449 lines
54 KiB
XML
449 lines
54 KiB
XML
<document id="FDAC248A55588BFFDAB4D8FC18FD429A" ID="10.11646/zootaxa.4117.1.7" ID-CLB-Dataset="38972" ID-DOI="10.11646/zootaxa.4117.1.7" ID-GBIF-Dataset="051656e2-ffab-4859-86d6-0bab91ca4d45" ID-ISSN="1175-5326" ID-Zenodo-Dep="262039" ID-ZooBank="79144BE2-86B1-4B46-80EA-1CA5AF366A06" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464591578104" checkinUser="plazi" docAuthor="Jirkov, Igor A." docDate="2016" docId="03A8EC00FF9080733B92F8FCD2C9FDE4" docLanguage="en" docName="data.336DE1B5E4A8B7FA11C576B5A5F314E7.cached" docOrigin="Zootaxa 4117 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lysippe nikiti Jirkov, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="131" masterDocId="FF919478FF9180753B05FF80D112FFD9" masterDocTitle="Lysippe nikiti sp. nov. (Polychaeta, Terebellomorpha) from the deep Pacific and Indian Oceans with an amendment to the generic diagnosis of Lysippe" masterLastPageNumber="134" masterPageNumber="125" pageNumber="126" updateTime="1698662646010" updateUser="ExternalLinkService">
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<mods:mods id="0340E742AAEC72E161F33E4B6C5EC8A6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="22E5BEAC60E05EFC4107620ACBE8CEE6">
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<mods:title id="81B8064B6025A77752587D7E4AEC2FC3">Lysippe nikiti sp. nov. (Polychaeta, Terebellomorpha) from the deep Pacific and Indian Oceans with an amendment to the generic diagnosis of Lysippe</mods:title>
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</mods:titleInfo>
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<mods:name id="B73AE14109CA10F7685095C337DCB199" type="personal">
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<mods:role id="F0968044899F663A39BFB4F5C161F036">
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<mods:roleTerm id="20CEEAE919FD1CC2391DE02621EC45AB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ADAC2EE4439B4E059B52F4F5D241CFF1">Jirkov, Igor A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5C9DCA19C9B329DD19B36FCAFCF51136">text</mods:typeOfResource>
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<mods:relatedItem id="BDF81EA6FFF12B17EA4679EC0DDCD0E1" type="host">
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<mods:titleInfo id="F838C24ABCE4C36700A084F5C1D7D3AD">
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<mods:title id="A03984C4F8E5E824E8D927798C8FB95D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3E7C1B0E6EC2E58B756DE4C682FCCEDD">
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<mods:date id="3D01779E408A226A080708113AD6EBED">2016</mods:date>
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<mods:detail id="3C3DFA03B84F0A79748FC194383B83A2" type="volume">
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<mods:number id="3938CA86B9C329C55CF342499438E744">4117</mods:number>
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</mods:detail>
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<mods:detail id="81E1E8B35EFD9FDF6B8AA96880055B70" type="issue">
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<mods:number id="3013ADE987CE75CE010434C22A030783">1</mods:number>
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</mods:detail>
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<mods:extent id="C6F2C750D86C0DE4C2474B9D52F3F79A" unit="page">
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<mods:start id="997FF4E2C16BD8F9F1B3A9BD92DB61CA">125</mods:start>
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<mods:end id="D4A42D58B037B8ADA69E26A5E69A1F57">134</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="6202212F7494FAB8C5125352A021D2FB">journal article</mods:classification>
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<mods:identifier id="5C4D791B3CD5A5600F136EF03CC18596" type="CLB-Dataset">38972</mods:identifier>
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<mods:identifier id="790A968A47C8E1A3C8010148F6299AD1" type="DOI">10.11646/zootaxa.4117.1.7</mods:identifier>
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<mods:identifier id="9ED60F279090ABAF29CCBFF61BDCDF8E" type="GBIF-Dataset">051656e2-ffab-4859-86d6-0bab91ca4d45</mods:identifier>
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<mods:identifier id="DABA966745E996A65B1E4192644C1C31" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0A168A99D38C5CE5ACB1686B6DE90516" type="Zenodo-Dep">262039</mods:identifier>
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<mods:identifier id="DA23E511A1B22A2DA3635C7799C39BC6" type="ZooBank">79144BE2-86B1-4B46-80EA-1CA5AF366A06</mods:identifier>
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</mods:mods>
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<treatment id="03A8EC00FF9080733B92F8FCD2C9FDE4" ID-DOI="http://doi.org/10.5281/zenodo.5680570" ID-GBIF-Taxon="120733358" ID-Zenodo-Dep="5680570" LSID="urn:lsid:plazi:treatment:03A8EC00FF9080733B92F8FCD2C9FDE4" httpUri="http://treatment.plazi.org/id/03A8EC00FF9080733B92F8FCD2C9FDE4" lastPageId="6" lastPageNumber="131" pageId="1" pageNumber="126">
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<subSubSection id="C31B0E9DFF9080743B92F8FCD08CF84F" box="[151,414,1916,1942]" pageId="1" pageNumber="126" type="nomenclature">
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<paragraph id="8BBE5D16FF9080743B92F8FCD08CF84F" blockId="1.[151,414,1916,1942]" box="[151,414,1916,1942]" pageId="1" pageNumber="126">
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<heading id="D0F6EA7AFF9080743B92F8FCD08CF84F" bold="true" box="[151,414,1916,1942]" fontSize="11" level="1" pageId="1" pageNumber="126" reason="1">
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<emphasis id="B9758104FF9080743B92F8FCD08CF84F" bold="true" box="[151,414,1916,1942]" pageId="1" pageNumber="126">
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<taxonomicName id="4C012695FF9080743B92F8FCD02BF84F" ID-CoL="6R2SQ" box="[151,313,1916,1942]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="126" phylum="Annelida" rank="species" species="nikiti" status="sp. nov.">
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<emphasis id="B9758104FF9080743B92F8FCD02BF84F" bold="true" box="[151,313,1916,1942]" italics="true" pageId="1" pageNumber="126">Lysippe nikiti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2463C7FFF9080743A44F8FCD08CF84F" box="[321,414,1916,1942]" pageId="1" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B0E9DFF9080773B92F843D364FEC4" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126" type="materials_examined">
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<paragraph id="8BBE5D16FF9080773B92F843D364FEC4" blockId="1.[151,1436,1987,2012]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126">
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<emphasis id="B9758104FF9080743B92F843D02DF805" bold="true" box="[151,319,1987,2012]" pageId="1" pageNumber="126">
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<typeStatus id="54BAE3B4FF9080743B92F843D1C0F805" box="[151,210,1987,2012]" pageId="1" pageNumber="126">Type</typeStatus>
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material
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</emphasis>
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. All from R/V
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<emphasis id="B9758104FF9080743AEAF844D33EF802" box="[495,556,1988,2011]" italics="true" pageId="1" pageNumber="126">Vitjaz</emphasis>
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.
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<typeStatus id="54BAE3B4FF9080743938F844D3C7F805" box="[573,725,1988,2012]" pageId="1" pageNumber="126" type="holotype">HOLOTYPE</typeStatus>
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: cruise 39, station 5625,
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<geoCoordinate id="EE353BD1FF90807438F4F843D51DF805" box="[1009,1039,1987,2012]" direction="north" orientation="latitude" pageId="1" pageNumber="126" precision="925" value="45.383335">45</geoCoordinate>
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°23' N
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<geoCoordinate id="EE353BD1FF9080743F5BF843D5C2F802" box="[1118,1232,1987,2012]" direction="east" orientation="longitude" pageId="1" pageNumber="126" precision="925" value="153.76666">153°46' E</geoCoordinate>
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,
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<quantity id="4CF9F0F3FF9080743FD8F843D427F802" box="[1245,1333,1987,2012]" metricMagnitude="3" metricUnit="m" metricValue="6.21" pageId="1" pageNumber="126" unit="m" value="6210.0">6210 m</quantity>
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, ‘Ocean’ grab,
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<specimenCount id="9D07969FFF9380773BD2FF18D049FF69" box="[215,347,151,176]" pageId="2" pageNumber="127" type="generic">1 specimen</specimenCount>
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;
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<typeStatus id="54BAE3B4FF9380773A63FF18D317FF69" box="[358,517,152,176]" pageId="2" pageNumber="127" type="paratype">PARATYPES</typeStatus>
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: cruise 29, station 4158,
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<geoCoordinate id="EE353BD1FF938077381AFF17D22FFF69" box="[799,829,151,176]" direction="north" orientation="latitude" pageId="2" pageNumber="127" precision="925" value="46.933334">46</geoCoordinate>
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°56' N
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<geoCoordinate id="EE353BD1FF938077388EFF17D511FF76" box="[907,1027,151,176]" direction="west" orientation="longitude" pageId="2" pageNumber="127" precision="925" value="-143.96666">143°58' W</geoCoordinate>
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,
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<quantity id="4CF9F0F3FF9380773F0AFF17D575FF76" box="[1039,1127,151,176]" metricMagnitude="3" metricUnit="m" metricValue="4.653" pageId="2" pageNumber="127" unit="m" value="4653.0">4653 m</quantity>
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, ‘Ocean’ grab,
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<specimenCount id="9D07969FFF9380773E13FF18D48AFF69" box="[1302,1432,151,176]" pageId="2" pageNumber="127" type="generic">1 specimen</specimenCount>
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, anterior fragment; cruise 31, station 4667,
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<geoCoordinate id="EE353BD1FF9380773973FF3CD391FF0D" box="[630,643,188,212]" direction="south" orientation="latitude" pageId="2" pageNumber="127" precision="925" value="-8.783334">8</geoCoordinate>
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°47' S
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<geoCoordinate id="EE353BD1FF93807739CAFF3CD220FF0D" box="[719,818,188,213]" direction="east" orientation="longitude" pageId="2" pageNumber="127" precision="925" value="52.4">52°24' E</geoCoordinate>
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,
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<quantity id="4CF9F0F3FF938077383BFF3CD287FF0D" box="[830,917,188,212]" metricMagnitude="3" metricUnit="m" metricValue="4.208" pageId="2" pageNumber="127" unit="m" value="4208.0">4208 m</quantity>
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, ‘Ocean’ grab,
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<specimenCount id="9D07969FFF9380773F46FF3CD5D4FF0C" box="[1091,1222,188,213]" pageId="2" pageNumber="127" type="generic">1 specimen</specimenCount>
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, anterior fragment; cruise 45, station 6136,
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<geoCoordinate id="EE353BD1FF9380773AA0FF5FD0D1FF21" box="[421,451,223,248]" direction="north" orientation="latitude" pageId="2" pageNumber="127" precision="925" value="53.416668">53</geoCoordinate>
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°25' N
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<geoCoordinate id="EE353BD1FF9380773914FF5FD398FF2E" box="[529,650,223,248]" direction="west" orientation="longitude" pageId="2" pageNumber="127" precision="925" value="-163.46666">163°28' W</geoCoordinate>
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,
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<quantity id="4CF9F0F3FF9380773993FF5FD3FFFF2E" box="[662,749,223,248]" metricMagnitude="3" metricUnit="m" metricValue="4.26" pageId="2" pageNumber="127" unit="m" value="4260.0">4260 m</quantity>
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, ‘Sigsbi’ trawl,
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<specimenCount id="9D07969FFF93807738A7FF60D53DFF21" box="[930,1071,223,248]" pageId="2" pageNumber="127" type="generic">3 specimens</specimenCount>
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, two anterior fragments and one complete specimen retained as SEM stubs.
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</paragraph>
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</subSubSection>
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<caption id="DF7E0D9EFF9380773B92FA26D2BEF9F9" httpUri="https://zenodo.org/record/262040/files/figure.png" pageId="2" pageNumber="127" targetBox="[153,1433,339,1422]" targetPageId="2">
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<paragraph id="8BBE5D16FF9380773B92FA26D2BEF9F9" blockId="2.[151,1436,1446,1568]" pageId="2" pageNumber="127">
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<emphasis id="B9758104FF9380773B92FA26D001FA62" bold="true" box="[151,275,1446,1468]" pageId="2" pageNumber="127">FIGURE 1.</emphasis>
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<taxonomicName id="4C012695FF9380773A1FFA27D0A4FA65" box="[282,438,1446,1468]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="127" phylum="Annelida" rank="species" species="labiata">
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<emphasis id="B9758104FF9380773A1FFA27D0A4FA65" box="[282,438,1446,1468]" italics="true" pageId="2" pageNumber="127">Lysippe labiata</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C012695FF9380773AEDFA27D378FA65" box="[488,618,1447,1468]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="127" phylum="Annelida" rank="species" species="sexcirrata">
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<emphasis id="B9758104FF9380773AEDFA27D378FA65" box="[488,618,1447,1468]" italics="true" pageId="2" pageNumber="127">L. sexcirrata</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C012695FF9380773972FA27D3C8FA65" box="[631,730,1446,1468]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="127" phylum="Annelida" rank="species" species="labiata">
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<emphasis id="B9758104FF9380773972FA27D3C8FA65" box="[631,730,1446,1468]" italics="true" pageId="2" pageNumber="127">L. labiata</emphasis>
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</taxonomicName>
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A, B
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<emphasis id="B9758104FF9380773814FA27D235FA65" box="[785,807,1447,1468]" italics="true" pageId="2" pageNumber="127">—</emphasis>
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anterior of worms with withdrawn and everted nuchal organs; C
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<emphasis id="B9758104FF9380773BADFA49D1ACFA07" box="[168,190,1481,1502]" italics="true" pageId="2" pageNumber="127">—</emphasis>
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posterior end left view; D–E
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<emphasis id="B9758104FF9380773AE1FA49D0E8FA07" box="[484,506,1481,1502]" italics="true" pageId="2" pageNumber="127">—</emphasis>
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pygidia. Specimens from R/V
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<emphasis id="B9758104FF9380773831FA47D26DFA04" box="[820,895,1479,1501]" italics="true" pageId="2" pageNumber="127">Maslov</emphasis>
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cruise 10 station 49 69°29’ N 56°00’ E depth 38 m, bottom temperature +1,19°C. F
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<emphasis id="B9758104FF9380773AE4FA6AD0E5FA26" box="[481,503,1514,1535]" italics="true" pageId="2" pageNumber="127">—</emphasis>
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pygidium of
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<taxonomicName id="4C012695FF938077397BFA6AD213FA26" box="[638,769,1514,1535]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="127" phylum="Annelida" rank="species" species="sexcirrata">
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<emphasis id="B9758104FF938077397BFA6AD213FA26" box="[638,769,1514,1535]" italics="true" pageId="2" pageNumber="127">L. sexcirrata</emphasis>
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</taxonomicName>
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R/V
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<emphasis id="B9758104FF938077383EFA69D2BBFA26" box="[827,937,1513,1535]" italics="true" pageId="2" pageNumber="127">Sevastopol</emphasis>
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cruise 5 station 1157 64°56’ N 24°25’ W depth 155 m, bottom temperature +8,17°C. no
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<emphasis id="B9758104FF9380773929F98BD350F9F9" box="[556,578,1547,1568]" italics="true" pageId="2" pageNumber="127">—</emphasis>
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nuchal organs; np
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<emphasis id="B9758104FF93807739F3F98BD21EF9F9" box="[758,780,1547,1568]" italics="true" pageId="2" pageNumber="127">—</emphasis>
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nephridiopores.
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</paragraph>
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</caption>
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<subSubSection id="C31B0E9DFF9380733BC2F9CFD323FE51" lastPageId="6" lastPageNumber="131" pageId="2" pageNumber="127" type="description">
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<paragraph id="8BBE5D16FF9380763BC2F9CFD015FEC4" blockId="2.[151,1437,1615,2036]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="128" pageId="2" pageNumber="127">
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<emphasis id="B9758104FF9380773BC2F9CFD0F7F9B1" bold="true" box="[199,485,1615,1640]" pageId="2" pageNumber="127">
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Description of
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<typeStatus id="54BAE3B4FF9380773A7BF9CFD0F7F9B1" box="[382,485,1615,1640]" pageId="2" pageNumber="127" type="holotype">holotype</typeStatus>
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</emphasis>
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. (
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<figureCitation id="133A4193FF9380773AFAF9CFD35AF9B1" box="[511,584,1615,1640]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="2" pageNumber="127">Fig. 4</figureCitation>
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). Complete specimen
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<quantity id="4CF9F0F3FF9380773849F9D0D28DF9BE" box="[844,927,1616,1640]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="127" unit="mm" value="30.0">30 mm</quantity>
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in length with no evidence of reproductive material (macerated internally). Prostomium with median lobe encircled by surrounding lobe (
|
||
<figureCitation id="133A4193FF9380773FE9F9F3D421F955" box="[1260,1331,1651,1676]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="2" pageNumber="127">Fig. 4</figureCitation>
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A–C, E), without glandular ridges or eyespots but with a pair of large, brown pigmented spots. Buccal tentacles not observed (lost or retracted). Lower lip large, extends laterally (as is typical for
|
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<taxonomicName id="4C012695FF93807738BDF93CD51CF90A" box="[952,1038,1724,1747]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="127" phylum="Annelida" rank="genus">
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<emphasis id="B9758104FF93807738BDF93CD51CF90A" box="[952,1038,1724,1747]" italics="true" pageId="2" pageNumber="127">Lysippe</emphasis>
|
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</taxonomicName>
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), ventral surface with longitudinal furrows and ridges; ridges terminate in papillae. Paleal chaetae very small, 2 to 3 times shorter and thinner than notochaetae of second thoracic chaetiger. 3 pairs of branchiae in two transverse groups, separated by a gap that is wider than the diameter of single branchiophore; branchiae visible only as scars in
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<typeStatus id="54BAE3B4FF9380773F33F8A7D589F899" box="[1078,1179,1831,1856]" pageId="2" pageNumber="127" type="holotype">holotype</typeStatus>
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, branchiostyles lost. In each group, the second from the outermost branchiae originate from segment II, the outermost originate from segment III and the innermost originate from segment IV. Nephridial papillae or nephridiopores clearly visible: one pair on each of segments IV and V, dorsally from the notopodia, and a third behind the notopodia of segment VI (
|
||
<figureCitation id="133A4193FF9380773B9AF837D1F5F809" box="[159,231,1975,2000]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="2" pageNumber="127">Fig. 4</figureCitation>
|
||
C). 17 thoracic chaetigers (excluding paleal segment), starting from segment
|
||
<date id="FFBF7BD6FF9380773F6DF838D5A3F809" box="[1128,1201,1976,2000]" pageId="2" pageNumber="127" value="2014-03">III. 14</date>
|
||
thoracic uncinigers, starting from segment VI. Elevated or modified notopodia absent. 10 abdominal uncinigers without rudimentary notopodia or glandular pads. Thoracic neuropodia are tori; abdominal neuropodia are pinnuli (uncini at the margin of neuropodia) (
|
||
<figureCitation id="133A4193FF9280763A51FF3CD08EFF0D" box="[340,412,188,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="3" pageNumber="128">Fig. 4</figureCitation>
|
||
D). Pinnules without dorsal cirrus. Pygidium of
|
||
<typeStatus id="54BAE3B4FF92807638D5FF3CD520FF0C" box="[976,1074,188,213]" pageId="3" pageNumber="128" type="holotype">holotype</typeStatus>
|
||
slightly macerated, so that the presence or absence of anal cirri is not clear. Methyl blue staining pattern absent, except for random spots. Tube unknown.
|
||
</paragraph>
|
||
<caption id="DF7E0D9EFF9280763B92FCD9D41AFCB6" box="[151,1288,854,879]" httpUri="https://zenodo.org/record/262041/files/figure.png" pageId="3" pageNumber="128" targetBox="[150,1436,337,833]" targetPageId="3">
|
||
<paragraph id="8BBE5D16FF9280763B92FCD9D41AFCB6" blockId="3.[151,1288,854,879]" box="[151,1288,854,879]" pageId="3" pageNumber="128">
|
||
<emphasis id="B9758104FF9280763B92FCD9D000FCB7" bold="true" box="[151,274,857,879]" pageId="3" pageNumber="128">FIGURE 2.</emphasis>
|
||
Position of branchiae of 5
|
||
<emphasis id="B9758104FF928076391BFCD6D33BFCBB" bold="true" box="[542,553,854,866]" pageId="3" pageNumber="128">th</emphasis>
|
||
segment (filled grey) in different ampharetid genera (after Jirkov 2001).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBE5D16FF9280763BC2FC16D5FAFBE6" blockId="3.[151,1436,918,1230]" pageId="3" pageNumber="128">
|
||
<emphasis id="B9758104FF9280763BC2FC16D36BFC76" bold="true" box="[199,633,918,943]" pageId="3" pageNumber="128">
|
||
Additional features from
|
||
<typeStatus id="54BAE3B4FF9280763AFEFC16D366FC76" box="[507,628,918,943]" pageId="3" pageNumber="128" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
All other specimens either have paleae or have paleal notopodia from which the paleae have been lost (
|
||
<figureCitation id="133A4193FF928076391FFC3AD34FFC0A" box="[538,605,954,979]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="3" pageNumber="128">Fig. 4</figureCitation>
|
||
E). A pair of large brown pigmented spots is usually present. Two large nuchal organs in posterior corners of the median lobe (
|
||
<figureCitation id="133A4193FF928076380BFC5ED241FC2E" box="[782,851,990,1015]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="3" pageNumber="128">Fig. 4</figureCitation>
|
||
H), though they may be completely withdrawn and invisible (
|
||
<figureCitation id="133A4193FF9280763A0DFB82D05FFBC2" box="[264,333,1026,1051]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="3" pageNumber="128">Fig. 4</figureCitation>
|
||
A, E). Uncini of thoracic uncinigers with numerous teeth in three to four rows, abdominal uncini with five to six rows but the number of teeth in each is less than in the thoracic uncini (
|
||
<figureCitation id="133A4193FF9280763F63FBA6D5BAFBE6" box="[1126,1192,1062,1087]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="3" pageNumber="128">Fig. 4</figureCitation>
|
||
F, G).
|
||
</paragraph>
|
||
<paragraph id="8BBE5D16FF9280763BC2FBCAD1FFFB17" blockId="3.[151,1436,918,1230]" pageId="3" pageNumber="128">
|
||
Despite obvious variation in presence/absence of paleae and shape of the lower lip, I consider all specimens to belong to a single species as they have the same number of branchiae and either characteristic brown pigmented spots or paleal notopodia without chaetae. Variation in lower lip shape does not seem to exceed that of
|
||
<taxonomicName id="4C012695FF9280763E40FB13D1F4FB17" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="labiata">
|
||
<emphasis id="B9758104FF9280763E40FB13D1F4FB17" italics="true" pageId="3" pageNumber="128">Lysippe labiata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF7E0D9EFF9280763B92FB7ED2AAFAEC" ID-Table-UUID="DF7E0D9EFF9280763B92FB7ED2AAFAEC" httpUri="http://table.plazi.org/id/DF7E0D9EFF9280763B92FB7ED2AAFAEC" pageId="3" pageNumber="128" targetBox="[159,1379,1402,1962]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="8BBE5D16FF9280763B92FB7ED2AAFAEC" blockId="3.[151,1436,1278,1333]" pageId="3" pageNumber="128">
|
||
<emphasis id="B9758104FF9280763B92FB7ED01BFACC" bold="true" box="[151,265,1278,1301]" pageId="3" pageNumber="128">TABLE 1.</emphasis>
|
||
Diagnostic characters of
|
||
<emphasis id="B9758104FF928076391DFB7FD362FACF" box="[536,624,1279,1302]" italics="true" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF928076391DFB7FD37EFACF" box="[536,620,1279,1302]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="genus">Lysippe</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Species are ordered firstly by the number of thoracic uncinigers, then by the number of pairs of branchiae, then by the number of abdominal uncinigers.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBE5D16FF9280733B9AFAD1D323FE51" blockId="3.[159,1379,1361,1383]" lastBlockId="6.[151,1436,151,573]" lastPageId="6" lastPageNumber="131" pageId="3" pageNumber="128">
|
||
species paleae thoracic uncinigers pairs of branchiae abdominal uncinigers
|
||
<emphasis id="B9758104FF9780733BC2FF17D0D4FF69" bold="true" box="[199,454,151,176]" pageId="6" pageNumber="131">Differential diagnosis</emphasis>
|
||
. The difference between the new species and previously described species is clear from
|
||
<tableCitation id="C68368ADFF9780733B92FF3CD1F9FF0D" box="[151,235,188,213]" captionStart="TABLE 1" captionStartId="3.[151,239,1278,1301]" captionTargetBox="[159,1379,1402,1962]" captionTargetPageId="3" captionText="TABLE 1. Diagnostic characters of Lysippe. Species are ordered firstly by the number of thoracic uncinigers, then by the number of pairs of branchiae, then by the number of abdominal uncinigers." httpUri="http://table.plazi.org/id/DF7E0D9EFF9280763B92FB7ED2AAFAEC" pageId="6" pageNumber="131" tableUuid="DF7E0D9EFF9280763B92FB7ED2AAFAEC">Table 1</tableCitation>
|
||
.
|
||
<taxonomicName id="4C012695FF9780733BFCFF3DD077FF0D" box="[249,357,188,212]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="131" phylum="Annelida" rank="species" species="storchi">
|
||
<emphasis id="B9758104FF9780733BFCFF3DD077FF0D" box="[249,357,188,212]" italics="true" pageId="6" pageNumber="131">L. storchi</emphasis>
|
||
</taxonomicName>
|
||
is the most morphologically similar species. It has the same number of thoracic uncinigers and pairs of branchiae but has 11 abdominal uncinigers, while the new species only 10. The difference between the species’ ranges makes it unlikely that they are a single species with wide geographic variation.
|
||
<taxonomicName id="4C012695FF9780733FD5FE85D429FEC5" box="[1232,1339,260,284]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="131" phylum="Annelida" rank="species" species="storchi">
|
||
<emphasis id="B9758104FF9780733FD5FE85D429FEC5" box="[1232,1339,260,284]" italics="true" pageId="6" pageNumber="131">L. storchi</emphasis>
|
||
</taxonomicName>
|
||
inhabits shallow, tropical waters, while the new species is limited to abyssal depths. Other species differ from the new one by at least two, sometimes three, of the following characters: number of thoracic uncinigers, number of abdominal uncinigers and number of branchiae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph id="8BBE5D16FF9280763B9AFAFBD5A3F873" pageId="3" pageNumber="128">
|
||
<table id="F901AFB6FF927F8A3B9AFAFAD471F873" box="[159,1379,1402,1962]" gridcols="4" gridrows="14" pageId="3" pageNumber="128">
|
||
<tr id="35315F54FF927F8A3B9AFAFAD471FA49" box="[159,1379,1402,1424]" gridrow="0" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AFAFAD040FA49" box="[159,338,1402,1424]" gridcol="0" gridrow="0" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AFAFBD009FA49" box="[159,283,1403,1424]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="mexicana">
|
||
<emphasis id="B9758104FF9280763B9AFAFBD009FA49" box="[159,283,1403,1424]" italics="true" pageId="3" pageNumber="128">L. mexicana</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76E03628FF927F8A3922FAFAD3F7FA49" box="[551,741,1402,1424]" gridcol="1" gridrow="0" pageId="3" pageNumber="128">+ 1 2</th>
|
||
<th id="76E03628FF927F8A38B5FAFAD2AFFA49" box="[944,957,1402,1424]" gridcol="2" gridrow="0" pageId="3" pageNumber="128">4</th>
|
||
<th id="76E03628FF927F8A3F8CFAFAD471FA49" box="[1161,1379,1402,1424]" gridcol="3" gridrow="0" pageId="3" pageNumber="128">1 1–1 2</th>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AFA24D471FA63" box="[159,1379,1444,1466]" gridrow="1" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AFA24D040FA63" box="[159,338,1444,1466]" gridcol="0" gridrow="1" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AFA25D02FFA63" box="[159,317,1444,1466]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="agulhasensis">
|
||
<emphasis id="B9758104FF9280763B9AFA25D02FFA63" box="[159,317,1444,1466]" italics="true" pageId="3" pageNumber="128">L. agulhasensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922FA24D3F7FA63" box="[551,741,1444,1466]" gridcol="1" gridrow="1" pageId="3" pageNumber="128">+ 1 2</td>
|
||
<td id="76E03628FF927F8A38B5FA24D2AFFA63" box="[944,957,1444,1466]" gridcol="2" gridrow="1" pageId="3" pageNumber="128">4</td>
|
||
<td id="76E03628FF927F8A3F8CFA24D471FA63" box="[1161,1379,1444,1466]" gridcol="3" gridrow="1" pageId="3" pageNumber="128">1 2</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AFA4DD471FA3A" box="[159,1379,1485,1507]" gridrow="2" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AFA4DD040FA3A" box="[159,338,1485,1507]" gridcol="0" gridrow="2" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AFA4ED010FA3A" box="[159,258,1485,1507]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="labiata">
|
||
<emphasis id="B9758104FF9280763B9AFA4ED010FA3A" box="[159,258,1485,1507]" italics="true" pageId="3" pageNumber="128">L. labiata</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922FA4DD3F7FA3A" box="[551,741,1485,1507]" gridcol="1" gridrow="2" pageId="3" pageNumber="128">+ 13</td>
|
||
<td id="76E03628FF927F8A38B5FA4DD2AFFA3A" box="[944,957,1485,1507]" gridcol="2" gridrow="2" pageId="3" pageNumber="128">4</td>
|
||
<td id="76E03628FF927F8A3F8CFA4DD471FA3A" box="[1161,1379,1485,1507]" gridcol="3" gridrow="2" pageId="3" pageNumber="128">14–15</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AFA76D471F9D4" box="[159,1379,1526,1549]" gridrow="3" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AFA76D040F9D4" box="[159,338,1526,1549]" gridcol="0" gridrow="3" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AFA78D040F9D5" box="[159,338,1526,1549]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="trichobranchia">
|
||
<emphasis id="B9758104FF9280763B9AFA78D040F9D5" box="[159,338,1526,1549]" italics="true" pageId="3" pageNumber="128">L. trichobranchia</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922FA76D3F7F9D4" box="[551,741,1526,1549]" gridcol="1" gridrow="3" pageId="3" pageNumber="128">– 13</td>
|
||
<td id="76E03628FF927F8A38B5FA76D2AFF9D4" box="[944,957,1526,1549]" gridcol="2" gridrow="3" pageId="3" pageNumber="128">4</td>
|
||
<td id="76E03628FF927F8A3F8CFA76D471F9D4" box="[1161,1379,1526,1549]" gridcol="3" gridrow="3" pageId="3" pageNumber="128">16–19</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF9A0D471F9EF" box="[159,1379,1568,1590]" gridrow="4" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF9A0D040F9EF" box="[159,338,1568,1590]" gridcol="0" gridrow="4" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF9A1D00DF9EF" box="[159,287,1569,1590]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="nipponica">
|
||
<emphasis id="B9758104FF9280763B9AF9A1D00DF9EF" box="[159,287,1569,1590]" italics="true" pageId="3" pageNumber="128">L. nipponica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F9A0D3F7F9EF" box="[551,741,1568,1590]" gridcol="1" gridrow="4" pageId="3" pageNumber="128">+ 1 3</td>
|
||
<td id="76E03628FF927F8A38B5F9A0D2AFF9EF" box="[944,957,1568,1590]" gridcol="2" gridrow="4" pageId="3" pageNumber="128">4</td>
|
||
<td id="76E03628FF927F8A3F8CF9A0D471F9EF" box="[1161,1379,1568,1590]" gridcol="3" gridrow="4" pageId="3" pageNumber="128">? (>5)</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF9C9D471F986" box="[159,1379,1609,1631]" gridrow="5" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF9C9D040F986" box="[159,338,1609,1631]" gridcol="0" gridrow="5" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF9CAD1FCF986" box="[159,238,1609,1631]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="nikiti">
|
||
<emphasis id="B9758104FF9280763B9AF9CAD1FCF986" box="[159,238,1609,1631]" italics="true" pageId="3" pageNumber="128">L. nikiti</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F9C9D3F7F986" box="[551,741,1609,1631]" gridcol="1" gridrow="5" pageId="3" pageNumber="128">± 1 4</td>
|
||
<td id="76E03628FF927F8A38B5F9C9D2AFF986" box="[944,957,1609,1631]" gridcol="2" gridrow="5" pageId="3" pageNumber="128">3</td>
|
||
<td id="76E03628FF927F8A3F8CF9C9D471F986" box="[1161,1379,1609,1631]" gridcol="3" gridrow="5" pageId="3" pageNumber="128">1 0</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF9F2D471F951" box="[159,1379,1650,1672]" gridrow="6" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF9F2D040F951" box="[159,338,1650,1672]" gridcol="0" gridrow="6" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF9F3D1EDF951" box="[159,255,1650,1672]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="storchi">
|
||
<emphasis id="B9758104FF9280763B9AF9F3D1EDF951" box="[159,255,1650,1672]" italics="true" pageId="3" pageNumber="128">L. storchi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F9F2D3F7F951" box="[551,741,1650,1672]" gridcol="1" gridrow="6" pageId="3" pageNumber="128">- 1 4</td>
|
||
<td id="76E03628FF927F8A38B5F9F2D2AFF951" box="[944,957,1650,1672]" gridcol="2" gridrow="6" pageId="3" pageNumber="128">3</td>
|
||
<td id="76E03628FF927F8A3F8CF9F2D471F951" box="[1161,1379,1650,1672]" gridcol="3" gridrow="6" pageId="3" pageNumber="128">1 1</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF91CD471F96B" box="[159,1379,1692,1714]" gridrow="7" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF91CD040F96B" box="[159,338,1692,1714]" gridcol="0" gridrow="7" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF91DD033F96B" box="[159,289,1693,1714]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="sexcirrata">
|
||
<emphasis id="B9758104FF9280763B9AF91DD033F96B" box="[159,289,1693,1714]" italics="true" pageId="3" pageNumber="128">L. sexcirrata</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F91CD3F7F96B" box="[551,741,1692,1714]" gridcol="1" gridrow="7" pageId="3" pageNumber="128">+ 1 4</td>
|
||
<td id="76E03628FF927F8A38B5F91CD2AFF96B" box="[944,957,1692,1714]" gridcol="2" gridrow="7" pageId="3" pageNumber="128">3</td>
|
||
<td id="76E03628FF927F8A3F8CF91CD471F96B" box="[1161,1379,1692,1714]" gridcol="3" gridrow="7" pageId="3" pageNumber="128">1 3</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF945D471F902" box="[159,1379,1733,1755]" gridrow="8" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF945D040F902" box="[159,338,1733,1755]" gridcol="0" gridrow="8" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF946D020F902" box="[159,306,1733,1755]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="annenkovae">
|
||
<emphasis id="B9758104FF9280763B9AF946D020F902" box="[159,306,1733,1755]" italics="true" pageId="3" pageNumber="128">L. annenkovae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F945D3F7F902" box="[551,741,1733,1755]" gridcol="1" gridrow="8" pageId="3" pageNumber="128">- 1 4</td>
|
||
<td id="76E03628FF927F8A38B5F945D2AFF902" box="[944,957,1733,1755]" gridcol="2" gridrow="8" pageId="3" pageNumber="128">3</td>
|
||
<td id="76E03628FF927F8A3F8CF945D471F902" box="[1161,1379,1733,1755]" gridcol="3" gridrow="8" pageId="3" pageNumber="128">1 5</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF96ED471F8DC" box="[159,1379,1774,1797]" gridrow="9" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF96ED040F8DC" box="[159,338,1774,1797]" gridcol="0" gridrow="9" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF970D03FF8DC" box="[159,301,1776,1797]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="gurjanovae">
|
||
<emphasis id="B9758104FF9280763B9AF970D03FF8DC" box="[159,301,1776,1797]" italics="true" pageId="3" pageNumber="128">L. gurjanovae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F96ED3F7F8DC" box="[551,741,1774,1797]" gridcol="1" gridrow="9" pageId="3" pageNumber="128">- 14</td>
|
||
<td id="76E03628FF927F8A38B5F96ED2AFF8DC" box="[944,957,1774,1797]" gridcol="2" gridrow="9" pageId="3" pageNumber="128">3</td>
|
||
<td id="76E03628FF927F8A3F8CF96ED471F8DC" box="[1161,1379,1774,1797]" gridcol="3" gridrow="9" pageId="3" pageNumber="128">17–19</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF898D471F8F7" box="[159,1379,1816,1838]" gridrow="10" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF898D040F8F7" box="[159,338,1816,1838]" gridcol="0" gridrow="10" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF899D1EDF8F7" box="[159,255,1816,1838]" class="Polychaeta" family="Ampharetidae" genus="Lysippides" higherTaxonomySource="CoL" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="hesslei">
|
||
<emphasis id="B9758104FF9280763B9AF899D1EDF8F7" box="[159,255,1816,1838]" italics="true" pageId="3" pageNumber="128">L. hesslei</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F898D3F7F8F7" box="[551,741,1816,1838]" gridcol="1" gridrow="10" pageId="3" pageNumber="128">- 14</td>
|
||
<td id="76E03628FF927F8A38B5F898D2AFF8F7" box="[944,957,1816,1838]" gridcol="2" gridrow="10" pageId="3" pageNumber="128">3</td>
|
||
<td id="76E03628FF927F8A3F8CF898D471F8F7" box="[1161,1379,1816,1838]" gridcol="3" gridrow="10" pageId="3" pageNumber="128">25–26</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF8C1D471F88E" box="[159,1379,1857,1879]" gridrow="11" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF8C1D040F88E" box="[159,338,1857,1879]" gridcol="0" gridrow="11" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF8C2D011F88E" box="[159,259,1857,1879]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="fragilis">
|
||
<emphasis id="B9758104FF9280763B9AF8C2D011F88E" box="[159,259,1857,1879]" italics="true" pageId="3" pageNumber="128">L. fragilis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F8C1D3F7F88E" box="[551,741,1857,1879]" gridcol="1" gridrow="11" pageId="3" pageNumber="128">+ 1 4</td>
|
||
<td id="76E03628FF927F8A38B5F8C1D2AFF88E" box="[944,957,1857,1879]" gridcol="2" gridrow="11" pageId="3" pageNumber="128">4</td>
|
||
<td id="76E03628FF927F8A3F8CF8C1D471F88E" box="[1161,1379,1857,1879]" gridcol="3" gridrow="11" pageId="3" pageNumber="128">8</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF8EBD471F858" box="[159,1379,1899,1921]" gridrow="12" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF8EBD040F858" box="[159,338,1899,1921]" gridcol="0" gridrow="12" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF8ECD1E7F858" box="[159,245,1900,1921]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="caeca">
|
||
<emphasis id="B9758104FF9280763B9AF8ECD1E7F858" box="[159,245,1900,1921]" italics="true" pageId="3" pageNumber="128">L. caeca</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F8EBD3F7F858" box="[551,741,1899,1921]" gridcol="1" gridrow="12" pageId="3" pageNumber="128">+ 1 4</td>
|
||
<td id="76E03628FF927F8A38B5F8EBD2AFF858" box="[944,957,1899,1921]" gridcol="2" gridrow="12" pageId="3" pageNumber="128">4</td>
|
||
<td id="76E03628FF927F8A3F8CF8EBD471F858" box="[1161,1379,1899,1921]" gridcol="3" gridrow="12" pageId="3" pageNumber="128">1 1</td>
|
||
</tr>
|
||
<tr id="35315F54FF927F8A3B9AF814D471F873" box="[159,1379,1940,1962]" gridrow="13" pageId="3" pageNumber="128">
|
||
<th id="76E03628FF927F8A3B9AF814D040F873" box="[159,338,1940,1962]" gridcol="0" gridrow="13" pageId="3" pageNumber="128">
|
||
<taxonomicName id="4C012695FF9280763B9AF815D00DF873" box="[159,287,1941,1962]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="128" phylum="Annelida" rank="species" species="annectens">
|
||
<emphasis id="B9758104FF9280763B9AF815D00DF873" box="[159,287,1941,1962]" italics="true" pageId="3" pageNumber="128">L. annectens</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76E03628FF927F8A3922F814D3F7F873" box="[551,741,1940,1962]" gridcol="1" gridrow="13" pageId="3" pageNumber="128">+ 1 4</td>
|
||
<td id="76E03628FF927F8A38B5F814D2AFF873" box="[944,957,1940,1962]" gridcol="2" gridrow="13" pageId="3" pageNumber="128">4</td>
|
||
<td id="76E03628FF927F8A3F8CF814D471F873" box="[1161,1379,1940,1962]" gridcol="3" gridrow="13" pageId="3" pageNumber="128">>1 0</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF7E0D9EFF9580713B92F8A5D3FAF8E3" box="[151,744,1828,1851]" httpUri="https://zenodo.org/record/262042/files/figure.png" pageId="4" pageNumber="129" targetBox="[185,1401,195,1807]" targetPageId="4">
|
||
<paragraph id="8BBE5D16FF9580713B92F8A5D3FAF8E3" blockId="4.[151,744,1828,1851]" box="[151,744,1828,1851]" pageId="4" pageNumber="129">
|
||
<emphasis id="B9758104FF9580713B92F8A5D000F8E3" bold="true" box="[151,274,1829,1851]" pageId="4" pageNumber="129">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C012695FF9580713A1CF8A6D0A7F8E3" box="[281,437,1828,1851]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="129" phylum="Annelida" rank="species" species="labiata">
|
||
<emphasis id="B9758104FF9580713A1CF8A6D0A7F8E3" box="[281,437,1828,1851]" italics="true" pageId="4" pageNumber="129">Lysippe labiata</emphasis>
|
||
</taxonomicName>
|
||
with everted buccal tentacles.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7E0D9EFF9480703B92F8ACD006F832" httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="5" pageNumber="130" targetBox="[250,1328,193,1815]" targetPageId="5">
|
||
<paragraph id="8BBE5D16FF9480703B92F8ACD006F832" blockId="5.[151,1436,1836,2027]" pageId="5" pageNumber="130">
|
||
<emphasis id="B9758104FF9480703B92F8ACD001F898" bold="true" box="[151,275,1836,1858]" pageId="5" pageNumber="130">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C012695FF9480703A1EF8ADD0B6F89B" box="[283,420,1836,1858]" class="Polychaeta" family="Ampharetidae" genus="Lysippe" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="130" phylum="Annelida" rank="species" species="nikiti" status="sp. nov.">
|
||
<emphasis id="B9758104FF9480703A1EF8ADD0B6F89B" box="[283,420,1836,1858]" italics="true" pageId="5" pageNumber="130">Lysippe nikiti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2463C7FFF9480703AAEF8ACD0EEF898" box="[427,508,1836,1857]" pageId="5" pageNumber="130" rank="species">
|
||
<emphasis id="B9758104FF9480703AAEF8ACD0EEF898" bold="true" box="[427,508,1836,1857]" pageId="5" pageNumber="130">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A
|
||
<emphasis id="B9758104FF9480703913F8ADD33EF89B" box="[534,556,1837,1858]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
anterior dorsal view perpendicular to prostomial dorsal surface; B
|
||
<emphasis id="B9758104FF9480703FCBF8ADD5F6F89B" box="[1230,1252,1837,1858]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
anterior left view; C
|
||
<emphasis id="B9758104FF9480703BADF8D1D1ACF8BF" box="[168,190,1873,1894]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
anterior dorso-lateral view; D
|
||
<emphasis id="B9758104FF9480703AF8F8D1D301F8BF" box="[509,531,1873,1894]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
last thoracic and 1
|
||
<emphasis id="B9758104FF94807039DAF8CDD3FAF880" bold="true" box="[735,744,1869,1881]" pageId="5" pageNumber="130">st</emphasis>
|
||
abdominal segments; E
|
||
<emphasis id="B9758104FF94807038EBF8D1D516F8BF" box="[1006,1028,1873,1894]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
dorsal view; F
|
||
<emphasis id="B9758104FF9480703FA7F8D1D5AAF8BF" box="[1186,1208,1873,1894]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
uncini of 5
|
||
<emphasis id="B9758104FF9480703E36F8CDD42CF880" bold="true" box="[1331,1342,1869,1881]" pageId="5" pageNumber="130">th</emphasis>
|
||
thoracic unciniger; G
|
||
<emphasis id="B9758104FF9480703A13F8F2D03EF85E" box="[278,300,1906,1927]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
uncini of mid abdomen; H
|
||
<emphasis id="B9758104FF9480703933F8F2D35EF85E" box="[566,588,1906,1927]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
anterior of worms with everted nuchal organs; I
|
||
<emphasis id="B9758104FF9480703F28F8F2D551F85E" box="[1069,1091,1906,1927]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
species range; A–D
|
||
<emphasis id="B9758104FF9480703E0CF8F2D40DF85E" box="[1289,1311,1906,1927]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
holotype, st. 5625; E
|
||
<emphasis id="B9758104FF9480703BECF814D1EDF870" box="[233,255,1940,1961]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
paratype, st. 4158; F–H
|
||
<emphasis id="B9758104FF9480703AF3F814D31EF870" box="[502,524,1940,1961]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
paratype, st. 6136. A–E
|
||
<emphasis id="B9758104FF9480703801F814D208F870" box="[772,794,1940,1961]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
methyl blue staining pattern. Numbers refer to segments: ll
|
||
<emphasis id="B9758104FF9480703E80F814D489F870" box="[1413,1435,1940,1961]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
lower lip, ml
|
||
<emphasis id="B9758104FF9480703A24F835D025F813" box="[289,311,1973,1994]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
median lobe of prostomium, ne —neuropodium; no
|
||
<emphasis id="B9758104FF948070385EF835D263F813" box="[859,881,1973,1994]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
nuchal organs; np
|
||
<emphasis id="B9758104FF9480703F28F835D551F813" box="[1069,1091,1973,1994]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
nephridiopores; nt—notopodium; pal
|
||
<emphasis id="B9758104FF9480703BB3F856D1DEF832" box="[182,204,2006,2027]" italics="true" pageId="5" pageNumber="130">—</emphasis>
|
||
paleae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31B0E9DFF9780733BC2FE14D47EFE2D" pageId="6" pageNumber="131" type="distribution">
|
||
<paragraph id="8BBE5D16FF9780733BC2FE14D47EFE2D" blockId="6.[151,1436,151,573]" pageId="6" pageNumber="131">
|
||
<emphasis id="B9758104FF9780733BC2FE14D04BFE74" bold="true" box="[199,345,404,429]" pageId="6" pageNumber="131">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="133A4193FF9780733A6FFE14D0BDFE75" box="[362,431,404,429]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[250,1328,193,1815]" captionTargetId="figure@5.[250,1328,193,1816]" captionTargetPageId="5" captionText="FIGURE 4. Lysippe nikiti sp. nov. A — anterior dorsal view perpendicular to prostomial dorsal surface; B — anterior left view; C — anterior dorso-lateral view; D — last thoracic and 1 st abdominal segments; E — dorsal view; F — uncini of 5 th thoracic unciniger; G — uncini of mid abdomen; H — anterior of worms with everted nuchal organs; I — species range; A – D — holotype, st. 5625; E — paratype, st. 4158; F – H — paratype, st. 6136. A – E — methyl blue staining pattern. Numbers refer to segments: ll — lower lip, ml — median lobe of prostomium, ne — neuropodium; no — nuchal organs; np — nephridiopores; nt—notopodium; pal — paleae." httpUri="https://zenodo.org/record/262043/files/figure.png" pageId="6" pageNumber="131">
|
||
Fig.
|
||
<date id="FFBF7BD6FF9780733A9AFE14D0BDFE75" box="[415,431,404,428]" pageId="6" pageNumber="131">4</date>
|
||
</figureCitation>
|
||
I). The species has one of the deepest ranges in the family (
|
||
<quantity id="4CF9F0F3FF9780733F51FE14D5E6FE75" box="[1108,1268,404,429]" metricMagnitude="3" metricUnit="m" metricValue="5.195" metricValueMax="6.21" metricValueMin="4.18" pageId="6" pageNumber="131" unit="m" value="5195.0" valueMax="6210.0" valueMin="4180.0">4180–6210 m</quantity>
|
||
) and is widely distributed in the North Pacific and Indian Oceans; its actual range is probably much wider, as such depths are not well known. Wide species distributions, such as this, are quite common amongst Polychaeta (
|
||
<bibRefCitation id="EF9020E7FF9780733FA9FE5CD472FE2D" author="Kucheruk" box="[1196,1376,476,501]" pageId="6" pageNumber="131" refString="Kucheruk, N. V. (1981) On the regularities of the geographycal distribution of the abyssal polychaetous annelids of the Eastern coast of Pacific Ocean. Transactions of P. P. Shirshov Institute of Oceanology of the Russian Academy of Science, 115, 37 - 52." type="journal article" year="1981">Kucheruk 1981</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B0E9DFF9780733BC2FE7FD2C9FDE4" pageId="6" pageNumber="131" type="etymology">
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<paragraph id="8BBE5D16FF9780733BC2FE7FD2C9FDE4" blockId="6.[151,1436,151,573]" pageId="6" pageNumber="131">
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<emphasis id="B9758104FF9780733BC2FE7FD054FDC1" bold="true" box="[199,326,511,536]" pageId="6" pageNumber="131">Etymology</emphasis>
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. The species is named after my late friend Nikita Kucheruk, who was, amongst his other specializations, an ampharetid taxonomist, describing several new species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |