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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.523.6066" ID-GBIF-Dataset="afcfa837-6ba8-4a8d-bf4f-618e7380e8f3" ID-PMC="PMC4602297" ID-Pensoft-Pub="1313-2970-523-63" ID-PubMed="26478699" ID-ZBK="1E0C16D8DD11429BA6BC65862AA7C42B" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-523-63" ModsDocOrigin="ZooKeys 523" ModsDocTitle="Five new cryptic freshwater gastropod species from New Caledonia (Caenogastropoda, Truncatelloidea, Tateidae)" checkinTime="1451243964501" checkinUser="pensoft" docAuthor="Haase, Martin &amp; Zielske, Susan" docDate="2015" docId="FC98D2760680B2D1D5AC237DB742E5A6" docLanguage="en" docName="ZooKeys 523: 63-87" docOrigin="ZooKeys 523" docSource="http://dx.doi.org/10.3897/zookeys.523.6066" docTitle="Hemistomia andreae Haase &amp; Zielske, 2015, sp. n." docType="treatment" docUuid="1C80E381-43F7-43EB-9853-425C5C6B925E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="66" masterDocId="FF8EE7237E0EFFF13504FFC5FFC9876C" masterDocTitle="Five new cryptic freshwater gastropod species from New Caledonia (Caenogastropoda, Truncatelloidea, Tateidae)" masterLastPageNumber="87" masterPageNumber="63" pageNumber="65" updateTime="1668160744127" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five new cryptic freshwater gastropod species from New Caledonia (Caenogastropoda, Truncatelloidea, Tateidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haase, Martin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zielske, Susan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>523</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>63</mods:start>
<mods:end>87</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.523.6066</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.523.6066</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-523-63</mods:identifier>
<mods:identifier type="ZBK">1E0C16D8DD11429BA6BC65862AA7C42B</mods:identifier>
<mods:identifier type="ZooBank">1E0C16D8DD11429BA6BC65862AA7C42B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152062493" LSID="urn:lsid:zoobank.org:act:1C80E381-43F7-43EB-9853-425C5C6B925E" httpUri="http://treatment.plazi.org/id/FC98D2760680B2D1D5AC237DB742E5A6" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65">
<subSubSection pageId="2" pageNumber="65" type="multiple">
<paragraph pageId="2" pageNumber="65">Taxon classification Animalia Littorinimorpha Tateidae</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="65" type="nomenclature">
<paragraph pageId="2" pageNumber="65">
<taxonomicName LSID="http://zoobank.org/1C80E381-43F7-43EB-9853-425C5C6B925E" class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="65">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="65" type="type material">
<paragraph pageId="2" pageNumber="65">Type material.</paragraph>
<paragraph pageId="2" pageNumber="65">Holotype MNHN IM 2000-27858; paratypes MNHN IM 2000-27859 (&gt; 50), NHMW 110181 (10).</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="65" type="type locality">
<paragraph pageId="2" pageNumber="65">Type locality.</paragraph>
<paragraph pageId="2" pageNumber="65">
NeCa 12, Bouloupari:
<normalizedToken originalValue="Ouaméni-valley">Ouameni-valley</normalizedToken>
, small stream on W-side of road in secondary forest,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-21.829695">21°49'46.9&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="165.94525">165°56'42.9&quot;E</geoCoordinate>
, 22 May 2012.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="65" type="etymology">
<paragraph pageId="2" pageNumber="65">Etymology.</paragraph>
<paragraph pageId="2" pageNumber="65">
The new species is dedicated to the senior
<normalizedToken originalValue="authors">author's</normalizedToken>
daughter on the occasion of her
<normalizedToken originalValue="quinceañera">'quinceanera'</normalizedToken>
, the 15th birthday.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="65" type="diagnosis">
<paragraph pageId="2" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="65">
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. is very similar to
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia cockerelli" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="cockerelli">Hemistomia cockerelli</taxonomicName>
and
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia nyo" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="nyo">Hemistomia nyo</taxonomicName>
. It differs from both in a clearer separation of the opercular pegs and a much more delicate penis. The protoconch of the new species has more whorls than
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia nyo" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="nyo">Hemistomia nyo</taxonomicName>
and the palatal denticle is further behind the outer lip.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="65" type="shell">
<paragraph pageId="2" pageNumber="65">Shell.</paragraph>
<paragraph pageId="2" pageNumber="65">Conical, 2.2 times higher than wide, 4.5-5.5 whorls, without colour, transparent; protoconch faintly pitted with 1-1.25 whorls; palatal denticle large, elongate, c. 1/3 whorl behind outer lip; with columellar fold in the body whorl; aperture slightly higher than wide (Figs 2A, 3A,B, 4A,B).</paragraph>
<caption pageId="2" pageNumber="65">
<paragraph pageId="2" pageNumber="65">
Figure 3. Shells (all paratypes). A, B
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. C, D
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. E, F
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium aremuum" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="aremuum">Leiorhagium aremuum</taxonomicName>
sp. n. G
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium clandestinum" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="clandestinum">Leiorhagium clandestinum</taxonomicName>
sp. n. H, I
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium neteae" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="neteae">Leiorhagium neteae</taxonomicName>
sp. n.
</paragraph>
</caption>
<caption pageId="2" pageNumber="65">
<paragraph pageId="2" pageNumber="65">
Figure 4. Protoconchs (left) and close-up views of apical microstructure (right). A, B
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. C, D
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. E, F
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium aremuum" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="aremuum">Leiorhagium aremuum</taxonomicName>
sp. n. G, H
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium clandestinum" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="clandestinum">Leiorhagium clandestinum</taxonomicName>
I, J
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium neteae" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="neteae">Leiorhagium neteae</taxonomicName>
sp. n. Scale bars 50
<normalizedToken originalValue="µm">µm</normalizedToken>
(A, C, E, G, I), 10
<normalizedToken originalValue="µm">µm</normalizedToken>
(B, D, F, H, J).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="65" type="operculum">
<paragraph pageId="2" pageNumber="65">Operculum.</paragraph>
<paragraph pageId="2" pageNumber="65">Elongate-ellipsoidal, paucisprial, nucleus submarginal, orange, one large and one small non-calcareous white peg, well separated from each other (N=5) (Fig. 5A,B).</paragraph>
<caption pageId="2" pageNumber="65">
<paragraph pageId="2" pageNumber="65">
Figure 5. Operculum. A, B
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. C, D
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. E, F
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium aremuum" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="aremuum">Leiorhagium aremuum</taxonomicName>
sp. n. G, H
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium neteae" order="Littorinimorpha" pageId="2" pageNumber="65" phylum="Mollusca" rank="species" species="neteae">Leiorhagium neteae</taxonomicName>
sp. n.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="65" type="external features">
<paragraph pageId="2" pageNumber="65">External features.</paragraph>
<paragraph pageId="2" pageNumber="65">Epidermis without pigment, eyes black.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="66" type="mantle cavity">
<paragraph pageId="3" pageNumber="66">
<pageBreakToken pageId="3" pageNumber="66" start="start">Mantle</pageBreakToken>
cavity.
</paragraph>
<paragraph pageId="3" pageNumber="66">Ctenidium with 24-26 (2 males) or 25-28 (3 females) filaments; osphradium kidney-shaped, behind middle of ctenidium.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="66" type="digestive system">
<paragraph pageId="3" pageNumber="66">Digestive system.</paragraph>
<paragraph pageId="3" pageNumber="66">Radula formula (N=3) (Fig. 6A): R (rhachis or central tooth): 3 1 3/2 2, L (lateral tooth): 3 1 5, M1 (inner marginal tooth): 21-25, M2 (outer marginal tooth): 27-32; stomach without caecum; rectum close to pallial oviduct in females and to prostate in males.</paragraph>
<caption pageId="3" pageNumber="66">
<paragraph pageId="3" pageNumber="66">
Figure 6. Radula. A
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. B
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. C
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium aremuum" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="aremuum">Leiorhagium aremuum</taxonomicName>
sp. n. D
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium neteae" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="neteae">Leiorhagium neteae</taxonomicName>
sp. n. Arrows indicate membranous junction of flank and face of lateral teeth typical for most Pacific tateid genera (partly dissolved in A and D).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="66" type="female genitalia">
<paragraph pageId="3" pageNumber="66">Female genitalia.</paragraph>
<paragraph pageId="3" pageNumber="66">Ovary without lobes, proximal end c. 1.25 whorls below apex, comprising 0.25-0.5 whorls, eventually reaching stomach; anterior capsule gland yellow-orange, posterior capsule gland opaque-white, albumen gland milky-white; proximal loop of renal oviduct upright comprising 180°, distal loop short; bursa copulatrix pear-shaped, reaching only slightly behind albumen gland; bursal duct long, entering anterior; seminal receptacle on ventral edge of and as long as bursa (N=3) (Fig. 7A).</paragraph>
<caption pageId="3" pageNumber="66">
<paragraph pageId="3" pageNumber="66">
Figure 7. Female genitalia. A
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. B
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. C
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium aremuum" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="aremuum">Leiorhagium aremuum</taxonomicName>
sp. n. D
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium neteae" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="neteae">Leiorhagium neteae</taxonomicName>
sp. n. ac anterior capsule gland, ag albumen gland, bc bursa copulatrix, bd bursal duct, go genital opening, od oviduct, pc posterior capsule gland, rs receptaculum seminis, vc vestibular capsule gland, ve ventral channel.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="66" type="male genitalia">
<paragraph pageId="3" pageNumber="66">Male genitalia.</paragraph>
<paragraph pageId="3" pageNumber="66">Proximal end of lobate testis 1-1.25 whorls below apex, comprising 0.75 whorls, covering proximal end of stomach; vesicula seminalis arising from anterior third of testis; penis fairly delicate with blunt end (N=2) (Fig. 8A,B).</paragraph>
<caption pageId="3" pageNumber="66">
<paragraph pageId="3" pageNumber="66">
Figure 8. Penis. A, B
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. C
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. D
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium aremuum" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="aremuum">Leiorhagium aremuum</taxonomicName>
sp. n. E
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium neteae" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="neteae">Leiorhagium neteae</taxonomicName>
sp. n. Scale bars = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="66" type="remarks">
<paragraph pageId="3" pageNumber="66">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="66">
This is
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="genus">Hemistomia</taxonomicName>
sp. n. 1 of
<bibRefCitation author="Zielske, S" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="16" pageNumber="79" pagination="171 - 181" title="Molecular phylogeny and a modified approach of character-based barcoding refining the taxonomy of New Caledonian freshwater gastropods (Caenogastropoda, Truncatelloidea, Tateidae)." url="10.1016/j.ympev.2015.04.020" volume="89" year="2015">Zielske and Haase (2015)</bibRefCitation>
. Both
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. and
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia cockerelli" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="cockerelli">Hemistomia cockerelli</taxonomicName>
do have the columellar fold in the body whorl assumed to be unique in
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia nyo" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="nyo">Hemistomia nyo</taxonomicName>
by
<bibRefCitation author="Haase, M" journalOrPublisher="Hydrobiologia" pageId="14" pageNumber="77" pagination="43 - 129" title="Radiation of crenobiontic gastropods on an ancient continental island: the Hemistomia-clade in New Caledonia (Gastropoda: Hydrobiidae)." url="10.1023/A:1003219931171" volume="367" year="1998">Haase and Bouchet (1998)</bibRefCitation>
.
<taxonomicName class="Gastropoda" family="Tateidae" genus="Hemistomia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemistomia andreae" order="Littorinimorpha" pageId="3" pageNumber="66" phylum="Mollusca" rank="species" species="andreae">Hemistomia andreae</taxonomicName>
sp. n. is only known from the type locality.
</paragraph>
</subSubSection>
</treatment>
</document>