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<document id="4367BDD6C85AE58612C7246427FFA503" ID-DOI="10.11646/zootaxa.5323.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8212306" ID-ZooBank="DAAF3E31-A245-4421-A41B-E9BE1538ABF9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691078968945" checkinUser="plazi" docAuthor="Noguera, Felipe A. &amp; Santos-Silva, Antonio" docDate="2023" docId="FE18879D9B0E830CFF2BFF63FAB5F802" docLanguage="en" docName="zootaxa.5323.4.1.pdf" docOrigin="Zootaxa 5323 (4)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynidius skillmani Noguera &amp; Santos-Silva 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="459" masterDocId="0221FFE59B098304FFBCFFF4FFD1FFDF" masterDocTitle="New species and records in Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae)" masterLastPageNumber="476" masterPageNumber="451" pageNumber="458" updateTime="1691079457083" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5822742F7A7730D1D19F2EEA486CC65A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A9C35236911F5DB1C7D16E21E023FEB1">
<mods:title id="43DF636813ACC493B8D5680320C2B013">New species and records in Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae)</mods:title>
</mods:titleInfo>
<mods:name id="8F0727E2FC7525E7DD968DA6EECB0E09" type="personal">
<mods:role id="1DDEB9FFDEEEC8DACEECDA4ADD162D22">
<mods:roleTerm id="EDA536721F60163A251FE513782B9004">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1826F1C6B81EC74E9CCA5E8480EDB4F5">Noguera, Felipe A.</mods:namePart>
<mods:affiliation id="23ECA48B693B28C374E435246B61D319">Estacioìn de Biologiìa Chamela, Instituto de Biologiìa, Universidad Nacional Autoìnoma de Meìxico, Apartado Postal 21, San Patricio, Jalisco 48980, Mexico.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="D1DB0DAD3FD40CC21B00DD0E71ED1F4F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F41D05B4C6A305FAB1C598426CECA90F">Santos-Silva, Antonio</mods:namePart>
<mods:affiliation id="B9925A9D0F4ECFA0650496916FC95310">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:titleInfo id="1A6D4FA9F037356233C897A19530E17C">
<mods:title id="FF0352ACCBFBE6950ED2F299ED803AF7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="17D60A0EC1CAA2446A4FD177E27560C2">
<mods:date id="CE4364AB7DE59B54A4D3976C057FE302">2023</mods:date>
<mods:detail id="74C3E6A5116B4D1515E6BAC1B07F7C11" type="pubDate">
<mods:number id="FEF22ABEDC3D72DF764F22349C5205F2">2023-08-03</mods:number>
</mods:detail>
<mods:detail id="AFC00446DC2056F60A02151434E05A7B" type="volume">
<mods:number id="6F95709430EED48450149B4A5E1B250A">5323</mods:number>
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<mods:detail id="2B66A55206FB50C56067ECA692B8D911" type="issue">
<mods:number id="F5142F1B61FD9FA9A9FC591DE28F74DC">4</mods:number>
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<mods:url id="313A82F4B9BDCBFAFA9EA94FE6F4F108">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification id="1F313D87458BA716544174CD479FEF67">journal article</mods:classification>
<mods:identifier id="693D3A9C138D21D0C8CFD51AF1F3D3FF" type="DOI">10.11646/zootaxa.5323.4.1</mods:identifier>
<mods:identifier id="0A2A8EB820A2E7401D84B2E68FB333CD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="521DB7E7B1C23C92C517D9ADE3609AC0" type="Zenodo-Dep">8212306</mods:identifier>
<mods:identifier id="5E6C677E1883A531378671F3E713416C" type="ZooBank">DAAF3E31-A245-4421-A41B-E9BE1538ABF9</mods:identifier>
</mods:mods>
<treatment id="FE18879D9B0E830CFF2BFF63FAB5F802" ID-DOI="http://doi.org/10.5281/zenodo.8212330" ID-Zenodo-Dep="8212330" LSID="urn:lsid:plazi:treatment:FE18879D9B0E830CFF2BFF63FAB5F802" httpUri="http://treatment.plazi.org/id/FE18879D9B0E830CFF2BFF63FAB5F802" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
<subSubSection id="3EAB65009B0E8303FF2BFF63FE2BFF6E" box="[151,506,151,178]" pageId="7" pageNumber="458" type="nomenclature">
<paragraph id="760E368B9B0E8303FF2BFF63FE2BFF6E" blockId="7.[151,506,151,213]" box="[151,506,151,178]" pageId="7" pageNumber="458">
<heading id="2D4681E79B0E8303FF2BFF63FE2BFF6E" bold="true" box="[151,506,151,178]" fontSize="11" level="1" pageId="7" pageNumber="458" reason="1">
<emphasis id="44C5EA999B0E8303FF2BFF63FE2BFF6E" bold="true" box="[151,506,151,178]" pageId="7" pageNumber="458">
<taxonomicName id="B1B14D089B0E8303FF2BFF63FE45FF6D" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[151,404,151,178]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="skillmani" status="sp. nov.">
<emphasis id="44C5EA999B0E8303FF2BFF63FE45FF6D" bold="true" box="[151,404,151,178]" italics="true" pageId="7" pageNumber="458">Phrynidius skillmani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FF657E29B0E8303FE20FF63FE2BFF6E" box="[412,506,151,177]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B0E8303FF2BFF4FFEF6FF0A" box="[151,295,187,213]" pageId="7" pageNumber="458" type="description">
<paragraph id="760E368B9B0E8303FF2BFF4FFEF6FF0A" blockId="7.[151,506,151,213]" box="[151,295,187,213]" pageId="7" pageNumber="458">
(
<figureCitation id="EE8A2A0E9B0E8303FF23FF4FFECEFF0A" box="[159,287,187,213]" captionStart="FIGURES 14–17" captionStartId="7.[152,270,1519,1543]" captionTargetBox="[152,1435,382,1495]" captionTargetId="figure-55@7.[152,1436,381,1495]" captionTargetPageId="7" captionText="FIGURES 14–17. Phrynidius skillmani sp. nov., holotype female: 14) Dorsal habitus; 15) Ventral habitus; 16) Lateral habitus; 17) Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.8212320" httpUri="https://zenodo.org/record/8212320/files/figure.png" pageId="7" pageNumber="458">Figs 14–17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B0E8303FF2BFEF7FE6AFEBA" pageId="7" pageNumber="458" type="materials_examined">
<paragraph id="760E368B9B0E8303FF2BFEF7FE6AFEBA" blockId="7.[151,1437,259,357]" pageId="7" pageNumber="458">
<emphasis id="44C5EA999B0E8303FF2BFEF7FE9DFEC2" bold="true" box="[151,332,259,285]" pageId="7" pageNumber="458">Type material.</emphasis>
<materialsCitation id="C6D93CD69B0E8303FEE6FEF7FE6AFEBA" collectingDate="2019-05-27" collectingDateMax="2019-05-28" collectingDateMin="2019-05-27" collectionCode="FSCA, FWSC" collectorName="de Caral &amp; Skillman &amp; Wappes &amp; Monzon" country="Guatemala" county="Morales" elevation="600" latitude="15.407148" location="Sierra de Caral" longLatPrecision="1" longitude="-88.69626" municipality="Finca la Firmeza" pageId="7" pageNumber="458" specimenCount="1" stateProvince="Izabal" typeStatus="holotype">
<typeStatus id="A90A88299B0E8303FEE6FEF7FE13FEC2" box="[346,450,259,285]" pageId="7" pageNumber="458">Holotype</typeStatus>
female from
<collectingCountry id="0EA6761B9B0E8303FDD3FEF7FCF1FEC2" box="[623,800,259,285]" name="Guatemala" pageId="7" pageNumber="458">GUATEMALA</collectingCountry>
,
<emphasis id="44C5EA999B0E8303FC8FFEF7FCA8FEC2" box="[819,889,259,285]" italics="true" pageId="7" pageNumber="458">
<collectingRegion id="B475F8699B0E8303FC8FFEF7FCA8FEC2" box="[819,889,259,285]" country="Guatemala" name="Izabal" pageId="7" pageNumber="458">Izabal</collectingRegion>
</emphasis>
:
<collectingCounty id="9F6F4E079B0E8303FC33FEF7FC3FFEC2" box="[911,1006,259,285]" pageId="7" pageNumber="458">Morales</collectingCounty>
,
<collectingMunicipality id="966AACF19B0E8303FC43FEF7FB1CFEC2" box="[1023,1229,259,285]" pageId="7" pageNumber="458">Finca la Firmeza</collectingMunicipality>
,
<location id="736E60509B0E8303FB63FEF7FA49FEC2" LSID="urn:lsid:plazi:treatment:FE18879D9B0E830CFF2BFF63FAB5F802:736E60509B0E8303FB63FEF7FA49FEC2" box="[1247,1432,259,285]" country="Guatemala" county="Morales" latitude="15.407148" longLatPrecision="1" longitude="-88.69626" municipality="Finca la Firmeza" name="Sierra de Caral" pageId="7" pageNumber="458" stateProvince="Izabal">
Sierra
<collectorName id="DB44535D9B0E8303FA8CFEF7FA49FEC2" box="[1328,1432,259,285]" pageId="7" pageNumber="458">de Caral</collectorName>
</location>
, FUNDAECO,
<quantity id="B1499B6E9B0E8303FE81FED3FE57FE9D" box="[317,390,295,322]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="458" unit="m" value="600.0">
<elevation id="FD9CD1B89B0E8303FE81FED3FE57FE9D" box="[317,390,295,322]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="458" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<geoCoordinate id="1385504C9B0E8303FE2EFED3FDD8FE9D" box="[402,521,295,322]" degrees="15.407148" direction="north" orientation="latitude" pageId="7" pageNumber="458" precision="1" value="15.407148">15.407148</geoCoordinate>
<geoCoordinate id="1385504C9B0E8303FDACFED3FD42FE9D" box="[528,659,295,322]" degrees="88.696255" direction="west" orientation="longitude" pageId="7" pageNumber="458" precision="1" value="-88.69626">-88.696255</geoCoordinate>
, beating along road,
<date id="020F104B9B0E8303FCC7FED3FBC7FE9E" box="[891,1046,295,321]" pageId="7" pageNumber="458" value="2019-05-27" valueMax="2019-05-28" valueMin="2019-05-27">
<collectingDate id="124BE9A39B0E8303FCC7FED3FBC7FE9E" box="[891,1046,295,321]" pageId="7" pageNumber="458" value="2019-05-27" valueMax="2019-05-28" valueMin="2019-05-27">27-28.V.2019</collectingDate>
</date>
,
<collectorName id="DB44535D9B0E8303FB9DFED3FB58FE9E" box="[1057,1161,295,321]" pageId="7" pageNumber="458">Skillman</collectorName>
,
<collectorName id="DB44535D9B0E8303FB28FEDCFB3DFE9D" box="[1172,1260,296,322]" pageId="7" pageNumber="458">Wappes</collectorName>
&amp;
<collectorName id="DB44535D9B0E8303FAB3FED3FABDFE9E" box="[1295,1388,295,321]" pageId="7" pageNumber="458">Monzón</collectorName>
leg. (
<collectionCode id="10A0AE4E9B0E8303FF23FEBFFF38FEBA" box="[159,233,331,357]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="7" pageNumber="458" type="Museum">FSCA</collectionCode>
, formerly
<collectionCode id="10A0AE4E9B0E8303FEE2FEBFFE7EFEBA" box="[350,431,331,357]" pageId="7" pageNumber="458">FWSC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="22CE66039B0E8303FF24FA1BFE53F9F4" ID-DOI="http://doi.org/10.5281/zenodo.8212320" ID-Zenodo-Dep="8212320" httpUri="https://zenodo.org/record/8212320/files/figure.png" pageId="7" pageNumber="458" startId="7.[152,270,1519,1543]" targetBox="[152,1435,382,1495]" targetPageId="7" targetType="figure">
<paragraph id="760E368B9B0E8303FF24FA1BFE53F9F4" blockId="7.[151,1436,1519,1579]" pageId="7" pageNumber="458">
<emphasis id="44C5EA999B0E8303FF24FA1BFE8BF9D8" bold="true" box="[152,346,1519,1543]" pageId="7" pageNumber="458">FIGURES 14–17.</emphasis>
<taxonomicName id="B1B14D089B0E8303FEE3FA1BFDE0F9D8" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[351,561,1519,1543]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="skillmani" status="sp. nov.">
<emphasis id="44C5EA999B0E8303FEE3FA1BFDE0F9D8" box="[351,561,1519,1543]" italics="true" pageId="7" pageNumber="458">Phrynidius skillmani</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B0E8303FD8BFA1BFD56F9D8" bold="true" box="[567,647,1519,1543]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="5FF657E29B0E8303FD8BFA1BFD56F9D8" box="[567,647,1519,1543]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female:
<emphasis id="44C5EA999B0E8303FCF8FA1BFCB7F9D8" bold="true" box="[836,870,1519,1543]" pageId="7" pageNumber="458">14)</emphasis>
Dorsal habitus;
<emphasis id="44C5EA999B0E8303FBB7FA1BFBFCF9D8" bold="true" box="[1035,1069,1519,1543]" pageId="7" pageNumber="458">15)</emphasis>
Ventral habitus;
<emphasis id="44C5EA999B0E8303FB6BFA1BFB28F9D8" bold="true" box="[1239,1273,1519,1543]" pageId="7" pageNumber="458">16)</emphasis>
Lateral habitus;
<emphasis id="44C5EA999B0E8303FF2BF9E7FF68F9F4" bold="true" box="[151,185,1555,1579]" pageId="7" pageNumber="458">17)</emphasis>
Head, frontal view.
</paragraph>
</caption>
<subSubSection id="3EAB65009B0E8303FF7BF989FE10F8DC" pageId="7" pageNumber="458" type="diagnosis">
<paragraph id="760E368B9B0E8303FF7BF989FE10F8DC" blockId="7.[151,1436,1661,2012]" pageId="7" pageNumber="458">
<emphasis id="44C5EA999B0E8303FF7BF989FE93F948" bold="true" box="[199,322,1661,1687]" pageId="7" pageNumber="458">Diagnosis.</emphasis>
<taxonomicName id="B1B14D089B0E8303FEF4F989FDE0F948" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[328,561,1661,1687]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="skillmani" status="sp. nov.">
<emphasis id="44C5EA999B0E8303FEF4F989FDE0F948" box="[328,561,1661,1687]" italics="true" pageId="7" pageNumber="458">Phrynidius skillmani</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B0E8303FD8BF989FD40F948" bold="true" box="[567,657,1661,1687]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="5FF657E29B0E8303FD8BF989FD40F948" box="[567,657,1661,1687]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="B1B14D089B0E8303FC98F98AFC41F948" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[804,912,1661,1687]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="obrieni" status="sp. nov.">
<emphasis id="44C5EA999B0E8303FC98F98AFC41F948" box="[804,912,1661,1687]" italics="true" pageId="7" pageNumber="458">P. obrieni</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B0E8303FC2BF989FC21F948" bold="true" box="[919,1008,1661,1687]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="5FF657E29B0E8303FC2BF989FC21F948" box="[919,1008,1661,1687]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="B1B14D089B0E8303FC42F98AFBA4F948" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1022,1141,1661,1687]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="wibmeri" status="sp. nov.">
<emphasis id="44C5EA999B0E8303FC42F98AFBA4F948" box="[1022,1141,1661,1687]" italics="true" pageId="7" pageNumber="458">P. wibmeri</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B0E8303FBC0F989FB04F948" bold="true" box="[1148,1237,1661,1687]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="5FF657E29B0E8303FBC0F989FB04F948" box="[1148,1237,1661,1687]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="B1B14D089B0E8303FAAEF98AFA4DF948" authorityName="Gutierrez, Toledo &amp; Noguera" authorityYear="2020" box="[1298,1436,1661,1687]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="diminutus">
<emphasis id="44C5EA999B0E8303FAAEF98AFA4DF948" box="[1298,1436,1661,1687]" italics="true" pageId="7" pageNumber="458">P. diminutus</emphasis>
</taxonomicName>
especially by the antennomere III twice length of IV (about 1.5 times longer than IV in these three species); from
<taxonomicName id="B1B14D089B0E8303FF2BF932FE85F900" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[151,340,1733,1759]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="guatemalensis" status="sp. nov.">
<emphasis id="44C5EA999B0E8303FF2BF932FE85F900" box="[151,340,1733,1759]" italics="true" pageId="7" pageNumber="458">P. guatemalensis</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B0E8303FEE0F931FE66F900" bold="true" box="[348,439,1733,1759]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="5FF657E29B0E8303FEE0F931FE66F900" box="[348,439,1733,1759]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
especially by the pronotal punctures distinctly coarser and more abundant. See remarks under
<taxonomicName id="B1B14D089B0E8303FF62F91EFEB1F8DC" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[222,352,1769,1795]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="poriferus" status="sp. nov.">
<emphasis id="44C5EA999B0E8303FF62F91EFEB1F8DC" box="[222,352,1769,1795]" italics="true" pageId="7" pageNumber="458">P. poriferus</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B0E8303FEDBF91DFE10F8DC" bold="true" box="[359,449,1769,1795]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="5FF657E29B0E8303FEDBF91DFE10F8DC" box="[359,449,1769,1795]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B0E830CFF7BF8F9FAD2F866" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458" type="description">
<paragraph id="760E368B9B0E8303FF7BF8F9FBA0F84C" blockId="7.[151,1436,1661,2012]" pageId="7" pageNumber="458">
<emphasis id="44C5EA999B0E8303FF7BF8F9FDE3F8F8" bold="true" box="[199,562,1805,1831]" pageId="7" pageNumber="458">
Description.
<typeStatus id="A90A88299B0E8303FEDCF8F9FE1CF8F8" box="[352,461,1805,1831]" pageId="7" pageNumber="458">Holotype</typeStatus>
Female.
</emphasis>
Integument mostly black; palpi mostly brownish, gradually yellowish toward apex on posterior third; pedicel dark brown basally, reddish brown on remaining surface; antennomere III reddish brown on basal quarter, dark reddish brown on remaining surface; antennomere IV reddish brown basally, dark brown on remaining surface; antennomeres V–XI dark brown. Legs mostly dark brown.
</paragraph>
<paragraph id="760E368B9B0E830CFF7BF869FA8CFC7A" blockId="7.[151,1436,1661,2012]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
<emphasis id="44C5EA999B0E8303FF7BF869FEDFF868" bold="true" box="[199,270,1949,1975]" pageId="7" pageNumber="458">Head.</emphasis>
Frons coarsely, somewhat rugose-punctate; with minute yellowish-brown pubescence not obscuring integument, with short, sparse, spatulate yellowish-brown setae interspersed laterally. Antennal tubercles coarsely punctate basally, punctures distinctly finer than on frons, except smooth apex; with abundant, minute, both yellowish and whitish pubescence partially obscuring integument, except glabrous smooth area, and a few short, erect, spatulate yellowish-brown setae interspersed frontally. Vertex coarsely, deeply punctate, punctures finer close to prothorax; with abundant yellowish-brown pubescence partially obscuring integument on some areas, except glabrous area close to prothorax. Area behind eyes coarsely, shallowly punctate, punctures finer than on anterior area of vertex; with minute yellowish-brown pubescence not obscuring integument. Genae coarsely, somewhat rugose-punctate; 2.5 times width of lower eye lobe; with yellowish-brown pubescence, denser frontally. Wide central area of postclypeus with fringe of minute, thick brownish setae close to frons, moderately sparse, whitish on remaining surface, and long, erect yellowish-brown setae directed forward close to anteclypeus. Labrum with sparse whitish pubescence posteriorly, and abundant yellowish-brown setae directed forward anteriorly. Eye lobes with a single ommatidium between them; distance between upper eye lobes 0.50 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.75 times distance between outer margins of eyes; lower eye lobes longer than wide (1.33 times). Antennae 1.25 times elytral length, almost reaching posterior quarter of elytra. Scape with abundant, bristly brownish pubescence partially obscuring integument, with minute yellowish setae interspersed, and a few very short brownish squamiform setae interspersed laterally. Pedicel mostly glabrous on basal half, with abundant yellowish pubescence not obscuring integument on posterior half. Antennomere III slightly arched; with yellowish pubescence on basal fifth, yellowish white on apical fifth, brownish on remaining surface; with short, erect, sparse, spatulate brownish setae interspersed throughout. Antennomere IV with yellowish-white pubescence basally, brownish on remaining surface, and short, arched brownish setae interspersed. Antennomeres V–VII with yellowish-brown pubescence not obscuring integument. Antennomeres VIII–XI with abundant yellowish-white pubescence almost obscuring integument. Antennal formula (ratio) based on length of antennomere III: scape = 0.78; pedicel = 0.16; IV = 0.50; V = 0.24; VI = 0.22; VII = 0.20; VIII = 0.18; IX = 0.18; X = 0.18; XI = 0.24.
</paragraph>
<paragraph id="760E368B9B01830CFF7BFC5BFCCEF8F6" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="459">
<emphasis id="44C5EA999B01830CFF7BFC5BFEF6FC16" bold="true" box="[199,295,943,969]" pageId="8" pageNumber="459">Thorax.</emphasis>
Prothorax slightly wider than long; sides slightly rounded. Pronotum coarsely rugose-punctate, punctures finer close to anterior and posterior margins; with sinuous, irregular longitudinal carina centrally, located about central third, one circular gibbosity on each side of anterior region of central carina, and irregular gibbosity on each side of posterior region of central carina; with minute yellowish-brown pubescence not obscuring integument, absent on some areas (probably rubbed), denser on wide longitudinal band on center of posterior third; with short, arched, spatulate yellowish-white setae interspersed. Sides of prothorax coarsely rugose-punctate; with minute yellowish-white pubescence not obscuring integument, absent on some areas, and a few short, arched, spatulate yellowish-white setae interspersed. Prosternum coarsely punctate, punctures finer and shallower than on sides of prothorax; with abundant yellowish pubescence obscuring integument laterally close to procoxal cavities. Prosternal process coarsely punctate, punctures as on prosternum; narrowest area, located near base, 0.28 times procoxal width; with abundant, bristly yellowish pubescence centrally, sparser, more whitish anteriorly and posteriorly. Sides of mesoventrite, mesanepisternum, and mesepimeron coarsely punctate; central area of mesoventrite nearly smooth; sides of mesoventrite and part of mesanepisternum with dense yellowish-brown pubescence obscuring integument; remaining surface of mesoventrite and mesanepisternum with whitish pubescence not obscuring integument. Mesoventral process slightly widened from middle; apex 0.35 times mesocoxal width; with yellowish-brown pubescence not obscuring integument. Metaventrite coarsely punctate, with abundant yellowish-white pubescence not obscuring integument laterally and centrally, obscuring integument between these areas.
<emphasis id="44C5EA999B01830CFB7FFA1BFAC7F9D6" bold="true" box="[1219,1302,1519,1545]" pageId="8" pageNumber="459">Elytra.</emphasis>
Sub-ovoid; abundantly, coarsely, deeply rugose-punctate; with irregular, moderately elevated tubercles with blunt apices from middle, especially laterally; with minute yellowish-brown pubescence not obscuring integument, absent on some areas, and long, erect, sub-spatulate, both brownish and yellowish setae throughout.
<emphasis id="44C5EA999B01830CFBF6F9AFFB58F9AA" bold="true" box="[1098,1161,1627,1653]" pageId="8" pageNumber="459">Legs.</emphasis>
Femora with dense pale yellowish-brown pubescence obscuring integument, and arched, spatulate yellowish setae interspersed. Tibiae with pubescence as on femora, except posterior quarter of ventral surface of protibiae with dense, bristly yellowish-brown pubescence, dorsal sulcus of mesotibiae with abundant, short, erect yellowish-brown setae, and dense fringe of thick yellowish-brown setae on apical margin of all tibiae; with short, arched brownish setae interspersed, and a few short yellowish-white squamiform setae interspersed.
</paragraph>
<paragraph id="760E368B9B01830CFF7BF8C6FCF6F8AE" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="459">
<emphasis id="44C5EA999B01830CFF7BF8C6FE92F893" bold="true" box="[199,323,1842,1868]" pageId="8" pageNumber="459">Abdomen.</emphasis>
Ventrites with somewhat abundant light yellowish-brown pubescence not obscuring integument, and short yellowish-brown squamiform setae interspersed.
</paragraph>
<paragraph id="760E368B9B01830CFF7BF88EFAD2F866" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="459">
<emphasis id="44C5EA999B01830CFF7BF88EFE63F84A" bold="true" box="[199,434,1914,1941]" pageId="8" pageNumber="459">Dimensions in mm.</emphasis>
Total length, 6.60; prothoracic length, 1.85; anterior prothoracic width, 1.65; posterior prothoracic width, 1.85; maximum prothoracic width, 2.00; humeral width, 2.05; elytral length, 4.40.
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B01830CFF7BF837FAB5F802" box="[199,1380,1987,2013]" pageId="8" pageNumber="459" type="etymology">
<paragraph id="760E368B9B01830CFF7BF837FAB5F802" blockId="8.[151,1437,151,2013]" box="[199,1380,1987,2013]" pageId="8" pageNumber="459">
<emphasis id="44C5EA999B01830CFF7BF837FE9FF802" bold="true" box="[199,334,1987,2013]" pageId="8" pageNumber="459">Etymology.</emphasis>
The new species is dedicated to Frederick W. Skillman, one of the collectors of the
<typeStatus id="A90A88299B01830CFB47F837FAB0F802" box="[1275,1377,1987,2013]" pageId="8" pageNumber="459">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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