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<document id="F02AED28A58C3E49BDC256229A66AF1E" ID-DOI="10.26107/RBZ-2019-0029" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577070" ID-ZooBank="0DBFFB4B-9418-4555-B593-A9EC4E8D70CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614783958724" checkinUser="carolina" docAuthor="Larson, Helen K. &amp; Hui, Tan Heok" docDate="2019" docId="FE268780FFF19A52E69AF822FD132BFB" docLanguage="en" docName="RafflesBZool.67.385-390.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Pogoneleotris heterolepis" docType="treatment" docVersion="3" lastPageNumber="389" masterDocId="021FFFF8FFF19A56E50CFFA2FFA52D03" masterDocTitle="A redescription of the bearded gudgeon, Pogoneleotris heterolepis (Günther, 1869) (Teleostei: Gobioidei: Butidae)" masterLastPageNumber="390" masterPageNumber="385" pageNumber="385" updateTime="1698878494446" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="FDD9E66A1E9894775CF5BDBF1667F673" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="801D5CE76BE0C8D09CEA68C93BCB9B9B">A redescription of the bearded gudgeon, Pogoneleotris heterolepis (Günther, 1869) (Teleostei: Gobioidei: Butidae)</mods:title>
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<mods:namePart id="6F5F64261DDCC9199CD2679B9ED27A64">Larson, Helen K.</mods:namePart>
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<mods:namePart id="DE552858DF9F930FF0957A62E7F40845">Hui, Tan Heok</mods:namePart>
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<mods:title id="F0850B0865787DD01610371E20F6E452">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="D9049EB579134D63C3D37D2EA5D1702D">
<mods:date id="F49590C5EC914320A7D032A99451AD1E">2019</mods:date>
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<mods:number id="E27684BC1F03A9E8F02093AC5CC53259">2019-07-09</mods:number>
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<mods:number id="0754005ABD16A7808444B6136DE4F0FB">67</mods:number>
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<mods:start id="E277EEF90607F02122130DEB59E964D3">385</mods:start>
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<mods:identifier id="118F2C52AD2E31B82AF9AD4F048E8A9B" type="DOI">10.26107/RBZ-2019-0029</mods:identifier>
<mods:identifier id="166C7742E24CA157680A8CDCEA03E7D5" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="09651540860F7093ED0AC2B41FA898B4" type="Zenodo-Dep">4577070</mods:identifier>
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<treatment id="FE268780FFF19A52E69AF822FD132BFB" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB" httpUri="http://treatment.plazi.org/id/FE268780FFF19A52E69AF822FD132BFB" lastPageId="4" lastPageNumber="389" pageId="0" pageNumber="385">
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<paragraph id="76303696FFF19A56E69AF822FACA2A9B" blockId="0.[918,1391,1920,1976]" box="[918,1391,1920,1944]" pageId="0" pageNumber="385">
<heading id="2D7881FAFFF19A56E69AF822FACA2A9B" bold="true" box="[918,1391,1920,1944]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="385" reason="2">
<taxonomicName id="B18F4D15FFF19A56E69AF822FACA2A9B" ID-CoL="784ZD" authority="(Gunther, 1869)" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[918,1391,1920,1944]" class="Actinopterygii" family="Eleotridae" genus="Pogoneleotris" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="385" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="44FBEA84FFF19A56E69AF822FACA2A9B" bold="true" box="[918,1391,1920,1944]" pageId="0" pageNumber="385">
<emphasis id="44FBEA84FFF19A56E69AF822FB0B2A9B" bold="true" box="[918,1198,1920,1944]" italics="true" pageId="0" pageNumber="385">Pogoneleotris heterolepis</emphasis>
(
<bibRefCitation id="121E4B67FFF19A56E1B2F822FACD2A9B" author="Gunther A" box="[1214,1384,1920,1944]" pageId="0" pageNumber="385" pagination="444 - 445" refId="ref3430" refString="Gunther A (1869) Descriptions of two new species of fishes discovered by the Marquis J. Doria. Annals and Magazine of Natural History, Series 4 (3): 444 - 445." type="journal article" year="1869">Günther, 1869</bibRefCitation>
)
</emphasis>
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<subSubSection id="3E95651DFFF19A56E14BF802FB1B2ABB" box="[1095,1214,1952,1976]" pageId="0" pageNumber="385" type="description">
<paragraph id="76303696FFF19A56E14BF802FB1B2ABB" blockId="0.[918,1391,1920,1976]" box="[1095,1214,1952,1976]" pageId="0" pageNumber="385">
(
<figureCitation id="EEB42A13FFF19A56E143F802FB132ABB" box="[1103,1206,1952,1976]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="1.[113,148,1173,1195]" captionStartId-1="2.[137,172,1512,1534]" captionStartId-2="2.[835,870,578,600]" captionStartId-3="4.[136,171,761,783]" captionTargetBox-0="[113,1451,189,1158]" captionTargetBox-1="[136,775,189,1496]" captionTargetBox-2="[835,1475,189,563]" captionTargetBox-3="[136,1474,189,746]" captionTargetId-0="figure-459@1.[113,1451,189,1158]" captionTargetId-1="figure-7@2.[136,775,189,1496]" captionTargetId-2="figure-558@2.[814,1508,166,590]" captionTargetId-3="figure-492@4.[136,1474,189,746]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="4" captionText-0="Fig. 1. Dorsal, lateral and ventral views of preserved male Pogoneleotris heterolepis (ZRC56949, 160.0 mm SL)." captionText-1="Fig. 2. Dorsal, lateral and ventral views (right side reversed) of head of fresh Pogoneleotris heterolepis (ZRC56949, 160.0 mm SL male); showing extent of scalation and sensory papillae." captionText-2="Fig. 3. Composite drawing of head showing lateral canals, sensory pores and papillae. Most papillae and flaps omitted for clarity, as are all scales." captionText-3="Fig. 4. Fresh Pogoneleotris heterolepis from Simunjan, Sarawak; ZRC56949, female above (166.7 mm SL), male below (160.0 mm SL)." figureDoi-0="http://doi.org/10.5281/zenodo.4577072" figureDoi-1="http://doi.org/10.5281/zenodo.4577076" figureDoi-2="http://doi.org/10.5281/zenodo.4577074" figureDoi-3="http://doi.org/10.5281/zenodo.4577078" httpUri-0="https://zenodo.org/record/4577072/files/figure.png" httpUri-1="https://zenodo.org/record/4577076/files/figure.png" httpUri-2="https://zenodo.org/record/4577074/files/figure.png" httpUri-3="https://zenodo.org/record/4577078/files/figure.png" pageId="0" pageNumber="385">Figs. 1–4</figureCitation>
)
</paragraph>
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<subSubSection id="3E95651DFFF19A57E64FF87DFE0D2884" lastPageId="1" lastPageNumber="386" pageId="0" pageNumber="385" type="reference_group">
<paragraph id="76303696FFF19A56E64FF87DFB852512" blockId="0.[835,1475,2014,2065]" pageId="0" pageNumber="385">
<treatmentCitationGroup id="569F11B8FFF19A56E64FF87DFB852512" pageId="0" pageNumber="385">
<treatmentCitation id="F72E1087FFF19A56E64FF87DFB6E2AF7" author="Gunther A" box="[835,1227,2014,2037]" page="445" pageId="0" pageNumber="385" year="1869">
<taxonomicName id="B18F4D15FFF19A56E64FF87DFB6E2AF7" ID-CoL="397ZW" authority="Gunther, 1869: 445" authorityName="Gunther" authorityPageNumber="445" authorityYear="1869" box="[835,1227,2014,2037]" class="Actinopterygii" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="385" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="44FBEA84FFF19A56E64FF87DFBA62AF7" box="[835,1027,2015,2036]" italics="true" pageId="0" pageNumber="385">Eleotris heterolepis</emphasis>
<materialsCitation id="C6E73CCBFFF19A56E107F87DFB6E2AF7" box="[1035,1227,2014,2037]" country="Malaysia" location="Borneo" pageId="0" pageNumber="385" specimenCount="1">
<bibRefCitation id="121E4B67FFF19A56E107F87DFB6E2AF7" author="Gunther A" box="[1035,1227,2014,2037]" pageId="0" pageNumber="385" pagination="444 - 445" refId="ref3430" refString="Gunther A (1869) Descriptions of two new species of fishes discovered by the Marquis J. Doria. Annals and Magazine of Natural History, Series 4 (3): 444 - 445." type="journal article" year="1869">Günther, 1869: 445</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="A9348834FFF19A56E1D5F87DFAA12AF7" box="[1241,1284,2015,2036]" pageId="0" pageNumber="385">type</typeStatus>
locality:
<collectingCountry id="0E987606FFF19A56E06FF87DFA1A2AF6" box="[1379,1471,2015,2037]" name="Malaysia" pageId="0" pageNumber="385">Malaysia</collectingCountry>
:
<location id="7350604DFFF19A56E665F85EFC162512" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB:7350604DFFF19A56E665F85EFC162512" box="[873,947,2044,2065]" country="Malaysia" name="Borneo" pageId="0" pageNumber="385">Borneo</location>
: Sarawak).
</treatmentCitationGroup>
</paragraph>
<caption id="22F0661EFFF09A57E57DFB37FB6829A8" ID-DOI="http://doi.org/10.5281/zenodo.4577072" ID-Zenodo-Dep="4577072" box="[113,1229,1173,1195]" httpUri="https://zenodo.org/record/4577072/files/figure.png" pageId="1" pageNumber="386" startId="1.[113,148,1173,1195]" targetBox="[113,1451,189,1158]" targetPageId="1">
<paragraph id="76303696FFF09A57E57DFB37FB6829A8" blockId="1.[113,1229,1173,1195]" box="[113,1229,1173,1195]" pageId="1" pageNumber="386">
Fig. 1. Dorsal, lateral and ventral views of preserved male
<taxonomicName id="B18F4D15FFF09A57E7BAFB37FC1529A9" ID-CoL="784ZD" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[694,944,1173,1194]" class="Actinopterygii" family="Eleotridae" genus="Pogoneleotris" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="386" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="44FBEA84FFF09A57E7BAFB37FC1529A9" box="[694,944,1173,1194]" italics="true" pageId="1" pageNumber="386">Pogoneleotris heterolepis</emphasis>
</taxonomicName>
(ZRC56949, 160.0 mm SL).
</paragraph>
</caption>
<paragraph id="76303696FFF09A57E57DFB40FE0D2884" blockId="1.[113,753,1250,1416]" pageId="1" pageNumber="386">
<treatmentCitationGroup id="569F11B8FFF09A57E57DFB40FE0D2884" pageId="1" pageNumber="386">
<taxonomicName id="B18F4D15FFF09A57E57DFB40FE2C29F4" ID-CoL="784ZD" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[113,393,1250,1271]" class="Actinopterygii" family="Eleotridae" genus="Pogoneleotris" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="386" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="44FBEA84FFF09A57E57DFB40FE2C29F4" box="[113,393,1250,1271]" italics="true" pageId="1" pageNumber="386">Pogoneleotris heterolepis</emphasis>
</taxonomicName>
–
<treatmentCitation id="F72E1087FFF09A57E4A7FB40FDD829F4" author="Bleeker P" box="[427,637,1250,1272]" page="107" pageId="1" pageNumber="386" year="1875">
<bibRefCitation id="121E4B67FFF09A57E4A7FB40FDD829F4" author="Bleeker P" box="[427,637,1250,1272]" pageId="1" pageNumber="386" pagination="101 - 112" refId="ref3350" refString="Bleeker P (1875) Notice sur les Eleotriformes et description de trois especes nouvelles. Archives Neerlandaises des Sciences Exactes et Naturelles, 10: 101 - 112." type="journal article" year="1875">Bleeker, 1875: 107</bibRefCitation>
</treatmentCitation>
(Borneo);
<treatmentCitation id="F72E1087FFF09A57E594FB5DFEDD2816" author="Vinciguerra D" box="[152,376,1279,1301]" page="543" pageId="1" pageNumber="386" year="1926">
<bibRefCitation id="121E4B67FFF09A57E594FB5DFEDD2816" author="Vinciguerra D" box="[152,376,1279,1301]" pageId="1" pageNumber="386" pagination="532 - 628" refId="ref3853" refString="Vinciguerra D (1926) Catalogo dei pesci raccolti a Borneo dais igg. Marchese G. Doria e dott. O. Beccari negli anni 1865 - 67. Annali del Museo Civico di Storia Naturale Giacoma Doria, Genova, (3) 10: 532 - 628, tavl. 1." type="journal article" year="1926">Vinciguerra, 1926: 543</bibRefCitation>
</treatmentCitation>
, pl. 1;
<treatmentCitation id="F72E1087FFF09A57E4BBFB5DFD272817" author="Koumans FP" box="[439,642,1279,1301]" page="323" pageId="1" pageNumber="386" year="1953">
<bibRefCitation id="121E4B67FFF09A57E4BBFB5DFD272817" author="Koumans FP" box="[439,642,1279,1301]" pageId="1" pageNumber="386" pagination="1 - 423" refId="ref3572" refString="Koumans FP (1953) Gobioidea. Vol. X. In: Weber M &amp; de Beaufort LF (eds.) The Fishes of the Indo-Australian archipelago. Brill, Leiden. Pp. 1 - 423." type="book chapter" year="1953">Koumans, 1953: 323</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F72E1087FFF09A57E787FB5DFF582832" author="Tortonese E" page="342" pageId="1" pageNumber="386" year="1963">
<bibRefCitation id="121E4B67FFF09A57E787FB5DFF582832" author="Tortonese E" pageId="1" pageNumber="386" pagination="333 - 350" refId="ref3794" refString="Tortonese E (1963) Catalogo dei tipi di pesci del Museo Civico di Storia Naturale di Genova (Parte III). Annali del Museo Civico di Storia Naturale Giacomo Doria, 73: 333 - 350." type="journal article" year="1963">Tortonese, 1963: 342</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F72E1087FFF09A57E406FABEFDB22832" author="Kottelat M &amp; Lim KKP" box="[266,535,1308,1330]" page="247" pageId="1" pageNumber="386" year="1995">
<bibRefCitation id="121E4B67FFF09A57E406FABEFDB22832" author="Kottelat M &amp; Lim KKP" box="[266,535,1308,1330]" pageId="1" pageNumber="386" pagination="227 - 256" refId="ref3538" refString="Kottelat M &amp; Lim KKP (1995) Freshwater fishes of Sarawak and Brunei Darussalam: a preliminary annotated checklist. The Sarawak Museum Journal, XLVIII, 69: 227 - 256." type="journal article" year="1995">Kottelat &amp; Lim, 1995: 247</bibRefCitation>
</treatmentCitation>
; Larson
<emphasis id="44FBEA84FFF09A57E762FABEFD242832" box="[622,641,1308,1329]" italics="true" pageId="1" pageNumber="386">in</emphasis>
<treatmentCitation id="F72E1087FFF09A57E786FABEFE9E284D" author="Randall JE &amp; Lim KKP" page="639" pageId="1" pageNumber="386" year="2000">
<bibRefCitation id="121E4B67FFF09A57E786FABEFE9E284D" author="Randall JE &amp; Lim KKP" pageId="1" pageNumber="386" pagination="569 - 667" refId="ref3718" refString="Randall JE &amp; Lim KKP (2000) A checklist of the fishes of the South China Sea. Raffles Bulletin of Zoology, Supplement 8: 569 - 667." type="journal article" year="2000">Randall &amp; Lim, 2000: 639</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F72E1087FFF09A57E445FA9BFDDA284D" author="Larson HK &amp; Murdy EO" box="[329,639,1336,1358]" page="3577" pageId="1" pageNumber="386" year="2001">
<bibRefCitation id="121E4B67FFF09A57E445FA9BFDDA284D" author="Larson HK &amp; Murdy EO" box="[329,639,1336,1358]" pageId="1" pageNumber="386" pagination="3574 - 3603" refId="ref3612" refString="Larson HK &amp; Murdy EO (2001) Families Eleotridae, Gobiidae. In: Carpenter KE &amp; Niem VH (eds.) Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Bony Fishes Part 4 (Labridae to Latimeriidae), Estuarine Crocodiles, Sea Turtles, Sea Snakes and Marine Mammals. Volume 6. FAO, Rome. Pp. 3574 - 3603." type="book chapter" year="2001">Larson &amp; Murdy, 2001: 3577</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F72E1087FFF09A57E780FA9AFE952869" author="Parenti LR &amp; Lim KKP" page="194" pageId="1" pageNumber="386" year="2005">
<bibRefCitation id="121E4B67FFF09A57E780FA9AFE952869" author="Parenti LR &amp; Lim KKP" pageId="1" pageNumber="386" pagination="175 - 208" refId="ref3688" refString="Parenti LR &amp; Lim KKP (2005) Fishes of the Rajang River, Sarawak, Malaysia. Raffles Bulletin of Zoology, Supplement 13: 175 - 208." type="journal article" year="2005">Parenti &amp; Lim 2005: 194</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F72E1087FFF09A57E431FAF7FE492868" author="Atack K" box="[317,492,1365,1387]" page="167" pageId="1" pageNumber="386" year="2006">
<bibRefCitation id="121E4B67FFF09A57E431FAF7FE492868" author="Atack K" box="[317,492,1365,1387]" pageId="1" pageNumber="386" refId="ref3320" refString="Atack K (2006) A Field Guide to the Fishes of Kuching Rivers, Sarawak, Malaysian Borneo. Natural History Publications, Kota Kinabalu, 200 pp." type="book" year="2006">Atack, 2006: 167</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F72E1087FFF09A57E4F5FAF7FD182868" author="Kottelat M" box="[505,701,1365,1387]" page="395" pageId="1" pageNumber="386" year="2013">
<bibRefCitation id="121E4B67FFF09A57E4F5FAF7FD182868" author="Kottelat M" box="[505,701,1365,1387]" pageId="1" pageNumber="386" pagination="1 - 663" refId="ref3493" refString="Kottelat M (2013) The fishes of the inland waters of Southeast Asia. A catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology, Supplement 27: 1 - 663." type="journal article" year="2013">Kottelat, 2013: 395</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F72E1087FFF09A57E7C6FAF4FE012884" author="Tan HH &amp; Grinang J" page="203" pageId="1" pageNumber="386" year="2018">
<bibRefCitation id="121E4B67FFF09A57E7C6FAF4FE012884" author="Tan HH &amp; Grinang J" pageId="1" pageNumber="386" pagination="1 - 8" refId="ref3749" refString="Tan HH &amp; Grinang J (2018) Pogoneleotris heterolepis (Teleostei: Eleotridae), a rare fish from Borneo: biological data and remarks on the type locality and type material. Ichthyological Exploration of Freshwaters, 1082: 1 - 8." type="journal article" year="2018">Tan &amp; Grinang, 2018: 203–210</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3E95651DFFF09A57E57DFA12FDED2AAB" pageId="1" pageNumber="386" type="materials_examined">
<paragraph id="76303696FFF09A57E57DFA12FDED2AAB" blockId="1.[112,752,1456,1961]" pageId="1" pageNumber="386">
<emphasis id="44FBEA84FFF09A57E57DFA12FEEA28CB" bold="true" box="[113,335,1456,1480]" pageId="1" pageNumber="386">Material examined.</emphasis>
<materialsCitation id="C6E73CCBFFF09A57E458FA12FF7C2B0B" collectionCode="BMNH" collectorName="G. Doria." country="Malaysia" location="Sarawak" pageId="1" pageNumber="386" specimenCode="BMNH 1868.6" specimenCount="1" stateProvince="Sarawak" typeStatus="holotype">
<collectingCountry id="0E987606FFF09A57E458FA12FE7A28CB" box="[340,479,1456,1480]" name="Malaysia" pageId="1" pageNumber="386">SARAWAK</collectingCountry>
:
<specimenCode id="26299EEDFFF09A57E4EBFA13FD2C28CB" box="[487,649,1456,1480]" collectionCode="BMNH" pageId="1" pageNumber="386">BMNH 1868.6</specimenCode>
.9.16,
<quantity id="B1779B73FFF09A57E7CBFA12FF3F28EB" metricMagnitude="-1" metricUnit="m" metricValue="1.3599999999999999" pageId="1" pageNumber="386" unit="mm" value="136.0">136 mm</quantity>
SL male,
<typeStatus id="A9348834FFF09A57E40FFA72FEC528EB" box="[259,352,1488,1512]" pageId="1" pageNumber="386" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B18F4D15FFF09A57E473FA72FDE828EB" authorityName="Gunther" authorityYear="1869" box="[383,589,1488,1512]" class="Actinopterygii" family="Eleotridae" genus="Eleotris" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="386" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="44FBEA84FFF09A57E473FA72FDE828EB" box="[383,589,1488,1512]" italics="true" pageId="1" pageNumber="386">Eleotris heterolepis</emphasis>
</taxonomicName>
,
<collectingRegion id="B44BF874FFF09A57E756FA72FD1D28EB" box="[602,696,1488,1512]" country="Malaysia" name="Sarawak" pageId="1" pageNumber="386">Sarawak</collectingRegion>
, coll.
<collectorName id="DB7A5340FFF09A57E57CFA52FF7C2B0B" box="[112,217,1520,1544]" pageId="1" pageNumber="386">G. Doria.</collectorName>
</materialsCitation>
<materialsCitation id="C6E73CCBFFF09A57E5EFFA52FD4E2B2B" accessionNumber="ZRC 54119" collectingDate="1982-07-17" collectionCode="ZRC" collectorName="Martin Abba" country="Malaysia" location="Kanowit" municipality="Rejang River" pageId="1" pageNumber="386" specimenCode="1" specimenCount="1" typeStatus="holotype">
<collectionCode id="109EAE53FFF09A57E5EFFA52FEB22B0B" box="[227,279,1520,1544]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
54119,
<specimenCode id="26299EEDFFF09A57E47AFA52FE252B0B" box="[374,384,1520,1544]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">1</specimenCode>
(186),
<collectingMunicipality id="9654ACECFFF09A57E4DCFA52FDC72B0B" box="[464,610,1520,1544]" pageId="1" pageNumber="386">Rejang River</collectingMunicipality>
at
<location id="7350604DFFF09A57E786FA52FD4F2B0B" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB:7350604DFFF09A57E786FA52FD4F2B0B" box="[650,746,1520,1544]" country="Malaysia" municipality="Rejang River" name="Kanowit" pageId="1" pageNumber="386">Kanowit</location>
, main river next to cemetery, coll.
<collectorName id="DB7A5340FFF09A57E4DCF9B2FDFC2B2B" box="[464,601,1552,1576]" pageId="1" pageNumber="386">Martin Abba</collectorName>
,
<date id="02311056FFF09A57E76EF9B2FD4E2B2B" box="[610,747,1552,1576]" pageId="1" pageNumber="386" value="1982-07-17">
<collectingDate id="1275E9BEFFF09A57E76EF9B2FD4E2B2B" box="[610,747,1552,1576]" pageId="1" pageNumber="386" value="1982-07-17">17 July 1982</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C6E73CCBFFF09A57E57DF992FDA62B6B" accessionNumber="ZRC 39733" collectingDate="1996-01-15" collectionCode="ZRC" collectorName="H. H. Tan &amp; S. H. Tan" country="Malaysia" location="Kota Samarahan" pageId="1" pageNumber="386" specimenCode="1" specimenCount="1" typeStatus="holotype">
<collectionCode id="109EAE53FFF09A57E57DF992FF002B4B" box="[113,165,1584,1608]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
39733,
<specimenCode id="26299EEDFFF09A57E40AF992FEB52B4B" box="[262,272,1584,1608]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">1</specimenCode>
(195),
<location id="7350604DFFF09A57E46CF993FDBD2B4B" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB:7350604DFFF09A57E46CF993FDBD2B4B" box="[352,536,1584,1608]" country="Malaysia" name="Kota Samarahan" pageId="1" pageNumber="386">Kota Samarahan</location>
market, coll.
<collectorName id="DB7A5340FFF09A57E7B6F993FF3E2B6B" pageId="1" pageNumber="386">H.H. Tan</collectorName>
and
<collectorName id="DB7A5340FFF09A57E5D9F9F2FE9E2B6B" box="[213,315,1616,1640]" pageId="1" pageNumber="386">S.H. Tan</collectorName>
,
<date id="02311056FFF09A57E445F9F2FDA62B6B" box="[329,515,1616,1641]" pageId="1" pageNumber="386" value="1996-01-15">
<collectingDate id="1275E9BEFFF09A57E445F9F2FDA62B6B" box="[329,515,1616,1641]" pageId="1" pageNumber="386" value="1996-01-15">15 January 1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C6E73CCBFFF09A57E71DF9F2FE902BAB" accessionNumber="ZRC 54118" collectingDate="1996-08-31" collectingDateMax="1996-09-05" collectingDateMin="1996-08-31" collectionCode="ZRC" collectorName="H. H. Tan" country="Malaysia" location="Serian" pageId="1" pageNumber="386" specimenCode="1" specimenCount="1" typeStatus="holotype">
<collectionCode id="109EAE53FFF09A57E71DF9F2FDE02B6B" box="[529,581,1616,1640]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
54118,
<specimenCode id="26299EEDFFF09A57E7AEF9F2FD092B6B" box="[674,684,1616,1640]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">1</specimenCode>
(156),
<location id="7350604DFFF09A57E57DF9D2FF102B8B" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB:7350604DFFF09A57E57DF9D2FF102B8B" box="[113,181,1648,1672]" country="Malaysia" name="Serian" pageId="1" pageNumber="386">Serian</location>
wet market, coll.
<collectorName id="DB7A5340FFF09A57E470F9D3FE412B8B" box="[380,484,1649,1672]" pageId="1" pageNumber="386">H.H. Tan</collectorName>
, between
<date id="02311056FFF09A57E75DF9D2FE902BAB" pageId="1" pageNumber="386" value="1996-08-31" valueMax="1996-09-05" valueMin="1996-08-31">
<collectingDate id="1275E9BEFFF09A57E75DF9D2FE902BAB" pageId="1" pageNumber="386" value="1996-08-31" valueMax="1996-09-05" valueMin="1996-08-31">31 August and 5 September 1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C6E73CCBFFF09A57E433F932FE922BEB" collectingDate="1996-08-31" collectingDateMax="1996-09-05" collectingDateMin="1996-08-31" collectionCode="ZRC" collectorName="Serian &amp; H. H. Tan" country="Malaysia" location="Museum" pageId="1" pageNumber="386" specimenCount="1" stateProvince="Sarawak" typeStatus="holotype">
<collectingRegion id="B44BF874FFF09A57E433F932FE3E2BAB" box="[319,411,1680,1704]" country="Malaysia" name="Sarawak" pageId="1" pageNumber="386">Sarawak</collectingRegion>
<location id="7350604DFFF09A57E4ACF933FE582BAB" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB:7350604DFFF09A57E4ACF933FE582BAB" box="[416,509,1681,1704]" country="Malaysia" name="Museum" pageId="1" pageNumber="386" stateProvince="Sarawak">Museum</location>
[held at
<collectionCode id="109EAE53FFF09A57E755F932FD372BAB" box="[601,658,1680,1704]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
], 1(152),
<collectorName id="DB7A5340FFF09A57E57DF912FF102BCB" box="[113,181,1712,1736]" pageId="1" pageNumber="386">Serian</collectorName>
wet market, coll.
<collectorName id="DB7A5340FFF09A57E470F913FE412BCB" box="[380,484,1713,1736]" pageId="1" pageNumber="386">H.H. Tan</collectorName>
, between
<date id="02311056FFF09A57E75DF912FE922BEB" pageId="1" pageNumber="386" value="1996-08-31" valueMax="1996-09-05" valueMin="1996-08-31">
<collectingDate id="1275E9BEFFF09A57E75DF912FE922BEB" pageId="1" pageNumber="386" value="1996-08-31" valueMax="1996-09-05" valueMin="1996-08-31">31 August and 5 September 1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C6E73CCBFFF09A57E44EF972FE232A0B" accessionNumber="ZRC 41853" collectingDate="1997-10-29" collectingDateMax="1997-10-31" collectingDateMin="1997-10-29" collectionCode="ZRC" collectorName="Serian." country="Malaysia" pageId="1" pageNumber="386" specimenCode="1" specimenCount="1" typeStatus="holotype">
<collectionCode id="109EAE53FFF09A57E44EF972FED32BEB" box="[322,374,1744,1768]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
41853,
<specimenCode id="26299EEDFFF09A57E4C0F972FE702BEB" box="[460,469,1744,1768]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">1</specimenCode>
(159.5),
<collectorName id="DB7A5340FFF09A57E739F972FD252BEB" box="[565,640,1744,1768]" pageId="1" pageNumber="386">Serian.</collectorName>
coll. H.H. Tan,
<date id="02311056FFF09A57E5A4F952FE232A0B" box="[168,390,1776,1800]" pageId="1" pageNumber="386" value="1997-10-29" valueMax="1997-10-31" valueMin="1997-10-29">
<collectingDate id="1275E9BEFFF09A57E5A4F952FE232A0B" box="[168,390,1776,1800]" pageId="1" pageNumber="386" value="1997-10-29" valueMax="1997-10-31" valueMin="1997-10-29">29–31 October 1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C6E73CCBFFF09A57E49DF952FD4E2A2B" accessionNumber="ZRC 56950" collectingDate="2017-09-07" collectionCode="ZRC" collectorName="J. Grinang" country="Malaysia" location="Sungei Simunjan" pageId="1" pageNumber="386" specimenCode="4" specimenCount="1" typeStatus="holotype">
<collectionCode id="109EAE53FFF09A57E49DF952FE602A0B" box="[401,453,1776,1800]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
56950,
<specimenCode id="26299EEDFFF09A57E710F952FD8F2A0B" box="[540,554,1776,1800]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">4</specimenCode>
(121–210),
<location id="7350604DFFF09A57E7A9F952FF7E2A2B" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB:7350604DFFF09A57E7A9F952FF7E2A2B" country="Malaysia" name="Sungei Simunjan" pageId="1" pageNumber="386">Sungei Simunjan</location>
, coll.
<collectorName id="DB7A5340FFF09A57E428F8B3FE3C2A2B" box="[292,409,1808,1832]" pageId="1" pageNumber="386">J. Grinang</collectorName>
and party,
<date id="02311056FFF09A57E710F8B3FD4E2A2B" box="[540,747,1808,1832]" pageId="1" pageNumber="386" value="2017-09-07">
<collectingDate id="1275E9BEFFF09A57E710F8B3FD4E2A2B" box="[540,747,1808,1832]" pageId="1" pageNumber="386" value="2017-09-07">7 September 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C6E73CCBFFF09A57E57EF892FEDC2A6B" accessionNumber="ZRC 56951" collectingDate="2017-09-07" collectionCode="ZRC" collectorName="J. Grinang" country="Malaysia" location="Sungei Ba" pageId="1" pageNumber="386" specimenCode="2" specimenCount="1" typeStatus="holotype">
<collectionCode id="109EAE53FFF09A57E57EF892FF032A4B" box="[114,166,1840,1864]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
56951,
<specimenCode id="26299EEDFFF09A57E40FF892FEB42A4B" box="[259,273,1840,1864]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">2</specimenCode>
(121–155),
<location id="7350604DFFF09A57E49CF892FDA02A4B" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB:7350604DFFF09A57E49CF892FDA02A4B" box="[400,517,1840,1864]" country="Malaysia" name="Sungei Ba" pageId="1" pageNumber="386">Sungei Ba</location>
, coll.
<collectorName id="DB7A5340FFF09A57E740F893FD1A2A4B" box="[588,703,1840,1864]" pageId="1" pageNumber="386">J. Grinang</collectorName>
and party,
<date id="02311056FFF09A57E5B9F8F3FEDC2A6B" box="[181,377,1872,1896]" pageId="1" pageNumber="386" value="2017-09-07">
<collectingDate id="1275E9BEFFF09A57E5B9F8F3FEDC2A6B" box="[181,377,1872,1896]" pageId="1" pageNumber="386" value="2017-09-07">7 September 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C6E73CCBFFF09A57E48FF8F2FEAC2A8B" accessionNumber="ZRC 56952" collectingDate="2017-10" collectionCode="ZRC" collectorName="Rejang &amp; Sibu" country="Malaysia" pageId="1" pageNumber="386" specimenCode="1" specimenCount="1" typeStatus="holotype">
<collectionCode id="109EAE53FFF09A57E48FF8F2FE122A6B" box="[387,439,1872,1896]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
56952,
<specimenCode id="26299EEDFFF09A57E700F8F2FDB32A6B" box="[524,534,1872,1896]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">1</specimenCode>
(171),
<collectorName id="DB7A5340FFF09A57E76CF8F2FD0B2A6B" box="[608,686,1872,1896]" pageId="1" pageNumber="386">Rejang</collectorName>
,
<collectorName id="DB7A5340FFF09A57E7B4F8F2FD4E2A6B" box="[696,747,1872,1896]" pageId="1" pageNumber="386">Sibu</collectorName>
,
<date id="02311056FFF09A57E57DF8D2FEAC2A8B" box="[113,265,1904,1928]" pageId="1" pageNumber="386" value="2017-10">
<collectingDate id="1275E9BEFFF09A57E57DF8D2FEAC2A8B" box="[113,265,1904,1928]" pageId="1" pageNumber="386" value="2017-10">October 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="C6E73CCBFFF09A57E419F8D2FDE02AAB" accessionNumber="ZRC 56949" collectingDate="2018-01-31" collectionCode="ZRC" collectorName="H. H. Tan" country="Malaysia" location="Sungei Simunjan" pageId="1" pageNumber="386" specimenCode="3" specimenCount="1" typeStatus="holotype">
<collectionCode id="109EAE53FFF09A57E419F8D2FEEC2A8B" box="[277,329,1904,1928]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">ZRC</collectionCode>
56949,
<specimenCode id="26299EEDFFF09A57E4A8F8D2FE152A8B" box="[420,432,1904,1928]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="386">3</specimenCode>
(150–167),
<location id="7350604DFFF09A57E73CF8D2FD4E2A8B" LSID="urn:lsid:plazi:treatment:FE268780FFF19A52E69AF822FD132BFB:7350604DFFF09A57E73CF8D2FD4E2A8B" box="[560,747,1904,1928]" country="Malaysia" name="Sungei Simunjan" pageId="1" pageNumber="386">Sungei Simunjan</location>
, coll.
<collectorName id="DB7A5340FFF09A57E5A5F833FEAB2AAB" box="[169,270,1937,1960]" pageId="1" pageNumber="386">H.H. Tan</collectorName>
and party,
<date id="02311056FFF09A57E481F832FDE02AAB" box="[397,581,1936,1961]" pageId="1" pageNumber="386" value="2018-01-31">
<collectingDate id="1275E9BEFFF09A57E481F832FDE02AAB" box="[397,581,1936,1961]" pageId="1" pageNumber="386" value="2018-01-31">31 January 2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3E95651DFFF09A57E57DF872FBC12A19" pageId="1" pageNumber="386" type="diagnosis">
<paragraph id="76303696FFF09A57E57DF872FBC12A19" blockId="1.[113,752,2000,2089]" lastBlockId="1.[812,1452,1250,1818]" pageId="1" pageNumber="386">
<emphasis id="44FBEA84FFF09A57E57DF872FF422AEB" bold="true" box="[113,231,2000,2024]" pageId="1" pageNumber="386">Diagnosis.</emphasis>
First dorsal fin VI; second dorsal fin I,11–12; anal fin I,10; pectoral fin 20–23, usually 22; 17 segmented caudal fin rays in 9/8 pattern, branched caudal fin rays 15- 17, usually
<quantity id="B1779B73FFF09A57E6BDFB40FC4E29F9" box="[945,1003,1250,1274]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="1" pageNumber="386" unit="in" value="15.0">15 in</quantity>
8/7 pattern, procurrent caudal elements 11–15 dorsally, 11–13 ventrally; lateral scales about 38–64, counting main rows only (difficult to see scale rows due to many auxiliary scales); TRB 15–25. Vertebrae 10+16=26 (n = 15); dorsal pterygiophore pattern 3-22110; epurals 2; 2 anal pteryiophores before first caudal vertebra. Pelvic fins separate, space between fin bases scaled. Body with large scales, ctenoid scales alternating with clusters of cycloid scales and auxiliary scales present. Teeth sharp, in about six rows in both jaws. Sensory canals and pores mostly obscured by skin and sensory papillae, but canal extends from pore just above upper lip continuously to rear of opercle; pores usually at ends of branches from main canal; four preopercular pores present, in canal continuous with oculoscapular canal. Sensory papillae in longitudinal pattern, papillae themselves are mostly elongate thin flaps or variously branched, others are seated in oval pits on cheek. Known only from estuarine rivers in
<collectingCountry id="0E987606FFF09A57E683F8A0FC4B2A19" box="[911,1006,1794,1818]" name="Malaysia" pageId="1" pageNumber="386">Sarawak</collectingCountry>
,
<collectingCountry id="0E987606FFF09A57E6F6F8A0FBFA2A19" box="[1018,1119,1794,1818]" name="Malaysia" pageId="1" pageNumber="386">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3E95651DFFF09A52E621F8E0FBF92E9C" lastPageId="4" lastPageNumber="389" pageId="1" pageNumber="386" type="description">
<paragraph id="76303696FFF09A57E621F8E0FAE12519" blockId="1.[812,1452,1858,2074]" pageId="1" pageNumber="386">
<emphasis id="44FBEA84FFF09A57E621F8E0FC1D2A59" bold="true" box="[813,952,1858,1882]" pageId="1" pageNumber="386">Description.</emphasis>
First dorsal fin VI; second dorsal fin I,11–12, usually I,11; anal fin I,10–11, usually I,10; pectoral fin 20–23, usually 22; 17 segmented caudal fin rays, branched caudal fin rays 15–17, usually
<quantity id="B1779B73FFF09A57E105F800FBE42AB9" box="[1033,1089,1954,1978]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="1" pageNumber="386" unit="in" value="15.0">15 in</quantity>
8/7 pattern; lateral scales 38–64, counting main rows only (difficult to see scale rows due to the many clusters of auxiliary scales); TRB 15–25; 42–67 predorsal scales; 14–22 circumpeduncular scales.
</paragraph>
<caption id="22F0661EFFF39A54E64FFDE0FC6C2F92" ID-DOI="http://doi.org/10.5281/zenodo.4577074" ID-Zenodo-Dep="4577074" httpUri="https://zenodo.org/record/4577074/files/figure.png" pageId="2" pageNumber="387" startId="2.[835,870,578,600]" targetBox="[835,1475,189,563]" targetPageId="2">
<paragraph id="76303696FFF39A54E64FFDE0FC6C2F92" blockId="2.[835,1474,578,657]" pageId="2" pageNumber="387">Fig. 3. Composite drawing of head showing lateral canals, sensory pores and papillae. Most papillae and flaps omitted for clarity, as are all scales.</paragraph>
</caption>
<paragraph id="76303696FFF39A54E64FFD6AFB7B2E62" blockId="2.[835,1474,712,865]" pageId="2" pageNumber="387">teeth large and bluntly pointed. Gill opening wide, reaching to below rear margin of preopercle or just anterior to it. Shoulder girdle with 2–5 large fleshy rather square knobs or lobes. Pectoral fin base with broad scaled fleshy area extending partway onto pectoral rays.</paragraph>
<caption id="22F0661EFFF39A54E585FA4AFD152B34" ID-DOI="http://doi.org/10.5281/zenodo.4577076" ID-Zenodo-Dep="4577076" httpUri="https://zenodo.org/record/4577076/files/figure.png" pageId="2" pageNumber="387" startId="2.[137,172,1512,1534]" targetBox="[136,775,189,1496]" targetPageId="2">
<paragraph id="76303696FFF39A54E585FA4AFD152B34" blockId="2.[136,776,1512,1591]" pageId="2" pageNumber="387">
Fig. 2. Dorsal, lateral and ventral views (right side reversed) of head of fresh
<taxonomicName id="B18F4D15FFF39A54E405F9A6FDA42B1A" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[265,513,1540,1561]" class="Actinopterygii" family="Eleotridae" genus="Pogoneleotris" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="387" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="44FBEA84FFF39A54E405F9A6FDA42B1A" box="[265,513,1540,1561]" italics="true" pageId="2" pageNumber="387">Pogoneleotris heterolepis</emphasis>
</taxonomicName>
(ZRC56949, 160.0 mm SL male); showing extent of scalation and sensory papillae.
</paragraph>
</caption>
<paragraph id="76303696FFF39A54E584F9CCFC7C28E3" blockId="2.[135,775,1646,2118]" lastBlockId="2.[834,1475,904,1504]" pageId="2" pageNumber="387">
Morphometric values of males and females are given in
<tableCitation id="3B0D032DFFF39A54E7C7F9CCFF302BA5" captionStart="Table 1" captionStartId="3.[113,168,189,211]" captionText="Table 1. Morphometrics of Pogoneleotris heterolepis." pageId="2" pageNumber="387">Table 1</tableCitation>
. See
<figureCitation id="EEB42A13FFF39A54E5DCF92CFEBC2BA5" box="[208,281,1678,1702]" captionStart="Fig" captionStartId="1.[113,148,1173,1195]" captionTargetBox="[113,1451,189,1158]" captionTargetId="figure-459@1.[113,1451,189,1158]" captionTargetPageId="1" captionText="Fig. 1. Dorsal, lateral and ventral views of preserved male Pogoneleotris heterolepis (ZRC56949, 160.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.4577072" httpUri="https://zenodo.org/record/4577072/files/figure.png" pageId="2" pageNumber="387">Figs. 1</figureCitation>
and
<figureCitation id="EEB42A13FFF39A54E45CF92CFEC52BA5" box="[336,352,1678,1702]" captionStart="Fig" captionStartId="4.[136,171,761,783]" captionTargetBox="[136,1474,189,746]" captionTargetId="figure-492@4.[136,1474,189,746]" captionTargetPageId="4" captionText="Fig. 4. Fresh Pogoneleotris heterolepis from Simunjan, Sarawak; ZRC56949, female above (166.7 mm SL), male below (160.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.4577078" httpUri="https://zenodo.org/record/4577078/files/figure.png" pageId="2" pageNumber="387">4</figureCitation>
. Body cylindrical anteriorly, becoming compressed and narrower posteriorly, with slender caudal peduncle (
<figureCitation id="EEB42A13FFF39A54E5FFF96CFE8B2BE5" box="[243,302,1742,1766]" captionStart="Fig" captionStartId="1.[113,148,1173,1195]" captionTargetBox="[113,1451,189,1158]" captionTargetId="figure-459@1.[113,1451,189,1158]" captionTargetPageId="1" captionText="Fig. 1. Dorsal, lateral and ventral views of preserved male Pogoneleotris heterolepis (ZRC56949, 160.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.4577072" httpUri="https://zenodo.org/record/4577072/files/figure.png" pageId="2" pageNumber="387">Fig. 1</figureCitation>
). Head broad, depressed anteriorly, with nape variably humped; with low concavity or groove at leading edge of scaled area behind interorbital. Eyes very small, facing dorsally. Interorbital very wide, about 30% of head length. Jaws large (about 40% of head length), oblique, at about 40° angle to body axis, chin tip anteriormost. Tongue large, tip rounded to blunt. Mouth terminal, oblique, lower jaw prominent, jaws reaching back to before very small eye; lip lining fimbriate, fimbriae often branched. Teeth in both jaws in 5–8 rows, with about six large stout curved teeth widely spaced across front of jaw, with 5–6 rows of small conical teeth crowded behind; and innermost row of Body with large vertically slender ctenoid scales (mostly posteriorly), with smaller ctenoid scales alternating with clusters of small cycloid and ctenoid scales and numerous auxiliary cycloid scales present at base of each large scale. Dorsal part of opercle and cheek behind eye with many very small cycloid scales in all but smallest specimens; patch of small cycloid scales on lower rear half of preopercle beneath the rows of branched cheek papillae; small scales on side of head and nape appear to sit in a concave base. Predorsal scales very small, reaching forward to close behind eyes, becoming more numerous and entering interorbital space in larger specimens; pectoral base, belly and breast with many small cycloid scales; broad area around anus naked, as is pectoral base axil and nearby area on body behind fin base (
<figureCitation id="EEB42A13FFF39A54E647FAEAFC352863" box="[843,912,1352,1376]" captionStart="Fig" captionStartId="2.[137,172,1512,1534]" captionTargetBox="[136,775,189,1496]" captionTargetId="figure-7@2.[136,775,189,1496]" captionTargetPageId="2" captionText="Fig. 2. Dorsal, lateral and ventral views (right side reversed) of head of fresh Pogoneleotris heterolepis (ZRC56949, 160.0 mm SL male); showing extent of scalation and sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.4577076" httpUri="https://zenodo.org/record/4577076/files/figure.png" pageId="2" pageNumber="387">Fig. 2</figureCitation>
). Extent of scales on chest variable, often reaching right up into isthmus either densely or scattered anteriorly. One specimen (ZRC 56952) with several rows of small scales on rear part of branchiostegal membrane, on curved lobes of skin.
</paragraph>
<paragraph id="76303696FFF39A54E64FF9AAFA662A03" blockId="2.[835,1475,1544,1792]" pageId="2" pageNumber="387">First dorsal fin low, triangular, does not quite reach second dorsal fin when adpressed. Second dorsal and anal fins pointed posteriorly. Pectoral fins with central rays longest, fins reaching back to below rear of first dorsal fin; bases of fin rays covered by thickened scaled flesh, with bases of rays obscured. Pelvic fins widely separate, pointed, but falling well short of anus; interspace between fins with 3–5 rows of cycloid scales. Caudal fin narrow, elongate and pointed.</paragraph>
<paragraph id="76303696FFF39A55E64FF88AFE0F2857" blockId="2.[835,1476,1832,2113]" lastBlockId="3.[113,753,1148,1364]" lastPageId="3" lastPageNumber="388" pageId="2" pageNumber="387">
Sensory head pores large, in distinctive arrangement (see
<figureCitation id="EEB42A13FFF39A54E64FF8EAFC212A63" box="[835,900,1864,1888]" captionStart="Fig" captionStartId="2.[835,870,578,600]" captionTargetBox="[835,1475,189,563]" captionTargetId="figure-558@2.[814,1508,166,590]" captionTargetPageId="2" captionText="Fig. 3. Composite drawing of head showing lateral canals, sensory pores and papillae. Most papillae and flaps omitted for clarity, as are all scales." figureDoi="http://doi.org/10.5281/zenodo.4577074" httpUri="https://zenodo.org/record/4577074/files/figure.png" pageId="2" pageNumber="387">Fig. 3</figureCitation>
); with a nasal pore beside each nostril tube (posterior nasal tube opening often with several flat filaments around opening), a very large wide pore below the eye, one small pore above preopercle (exact position variable) and usually four (occasionally three) pores along canal above opercle, 3–4 canals of which extend well on to upper part of opercle, ultimate canal pore enlarged; and four preopercular pores, the canals of which may extend on to interopercle and anterior part of opercle, preopercular canal continuous with oculoscapular canal (
<figureCitation id="EEB42A13FFF29A55E46FFB3EFE0C29B7" box="[355,425,1180,1204]" captionStart="Fig" captionStartId="2.[835,870,578,600]" captionTargetBox="[835,1475,189,563]" captionTargetId="figure-558@2.[814,1508,166,590]" captionTargetPageId="2" captionText="Fig. 3. Composite drawing of head showing lateral canals, sensory pores and papillae. Most papillae and flaps omitted for clarity, as are all scales." figureDoi="http://doi.org/10.5281/zenodo.4577074" httpUri="https://zenodo.org/record/4577074/files/figure.png" pageId="3" pageNumber="388">Fig. 3</figureCitation>
). In three specimens, a very small pore visible in middle of interorbital space; may be present in more specimens but could not be discerned due to scale covering. Pores generally obscured by thickened skin, crowded filaments and mucous coat, so that pores often could not be observed.
</paragraph>
<caption id="22F0661EFFF29A55E57DFF1FFD262DD1" box="[113,643,189,211]" pageId="3" pageNumber="388">
<paragraph id="76303696FFF29A55E57DFF1FFD262DD1" blockId="3.[113,643,189,211]" box="[113,643,189,211]" pageId="3" pageNumber="388">
Table 1. Morphometrics of
<taxonomicName id="B18F4D15FFF29A55E48FFF1FFDDA2DD1" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[387,639,189,210]" class="Actinopterygii" family="Eleotridae" genus="Pogoneleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="388" phylum="Chordata" rank="species" species="heterolepis">Pogoneleotris heterolepis</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="76303696FFF29A55E4EEFEA7FAC3292F" pageId="3" pageNumber="388">
<table id="048FC436FFF265A9E58CFEA7FA32292E" box="[128,1431,261,1069]" gridcols="7" gridrows="17" pageId="3" pageNumber="388">
<tr id="C8BF34D4FFF265A9E58CFEA7FA322C34" box="[128,1431,261,311]" gridrow="0" pageId="3" pageNumber="388" rowspan-0="1">
<th id="8B6E5DA8FFF265A9E4EEFEA7FD9A2C34" box="[482,575,261,311]" gridcol="1" gridrow="0" pageId="3" pageNumber="388">
<emphasis id="44FBEA84FFF29A55E4EEFEA7FD972C34" bold="true" box="[482,575,261,311]" pageId="3" pageNumber="388">Holotype (male)</emphasis>
</th>
<th id="8B6E5DA8FFF265A9E774FEA7FD5D2C34" box="[632,760,261,311]" gridcol="2" gridrow="0" pageId="3" pageNumber="388">
<emphasis id="44FBEA84FFF29A55E774FEA7FD7B2C34" bold="true" pageId="3" pageNumber="388">Males range (n= 11)</emphasis>
</th>
<th id="8B6E5DA8FFF265A9E64CFEA7FCDE2C34" box="[832,891,261,311]" gridcol="3" gridrow="0" pageId="3" pageNumber="388">
<emphasis id="44FBEA84FFF29A55E64CFEB1FCDE2C2B" bold="true" box="[832,891,275,296]" pageId="3" pageNumber="388">Mean</emphasis>
</th>
<th id="8B6E5DA8FFF265A9E6B5FEA7FBF52C34" box="[953,1104,261,311]" gridcol="4" gridrow="0" pageId="3" pageNumber="388">
<emphasis id="44FBEA84FFF29A55E6B5FEA7FB802C34" bold="true" pageId="3" pageNumber="388">Females range (n= 6)</emphasis>
</th>
<th id="8B6E5DA8FFF265A9E181FEA7FB6D2C34" box="[1165,1224,261,311]" gridcol="5" gridrow="0" pageId="3" pageNumber="388">
<emphasis id="44FBEA84FFF29A55E181FEB1FB6D2C2B" bold="true" box="[1165,1224,275,296]" pageId="3" pageNumber="388">Mean</emphasis>
</th>
<th id="8B6E5DA8FFF265A9E006FEA7FA322C34" box="[1290,1431,261,311]" gridcol="6" gridrow="0" pageId="3" pageNumber="388">
<emphasis id="44FBEA84FFF29A55E006FEA7FAC92C34" bold="true" pageId="3" pageNumber="388">Mean of both sexes</emphasis>
</th>
</tr>
<tr id="C8BF34D4FFF265A9E58CFEFAFA322C6D" box="[128,1431,344,366]" gridrow="1" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFEFAFA322C6D" box="[128,1431,344,366]" colspan="7" colspanRight="6" gridcol="0" gridrow="1" pageId="3" pageNumber="388">Percentage of standard length</th>
</tr>
<tr id="C8BF34D4FFF265A9E58CFE25FA322C9E" box="[128,1431,391,413]" gridrow="2" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFE25FE202C9E" box="[128,389,391,413]" gridcol="0" gridrow="2" pageId="3" pageNumber="388">Head length</th>
<td id="8B6E5DA8FFF265A9E4EEFE25FD9A2C9E" box="[482,575,391,413]" gridcol="1" gridrow="2" pageId="3" pageNumber="388">29.0</td>
<td id="8B6E5DA8FFF265A9E774FE25FD5D2C9E" box="[632,760,391,413]" gridcol="2" gridrow="2" pageId="3" pageNumber="388">28.2–30.1</td>
<td id="8B6E5DA8FFF265A9E64CFE25FCDE2C9E" box="[832,891,391,413]" gridcol="3" gridrow="2" pageId="3" pageNumber="388">28.9</td>
<td id="8B6E5DA8FFF265A9E6B5FE25FBF52C9E" box="[953,1104,391,413]" gridcol="4" gridrow="2" pageId="3" pageNumber="388">28.8–29.9</td>
<td id="8B6E5DA8FFF265A9E181FE25FB6D2C9E" box="[1165,1224,391,413]" gridcol="5" gridrow="2" pageId="3" pageNumber="388">29.5</td>
<td id="8B6E5DA8FFF265A9E006FE25FA322C9E" box="[1290,1431,391,413]" gridcol="6" gridrow="2" pageId="3" pageNumber="388">29.1</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFE14FA322CCF" box="[128,1431,438,460]" gridrow="3" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFE14FE202CCF" box="[128,389,438,460]" gridcol="0" gridrow="3" pageId="3" pageNumber="388">Body depth</th>
<td id="8B6E5DA8FFF265A9E4EEFE14FD9A2CCF" box="[482,575,438,460]" gridcol="1" gridrow="3" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FE14FD5D2CCF" box="[632,760,438,460]" gridcol="2" gridrow="3" pageId="3" pageNumber="388">17.1–21.5</td>
<td id="8B6E5DA8FFF265A9E64CFE14FCDE2CCF" box="[832,891,438,460]" gridcol="3" gridrow="3" pageId="3" pageNumber="388">19.0</td>
<td id="8B6E5DA8FFF265A9E6B5FE14FBF52CCF" box="[953,1104,438,460]" gridcol="4" gridrow="3" pageId="3" pageNumber="388">18.0–18.9</td>
<td id="8B6E5DA8FFF265A9E181FE14FB6D2CCF" box="[1165,1224,438,460]" gridcol="5" gridrow="3" pageId="3" pageNumber="388">18.5</td>
<td id="8B6E5DA8FFF265A9E006FE14FA322CCF" box="[1290,1431,438,460]" gridcol="6" gridrow="3" pageId="3" pageNumber="388">18.8</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFE47FA322CF8" box="[128,1431,485,507]" gridrow="4" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFE47FE202CF8" box="[128,389,485,507]" gridcol="0" gridrow="4" pageId="3" pageNumber="388">Length of caudal peduncle</th>
<td id="8B6E5DA8FFF265A9E4EEFE47FD9A2CF8" box="[482,575,485,507]" gridcol="1" gridrow="4" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FE47FD5D2CF8" box="[632,760,485,507]" gridcol="2" gridrow="4" pageId="3" pageNumber="388">21.0–25.5</td>
<td id="8B6E5DA8FFF265A9E64CFE47FCDE2CF8" box="[832,891,485,507]" gridcol="3" gridrow="4" pageId="3" pageNumber="388">22.9</td>
<td id="8B6E5DA8FFF265A9E6B5FE47FBF52CF8" box="[953,1104,485,507]" gridcol="4" gridrow="4" pageId="3" pageNumber="388">22.2–24.8</td>
<td id="8B6E5DA8FFF265A9E181FE47FB6D2CF8" box="[1165,1224,485,507]" gridcol="5" gridrow="4" pageId="3" pageNumber="388">23.1</td>
<td id="8B6E5DA8FFF265A9E006FE47FA322CF8" box="[1290,1431,485,507]" gridcol="6" gridrow="4" pageId="3" pageNumber="388">23.0</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFDB1FA322F29" box="[128,1431,531,554]" gridrow="5" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFDB1FE202F29" box="[128,389,531,554]" gridcol="0" gridrow="5" pageId="3" pageNumber="388">Depth of caudal peduncle</th>
<td id="8B6E5DA8FFF265A9E4EEFDB1FD9A2F29" box="[482,575,531,554]" gridcol="1" gridrow="5" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FDB1FD5D2F29" box="[632,760,531,554]" gridcol="2" gridrow="5" pageId="3" pageNumber="388">6.9–11.2</td>
<td id="8B6E5DA8FFF265A9E64CFDB1FCDE2F29" box="[832,891,531,554]" gridcol="3" gridrow="5" pageId="3" pageNumber="388">10.1</td>
<td id="8B6E5DA8FFF265A9E6B5FDB1FBF52F29" box="[953,1104,531,554]" gridcol="4" gridrow="5" pageId="3" pageNumber="388">8.7–10.3</td>
<td id="8B6E5DA8FFF265A9E181FDB1FB6D2F29" box="[1165,1224,531,554]" gridcol="5" gridrow="5" pageId="3" pageNumber="388">9.6</td>
<td id="8B6E5DA8FFF265A9E006FDB1FA322F29" box="[1290,1431,531,554]" gridcol="6" gridrow="5" pageId="3" pageNumber="388">9.9</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFDE0FA322F5B" box="[128,1431,578,600]" gridrow="6" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFDE0FE202F5B" box="[128,389,578,600]" gridcol="0" gridrow="6" pageId="3" pageNumber="388">Length of pectoral fin</th>
<td id="8B6E5DA8FFF265A9E4EEFDE0FD9A2F5B" box="[482,575,578,600]" gridcol="1" gridrow="6" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FDE0FD5D2F5B" box="[632,760,578,600]" gridcol="2" gridrow="6" pageId="3" pageNumber="388">14.9–21.5</td>
<td id="8B6E5DA8FFF265A9E64CFDE0FCDE2F5B" box="[832,891,578,600]" gridcol="3" gridrow="6" pageId="3" pageNumber="388">21.5</td>
<td id="8B6E5DA8FFF265A9E6B5FDE0FBF52F5B" box="[953,1104,578,600]" gridcol="4" gridrow="6" pageId="3" pageNumber="388">16.8–22.7</td>
<td id="8B6E5DA8FFF265A9E181FDE0FB6D2F5B" box="[1165,1224,578,600]" gridcol="5" gridrow="6" pageId="3" pageNumber="388">20.4</td>
<td id="8B6E5DA8FFF265A9E006FDE0FA322F5B" box="[1290,1431,578,600]" gridcol="6" gridrow="6" pageId="3" pageNumber="388">19.1</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFDD3FA322F84" box="[128,1431,625,647]" gridrow="7" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFDD3FE202F84" box="[128,389,625,647]" gridcol="0" gridrow="7" pageId="3" pageNumber="388">Length of pelvic fin</th>
<td id="8B6E5DA8FFF265A9E4EEFDD3FD9A2F84" box="[482,575,625,647]" gridcol="1" gridrow="7" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FDD3FD5D2F84" box="[632,760,625,647]" gridcol="2" gridrow="7" pageId="3" pageNumber="388">15.9–21.3</td>
<td id="8B6E5DA8FFF265A9E64CFDD3FCDE2F84" box="[832,891,625,647]" gridcol="3" gridrow="7" pageId="3" pageNumber="388">21.3</td>
<td id="8B6E5DA8FFF265A9E6B5FDD3FBF52F84" box="[953,1104,625,647]" gridcol="4" gridrow="7" pageId="3" pageNumber="388">19.0–22.3</td>
<td id="8B6E5DA8FFF265A9E181FDD3FB6D2F84" box="[1165,1224,625,647]" gridcol="5" gridrow="7" pageId="3" pageNumber="388">20.3</td>
<td id="8B6E5DA8FFF265A9E006FDD3FA322F84" box="[1290,1431,625,647]" gridcol="6" gridrow="7" pageId="3" pageNumber="388">19.6</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFD02FA322FB5" box="[128,1431,672,694]" gridrow="8" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFD02FE202FB5" box="[128,389,672,694]" gridcol="0" gridrow="8" pageId="3" pageNumber="388">Length of caudal fin</th>
<td id="8B6E5DA8FFF265A9E4EEFD02FD9A2FB5" box="[482,575,672,694]" gridcol="1" gridrow="8" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FD02FD5D2FB5" box="[632,760,672,694]" gridcol="2" gridrow="8" pageId="3" pageNumber="388">25.0–30.1</td>
<td id="8B6E5DA8FFF265A9E64CFD02FCDE2FB5" box="[832,891,672,694]" gridcol="3" gridrow="8" pageId="3" pageNumber="388">27.8</td>
<td id="8B6E5DA8FFF265A9E6B5FD02FBF52FB5" box="[953,1104,672,694]" gridcol="4" gridrow="8" pageId="3" pageNumber="388">27.8–31.3</td>
<td id="8B6E5DA8FFF265A9E181FD02FB6D2FB5" box="[1165,1224,672,694]" gridcol="5" gridrow="8" pageId="3" pageNumber="388">29.7</td>
<td id="8B6E5DA8FFF265A9E006FD02FA322FB5" box="[1290,1431,672,694]" gridcol="6" gridrow="8" pageId="3" pageNumber="388">28.5</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFD6DFA322FE6" box="[128,1431,719,741]" gridrow="9" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFD6DFE202FE6" box="[128,389,719,741]" gridcol="0" gridrow="9" pageId="3" pageNumber="388">Depressed D1 fin</th>
<td id="8B6E5DA8FFF265A9E4EEFD6DFD9A2FE6" box="[482,575,719,741]" gridcol="1" gridrow="9" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FD6DFD5D2FE6" box="[632,760,719,741]" gridcol="2" gridrow="9" pageId="3" pageNumber="388">14.5–19.3</td>
<td id="8B6E5DA8FFF265A9E64CFD6DFCDE2FE6" box="[832,891,719,741]" gridcol="3" gridrow="9" pageId="3" pageNumber="388">17.9</td>
<td id="8B6E5DA8FFF265A9E6B5FD6DFBF52FE6" box="[953,1104,719,741]" gridcol="4" gridrow="9" pageId="3" pageNumber="388">17.0–20.1</td>
<td id="8B6E5DA8FFF265A9E181FD6DFB6D2FE6" box="[1165,1224,719,741]" gridcol="5" gridrow="9" pageId="3" pageNumber="388">18.3</td>
<td id="8B6E5DA8FFF265A9E006FD6DFA322FE6" box="[1290,1431,719,741]" gridcol="6" gridrow="9" pageId="3" pageNumber="388">18.1</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFD5CFA322E17" box="[128,1431,766,788]" gridrow="10" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFD5CFA322E17" box="[128,1431,766,788]" colspan="7" colspanRight="6" gridcol="0" gridrow="10" pageId="3" pageNumber="388">Percentage of head length</th>
</tr>
<tr id="C8BF34D4FFF265A9E58CFC8FFA322E40" box="[128,1431,813,835]" gridrow="11" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFC8FFE202E40" box="[128,389,813,835]" gridcol="0" gridrow="11" pageId="3" pageNumber="388">Head depth</th>
<td id="8B6E5DA8FFF265A9E4EEFC8FFD9A2E40" box="[482,575,813,835]" gridcol="1" gridrow="11" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FC8FFD5D2E40" box="[632,760,813,835]" gridcol="2" gridrow="11" pageId="3" pageNumber="388">51.8–66.5</td>
<td id="8B6E5DA8FFF265A9E64CFC8FFCDE2E40" box="[832,891,813,835]" gridcol="3" gridrow="11" pageId="3" pageNumber="388">57.2</td>
<td id="8B6E5DA8FFF265A9E6B5FC8FFBF52E40" box="[953,1104,813,835]" gridcol="4" gridrow="11" pageId="3" pageNumber="388">48.3–57.5</td>
<td id="8B6E5DA8FFF265A9E181FC8FFB6D2E40" box="[1165,1224,813,835]" gridcol="5" gridrow="11" pageId="3" pageNumber="388">52.8</td>
<td id="8B6E5DA8FFF265A9E006FC8FFA322E40" box="[1290,1431,813,835]" gridcol="6" gridrow="11" pageId="3" pageNumber="388">55.4</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFCFEFA322E71" box="[128,1431,860,882]" gridrow="12" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFCFEFE202E71" box="[128,389,860,882]" gridcol="0" gridrow="12" pageId="3" pageNumber="388">Head width</th>
<td id="8B6E5DA8FFF265A9E4EEFCFEFD9A2E71" box="[482,575,860,882]" gridcol="1" gridrow="12" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FCFEFD5D2E71" box="[632,760,860,882]" gridcol="2" gridrow="12" pageId="3" pageNumber="388">68.5–83.6</td>
<td id="8B6E5DA8FFF265A9E64CFCFEFCDE2E71" box="[832,891,860,882]" gridcol="3" gridrow="12" pageId="3" pageNumber="388">74.8</td>
<td id="8B6E5DA8FFF265A9E6B5FCFEFBF52E71" box="[953,1104,860,882]" gridcol="4" gridrow="12" pageId="3" pageNumber="388">68.6–72.3</td>
<td id="8B6E5DA8FFF265A9E181FCFEFB6D2E71" box="[1165,1224,860,882]" gridcol="5" gridrow="12" pageId="3" pageNumber="388">70.3</td>
<td id="8B6E5DA8FFF265A9E006FCFEFA322E71" box="[1290,1431,860,882]" gridcol="6" gridrow="12" pageId="3" pageNumber="388">73.1</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFC28FA322EA2" box="[128,1431,906,929]" gridrow="13" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFC28FE202EA2" box="[128,389,906,929]" gridcol="0" gridrow="13" pageId="3" pageNumber="388">Snout length</th>
<td id="8B6E5DA8FFF265A9E4EEFC28FD9A2EA2" box="[482,575,906,929]" gridcol="1" gridrow="13" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FC28FD5D2EA2" box="[632,760,906,929]" gridcol="2" gridrow="13" pageId="3" pageNumber="388">24.0–36.8</td>
<td id="8B6E5DA8FFF265A9E64CFC28FCDE2EA2" box="[832,891,906,929]" gridcol="3" gridrow="13" pageId="3" pageNumber="388">28.1</td>
<td id="8B6E5DA8FFF265A9E6B5FC28FBF52EA2" box="[953,1104,906,929]" gridcol="4" gridrow="13" pageId="3" pageNumber="388">24.7–29.3</td>
<td id="8B6E5DA8FFF265A9E181FC28FB6D2EA2" box="[1165,1224,906,929]" gridcol="5" gridrow="13" pageId="3" pageNumber="388">26.7</td>
<td id="8B6E5DA8FFF265A9E006FC28FA322EA2" box="[1290,1431,906,929]" gridcol="6" gridrow="13" pageId="3" pageNumber="388">27.6</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFC1BFA322ECC" box="[128,1431,953,975]" gridrow="14" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFC1BFE202ECC" box="[128,389,953,975]" gridcol="0" gridrow="14" pageId="3" pageNumber="388">Eye width</th>
<td id="8B6E5DA8FFF265A9E4EEFC1BFD9A2ECC" box="[482,575,953,975]" gridcol="1" gridrow="14" pageId="3" pageNumber="388">5.3</td>
<td id="8B6E5DA8FFF265A9E774FC1BFD5D2ECC" box="[632,760,953,975]" gridcol="2" gridrow="14" pageId="3" pageNumber="388">3.0–5.8</td>
<td id="8B6E5DA8FFF265A9E64CFC1BFCDE2ECC" box="[832,891,953,975]" gridcol="3" gridrow="14" pageId="3" pageNumber="388">4.1</td>
<td id="8B6E5DA8FFF265A9E6B5FC1BFBF52ECC" box="[953,1104,953,975]" gridcol="4" gridrow="14" pageId="3" pageNumber="388">2.5–4.8</td>
<td id="8B6E5DA8FFF265A9E181FC1BFB6D2ECC" box="[1165,1224,953,975]" gridcol="5" gridrow="14" pageId="3" pageNumber="388">3.9</td>
<td id="8B6E5DA8FFF265A9E006FC1BFA322ECC" box="[1290,1431,953,975]" gridcol="6" gridrow="14" pageId="3" pageNumber="388">4.0</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFC4AFA322EFD" box="[128,1431,1000,1022]" gridrow="15" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFC4AFE202EFD" box="[128,389,1000,1022]" gridcol="0" gridrow="15" pageId="3" pageNumber="388">Jaw length</th>
<td id="8B6E5DA8FFF265A9E4EEFC4AFD9A2EFD" box="[482,575,1000,1022]" gridcol="1" gridrow="15" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FC4AFD5D2EFD" box="[632,760,1000,1022]" gridcol="2" gridrow="15" pageId="3" pageNumber="388">37.5–42.4</td>
<td id="8B6E5DA8FFF265A9E64CFC4AFCDE2EFD" box="[832,891,1000,1022]" gridcol="3" gridrow="15" pageId="3" pageNumber="388">40.2</td>
<td id="8B6E5DA8FFF265A9E6B5FC4AFBF52EFD" box="[953,1104,1000,1022]" gridcol="4" gridrow="15" pageId="3" pageNumber="388">36.2–41.6</td>
<td id="8B6E5DA8FFF265A9E181FC4AFB6D2EFD" box="[1165,1224,1000,1022]" gridcol="5" gridrow="15" pageId="3" pageNumber="388">38.6</td>
<td id="8B6E5DA8FFF265A9E006FC4AFA322EFD" box="[1290,1431,1000,1022]" gridcol="6" gridrow="15" pageId="3" pageNumber="388">39.6</td>
</tr>
<tr id="C8BF34D4FFF265A9E58CFBB5FA32292E" box="[128,1431,1047,1069]" gridrow="16" pageId="3" pageNumber="388">
<th id="8B6E5DA8FFF265A9E58CFBB5FE20292E" box="[128,389,1047,1069]" gridcol="0" gridrow="16" pageId="3" pageNumber="388">Interorbital width</th>
<td id="8B6E5DA8FFF265A9E4EEFBB5FD9A292E" box="[482,575,1047,1069]" gridcol="1" gridrow="16" pageId="3" pageNumber="388">–</td>
<td id="8B6E5DA8FFF265A9E774FBB5FD5D292E" box="[632,760,1047,1069]" gridcol="2" gridrow="16" pageId="3" pageNumber="388">27.7–32.0</td>
<td id="8B6E5DA8FFF265A9E64CFBB5FCDE292E" box="[832,891,1047,1069]" gridcol="3" gridrow="16" pageId="3" pageNumber="388">30.1</td>
<td id="8B6E5DA8FFF265A9E6B5FBB5FBF5292E" box="[953,1104,1047,1069]" gridcol="4" gridrow="16" pageId="3" pageNumber="388">28.1–30.4</td>
<td id="8B6E5DA8FFF265A9E181FBB5FB6D292E" box="[1165,1224,1047,1069]" gridcol="5" gridrow="16" pageId="3" pageNumber="388">29.3</td>
<td id="8B6E5DA8FFF265A9E006FBB5FA32292E" box="[1290,1431,1047,1069]" gridcol="6" gridrow="16" pageId="3" pageNumber="388">29.8</td>
</tr>
</table>
</paragraph>
<paragraph id="76303696FFF29A55E57DFADEFEE02A77" blockId="3.[113,754,1404,1908]" pageId="3" pageNumber="388">
Sensory papillae mostly filamentous, in roughly longitudinal arrangement (see
<figureCitation id="EEB42A13FFF29A55E43EFA3EFED628B7" box="[306,371,1436,1460]" captionStart="Fig" captionStartId="2.[137,172,1512,1534]" captionTargetBox="[136,775,189,1496]" captionTargetId="figure-7@2.[136,775,189,1496]" captionTargetPageId="2" captionText="Fig. 2. Dorsal, lateral and ventral views (right side reversed) of head of fresh Pogoneleotris heterolepis (ZRC56949, 160.0 mm SL male); showing extent of scalation and sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.4577076" httpUri="https://zenodo.org/record/4577076/files/figure.png" pageId="3" pageNumber="388">Fig. 2</figureCitation>
), a few papillae on top of head are short and bulbous, most are tentacle-like, branched 2–4 times on side and underside of head, fewer to no branches in papillae on top of head; on cheek, branched papillae arranged in at least four longitudinal rows; in between rows are 2–4 distinct lines occupied by row of widely spaced small papillae set within an oval “cell”; in larger specimens, these may be replaced by many-branched papillae (
<figureCitation id="EEB42A13FFF29A55E747F9DEFD2F2B97" box="[587,650,1660,1684]" captionStart="Fig" captionStartId="2.[835,870,578,600]" captionTargetBox="[835,1475,189,563]" captionTargetId="figure-558@2.[814,1508,166,590]" captionTargetPageId="2" captionText="Fig. 3. Composite drawing of head showing lateral canals, sensory pores and papillae. Most papillae and flaps omitted for clarity, as are all scales." figureDoi="http://doi.org/10.5281/zenodo.4577074" httpUri="https://zenodo.org/record/4577074/files/figure.png" pageId="3" pageNumber="388">Fig. 3</figureCitation>
). Opercle with fewer filamentous papillae. Preserved, the branched and filamentous papillae are stiff and almost bristle-like in large specimens. Large slender scales along mid-side of body each with a vertical row of sensory papillae. Caudal fin with two rows of papillae, one on either side of fin mid-base on membrane between segmented fin rays 6 and 7 and rays 8 and 9.
</paragraph>
<paragraph id="76303696FFF29A55E57DF83EFA092817" blockId="3.[113,752,1948,2100]" lastBlockId="3.[812,1452,1148,1300]" pageId="3" pageNumber="388">
When fresh, body and fins plain dark reddish brown, pink, purple to black (see
<figureCitation id="EEB42A13FFF29A55E444F81EFE2D2AD7" box="[328,392,1980,2004]" captionStart="Fig" captionStartId="4.[136,171,761,783]" captionTargetBox="[136,1474,189,746]" captionTargetId="figure-492@4.[136,1474,189,746]" captionTargetPageId="4" captionText="Fig. 4. Fresh Pogoneleotris heterolepis from Simunjan, Sarawak; ZRC56949, female above (166.7 mm SL), male below (160.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.4577078" httpUri="https://zenodo.org/record/4577078/files/figure.png" pageId="3" pageNumber="388">Fig. 4</figureCitation>
). The present specimens are pink, with dark pink to purplish scales; side of head pale pink, underside of head whitish (
<figureCitation id="EEB42A13FFF29A55E490F85EFE442517" box="[412,481,2044,2068]" captionStart="Fig" captionStartId="4.[136,171,761,783]" captionTargetBox="[136,1474,189,746]" captionTargetId="figure-492@4.[136,1474,189,746]" captionTargetPageId="4" captionText="Fig. 4. Fresh Pogoneleotris heterolepis from Simunjan, Sarawak; ZRC56949, female above (166.7 mm SL), male below (160.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.4577078" httpUri="https://zenodo.org/record/4577078/files/figure.png" pageId="3" pageNumber="388">Fig. 4</figureCitation>
). Males darker in colour than females. Dorsal fins pinkish white with spines darker pink to grey, slightly more grey in males; anal fin dull pink, slightly darker in males; pectoral fin pink in females, pale grey in males; pelvic fins pale pink in females, pinkish white with purplish tips in males; caudal fin pink to pinkish grey in females, caudal fin rays grey with pink membranes in males.
</paragraph>
<paragraph id="76303696FFF29A55E620FA9EFA8E2B17" blockId="3.[812,1452,1340,1556]" pageId="3" pageNumber="388">
Preserved, head and body yellowish brown to dark greyish brown, with usually only skin behind scales dark-pigmented, giving fish a pale-headed appearance (
<figureCitation id="EEB42A13FFF29A55E1DDFADEFAB62897" box="[1233,1299,1404,1428]" captionStart="Fig" captionStartId="1.[113,148,1173,1195]" captionTargetBox="[113,1451,189,1158]" captionTargetId="figure-459@1.[113,1451,189,1158]" captionTargetPageId="1" captionText="Fig. 1. Dorsal, lateral and ventral views of preserved male Pogoneleotris heterolepis (ZRC56949, 160.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.4577072" httpUri="https://zenodo.org/record/4577072/files/figure.png" pageId="3" pageNumber="388">Fig. 1</figureCitation>
). Some small specimens with head dark grey and body almost black but underside of head dull whitish. Fins usually plain pale yellow to white, with pectoral, pelvic and caudal fins variably dark grey in some specimens. Reaches
<quantity id="B1779B73FFF29A55E1AEFA5EFB592B10" box="[1186,1276,1532,1556]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="388" unit="mm" value="210.0">210 mm</quantity>
SL.
</paragraph>
<paragraph id="76303696FFF29A55E620F999FACC2BB7" blockId="3.[812,1452,1595,1716]" pageId="3" pageNumber="388">
<emphasis id="44FBEA84FFF29A55E620F999FC2F2B50" bold="true" box="[812,906,1595,1619]" pageId="3" pageNumber="388">Habitat.</emphasis>
Known only from estuarine habitats; found in turbid rivers with mud substrate (
<figureCitation id="EEB42A13FFF29A55E159F9FEFB3C2B77" box="[1109,1177,1628,1652]" captionStart="Fig" captionStartId="4.[135,170,1334,1356]" captionTargetBox="[135,774,838,1318]" captionTargetId="figure-513@4.[135,775,838,1318]" captionTargetPageId="4" captionText="Fig. 5. Simunjan River habitat of this species; showing fishers’ boat moored beside lines onto bank from gill net." figureDoi="http://doi.org/10.5281/zenodo.4577080" httpUri="https://zenodo.org/record/4577080/files/figure.png" pageId="3" pageNumber="388">Fig. 5</figureCitation>
).
<bibRefCitation id="121E4B67FFF29A55E1BDF9FEFA092B77" author="Tan HH &amp; Grinang J" box="[1201,1452,1628,1652]" pageId="3" pageNumber="388" pagination="1 - 8" refId="ref3749" refString="Tan HH &amp; Grinang J (2018) Pogoneleotris heterolepis (Teleostei: Eleotridae), a rare fish from Borneo: biological data and remarks on the type locality and type material. Ichthyological Exploration of Freshwaters, 1082: 1 - 8." type="journal article" year="2018">Tan &amp; Grinang (2018)</bibRefCitation>
describe the habitat from which these came, and dissected one specimen (which contained crustacean remains).
</paragraph>
<paragraph id="76303696FFF29A55E620F979FB412510" blockId="3.[811,1453,1755,2068]" pageId="3" pageNumber="388">
<emphasis id="44FBEA84FFF29A55E620F979FC6F2BF0" bold="true" box="[812,970,1755,1779]" pageId="3" pageNumber="388">Comparisons.</emphasis>
<taxonomicName id="B18F4D15FFF29A55E6DCF97EFB472BF7" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[976,1250,1756,1780]" class="Actinopterygii" family="Eleotridae" genus="Pogoneleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="388" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="44FBEA84FFF29A55E6DCF97EFB472BF7" box="[976,1250,1756,1780]" italics="true" pageId="3" pageNumber="388">Pogoneleotris heterolepis</emphasis>
</taxonomicName>
is unlike any other species of gudgeon, with its complex branched elongate sensory papillae, very small eyes, many multiple auxiliary scales on the body, distinctive sensory pore pattern and the stout fleshy lobes on the shoulder. It is the only butid and one of very few gobioid fishes (other than
<taxonomicName id="B18F4D15FFF29A55E1FCF8DEFADB2A97" authorityName="Boulenger" authorityYear="1901" box="[1264,1406,1916,1940]" class="Actinopterygii" family="Rhyacichthyidae" genus="Rhyacichthys" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="388" phylum="Chordata" rank="genus">
<emphasis id="44FBEA84FFF29A55E1FCF8DEFADB2A97" box="[1264,1406,1916,1940]" italics="true" pageId="3" pageNumber="388">Rhyacichthys</emphasis>
</taxonomicName>
and
<taxonomicName id="B18F4D15FFF29A55E620F83EFC122AB7" authorityName="Shibukawa, Iwata &amp; Viravong" authorityYear="2001" box="[812,951,1948,1972]" class="Actinopterygii" family="Odontobutidae" genus="Terateleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="388" phylum="Chordata" rank="genus">
<emphasis id="44FBEA84FFF29A55E620F83EFC122AB7" box="[812,951,1948,1972]" italics="true" pageId="3" pageNumber="388">Terateleotris</emphasis>
</taxonomicName>
) known to possess a preopercular canal that is joined to the oculoscapular canal, a condition first noted by Prachit Wongrat (unpublished thesis). It is also the only butid with a longitudinal papilla pattern.
</paragraph>
<paragraph id="76303696FFF59A52E73AFFD0FBB62D89" blockId="4.[566,1043,114,138]" box="[566,1043,114,138]" pageId="4" pageNumber="389">
<emphasis id="44FBEA84FFF59A52E73AFFD0FC702D8A" box="[566,981,114,137]" italics="true" pageId="4" pageNumber="389">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
2019
</paragraph>
<caption id="22F0661EFFF59A52E584FD5BFA662E13" ID-DOI="http://doi.org/10.5281/zenodo.4577078" ID-Zenodo-Dep="4577078" box="[136,1475,761,784]" httpUri="https://zenodo.org/record/4577078/files/figure.png" pageId="4" pageNumber="389" startId="4.[136,171,761,783]" targetBox="[136,1474,189,746]" targetPageId="4">
<paragraph id="76303696FFF59A52E584FD5BFA662E13" blockId="4.[136,1475,761,784]" box="[136,1475,761,784]" pageId="4" pageNumber="389">
Fig. 4. Fresh
<taxonomicName id="B18F4D15FFF59A52E407FD5BFDA12E0D" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[267,516,761,782]" class="Actinopterygii" family="Eleotridae" genus="Pogoneleotris" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="389" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="44FBEA84FFF59A52E407FD5BFDA12E0D" box="[267,516,761,782]" italics="true" pageId="4" pageNumber="389">Pogoneleotris heterolepis</emphasis>
</taxonomicName>
from Simunjan, Sarawak; ZRC56949, female above (166.7 mm SL), male below (160.0 mm SL).
</paragraph>
</caption>
<paragraph id="76303696FFF59A52E64FFCE5FBF92E9C" blockId="4.[835,1474,839,927]" pageId="4" pageNumber="389">
for field assistance; P.S.Y. Ng, for providing
<figureCitation id="EEB42A13FFF59A52E02BFCE5FACD2E5C" box="[1319,1384,839,863]" captionStart="Fig" captionStartId="2.[137,172,1512,1534]" captionTargetBox="[136,775,189,1496]" captionTargetId="figure-7@2.[136,775,189,1496]" captionTargetPageId="2" captionText="Fig. 2. Dorsal, lateral and ventral views (right side reversed) of head of fresh Pogoneleotris heterolepis (ZRC56949, 160.0 mm SL male); showing extent of scalation and sensory papillae." figureDoi="http://doi.org/10.5281/zenodo.4577076" httpUri="https://zenodo.org/record/4577076/files/figure.png" pageId="4" pageNumber="389">Fig. 2</figureCitation>
; and the Lee Kong Chian Natural History Museum, for supporting research and travel funds.
</paragraph>
</subSubSection>
<caption id="22F0661EFFF59A52E58BFA94FDCA286B" ID-DOI="http://doi.org/10.5281/zenodo.4577080" ID-Zenodo-Dep="4577080" httpUri="https://zenodo.org/record/4577080/files/figure.png" pageId="4" pageNumber="389" startId="4.[135,170,1334,1356]" targetBox="[135,774,838,1318]" targetPageId="4">
<paragraph id="76303696FFF59A52E58BFA94FDCA286B" blockId="4.[135,774,1334,1385]" pageId="4" pageNumber="389">Fig. 5. Simunjan River habitat of this species; showing fishers’ boat moored beside lines onto bank from gill net.</paragraph>
</caption>
<subSubSection id="3E95651DFFF59A52E584FA02FD132BFB" pageId="4" pageNumber="389" type="discussion">
<paragraph id="76303696FFF59A52E584FA02FE602B3B" blockId="4.[136,775,1440,1592]" pageId="4" pageNumber="389">
<emphasis id="44FBEA84FFF59A52E584FA02FF5228BB" bold="true" box="[136,247,1440,1464]" pageId="4" pageNumber="389">Remarks.</emphasis>
It is commonly known as the bearded gudgeon,
<emphasis id="44FBEA84FFF59A52E584FA62FEED28DB" box="[136,328,1472,1496]" italics="true" pageId="4" pageNumber="389">Ikan buccat butta</emphasis>
or
<emphasis id="44FBEA84FFF59A52E47CFA63FE4B28DB" box="[368,494,1472,1496]" italics="true" pageId="4" pageNumber="389">Batu betutu</emphasis>
, and has been referred to as the
<collectingCountry id="0E987606FFF59A52E5C7FA42FE8228FB" box="[203,295,1504,1528]" name="Malaysia" pageId="4" pageNumber="389">Sarawak</collectingCountry>
mud gudgeon. At the Rajang River delta, the local name in the Melanau dialect is
<emphasis id="44FBEA84FFF59A52E729F9A2FD0B2B1B" box="[549,686,1536,1560]" italics="true" pageId="4" pageNumber="389">Ilong mapak</emphasis>
[=blind fish] (
<bibRefCitation id="121E4B67FFF59A52E5CBF982FE1C2B3B" author="Tan HH &amp; Grinang J" box="[199,441,1568,1592]" pageId="4" pageNumber="389" pagination="1 - 8" refId="ref3749" refString="Tan HH &amp; Grinang J (2018) Pogoneleotris heterolepis (Teleostei: Eleotridae), a rare fish from Borneo: biological data and remarks on the type locality and type material. Ichthyological Exploration of Freshwaters, 1082: 1 - 8." type="journal article" year="2018">Tan &amp; Grinang, 2018</bibRefCitation>
).
</paragraph>
<paragraph id="76303696FFF59A52E584F9C2FD132BFB" blockId="4.[136,776,1632,1784]" pageId="4" pageNumber="389">This species’ conservation status has not been assessed by the IUCN. As it is regularly consumed by the local people and is restricted to mangrove habitats (which are generally threatened throughout South-east Asia), its conservation status deserves assessment sooner rather than later.</paragraph>
</subSubSection>
</treatment>
</document>
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