.[151,1436,884,935]" box="[151,1436,884,906]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFCB6FA23458C" box="[151,1436,884,906]" pageId="1" pageNumber="442">
2. Ventral part of occipital carina complete, joining hypostomal carina above base of mandible (
<figureCitation id="15B12A022125FFB9FBBBFCB6FB26458C" box="[1121,1177,884,906]" captionStart="FIGURES 1–2" captionStartId="2.[152,270,1944,1968]" captionTargetBox="[448,1140,187,1915]" captionTargetId="figure-21@2.[438,1150,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–2. Epirhyssa spp., head, posterolateral view showing occipital and hypostomal carinae (modified from Gauld 1991). 1. Occipital carina complete. 2. Occipital carina with ventral part absent." figureDoi="http://doi.org/10.5281/zenodo.10628034" httpUri="https://zenodo.org/record/10628034/files/figure.png" pageId="1" pageNumber="442">Fig. 1</figureCitation>
)................... 3
</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DFC50FA2345A2" blockId="1.[151,1436,884,935]" box="[151,1436,913,935]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFC50FA2345A2" box="[151,1436,913,935]" pageId="1" pageNumber="442">
- Ventral part of occipital carina absent, thus this carina does not join the hypostomal carina (
<figureCitation id="15B12A022125FFB9FB8AFC53FB3645A2" box="[1104,1161,913,935]" captionStart="FIGURES 1–2" captionStartId="2.[152,270,1944,1968]" captionTargetBox="[448,1140,187,1915]" captionTargetId="figure-21@2.[438,1150,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–2. Epirhyssa spp., head, posterolateral view showing occipital and hypostomal carinae (modified from Gauld 1991). 1. Occipital carina complete. 2. Occipital carina with ventral part absent." figureDoi="http://doi.org/10.5281/zenodo.10628034" httpUri="https://zenodo.org/record/10628034/files/figure.png" pageId="1" pageNumber="442">Fig. 2</figureCitation>
).................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DFC0EFA234219" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DFC0EFA2245E7" blockId="1.[151,1437,972,1053]" box="[151,1437,972,994]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFC0EFA2245E7" box="[151,1437,972,994]" pageId="1" pageNumber="442">
3. Tergite II of metasoma extensively smooth and polished, with scattered hairs only anteriorly and laterally.
<taxonomicName id="4A8A4D042125FFB9FB2AFC0EFA2245E7" ID-CoL="6G8BZ" authority="Porter" authorityName="Porter" authorityYear="1978" box="[1264,1437,972,994]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="prolasia">
<emphasis id="BFFEEA952125FFB9FB2AFC0EFAE345E7" bold="true" box="[1264,1372,972,994]" italics="true" pageId="1" pageNumber="442">E. prolasia</emphasis>
Porter
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DFC28FA234219" blockId="1.[151,1437,972,1053]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFC28FA234219" pageId="1" pageNumber="442">- Tergite II of metasoma with alutaceous sculpture (fine wrinkling) over entire surface, and more or less uniformly covered with short hairs.......................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DFB83FA234297" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DFB83FA224270" blockId="1.[151,1437,1089,1170]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFB83FA224270" pageId="1" pageNumber="442">
4. Head, mesosoma and metasoma predominantly blackish brown (
<figureCitation id="15B12A022125FFB9FC8EFB83FC304253" box="[852,911,1089,1111]" captionStart="FIGURES 3–4" captionStartId="4.[152,270,1883,1907]" captionTargetBox="[354,1233,597,1860]" captionTargetId="figure-182@4.[354,1233,597,1860]" captionTargetPageId="4" captionText="FIGURES 3–4. Epirhyssa spp., lateral habitus. 3. E. curtisi Gauld. 4. E. leonoreae Zuñiga &amp; Hanson." figureDoi="http://doi.org/10.5281/zenodo.10628036" httpUri="https://zenodo.org/record/10628036/files/figure.png" pageId="1" pageNumber="442">Fig. 3</figureCitation>
). [Mesoscutum with closely spaced, coarse short dark pubescence.]............................................................................
<taxonomicName id="4A8A4D042125FFB9FADBFB9DFA224270" ID-CoL="6G8QV" authority="Gauld" authorityName="Gauld" box="[1281,1437,1119,1141]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="curtisi">
<emphasis id="BFFEEA952125FFB9FADBFB9DFAE34271" bold="true" box="[1281,1372,1119,1140]" italics="true" pageId="1" pageNumber="442">E. curtisi</emphasis>
Gauld
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DFBBFFA234297" blockId="1.[151,1437,1089,1170]" box="[151,1436,1148,1170]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFBBFFA234297" box="[151,1436,1148,1170]" pageId="1" pageNumber="442">
- Head, mesosoma and metasoma predominantly yellowish with extensive black marks (
<figureCitation id="15B12A022125FFB9FBFEFBBEFBE14294" box="[1060,1118,1148,1170]" captionStart="FIGURES 3–4" captionStartId="4.[152,270,1883,1907]" captionTargetBox="[354,1233,597,1860]" captionTargetId="figure-182@4.[354,1233,597,1860]" captionTargetPageId="4" captionText="FIGURES 3–4. Epirhyssa spp., lateral habitus. 3. E. curtisi Gauld. 4. E. leonoreae Zuñiga &amp; Hanson." figureDoi="http://doi.org/10.5281/zenodo.10628036" httpUri="https://zenodo.org/record/10628036/files/figure.png" pageId="1" pageNumber="442">Fig. 4</figureCitation>
)........................ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DFB75FA224321" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DFB75FA2242EF" blockId="1.[151,1437,1207,1317]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFB75FA2242EF" pageId="1" pageNumber="442">
5. Tergite I at least 1.8 × as long as posterior width; propodeum with a narrow posteromedian black mark, occupying less than 1/3 the width of dorsal surface (
<figureCitation id="15B12A022125FFB9FE35FB16FD9742EF" box="[495,552,1236,1258]" captionStart="FIGURES 5–6" captionStartId="5.[152,270,1380,1404]" captionTargetBox="[357,1231,181,1357]" captionTargetId="figure-17@5.[357,1231,181,1357]" captionTargetPageId="5" captionText="FIGURES 5–6. Epirhyssa spp., propodeum and tergite I, dorsal view. 5. E. oranensis Gauld. 6. E. leonareae Zuñiga &amp; Hanson." figureDoi="http://doi.org/10.5281/zenodo.10628038" httpUri="https://zenodo.org/record/10628038/files/figure.png" pageId="1" pageNumber="442">Fig. 5</figureCitation>
)........................................................
<taxonomicName id="4A8A4D042125FFB9FB3AFB16FA2242EF" ID-CoL="6G8BV" authority="Gauld" authorityName="Gauld" box="[1248,1437,1236,1258]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="oranensis">
<emphasis id="BFFEEA952125FFB9FB3AFB16FAE342EC" bold="true" box="[1248,1372,1236,1257]" italics="true" pageId="1" pageNumber="442">E. oranensis</emphasis>
Gauld
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DFB30FA224321" blockId="1.[151,1437,1207,1317]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFB30FA224321" pageId="1" pageNumber="442">
- Tergite I less than 1.8 × as long as posterior width; propodeum with a broad posteromedian black mark, occupying more than 1/2 the width of dorsal surface (
<figureCitation id="15B12A022125FFB9FDCEFACDFDF24320" box="[532,589,1295,1317]" captionStart="FIGURES 5–6" captionStartId="5.[152,270,1380,1404]" captionTargetBox="[357,1231,181,1357]" captionTargetId="figure-17@5.[357,1231,181,1357]" captionTargetPageId="5" captionText="FIGURES 5–6. Epirhyssa spp., propodeum and tergite I, dorsal view. 5. E. oranensis Gauld. 6. E. leonareae Zuñiga &amp; Hanson." figureDoi="http://doi.org/10.5281/zenodo.10628038" httpUri="https://zenodo.org/record/10628038/files/figure.png" pageId="1" pageNumber="442">Fig. 6</figureCitation>
)......................................
<taxonomicName id="4A8A4D042125FFB9FBEBFACDFADC4321" authority="Zuniga &amp; Hanson" authorityName="Zuniga &amp; Hanson" authorityYear="2024" box="[1073,1379,1294,1317]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="leonoreae" status="sp.n.">
<emphasis id="BFFEEA952125FFB9FBEBFACDFB114321" bold="true" box="[1073,1198,1294,1316]" italics="true" pageId="1" pageNumber="442">E. leonoreae</emphasis>
Zuñiga &amp; Hanson
</taxonomicName>
,
<emphasis id="BFFEEA952125FFB9FAB6FACDFA224321" bold="true" box="[1388,1437,1295,1316]" pageId="1" pageNumber="442">
<taxonomicNameLabel id="A4CD57EE2125FFB9FAB6FACDFA224321" box="[1388,1437,1295,1316]" pageId="1" pageNumber="442" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DFA8BFA23439C" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DFA8BFA23435A" blockId="1.[151,1436,1353,1434]" box="[151,1436,1353,1375]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFA8BFA23435A" box="[151,1436,1353,1375]" pageId="1" pageNumber="442">
6. Mesoscutum with three complete longitudinal black stripes, two lateral stripes not reaching lateral margin (
<figureCitation id="15B12A022125FFB9FB3CFA8BFA9F435B" box="[1254,1312,1353,1375]" captionStart="FIGURES 7–8" captionStartId="6.[152,270,1737,1761]" captionTargetBox="[350,1237,489,1713]" captionTargetId="figure-180@6.[350,1237,489,1713]" captionTargetPageId="6" captionText="FIGURES 7–8. Epirhyssa spp., mesoscutum, dorsal view. 7. E. lewisi Zuñiga &amp; Hanson. 8. E. praecincta Porter." figureDoi="http://doi.org/10.5281/zenodo.10628040" httpUri="https://zenodo.org/record/10628040/files/figure.png" pageId="1" pageNumber="442">Fig. 7</figureCitation>
)........ 7
</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DFAA5FA23439C" blockId="1.[151,1436,1353,1434]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFAA5FA23439C" pageId="1" pageNumber="442">
- Mesoscutum with complete longitudinal black stripe in middle and with two lateral black marks anteriorly that reach the lateral margins but do not extend posteriorly (
<figureCitation id="15B12A022125FFB9FD8DFA46FD2F439C" box="[599,656,1412,1434]" captionStart="FIGURES 7–8" captionStartId="6.[152,270,1737,1761]" captionTargetBox="[350,1237,489,1713]" captionTargetId="figure-180@6.[350,1237,489,1713]" captionTargetPageId="6" captionText="FIGURES 7–8. Epirhyssa spp., mesoscutum, dorsal view. 7. E. lewisi Zuñiga &amp; Hanson. 8. E. praecincta Porter." figureDoi="http://doi.org/10.5281/zenodo.10628040" httpUri="https://zenodo.org/record/10628040/files/figure.png" pageId="1" pageNumber="442">Fig. 8</figureCitation>
).............................................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DFA7DFA224029" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DFA7DFA2243F7" blockId="1.[151,1437,1471,1581]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFA7DFA2243F7" pageId="1" pageNumber="442">
7. Antennal flagellum black, except segments 5–7 or 6–7 yellowish orange; metasomal tergites VI–VII at least partially yellow (
<figureCitation id="15B12A022125FFB9FF33FA1EFE9C43F4" box="[233,291,1500,1522]" captionStart="FIGURES 9–10" captionStartId="7.[152,270,1840,1864]" captionTargetBox="[356,1231,561,1817]" captionTargetId="figure-161@7.[356,1231,561,1817]" captionTargetPageId="7" captionText="FIGURES 9–10. Epirhyssa spp., lateral habitus. 9. E. quirosi Gauld. 10. E. lewisi Zuñiga &amp; Hanson" figureDoi="http://doi.org/10.5281/zenodo.10628042" httpUri="https://zenodo.org/record/10628042/files/figure.png" pageId="1" pageNumber="442">Fig. 9</figureCitation>
).................................................................................
<taxonomicName id="4A8A4D042125FFB9FB20FA1EFA2243F7" ID-CoL="6G8BW" authority="Gauld" authorityName="Gauld" box="[1274,1437,1500,1522]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="quirosi">
<emphasis id="BFFEEA952125FFB9FB20FA1EFAE443F4" bold="true" box="[1274,1371,1500,1521]" italics="true" pageId="1" pageNumber="442">E. quirosi</emphasis>
Gauld
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DFA38FA224029" blockId="1.[151,1437,1471,1581]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DFA38FA224029" pageId="1" pageNumber="442">
- Flagellum dark brown, except segments 2–5 slightly paler; metasomal tergites VI–VII entirely black (
<figureCitation id="15B12A022125FFB9FB69FA3BFB47400A" box="[1203,1272,1529,1551]" captionStart="FIGURES 9–10" captionStartId="7.[152,270,1840,1864]" captionTargetBox="[356,1231,561,1817]" captionTargetId="figure-161@7.[356,1231,561,1817]" captionTargetPageId="7" captionText="FIGURES 9–10. Epirhyssa spp., lateral habitus. 9. E. quirosi Gauld. 10. E. lewisi Zuñiga &amp; Hanson" figureDoi="http://doi.org/10.5281/zenodo.10628042" httpUri="https://zenodo.org/record/10628042/files/figure.png" pageId="1" pageNumber="442">Fig. 10</figureCitation>
).....................................................................................
<taxonomicName id="4A8A4D042125FFB9FB80F9D5FADC4029" authority="Zuniga &amp; Hanson" authorityName="Zuniga &amp; Hanson" authorityYear="2024" box="[1114,1379,1558,1581]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="lewisi" status="sp.n.">
<emphasis id="BFFEEA952125FFB9FB80F9D5FB114029" bold="true" box="[1114,1198,1558,1580]" italics="true" pageId="1" pageNumber="442">E. lewisi</emphasis>
Zuñiga &amp; Hanson
</taxonomicName>
,
<emphasis id="BFFEEA952125FFB9FAB6F9D5FA224029" bold="true" box="[1388,1437,1559,1580]" pageId="1" pageNumber="442">
<taxonomicNameLabel id="A4CD57EE2125FFB9FAB6F9D5FA224029" box="[1388,1437,1559,1580]" pageId="1" pageNumber="442" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DF993FA2240BA" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DF993FA224080" blockId="1.[151,1437,1617,1727]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DF993FA224080" pageId="1" pageNumber="442">
8. Antennal flagellum with pale band on segments 9–13 (or 15); face usually with a blackish median vertical stripe; apical margin of clypeus with a small nipple-like median tubercle (
<figureCitation id="15B12A022125FFB9FD06F9ADFCA04081" box="[732,799,1647,1669]" captionStart="FIGURES 11–12" captionStartId="8.[152,270,1495,1519]" captionTargetBox="[357,1230,198,1454]" captionTargetId="figure-21@8.[340,1247,181,1471]" captionTargetPageId="8" captionText="FIGURES 11–12. Epirhyssa spp., head, anterior view. 11. E. theloides Gauld. 12. E. praecincta Porter." figureDoi="http://doi.org/10.5281/zenodo.10628046" httpUri="https://zenodo.org/record/10628046/files/figure.png" pageId="1" pageNumber="442">Fig. 11</figureCitation>
)....................................
<taxonomicName id="4A8A4D042125FFB9FB32F9ADFA224080" ID-CoL="6FWC2" authority="Gauld" authorityName="Gauld" box="[1256,1437,1646,1669]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="theloides">
<emphasis id="BFFEEA952125FFB9FB32F9ADFAE34081" bold="true" box="[1256,1372,1646,1668]" italics="true" pageId="1" pageNumber="442">E. theloides</emphasis>
Gauld
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DF94FFA2240BA" blockId="1.[151,1437,1617,1727]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DF94FFA2240BA" pageId="1" pageNumber="442">
- Flagellum with pale band on segments (5) 6–9 (or 10); face uniformly yellow; apical margin of clypeus with broad rounded median tubercle (
<figureCitation id="15B12A022125FFB9FE53F96BFE7040BA" box="[393,463,1705,1727]" captionStart="FIGURES 11–12" captionStartId="8.[152,270,1495,1519]" captionTargetBox="[357,1230,198,1454]" captionTargetId="figure-21@8.[340,1247,181,1471]" captionTargetPageId="8" captionText="FIGURES 11–12. Epirhyssa spp., head, anterior view. 11. E. theloides Gauld. 12. E. praecincta Porter." figureDoi="http://doi.org/10.5281/zenodo.10628046" httpUri="https://zenodo.org/record/10628046/files/figure.png" pageId="1" pageNumber="442">Fig. 12</figureCitation>
)...............................................................
<taxonomicName id="4A8A4D042125FFB9FB02F968FA2240BA" ID-CoL="6FWBZ" authority="Porter" authorityName="Porter" box="[1240,1437,1705,1727]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="praecincta">
<emphasis id="BFFEEA952125FFB9FB02F968FAE340BB" bold="true" box="[1240,1372,1705,1727]" italics="true" pageId="1" pageNumber="442">E. praecincta</emphasis>
Porter
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DF926FA234154" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DF926FA234112" blockId="1.[151,1436,1764,1874]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DF926FA234112" pageId="1" pageNumber="442">9. Antennal flagellum more or less entirely black basally, without a pale submedian band and with only extreme apex slightly brownish; tergite I of metasoma moderately slender, 1.6–2.2 × as long as posteriorly broad......................... 10</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DF8DDFA234154" blockId="1.[151,1436,1764,1874]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DF8DDFA234154" pageId="1" pageNumber="442">- Flagellum basally pale yellowish, sometimes with a submedian pale band; tergite I of metasoma very slender, 2.4 or more × as long as posteriorly broad.............................................................................. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222125FFB9FF4DF8B5FA2341E1" pageId="1" pageNumber="442">
<paragraph id="8D3536872125FFB9FF4DF8B5FA2241AF" blockId="1.[151,1437,1911,2021]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DF8B5FA2241AF" pageId="1" pageNumber="442">
10. Mesosoma and metasoma uniformly orange, face subquadrate, as broad as medially high (from clypeofacial suture to base of antenna)................................................................................
<taxonomicName id="4A8A4D042125FFB9FB27F856FA2241AF" ID-CoL="6G8BY" authority="Gauld" authorityName="Gauld" authorityYear="1991" box="[1277,1437,1940,1962]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="porteri">
<emphasis id="BFFEEA952125FFB9FB27F856FAE341AC" bold="true" box="[1277,1372,1940,1961]" italics="true" pageId="1" pageNumber="442">E. porteri</emphasis>
Gauld
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872125FFB9FF4DF870FA2341E1" blockId="1.[151,1437,1911,2021]" pageId="1" pageNumber="442">
<keyLead id="367B95B22125FFB9FF4DF870FA2341E1" pageId="1" pageNumber="442">- Mesosoma and metasoma yellow with profuse black markings; face transverse 1.1–1.3 × as broad as medially high (from clypeofacial suture to base of antenna)................................................................... 11</keyLead>
</paragraph>
</keyStep>
<caption id="D9F5660F2126FFBAFF42F85AFC7F41D1" ID-DOI="http://doi.org/10.5281/zenodo.10628034" ID-Zenodo-Dep="10628034" httpUri="https://zenodo.org/record/10628034/files/figure.png" pageId="2" pageNumber="443" startId="2.[152,270,1944,1968]" targetBox="[448,1140,187,1915]" targetPageId="2" targetType="figure">
<paragraph id="8D3536872126FFBAFF42F85AFC7F41D1" blockId="2.[151,1436,1944,2004]" pageId="2" pageNumber="443">
<emphasis id="BFFEEA952126FFBAFF42F85AFEFC41B5" bold="true" box="[152,323,1944,1968]" pageId="2" pageNumber="443">FIGURES 1–2.</emphasis>
<taxonomicName id="4A8A4D042126FFBAFE91F85AFE5A41B5" box="[331,485,1944,1968]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BFFEEA952126FFBAFE91F85AFE0F41B5" box="[331,432,1944,1968]" italics="true" pageId="2" pageNumber="443">Epirhyssa</emphasis>
spp.
</taxonomicName>
, head, posterolateral view showing occipital and hypostomal carinae (modified from
<bibRefCitation id="E91B4B762126FFBAFA84F85AFF7241D1" author="Gauld, I." pageId="2" pageNumber="443" pagination="1 - 589" refId="ref5115" refString="Gauld, I. (1991) The Ichneumonidae of Costa Rica, 1. Memoirs of the American Entomological Institute, 47, 1 - 589." type="journal article" year="1991">Gauld 1991</bibRefCitation>
). 1. Occipital carina complete. 2. Occipital carina with ventral part absent.
</paragraph>
</caption>
<keyStep id="367E2E222127FFBBFF4DFF5AFA234700" pageId="3" pageNumber="444">
<paragraph id="8D3536872127FFBBFF4DFF5AFA2346CF" blockId="3.[151,1437,151,261]" pageId="3" pageNumber="444">
<keyLead id="367B95B22127FFBBFF4DFF5AFA2346CF" pageId="3" pageNumber="444">
11. Lateral lobe of mesoscutum entirely yellow, or with a faint yellowish brown stripe, fore wing with first subdiscal cell very strongly widened distally, its distal breadth 2.4–2.7 × its basal breadth..........................
<taxonomicName id="4A8A4D042127FFBBFB17FF77FA2346CF" ID-CoL="6G8R2" authority="Cresson" authorityName="Cresson" authorityYear="1874" box="[1229,1436,181,202]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="BFFEEA952127FFBBFB17FF77FAF646CF" bold="true" box="[1229,1353,181,202]" italics="true" pageId="3" pageNumber="444">E. mexicana</emphasis>
Cresson
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872127FFBBFF4DFF11FA234700" blockId="3.[151,1437,151,261]" pageId="3" pageNumber="444">
<keyLead id="367B95B22127FFBBFF4DFF11FA234700" pageId="3" pageNumber="444">- Lateral lobe of mesoscutum with a longitudinal black stripe; fore wing with first subdiscal cell moderately widened distally, its distal breadth 1.5–2.1 × its basal breadth................................................................. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222127FFBBFF4DFEE8FA2347B0" pageId="3" pageNumber="444">
<paragraph id="8D3536872127FFBBFF4DFEE8FA224758" blockId="3.[151,1437,298,437]" pageId="3" pageNumber="444">
<keyLead id="367B95B22127FFBBFF4DFEE8FA224758" pageId="3" pageNumber="444">
12. Apex of clypeus with a strong median tubercle; fore wing with a triangular black mark centered around distal end of Rs; distal apex of antenna slender, subapical flagellomeres about 1.2 × as broad as the central flagellomeres.......
<taxonomicName id="4A8A4D042127FFBBFB38FE8AFA224758" ID-CoL="6FWQL" authority="Gauld" authorityName="Gauld" authorityYear="1991" box="[1250,1437,327,349]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="frohbergi">
<emphasis id="BFFEEA952127FFBBFB38FE8AFAE44758" bold="true" box="[1250,1371,327,349]" italics="true" pageId="3" pageNumber="444">E. frohbergi</emphasis>
Gauld
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872127FFBBFF4DFEA7FA2347B0" blockId="3.[151,1437,298,437]" pageId="3" pageNumber="444">
<keyLead id="367B95B22127FFBBFF4DFEA7FA2347B0" pageId="3" pageNumber="444">
- Apex of clypeus without any trace of a median tubercle; fore wing with entire distal apex blackish; distal end of flagellum slightly clavate, subapical flagellomeres about 1.5 or more × as broad as the central flagellomeres...................................................................................................
<taxonomicName id="4A8A4D042127FFBBFB5DFE62FA2347B0" ID-CoL="6G8BC" authority="Gauld and Ward" authorityName="Gauld and Ward" box="[1159,1436,415,437]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="zurquiae">
<emphasis id="BFFEEA952127FFBBFB5DFE62FB4547B1" bold="true" box="[1159,1274,415,437]" italics="true" pageId="3" pageNumber="444">E. zurquiae</emphasis>
Gauld and Ward
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="367E2E222127FFBBFF4DFE18FA22444D" pageId="3" pageNumber="444">
<paragraph id="8D3536872127FFBBFF4DFE18FA224409" blockId="3.[151,1437,474,584]" pageId="3" pageNumber="444">
<keyLead id="367B95B22127FFBBFF4DFE18FA224409" pageId="3" pageNumber="444">
13. Hind coxa yellowish orange, very long and slender, in profile about 3.4 × as long as broad; metasoma entirely orange; antennal flagellum without a submedian pale band..........................................................
<taxonomicName id="4A8A4D042127FFBBFAF4FE3AFA224409" ID-CoL="6FWBS" authorityName="Gauld &amp; Ward" authorityYear="1997" box="[1326,1437,503,525]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="BFFEEA952127FFBBFAF4FE3AFA224409" bold="true" box="[1326,1437,503,525]" italics="true" pageId="3" pageNumber="444">E. osaensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8D3536872127FFBBFF4DFDD7FA22444D" blockId="3.[151,1437,474,584]" pageId="3" pageNumber="444">
<keyLead id="367B95B22127FFBBFF4DFDD7FA22444D" pageId="3" pageNumber="444">
- Hind coxa yellowish dorsally and black ventrally, moderately slender, in profile 2.7–2.9 × as long as broad; metasoma yellow and black banded; flagellum with a submedian pale band........................................
<taxonomicName id="4A8A4D042127FFBBFB23FDF0FA22444D" ID-CoL="6FWC7" authority="Porter" authorityName="Porter" authorityYear="1978" box="[1273,1437,562,584]" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="oaxaca">
<emphasis id="BFFEEA952127FFBBFB23FDF0FAE34442" bold="true" box="[1273,1372,562,583]" italics="true" pageId="3" pageNumber="444">E. oaxaca</emphasis>
Porter
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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