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<document id="6DC682B6868F2C45EA37650972C52A02" ID-DOI="10.1051/acarologia/20132081" ID-ISSN="2107-7207" ID-Zenodo-Dep="5393111" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630318087619" checkinUser="marcus" docAuthor="Nakamura, K., Nakamura, Y. - N. &amp; Fujikawa, T." docDate="2013" docId="03F5672C4E33C15FFC43F256A186BD1B" docLanguage="en" docName="Acarologia.2013.53.1.41-76.pdf" docOrigin="Acarologia 53 (1)" docSource="http://dx.doi.org/10.1051/acarologia/20132081" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Tectocepheus acutus Nakamura &amp; Nakamura &amp; Fujikawa 2013, n.sp." docType="treatment" docVersion="3" lastPageNumber="63" masterDocId="FFCC1F544E27C147FF93F467A319BE21" masterDocTitle="Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011" masterLastPageNumber="76" masterPageNumber="41" pageNumber="60" updateTime="1699225018176" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
<mods:mods id="9BBF2521154AF8B4BC1EA8F78FF538FA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="65CE584B1E7A345188CDF451B5AFC5FE">Oribatid Mites (Acari, Oribatida) From Tohoku (Northeast Japan), Collected After A Tidal Wave In 2011</mods:title>
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<mods:name id="5E460EF9A3109E0DF8FF2D3C1E8FF67C" type="personal">
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<mods:namePart id="158991E77F0AEF2A064E68A1FD082310">Nakamura, K.</mods:namePart>
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<mods:namePart id="786A46496D168AB48410A4429E18C634">Nakamura, Y. - N.</mods:namePart>
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<mods:namePart id="1A8F453DD27386C3365809BA4D571125">Fujikawa, T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A8DBFBE1EB7CB91FD4B13043932F37D5">text</mods:typeOfResource>
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<mods:title id="467010F3CDF601679E34C6B084528DFB">Acarologia</mods:title>
</mods:titleInfo>
<mods:part id="3732CD0D261C9156166EB870F9D928A9">
<mods:date id="2937F4597FB6AA92517B8DF2323C2926">2013</mods:date>
<mods:detail id="5967AF0F2040DB354422C91A0BE74849" type="pubDate">
<mods:number id="89267A198686C198CFDBD7A733E1BA72">2013-03-29</mods:number>
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<mods:detail id="178DA6612100978CDED78C5C57D52590" type="volume">
<mods:number id="BB940CBB8151DF5DF39EEE003003C303">53</mods:number>
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<mods:number id="7FAB50F9EF32DEE0920E70BA7751403A">1</mods:number>
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<mods:url id="0370A202CD5F98BA2941618147916EDC">http://dx.doi.org/10.1051/acarologia/20132081</mods:url>
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<mods:classification id="5E331715B63285D5E5927870097E6DE3">journal article</mods:classification>
<mods:identifier id="091AE50FF5337A558BB4C8E1BD72180C" type="DOI">10.1051/acarologia/20132081</mods:identifier>
<mods:identifier id="19F6409EC93C5AAE0B2658E635A626E1" type="ISSN">2107-7207</mods:identifier>
<mods:identifier id="084F00B019E704D22882A28EF2DCFCE1" type="Zenodo-Dep">5393111</mods:identifier>
</mods:mods>
<treatment id="03F5672C4E33C15FFC43F256A186BD1B" LSID="urn:lsid:plazi:treatment:03F5672C4E33C15FFC43F256A186BD1B" httpUri="http://treatment.plazi.org/id/03F5672C4E33C15FFC43F256A186BD1B" lastPageId="24" lastPageNumber="63" pageId="20" pageNumber="60">
<subSubSection id="C34685B14E33C153FC43F256A7F8B8B1" pageId="20" pageNumber="60" type="nomenclature">
<paragraph id="8BE3D63A4E33C153FC43F256A7E6B86B" blockId="20.[899,1355,1585,1680]" box="[976,1279,1585,1610]" pageId="20" pageNumber="60">
<heading id="D0AB61564E33C153FC43F256A7E6B86B" bold="true" box="[976,1279,1585,1610]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="60" reason="2">
<emphasis id="B9280A284E33C153FC43F256A7E6B86B" bold="true" box="[976,1279,1585,1610]" pageId="20" pageNumber="60">
<taxonomicName id="4C5CADB94E33C153FC43F256A7A7B868" ID-CoL="8T8P6" authority="Nakamura &amp; Nakamura &amp; Fujikawa, 2013" authorityName="Nakamura &amp; Nakamura &amp; Fujikawa" authorityYear="2013" box="[976,1214,1585,1609]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="60" phylum="Arthropoda" rank="species" species="acutus" status="sp. nov.">
<emphasis id="B9280A284E33C153FC43F256A7A7B868" bold="true" box="[976,1214,1585,1609]" italics="true" pageId="20" pageNumber="60">Tectocepheus acutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BB7534E33C153FB56F255A7E6B86B" box="[1221,1279,1586,1610]" pageId="20" pageNumber="60" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE3D63A4E33C153FC10F232A7F8B8B1" blockId="20.[899,1355,1585,1680]" pageId="20" pageNumber="60">
[Japanese name: Togari-kuwagatadani]
<emphasis id="B9280A284E33C153FC7DF21FA7F8B8B1" bold="true" box="[1006,1249,1656,1680]" pageId="20" pageNumber="60">
(
<figureCitation id="1367CABF4E33C153FC66F21FA768B8B1" box="[1013,1137,1656,1680]" captionStart="FIGURE" captionStartId="21.[159,171,1829,1848]" captionTargetBox="[268,1337,405,1765]" captionTargetId="figure-4@21.[242,1346,338,1798]" captionTargetPageId="21" captionText="FIGURE 15: A – Left of lyrifissure im region of Tectocepheus velatus velatus (Michael, 1880) (NSMT-Ac 13633); B-G: Tectocepheus acutus sp. nov. B – Dorsal view; C – Prodorsum; D – Solenidial region on left tarsus I and tibia I; E – Solenidial region on left genu I;" figureDoi="http://doi.org/10.5281/zenodo.5393145" httpUri="https://zenodo.org/record/5393145/files/figure.png" pageId="20" pageNumber="60">Figures 15</figureCitation>
B-G – 16)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34685B14E33C15FFCA9F2D5A186BD1B" lastPageId="24" lastPageNumber="64" pageId="20" pageNumber="60" type="diagnosis">
<paragraph id="8BE3D63A4E33C153FCA9F2D5A68CB9BD" blockId="20.[826,1429,1714,1948]" pageId="20" pageNumber="60">
Diagnosis — Average body length 282 µm; width 171 µm. Rostral anterior margin without incision, broadly rounded with rostral trowel. Lamellar cuspis with dents bilaterally to lamellar seta, not extending to level of rostral anterior margin. Setae
<emphasis id="B9280A284E33C153FAECF358A68CB977" box="[1407,1429,1855,1878]" italics="true" pageId="20" pageNumber="60">in</emphasis>
roughened rod-like. Sensilli with spinose, clavate head and smooth, thin stem. Bothridia with deep
</paragraph>
<caption id="DF2386B24E32C152FF0CF342A637B975" ID-DOI="http://doi.org/10.5281/zenodo.5393145" ID-Zenodo-Dep="5393145" httpUri="https://zenodo.org/record/5393145/files/figure.png" pageId="21" pageNumber="61" startId="21.[159,171,1829,1848]" targetBox="[268,1337,405,1765]" targetPageId="21">
<paragraph id="8BE3D63A4E32C152FF0CF342A637B975" blockId="21.[159,1429,1829,1904]" pageId="21" pageNumber="61">
FIGURE 15: A – Left of lyrifissure
<emphasis id="B9280A284E32C152FE73F341A2EFB919" box="[480,502,1830,1848]" italics="true" pageId="21" pageNumber="61">im</emphasis>
region of
<taxonomicName id="4C5CADB94E32C152FDC5F342A0C6B919" authority="velatus (Michael, 1880)" authorityName="velatus (Michael" authorityYear="1880" baseAuthorityName="Michael" baseAuthorityYear="1880" box="[598,991,1829,1848]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="velatus" subSpecies="velatus">
<emphasis id="B9280A284E32C152FDC5F342A053B919" box="[598,842,1829,1848]" italics="true" pageId="21" pageNumber="61">Tectocepheus velatus velatus</emphasis>
(
<bibRefCitation id="EFCDABCB4E32C152FCC4F342A0C3B919" author="Michael A. D." box="[855,986,1829,1848]" pageId="21" pageNumber="61" pagination="177 - 201" refId="ref15743" refString="Michael A. D. 1880 - A further contribution to the knowledgeme of British Oribatidae. (Part II.) - Journ. R. Micr. Soc. Trans. Soc., III (5): 177 - 201, pls. 5 &amp; 6." type="journal article" year="1880">Michael, 1880</bibRefCitation>
)
</taxonomicName>
(NSMT-Ac 13633); B-G:
<taxonomicName id="4C5CADB94E32C152FB55F342A66DB919" authority="Nakamura &amp; Nakamura &amp; Fujikawa, 2013" authorityName="Nakamura &amp; Nakamura &amp; Fujikawa" authorityYear="2013" box="[1222,1396,1829,1848]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="61" phylum="Arthropoda" rank="species" species="acutus" status="sp. nov.">
<emphasis id="B9280A284E32C152FB55F342A66DB919" box="[1222,1396,1829,1848]" italics="true" pageId="21" pageNumber="61">Tectocepheus acutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BB7534E32C152FAE9F342A3F0B975" pageId="21" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
B – Dorsal view; C – Prodorsum; D – Solenidial region on left tarsus I and tibia I; E – Solenidial region on left genu I;
</paragraph>
</caption>
<paragraph id="8BE3D63A4E32C152FF52F33AA653B94E" blockId="21.[159,1429,1829,1904]" box="[193,1354,1884,1904]" pageId="21" pageNumber="61">
<materialsCitation id="3B34DC674E32C152FF52F33AA65FB94E" box="[193,1350,1884,1904]" collectionCode="III" location="Solenidial region" pageId="21" pageNumber="61" specimenCount="2" typeStatus="holotype">
F –
<location id="8E8380E14E32C152FF71F33BA298B94E" LSID="urn:lsid:plazi:treatment:03F5672C4E33C15FFC43F256A186BD1B:8E8380E14E32C152FF71F33BA298B94E" box="[226,385,1884,1903]" name="Solenidial region" pageId="21" pageNumber="61">Solenidial region</location>
on right genu
<collectionCode id="ED4D4EFF4E32C152FD98F33AA13BB951" box="[523,546,1885,1904]" pageId="21" pageNumber="61">III</collectionCode>
; G – Chelicera (B, F:
<typeStatus id="54E768984E32C152FD77F33BA024B94E" box="[740,829,1884,1903]" pageId="21" pageNumber="61" type="holotype">Holotype</typeStatus>
NSMT-Ac 13630; C, C, E, G:
<typeStatus id="54E768984E32C152FBD8F33AA786B951" box="[1099,1183,1885,1904]" pageId="21" pageNumber="61" type="paratype">Paratype</typeStatus>
NSMT-Ac 13631)
</materialsCitation>
.
</paragraph>
<caption id="DF2386B24E31C151FE38F33CA792B94F" ID-DOI="http://doi.org/10.5281/zenodo.5393149" ID-Zenodo-Dep="5393149" box="[427,1163,1883,1902]" httpUri="https://zenodo.org/record/5393149/files/figure.png" pageId="22" pageNumber="62" startId="22.[427,439,1883,1902]" targetBox="[350,1229,350,1843]" targetPageId="22">
<paragraph id="8BE3D63A4E31C151FE38F33CA792B94F" blockId="22.[427,1163,1883,1902]" box="[427,1163,1883,1902]" pageId="22" pageNumber="62">
FIGURE 16:
<taxonomicName id="4C5CADB94E31C151FD8FF33CA1D1B94C" authority="Nakamura &amp; Nakamura &amp; Fujikawa, 2013" authorityName="Nakamura &amp; Nakamura &amp; Fujikawa" authorityYear="2013" box="[540,712,1883,1902]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="62" phylum="Arthropoda" rank="species" species="acutus" status="sp. nov.">
<emphasis id="B9280A284E31C151FD8FF33CA1D1B94C" box="[540,712,1883,1902]" italics="true" pageId="22" pageNumber="62">Tectocepheus acutus</emphasis>
</taxonomicName>
<emphasis id="B9280A284E31C151FD5DF33CA1E2B94F" bold="true" box="[718,763,1883,1902]" pageId="22" pageNumber="62">
<taxonomicNameLabel id="A21BB7534E31C151FD5DF33CA1E2B94F" box="[718,763,1883,1902]" pageId="22" pageNumber="62" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
: Ventral view (Paratype NSMT-Ac 13631).
</paragraph>
</caption>
<paragraph id="8BE3D63A4E30C150FF0CF550A17FBC65" blockId="23.[159,762,311,1951]" pageId="23" pageNumber="63">
incision, without swelling or projection. Depressions and dorsosejugal scissure absent. Humeral region with small triangular projection. Ten pairs of notogastral setae. Genito-anal setal formula: 6- 1-2-3. Lyrifissures
<emphasis id="B9280A284E30C150FE10F5A4A2BABFFA" box="[387,419,451,475]" italics="true" pageId="23" pageNumber="63">iad</emphasis>
located along, near anterior margin of anal aperture. Epimeral setal formula: 3- 1-3-3. Trochantera III and IV bearing carina with sharply pointed apex. Monodactylous.
</paragraph>
<paragraph id="8BE3D63A4E30C150FF56F63AA12ABC9D" blockId="23.[159,762,311,1951]" pageId="23" pageNumber="63">
<materialsCitation id="3B34DC674E30C150FF56F63AA12ABC9D" collectionCode="AC" location="Solenidial region" pageId="23" pageNumber="63" specimenCode="AC 13631" specimenCount="1" typeStatus="holotype">
Material examined —
<typeStatus id="54E768984E30C150FD9AF63AA161BC54" box="[521,632,605,629]" pageId="23" pageNumber="63" type="holotype">Holotype</typeStatus>
(Female) (NSMT-Ac 13630) from point B;
<specimenCount id="9D5A1DB34E30C150FD8BF6E6A18CBCB8" box="[536,661,641,665]" pageId="23" pageNumber="63" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(NSMT-
<specimenCode id="DBFA7E414E30C150FF0CF6C3A20DBC9D" box="[159,276,676,700]" collectionCode="AC" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="23" pageNumber="63" type="University or college">AC 13631</specimenCode>
): same data as holotype.
</materialsCitation>
</paragraph>
<paragraph id="8BE3D63A4E30C150FF56F6B2A27CBD2E" blockId="23.[159,762,311,1951]" pageId="23" pageNumber="63">Etymology — After long, sharply pointed apex on trochanter III.</paragraph>
<paragraph id="8BE3D63A4E30C150FF56F74DA20FBDEC" blockId="23.[159,762,311,1951]" pageId="23" pageNumber="63">Measurements and body appearance — Body length 279 – 286 µm; width 164 – 179 µm. Body colour light brown. Whole body surface covered with cerotegument; cerotegument irregularly dark granulate.</paragraph>
<paragraph id="8BE3D63A4E30C150FF56F780A2BFB82C" blockId="23.[159,762,311,1951]" pageId="23" pageNumber="63">
Prodorsum — Rostral anterior margin without incision, broadly rounded with rostral trowel (
<figureCitation id="1367CABF4E30C150FD5EF06DA3C9BA64" captionStart="FIGURE" captionStartId="21.[159,171,1829,1848]" captionTargetBox="[268,1337,405,1765]" captionTargetId="figure-4@21.[242,1346,338,1798]" captionTargetPageId="21" captionText="FIGURE 15: A – Left of lyrifissure im region of Tectocepheus velatus velatus (Michael, 1880) (NSMT-Ac 13633); B-G: Tectocepheus acutus sp. nov. B – Dorsal view; C – Prodorsum; D – Solenidial region on left tarsus I and tibia I; E – Solenidial region on left genu I;" figureDoi="http://doi.org/10.5281/zenodo.5393145" httpUri="https://zenodo.org/record/5393145/files/figure.png" pageId="23" pageNumber="63">Fig. 15C</figureCitation>
). Rostral setae (16 µm) consisting of unilaterally spinose head and thick, smooth stem, inserting dorsally under lamellar cuspis. Lamellar cuspis with dents bilaterally to lamellar seta, not extending to level of rostral anterior margin; outer dent longer than inner dent. Inner sides of cuspides straight. Setae
<emphasis id="B9280A284E30C150FE31F098A2ADBB36" box="[418,436,1279,1303]" italics="true" pageId="23" pageNumber="63">le</emphasis>
(33 µm) setiform minutely roughened, strongly curved inside. Setae
<emphasis id="B9280A284E30C150FD2CF144A1CCBB1B" box="[703,725,1315,1338]" italics="true" pageId="23" pageNumber="63">in</emphasis>
(7 µm) minutely roughened rod-like. Sensilli (46 µm) composed of thin, roughened stem and globular head bearing dark coloured spines. Setae
<emphasis id="B9280A284E30C150FD2DF1EBA1CFBB82" box="[702,726,1420,1443]" italics="true" pageId="23" pageNumber="63">ex</emphasis>
(4 µm) smooth, short bacilliform. Bothridia opening antero-laterally, with deep incision, without swelling or projection.
</paragraph>
<paragraph id="8BE3D63A4E30C150FF56F241A114B9BC" blockId="23.[159,762,311,1951]" pageId="23" pageNumber="63">
Notogaster — Depressions and dorsosejugal scissure absent. Dorsophragmatic apophysis (hy) conspicuously long. Humeral region with small triangular projection (
<figureCitation id="1367CABF4E30C150FE03F2E8A11AB886" box="[400,515,1679,1703]" captionStart="FIGURE" captionStartId="21.[159,171,1829,1848]" captionTargetBox="[268,1337,405,1765]" captionTargetId="figure-4@21.[242,1346,338,1798]" captionTargetPageId="21" captionText="FIGURE 15: A – Left of lyrifissure im region of Tectocepheus velatus velatus (Michael, 1880) (NSMT-Ac 13633); B-G: Tectocepheus acutus sp. nov. B – Dorsal view; C – Prodorsum; D – Solenidial region on left tarsus I and tibia I; E – Solenidial region on left genu I;" figureDoi="http://doi.org/10.5281/zenodo.5393145" httpUri="https://zenodo.org/record/5393145/files/figure.png" pageId="23" pageNumber="63">Fig. 15B</figureCitation>
). Ten pairs of notogastral setae short (4 – 7µm) setiform, roughened throughout length. Lyrifissures
<emphasis id="B9280A284E30C150FD8CF2B1A12BB8CC" box="[543,562,1750,1773]" italics="true" pageId="23" pageNumber="63">ia</emphasis>
aligned longitudinally or obliquely, lateral to setae
<emphasis id="B9280A284E30C150FDDAF29EA14DB931" box="[585,596,1785,1808]" italics="true" pageId="23" pageNumber="63">c</emphasis>
;
<emphasis id="B9280A284E30C150FDF0F29EA166B931" box="[611,639,1785,1808]" italics="true" pageId="23" pageNumber="63">im</emphasis>
obliquely, lateral to
<emphasis id="B9280A284E30C150FE80F37CA22AB912" box="[275,307,1819,1843]" italics="true" pageId="23" pageNumber="63">gla</emphasis>
;
<emphasis id="B9280A284E30C150FED5F37CA242B912" box="[326,347,1819,1843]" italics="true" pageId="23" pageNumber="63">ih</emphasis>
,
<emphasis id="B9280A284E30C150FEFFF37BA298B912" box="[364,385,1820,1843]" italics="true" pageId="23" pageNumber="63">ip</emphasis>
and
<emphasis id="B9280A284E30C150FE52F37BA2F9B912" box="[449,480,1820,1843]" italics="true" pageId="23" pageNumber="63">psi</emphasis>
perpendicular to notogastral outline. Relative distances: (
<emphasis id="B9280A284E30C150FDDDF359A142B977" box="[590,603,1854,1878]" italics="true" pageId="23" pageNumber="63">h</emphasis>
<subScript id="17D8D47F4E30C150FDCFF32FA17CB978" attach="left" box="[604,613,1864,1881]" fontSize="7" pageId="23" pageNumber="63">3</subScript>
-
<emphasis id="B9280A284E30C150FDFCF359A165B977" box="[623,636,1854,1878]" italics="true" pageId="23" pageNumber="63">h</emphasis>
<subScript id="17D8D47F4E30C150FDEFF32FA19CB978" attach="left" box="[636,645,1864,1881]" fontSize="7" pageId="23" pageNumber="63">3</subScript>
: 120 µm)&gt; (
<emphasis id="B9280A284E30C150FF53F306A3D4B958" box="[192,205,1889,1913]" italics="true" pageId="23" pageNumber="63">h</emphasis>
<subScript id="17D8D47F4E30C150FF5DF30CA3CEB95D" attach="left" box="[206,215,1899,1916]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
-
<emphasis id="B9280A284E30C150FF72F306A3F7B958" box="[225,238,1889,1913]" italics="true" pageId="23" pageNumber="63">h</emphasis>
<subScript id="17D8D47F4E30C150FF7DF30CA3EEB95D" attach="left" box="[238,247,1899,1916]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
: 95 µm)&gt; (
<emphasis id="B9280A284E30C150FE1AF306A2A2B958" box="[393,443,1889,1913]" italics="true" pageId="23" pageNumber="63">lp -lp</emphasis>
: 88 µm)&gt; (
<emphasis id="B9280A284E30C150FDD1F306A156B958" box="[578,591,1889,1913]" italics="true" pageId="23" pageNumber="63">h</emphasis>
<subScript id="17D8D47F4E30C150FDC3F30CA140B95D" attach="left" box="[592,601,1899,1916]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
-
<emphasis id="B9280A284E30C150FDF0F306A169B958" box="[611,624,1889,1913]" italics="true" pageId="23" pageNumber="63">h</emphasis>
<subScript id="17D8D47F4E30C150FDE3F30CA160B95D" attach="left" box="[624,633,1899,1916]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
: 43 µm)&gt; (
<emphasis id="B9280A284E30C150FF3BF3E3A3F1B9BD" box="[168,232,1924,1948]" italics="true" pageId="23" pageNumber="63">lm -lm</emphasis>
: 34 µm)&gt; (
<emphasis id="B9280A284E30C150FEFDF3E2A262B9BD" box="[366,379,1925,1948]" italics="true" pageId="23" pageNumber="63">p</emphasis>
<subScript id="17D8D47F4E30C150FEEFF3E9A29CB9BE" attach="left" box="[380,389,1934,1951]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
-
<emphasis id="B9280A284E30C150FE1CF3E2A285B9BD" box="[399,412,1925,1948]" italics="true" pageId="23" pageNumber="63">p</emphasis>
<subScript id="17D8D47F4E30C150FE0FF3E9A2BCB9BE" attach="left" box="[412,421,1934,1951]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
: 18 µm).
</paragraph>
<paragraph id="8BE3D63A4E30C150FCF3F550A62ABDC8" blockId="23.[826,1429,311,1002]" pageId="23" pageNumber="63">
Ventral region — Genital (31 µm) and anal (55 µm) apertures almost pentagonal in shape; distance (23 µm) between them appreciably shorter than half length of anal aperture. Genito-anal setal formula: 6-1-2-3; all setae smooth setiform (
<figureCitation id="1367CABF4E30C150FB4EF5A4A621BFFA" box="[1245,1336,451,475]" captionStart="FIGURE" captionStartId="22.[427,439,1883,1902]" captionTargetBox="[350,1229,350,1843]" captionTargetId="figure-4@22.[344,1243,339,1851]" captionTargetPageId="22" captionText="FIGURE 16: Tectocepheus acutus n.sp.: Ventral view (Paratype NSMT-Ac 13631)." figureDoi="http://doi.org/10.5281/zenodo.5393149" httpUri="https://zenodo.org/record/5393149/files/figure.png" pageId="23" pageNumber="63">Fig. 16</figureCitation>
). Setae
<emphasis id="B9280A284E30C150FCA9F580A05EBFDF" box="[826,839,487,510]" italics="true" pageId="23" pageNumber="63">g</emphasis>
<subScript id="17D8D47F4E30C150FCD4F597A049BC20" attach="left" box="[839,848,496,513]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
and
<emphasis id="B9280A284E30C150FC02F580A087BFDF" box="[913,926,487,510]" italics="true" pageId="23" pageNumber="63">g</emphasis>
<subScript id="17D8D47F4E30C150FC0DF597A0BEBC20" attach="left" box="[926,935,496,513]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
(9 µm) not inserting at the same level at anterior margin of plates. Setae
<emphasis id="B9280A284E30C150FB24F66DA7C9BC00" box="[1207,1232,522,545]" italics="true" pageId="23" pageNumber="63">ag</emphasis>
(8 µm) inserting latero-posterior to genital aperture. Setae
<emphasis id="B9280A284E30C150FAA9F64BA64ABC65" box="[1338,1363,556,580]" italics="true" pageId="23" pageNumber="63">ad</emphasis>
<subScript id="17D8D47F4E30C150FAC0F651A645BC66" attach="left" box="[1363,1372,566,583]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
and
<emphasis id="B9280A284E30C150FCA9F628A04ABC46" box="[826,851,591,615]" italics="true" pageId="23" pageNumber="63">ad</emphasis>
<subScript id="17D8D47F4E30C150FCC0F63EA045BC4B" attach="left" box="[851,860,601,618]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
(10 µm) aligned in postanal position,
<emphasis id="B9280A284E30C150FA8EF628A62FBC46" box="[1309,1334,591,615]" italics="true" pageId="23" pageNumber="63">ad</emphasis>
<subScript id="17D8D47F4E30C150FAA5F63EA626BC4B" attach="left" box="[1334,1343,601,618]" fontSize="7" pageId="23" pageNumber="63">3</subScript>
(7 µm) in adanal;
<emphasis id="B9280A284E30C150FC29F615A0CABCAB" box="[954,979,626,650]" italics="true" pageId="23" pageNumber="63">ad</emphasis>
<subScript id="17D8D47F4E30C150FC40F61BA0C5BCAC" attach="left" box="[979,988,636,653]" fontSize="7" pageId="23" pageNumber="63">3</subScript>
inserting almost at mid-distance between
<emphasis id="B9280A284E30C150FC18F6F1A0BFBC8C" box="[907,934,662,685]" italics="true" pageId="23" pageNumber="63">an</emphasis>
<subScript id="17D8D47F4E30C150FC35F6F8A0B6BC91" attach="left" box="[934,943,671,688]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
and
<emphasis id="B9280A284E30C150FC63F6F1A712BC8C" box="[1008,1035,662,685]" italics="true" pageId="23" pageNumber="63">an</emphasis>
<subScript id="17D8D47F4E30C150FB99F6F8A70ABC91" attach="left" box="[1034,1043,671,688]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
(7 µm). Lyrifissures
<emphasis id="B9280A284E30C150FA85F6F2A62FBC8C" box="[1302,1334,661,685]" italics="true" pageId="23" pageNumber="63">iad</emphasis>
located along, near anterior margin of anal aperture with enclosed angle 120-140º. Sternal ridge indistinct. Epimeral borders 1-4, sj distinct. Epimeral setal formula: 3-1-3-3; setae smooth, short setiform (3 – 7 µm). Subcapitulum diarthric, subcapitular setae 3 pairs,
<emphasis id="B9280A284E30C150FC15F70FA080BD5E" box="[902,921,872,895]" italics="true" pageId="23" pageNumber="63">A</emphasis>
(14 µm),
<emphasis id="B9280A284E30C150FB82F70EA73FBDA1" box="[1041,1062,873,896]" italics="true" pageId="23" pageNumber="63">m</emphasis>
(14 µm), and
<emphasis id="B9280A284E30C150FB47F70FA7F8BDA1" box="[1236,1249,872,896]" italics="true" pageId="23" pageNumber="63">h</emphasis>
(11 µm); setae thin smooth setiform. Cheliceral setae
<emphasis id="B9280A284E30C150FA94F7ECA632BD82" box="[1287,1323,907,931]" italics="true" pageId="23" pageNumber="63">cha</emphasis>
(23 µm),
<emphasis id="B9280A284E30C150FCA9F7C9A047BDE7" box="[826,862,942,966]" italics="true" pageId="23" pageNumber="63">chb</emphasis>
(16 µm) thin, barbed setiform. Trägårdh’s organ short (14 µm), with a blunt apex (
<figureCitation id="1367CABF4E30C150FB53F7B6A63EBDC8" box="[1216,1319,977,1001]" captionStart="FIGURE" captionStartId="21.[159,171,1829,1848]" captionTargetBox="[268,1337,405,1765]" captionTargetId="figure-4@21.[242,1346,338,1798]" captionTargetPageId="21" captionText="FIGURE 15: A – Left of lyrifissure im region of Tectocepheus velatus velatus (Michael, 1880) (NSMT-Ac 13633); B-G: Tectocepheus acutus sp. nov. B – Dorsal view; C – Prodorsum; D – Solenidial region on left tarsus I and tibia I; E – Solenidial region on left genu I;" figureDoi="http://doi.org/10.5281/zenodo.5393145" httpUri="https://zenodo.org/record/5393145/files/figure.png" pageId="23" pageNumber="63">Fig. 15G</figureCitation>
).
</paragraph>
<paragraph id="8BE3D63A4E30C150FCF3F061A68CB8FA" blockId="23.[826,1429,1030,1756]" pageId="23" pageNumber="63">
<materialsCitation id="3B34DC674E30C150FCF3F061A688B8FA" collectionCode="II, III, IV" pageId="23" pageNumber="63" specimenCount="1" typeStatus="paratype">
Legs — Monodactylous; claw (21 µm) sparsely dentate dorsally. Setal formula: I (1-5-3-4-18),
<collectionCode id="ED4D4EFF4E30C150FA10F04DA68CBA60" box="[1411,1429,1066,1089]" pageId="23" pageNumber="63">II</collectionCode>
(1-5-3-4-14),
<collectionCode id="ED4D4EFF4E30C150FC41F02AA0F4BA45" box="[978,1005,1101,1124]" pageId="23" pageNumber="63">III</collectionCode>
(2-4-1-3-14),
<collectionCode id="ED4D4EFF4E30C150FB03F02AA7B5BA45" box="[1168,1196,1101,1124]" pageId="23" pageNumber="63">IV</collectionCode>
(1-2-2-3-10). Measurements (µm) of right segments according to depressed
<typeStatus id="54E768984E30C150FC50F0F4A736BA8A" box="[963,1071,1171,1195]" pageId="23" pageNumber="63" type="paratype">paratype</typeStatus>
: I (25-40-16-31-35),
<collectionCode id="ED4D4EFF4E30C150FAD7F0F4A64FBA8B" box="[1348,1366,1171,1194]" pageId="23" pageNumber="63">II</collectionCode>
(21- 46-14-25-28),
<collectionCode id="ED4D4EFF4E30C150FC49F0D1A0ECBAEC" box="[986,1013,1206,1229]" pageId="23" pageNumber="63">III</collectionCode>
(50-36-14-26-23),
<collectionCode id="ED4D4EFF4E30C150FB58F0D1A7FEBAEC" box="[1227,1255,1206,1229]" pageId="23" pageNumber="63">IV</collectionCode>
(51-43-21-?-?). Trochantera
<collectionCode id="ED4D4EFF4E30C150FC5CF0BEA0F3BAD1" box="[975,1002,1241,1264]" pageId="23" pageNumber="63">III</collectionCode>
–
<collectionCode id="ED4D4EFF4E30C150FB9FF0BEA731BAD1" box="[1036,1064,1241,1264]" pageId="23" pageNumber="63">IV</collectionCode>
and femora III-IV bearing carina; trochantera
<collectionCode id="ED4D4EFF4E30C150FB92F09BA705BB32" box="[1025,1052,1276,1299]" pageId="23" pageNumber="63">III</collectionCode>
with long, sharply pointed apex (21 µm) bending to rostral side (
<figureCitation id="1367CABF4E30C150FB25F179A613BB17" box="[1206,1290,1310,1334]" captionStart="FIGURE" captionStartId="22.[427,439,1883,1902]" captionTargetBox="[350,1229,350,1843]" captionTargetId="figure-4@22.[344,1243,339,1851]" captionTargetPageId="22" captionText="FIGURE 16: Tectocepheus acutus n.sp.: Ventral view (Paratype NSMT-Ac 13631)." figureDoi="http://doi.org/10.5281/zenodo.5393149" httpUri="https://zenodo.org/record/5393149/files/figure.png" pageId="23" pageNumber="63">Fig. 16</figureCitation>
). On tarsus I, famulus
<emphasis id="B9280A284E30C150FC2FF125A0D1BB7B" box="[956,968,1346,1370]" italics="true" pageId="23" pageNumber="63">ε</emphasis>
(9 µm) consisting of a fine tip and expanded basal portion, situated antero-laterally contiguous to
<emphasis id="B9280A284E30C150FC2BF1EFA0D0BB81" box="[952,969,1416,1440]" italics="true" pageId="23" pageNumber="63">ω</emphasis>
<subScript id="17D8D47F4E30C150FC59F1F6A0CABB83" attach="left" box="[970,979,1425,1442]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
;
<emphasis id="B9280A284E30C150FC71F1EFA0EABB81" box="[994,1011,1416,1440]" italics="true" pageId="23" pageNumber="63">ω</emphasis>
<subScript id="17D8D47F4E30C150FC67F1F6A0E4BB83" attach="left" box="[1012,1021,1425,1442]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
(31 µm) terminating in fine tip;
<emphasis id="B9280A284E30C150FAEAF1EFA693BB81" box="[1401,1418,1416,1440]" italics="true" pageId="23" pageNumber="63">ω</emphasis>
<subScript id="17D8D47F4E30C150FA19F1F6A68ABB83" attach="left" box="[1418,1427,1425,1442]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
(21 µm) bacilliform situated posterior to
<emphasis id="B9280A284E30C150FABEF1CCA627BBE2" box="[1325,1342,1451,1475]" italics="true" pageId="23" pageNumber="63">ω</emphasis>
<subScript id="17D8D47F4E30C150FAADF1D3A65EBBE4" attach="left" box="[1342,1351,1460,1477]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
;
<emphasis id="B9280A284E30C150FACFF1CCA675BBE2" box="[1372,1388,1451,1475]" italics="true" pageId="23" pageNumber="63">ft</emphasis>
’ (8 µm), inserting posterior contiguous to
<emphasis id="B9280A284E30C150FB6BF1A9A610BBC7" box="[1272,1289,1486,1510]" italics="true" pageId="23" pageNumber="63">ω</emphasis>
<subScript id="17D8D47F4E30C150FA9AF1B0A60BBBC9" attach="left" box="[1289,1298,1495,1512]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
(
<figureCitation id="1367CABF4E30C150FAB1F1A9A691BBC7" box="[1314,1416,1486,1510]" captionStart="FIGURE" captionStartId="21.[159,171,1829,1848]" captionTargetBox="[268,1337,405,1765]" captionTargetId="figure-4@21.[242,1346,338,1798]" captionTargetPageId="21" captionText="FIGURE 15: A – Left of lyrifissure im region of Tectocepheus velatus velatus (Michael, 1880) (NSMT-Ac 13633); B-G: Tectocepheus acutus sp. nov. B – Dorsal view; C – Prodorsum; D – Solenidial region on left tarsus I and tibia I; E – Solenidial region on left genu I;" figureDoi="http://doi.org/10.5281/zenodo.5393145" httpUri="https://zenodo.org/record/5393145/files/figure.png" pageId="23" pageNumber="63">Fig. 15D</figureCitation>
). Solenidion
<emphasis id="B9280A284E30C150FC55F196A0CEB828" box="[966,983,1521,1545]" italics="true" pageId="23" pageNumber="63">’</emphasis>
<subScript id="17D8D47F4E30C150FC4BF19DA0F8B82A" attach="left" box="[984,993,1530,1547]" fontSize="7" pageId="23" pageNumber="63">2</subScript>
(15 µm) originating from a small apophysis on the tip of tibia I;
<emphasis id="B9280A284E30C150FB3EF273A7A7B80D" box="[1197,1214,1556,1580]" italics="true" pageId="23" pageNumber="63">’</emphasis>
<subScript id="17D8D47F4E30C150FB2CF279A7D1B80E" attach="left" box="[1215,1224,1566,1583]" fontSize="7" pageId="23" pageNumber="63">1</subScript>
(65 µm) situated at the base of apophysis. On genu I, solenidion
<emphasis id="B9280A284E30C150FA16F250A68DB86E" box="[1413,1428,1591,1615]" italics="true" pageId="23" pageNumber="63">σ</emphasis>
(18 µm) terminating in a fine tip, longer than seta
<emphasis id="B9280A284E30C150FCA9F21AA05EB8B4" box="[826,839,1661,1685]" italics="true" pageId="23" pageNumber="63">d</emphasis>
(8 µm) (
<figureCitation id="1367CABF4E30C150FC22F21AA705B8B4" box="[945,1052,1661,1685]" captionStart="FIGURE" captionStartId="21.[159,171,1829,1848]" captionTargetBox="[268,1337,405,1765]" captionTargetId="figure-4@21.[242,1346,338,1798]" captionTargetPageId="21" captionText="FIGURE 15: A – Left of lyrifissure im region of Tectocepheus velatus velatus (Michael, 1880) (NSMT-Ac 13633); B-G: Tectocepheus acutus sp. nov. B – Dorsal view; C – Prodorsum; D – Solenidial region on left tarsus I and tibia I; E – Solenidial region on left genu I;" figureDoi="http://doi.org/10.5281/zenodo.5393145" httpUri="https://zenodo.org/record/5393145/files/figure.png" pageId="23" pageNumber="63">Fig. 15E</figureCitation>
). All solenidia on tarsus
<collectionCode id="ED4D4EFF4E30C150FAD8F219A646B8B4" box="[1355,1375,1662,1685]" pageId="23" pageNumber="63">II</collectionCode>
, tibiae II-IV and genua II-III short rod-like (
<emphasis id="B9280A284E30C150FA82F2C7A639B899" box="[1297,1312,1696,1720]" italics="true" pageId="23" pageNumber="63">σ</emphasis>
<collectionCode id="ED4D4EFF4E30C150FAB2F2C6A626B899" box="[1313,1343,1697,1720]" pageId="23" pageNumber="63">III</collectionCode>
: 7 µm), shorter than each dorsal seta (
<emphasis id="B9280A284E30C150FB06F2A4A7BBB8FA" box="[1173,1186,1731,1755]" italics="true" pageId="23" pageNumber="63">d</emphasis>
<collectionCode id="ED4D4EFF4E30C150FB31F2A3A7D9B8FA" box="[1186,1216,1732,1755]" pageId="23" pageNumber="63">III</collectionCode>
: 11 µm) (
<figureCitation id="1367CABF4E30C150FAB4F2A4A690B8FA" box="[1319,1417,1731,1755]" captionStart="FIGURE" captionStartId="21.[159,171,1829,1848]" captionTargetBox="[268,1337,405,1765]" captionTargetId="figure-4@21.[242,1346,338,1798]" captionTargetPageId="21" captionText="FIGURE 15: A – Left of lyrifissure im region of Tectocepheus velatus velatus (Michael, 1880) (NSMT-Ac 13633); B-G: Tectocepheus acutus sp. nov. B – Dorsal view; C – Prodorsum; D – Solenidial region on left tarsus I and tibia I; E – Solenidial region on left genu I;" figureDoi="http://doi.org/10.5281/zenodo.5393145" httpUri="https://zenodo.org/record/5393145/files/figure.png" pageId="23" pageNumber="63">Fig. 15F</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE3D63A4E30C15FFCF3F29FA186BD1B" blockId="23.[826,1429,1784,1948]" lastBlockId="24.[159,762,311,826]" lastPageId="24" lastPageNumber="64" pageId="23" pageNumber="63">
Remarks — The new species is similar to
<taxonomicName id="4C5CADB94E30C150FAC6F29EA76DB912" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="63" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B9280A284E30C150FAC6F29EA0CFB912" italics="true" pageId="23" pageNumber="63">Tectocepheus minor</emphasis>
<bibRefCitation id="EFCDABCB4E30C150FC4EF37CA76DB912" author="Berlese A." box="[989,1140,1819,1843]" pageId="23" pageNumber="63" pagination="235 - 252" refId="ref14902" refString="Berlese A. 1903 (1904) - Spicilegia Zoologica. Acari Nuovi - Redia, 1: 235 - 252." type="journal article" year="1903">
Berlese,
<quantity id="4CA47BDF4E30C150FBACF37CA76DB912" box="[1087,1140,1819,1843]" metricMagnitude="1" metricUnit="m" metricValue="4.83362" pageId="23" pageNumber="63" unit="in" value="1903.0">1903</quantity>
</bibRefCitation>
</taxonomicName>
in shape of dentate cuspis and small enclosed angle of lyrifissure pair
<emphasis id="B9280A284E30C150FAFDF359A697B977" box="[1390,1422,1854,1878]" italics="true" pageId="23" pageNumber="63">iad</emphasis>
, however, the former is different from the latter in shape of bothridial ventral extension, long sharply pointed carina on trochantera III, arrangement of genital setae, and solenidia of legs (
<bibRefCitation id="EFCDABCB4E3FC15FFDD7F53DA3CEBFB4" author="Laumann M. &amp; Norton R. A. &amp; Weigmann G. &amp; Scheu S. &amp; Maraun M. &amp; Heethoff M." pageId="24" pageNumber="64" pagination="111 - 122" refId="ref15614" refString="Laumann M., Norton R. A., Weigmann G., Scheu S., Maraun M., Heethoff M. 2007 - Speciation in the parthenogenetic oribatid mite genus Tectocepheus (Acari, Oribatida) as indicated by molecular phylogeny - Pedobiologia, 51: 111 - 122." type="journal article" year="2007">
Laumann
<emphasis id="B9280A284E3FC15FFD2DF53CA1EDBF53" box="[702,756,346,370]" italics="true" pageId="24" pageNumber="64">et al.</emphasis>
, 2007
</bibRefCitation>
;
<bibRefCitation id="EFCDABCB4E3FC15FFF77F51AA118BFB4" author="Nubel-Reidelbach E." box="[228,513,381,405]" pageId="24" pageNumber="64" pagination="3 - 94" refId="ref15914" refString="Nubel-Reidelbach E. 1994 - Taxonomie und Systematik der Gattung Tectocepheus Berlese, 1895 (Acari, Oribatei) - Aandrias, Staatliches Museum fur Naturkunde Karlsrulhe, 12: 3 - 94." type="journal article" year="1994">Nübel-Reidelbach, 1994</bibRefCitation>
). The new species is similar in shape of long sharply pointed carina on trochantera III, and shape, length and situation of solenidia, famulus and seta
<emphasis id="B9280A284E3FC15FFE7CF581A2E6BFDF" box="[495,511,486,510]" italics="true" pageId="24" pageNumber="64">ft</emphasis>
’ on tarsus I to
<taxonomicName id="4C5CADB94E3FC15FFD2BF580A10CBC00" authority="Nakamura et al., 2010" authorityName="Nakamura" authorityYear="2010" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="64" phylum="Arthropoda" rank="species" species="kumayaensis">
<emphasis id="B9280A284E3FC15FFD2BF580A217BC00" italics="true" pageId="24" pageNumber="64">T. kumayaensis</emphasis>
<bibRefCitation id="EFCDABCB4E3FC15FFE86F66EA10CBC00" author="Nakamura Y. - N. &amp; Fukumori S. &amp; Fujikawa T." box="[277,533,521,545]" pageId="24" pageNumber="64" pagination="439 - 477" refId="ref15794" refString="Nakamura Y. - N., Fukumori S., Fujikawa T. 2010 - Oribatid fauna (Acari, Oribatida) from the Kumaya Cave of Iheya village in Central Ryukyu Arc, South Japan, with a description of several new species - Acarologia, 50 (4): 439 - 477. doi: 10.1051 / acarologia / 20101988" type="journal article" year="2010">
Nakamura
<emphasis id="B9280A284E3FC15FFE09F66DA2D6BC00" box="[410,463,521,545]" italics="true" pageId="24" pageNumber="64">et al.</emphasis>
, 2010
</bibRefCitation>
</taxonomicName>
. However, the new species differs from the latter in having a round rostrum, dentate cuspides, lyrifissures
<emphasis id="B9280A284E3FC15FFDDCF637A172BC46" box="[591,619,592,615]" italics="true" pageId="24" pageNumber="64">im</emphasis>
located laterally to opisthonotal gland
<emphasis id="B9280A284E3FC15FFE6DF615A107BCAB" box="[510,542,626,650]" italics="true" pageId="24" pageNumber="64">gla</emphasis>
and genital setae aligned almost in a line. The new species is distinguished from its congeners by shape of rostral region, cuspides, dorsosejugal region with long dorsophragmatic apophysis, humeral region, carina on trochantera III and IV, and solenidia of legs.
</paragraph>
</subSubSection>
</treatment>
</document>
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