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<document id="3C4B40CC9D9F85C6F289276426B3749F" ID-CLB-Dataset="99512" ID-DOI="10.5252/geodiversitas2022v44a23" ID-GBIF-Dataset="1818f0c9-dbee-4fa4-b69a-ad6d3712913b" ID-ISSN="1638-9395" ID-Zenodo-Dep="6819412" ID-ZooBank="urn:lsid:zoobank.org:pub:999E7A6A-3781-4746-8519-62E1010D3880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657550474273" checkinUser="felipe" docAuthor="Klopfstein, Seraina" docDate="2022" docId="FC1A686EFFB6FFED8F0A473CFC910921" docLanguage="en" docName="Geodiversitas.44.23.645-664.pdf.imf" docOrigin="Geodiversitas 44 (23)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Crusopimpla minuta Klopfstein 2022, n. sp." docType="treatment" docUuid="B00D3333-29A0-4116-A6A6-073E6B10D466" docUuidSource="ZooBank" docVersion="8" lastPageNumber="653" masterDocId="00231016FFBFFFE78E394169FF940D6F" masterDocTitle="High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)" masterLastPageNumber="664" masterPageNumber="645" pageNumber="652" updateTime="1699358479615" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="E7E7B08E23E9BE1B8EDBF49CB8137102">High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)</mods:title>
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<mods:namePart id="B8249F8377342D0265EEA8D154224D97">Klopfstein, Seraina</mods:namePart>
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<mods:title id="1FE380BCBFAB99CE151AD7B61C0FB1A9">Geodiversitas</mods:title>
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<mods:date id="97E69CA39D8A69FF0D9996B852124321">2022</mods:date>
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<mods:number id="80D669FC7CCD7D3D05CECBC8526F7E09">2022-07-07</mods:number>
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<treatment id="FC1A686EFFB6FFED8F0A473CFC910921" ID-DOI="http://doi.org/10.5281/zenodo.6857798" ID-GBIF-Taxon="197821802" ID-Zenodo-Dep="6857798" LSID="urn:lsid:zoobank.org:act:B00D3333-29A0-4116-A6A6-073E6B10D466" httpUri="http://treatment.plazi.org/id/FC1A686EFFB6FFED8F0A473CFC910921" lastPageId="10" lastPageNumber="653" pageId="9" pageNumber="652">
<subSubSection id="3CA98AF3FFB6FFEE8F0A473CFDCC0B1E" box="[307,600,1621,1649]" pageId="9" pageNumber="652" type="nomenclature">
<paragraph id="740CD978FFB6FFEE8F0A473CFDCC0B1E" blockId="9.[307,600,1621,1649]" box="[307,600,1621,1649]" pageId="9" pageNumber="652">
<heading id="2F446E14FFB6FFEE8F0A473CFDCC0B1E" box="[307,600,1621,1649]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="652" reason="2">
<taxonomicName id="B3B3A2FBFFB6FFEE8F0A473CFD820B1F" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[307,534,1621,1648]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="652" phylum="Arthropoda" rank="species" species="minuta" status="sp. nov.">
<emphasis id="46C7056AFFB6FFEE8F0A473CFD820B1F" bold="true" box="[307,534,1621,1648]" italics="true" pageId="9" pageNumber="652">Crusopimpla minuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFB6FFEE8C24473EFDCC0B1E" box="[541,600,1623,1649]" pageId="9" pageNumber="652" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="3CA98AF3FFB6FFEE8FA7471FFE790BFF" box="[414,493,1654,1680]" pageId="9" pageNumber="652" type="description">
<paragraph id="740CD978FFB6FFEE8FA7471FFE790BFF" blockId="9.[414,493,1654,1680]" box="[414,493,1654,1680]" pageId="9" pageNumber="652">
(
<figureCitation id="EC88C5FDFFB6FFEE8F9E471FFE700BFF" box="[423,484,1654,1680]" captionStart="FIG" captionStartId="9.[132,143,704,721]" captionTargetBox="[134,1454,213,663]" captionTargetId="figure-634@9.[134,889,213,663]" captionTargetPageId="9" captionText="FIG. 4. — Holotype of Crusopimpla minuta n. sp. (FUR-13076), photograph (A), detail (B) and interpretative drawing (C), where dotted lines represent uncertain and/or interpolated interpretations. Scale bars: A, C, 1 mm; B, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6819420" httpUri="https://zenodo.org/record/6819420/files/figure.png" pageId="9" pageNumber="652">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB6FFEE8EBD47D0FC930BA5" box="[132,775,1721,1738]" pageId="9" pageNumber="652" type="description">
<paragraph id="740CD978FFB6FFEE8EBD47D0FC930BA5" blockId="9.[132,775,1721,1738]" box="[132,775,1721,1738]" pageId="9" pageNumber="652">
<uri id="0022D57AFFB6FFEE8EBD47D0FC930BA5" box="[132,775,1721,1738]" pageId="9" pageNumber="652">
urn:lsid:zoobank.org:act:
<uuid id="0015E3ADFFB6FFEE8F5847D0FC930BA5" box="[353,775,1721,1738]" pageId="9" pageNumber="652">B00D3333-29A0-4116-A6A6-073E6B10D466</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB6FFEE8EBD4793FEF50A43" pageId="9" pageNumber="652" type="materials_examined">
<paragraph id="740CD978FFB6FFEE8EBD4793FEF50A43" blockId="9.[132,775,1785,1836]" pageId="9" pageNumber="652">
TYPE MATERIAL. —
<materialsCitation id="C4DBD325FFB6FFEE8F7A4790FEF50A43" ID-GBIF-Occurrence="3863921309" collectionCode="FUR" collectorName="E. Rettig" country="Denmark" location="MorsO Kommune" municipality="Jutland" pageId="9" pageNumber="652" specimenCode="FUR-13076" specimenCount="1" specimenCount-female="1" stateProvince="Limfjord" typeStatus="holotype">
<typeStatus id="AB0867DAFFB6FFEE8F7A4790FE340A7E" box="[323,416,1785,1809]" pageId="9" pageNumber="652" type="holotype">Holotype</typeStatus>
<specimenCount id="62B512F1FFB6FFEE8F9F4790FE7D0A7E" box="[422,489,1785,1809]" pageId="9" pageNumber="652" type="female">female</specimenCount>
:
<specimenCode id="24157103FFB6FFEE8FC84790FDFE0A7E" box="[497,618,1785,1809]" pageId="9" pageNumber="652">FUR-13076</specimenCode>
(part and counterpart, leg.
<collectorName id="D946BCAEFFB6FFEE8EC2467DFEC20A43" box="[251,342,1812,1836]" pageId="9" pageNumber="652">E. Rettig</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB6FFEE8EBD462BFD660A35" box="[132,754,1858,1882]" pageId="9" pageNumber="652" type="etymology">
<paragraph id="740CD978FFB6FFEE8EBD462BFD660A35" blockId="9.[132,754,1858,1882]" box="[132,754,1858,1882]" pageId="9" pageNumber="652">ETYMOLOGY. — The name of this species refers to its small size.</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB6FFEE8EBD4618FD310A84" pageId="9" pageNumber="652" type="materials_examined">
<paragraph id="740CD978FFB6FFEE8EBD4618FDFB0ACD" blockId="9.[132,777,1904,1955]" pageId="9" pageNumber="652">
TYPE HORIZON AND LOCALITY. —
<collectingCountry id="0CA499E8FFB6FFEE8FD14619FDDF0AE7" box="[488,587,1904,1928]" name="Denmark" pageId="9" pageNumber="652">Denmark</collectingCountry>
,
<collectingMunicipality id="94684302FFB6FFEE8C6E4619FD370AE7" box="[599,675,1904,1928]" pageId="9" pageNumber="652">Jutland</collectingMunicipality>
,
<collectingRegion id="B677179AFFB6FFEE8C894619FC9D0AE7" box="[688,777,1904,1928]" pageId="9" pageNumber="652">Limfjord</collectingRegion>
region,
<location id="716C8FA3FFB6FFEE8EF646E0FE150ACD" LSID="urn:lsid:plazi:treatment:FC1A686EFFB6FFED8F0A473CFC910921:716C8FA3FFB6FFEE8EF646E0FE150ACD" box="[207,385,1929,1954]" country="Denmark" municipality="Jutland" name="MorsO Kommune" pageId="9" pageNumber="652" stateProvince="Limfjord">MorsØ Kommune</location>
,
<location id="716C8FA3FFB6FFEE8FB346E2FE410ACC" LSID="urn:lsid:plazi:treatment:FC1A686EFFB6FFED8F0A473CFC910921:716C8FA3FFB6FFEE8FB346E2FE410ACC" box="[394,469,1931,1955]" country="Denmark" municipality="Jutland" name="Svalklit" pageId="9" pageNumber="652" stateProvince="Limfjord">Svalklit</location>
. Cement stone.
</paragraph>
<paragraph id="740CD978FFB6FFEE8EBD46D0FD310A84" blockId="9.[132,775,1977,2027]" pageId="9" pageNumber="652">
MATERIAL EXAMINED. — Tentatively placed in this species:
<materialsCitation id="C4DBD325FFB6FFEE8CF746D0FEE00A84" ID-GBIF-Occurrence="3863921322" collectionCode="FUR" collectorName="M. Breiner" pageId="9" pageNumber="652" specimenCode="FUR- 14944" specimenCount="1">
<specimenCode id="24157103FFB6FFEE8CF746D0FF570A84" pageId="9" pageNumber="652">FUR- 14944</specimenCode>
(leg.
<collectorName id="D946BCAEFFB6FFEE8EC346BDFEFE0A84" box="[250,362,2003,2027]" pageId="9" pageNumber="652">M. Breiner</collectorName>
),
</materialsCitation>
<materialsCitation id="C4DBD325FFB6FFEE8F4346BAFD310A84" ID-GBIF-Occurrence="3863921325" box="[378,677,2003,2027]" collectionCode="FUR" collectorName="O. Burholt" pageId="9" pageNumber="652" specimenCode="FUR-16636" specimenCount="1">
<specimenCode id="24157103FFB6FFEE8F4346BAFE670A84" box="[378,499,2003,2027]" pageId="9" pageNumber="652">FUR-16636</specimenCode>
(leg.
<collectorName id="D946BCAEFFB6FFEE8C1046BAFD0E0A84" box="[553,666,2003,2027]" pageId="9" pageNumber="652">O. Burholt</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB6FFEE8D14425FFA090882" pageId="9" pageNumber="652" type="diagnosis">
<paragraph id="740CD978FFB6FFEE8D14425FFA25094E" blockId="9.[810,1457,821,1058]" pageId="9" pageNumber="652">
DIAGNOSIS. — Taxonomic placement:this species is very similar to the other species of
<taxonomicName id="B3B3A2FBFFB6FFEE8D8B4239FBB40E07" authorityName="Kopylov, Spasojevic &amp; Klopfstein" authorityYear="2018" box="[946,1056,848,872]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="652" phylum="Arthropoda" rank="genus">
<emphasis id="46C7056AFFB6FFEE8D8B4239FBB40E07" box="[946,1056,848,872]" italics="true" pageId="9" pageNumber="652">Crusopimpla</emphasis>
</taxonomicName>
described herein.In terms of typical pimpline characters, the only somewhat equivocal one is the shape of vein 2m-cu, which is less outward-bowed in this species than in any other pimpline described her from the Fur Formation. However, this might just be a consequence of its small size. Placement in
<taxonomicName id="B3B3A2FBFFB6FFEE8A9542D3FA890EBD" authorityName="Kopylov, Spasojevic &amp; Klopfstein" authorityYear="2018" box="[1196,1309,954,978]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="652" phylum="Arthropoda" rank="genus">
<emphasis id="46C7056AFFB6FFEE8A9542D3FA890EBD" box="[1196,1309,954,978]" italics="true" pageId="9" pageNumber="652">Crusopimpla</emphasis>
</taxonomicName>
then is not only supported by the very similar colour pattern to
<taxonomicName id="B3B3A2FBFFB6FFEE8AEE42BCFAB00E83" authorityName="Klopfstein" authorityYear="2022" box="[1239,1316,981,1004]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="652" phylum="Arthropoda" rank="species" species="rettigi">
<emphasis id="46C7056AFFB6FFEE8AEE42BCFAB00E83" box="[1239,1316,981,1004]" italics="true" pageId="9" pageNumber="652">C. rettigi</emphasis>
</taxonomicName>
and
<taxonomicName id="B3B3A2FBFFB6FFEE8B6942BCFA3C0E83" authorityName="Klopfstein" authorityYear="2022" box="[1360,1448,980,1004]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="652" phylum="Arthropoda" rank="species" species="violina">
<emphasis id="46C7056AFFB6FFEE8B6942BCFA3C0E83" box="[1360,1448,980,1004]" italics="true" pageId="9" pageNumber="652">C. violina</emphasis>
</taxonomicName>
, but also by the clearly visible propodeal carination.
<taxonomicName id="B3B3A2FBFFB6FFEE8ACE4286FA3A0969" authorityName="Klopfstein" authorityYear="2022" box="[1271,1454,1007,1031]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="652" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="46C7056AFFB6FFEE8ACE4286FA3A0969" box="[1271,1454,1007,1031]" italics="true" pageId="9" pageNumber="652">Crusopimpla minuta</emphasis>
</taxonomicName>
can be distinguished from any other species of the genus by its small size.
</paragraph>
<paragraph id="740CD978FFB6FFEE8D13453CFC290900" blockId="9.[810,1457,1109,1358]" box="[810,957,1109,1135]" pageId="9" pageNumber="652">DESCRIPTION</paragraph>
<paragraph id="740CD978FFB6FFEE8D14451CFC3F09E1" blockId="9.[810,1457,1109,1358]" box="[813,939,1141,1166]" pageId="9" pageNumber="652">
<emphasis id="46C7056AFFB6FFEE8D14451CFC3F09E1" box="[813,939,1141,1166]" italics="true" pageId="9" pageNumber="652">Preservation</emphasis>
</paragraph>
<paragraph id="740CD978FFB6FFEE8D1445FDFB280821" blockId="9.[810,1457,1109,1358]" pageId="9" pageNumber="652">
<typeStatus id="AB0867DAFFB6FFEE8D1445FDFC0109C1" box="[813,917,1172,1198]" pageId="9" pageNumber="652" type="holotype">Holotype</typeStatus>
in lateral view. Head and base of antennae preserved. Mesosoma rather well preserved, including details of metanotal area and part of propodeal carination; fore wings nearly complete, one partial hind wings partly discernible. Metasoma complete with ovipositor sheaths, the latter maybe broken off and only partial (unclear).
</paragraph>
<paragraph id="740CD978FFB6FFEE8D14441AFA090882" blockId="9.[810,1456,1395,1517]" pageId="9" pageNumber="652">
<emphasis id="46C7056AFFB6FFEE8D14441AFC2908E2" bold="true" box="[813,957,1395,1421]" pageId="9" pageNumber="652">
Body
<quantity id="B34B749DFFB6FFEE8D53441AFC2908E2" box="[874,957,1395,1421]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="652" unit="mm" value="5.3">5.3 mm</quantity>
</emphasis>
. Head and mesosoma black or dark brown, wing venation orange-brown.Antennae entirely orange (
<typeStatus id="AB0867DAFFB6FFEE8B1244FAFA1E08C1" box="[1323,1418,1427,1454]" pageId="9" pageNumber="652" type="holotype">holotype</typeStatus>
), as are the legs.
<specimenCode id="24157103FFB6FFEE8D9F44DAFC5308A2" box="[934,967,1459,1485]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="652">T1</specimenCode>
dark only at the very base, rest orange, remaining tergites entirely orange; ovipositor sheaths dark brown.
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB6FFED8D14477AFC910921" lastPageId="10" lastPageNumber="653" pageId="9" pageNumber="652" type="description">
<paragraph id="740CD978FFB6FFEE8D14477AFA9D0B23" blockId="9.[813,1455,1554,1613]" pageId="9" pageNumber="652">
<emphasis id="46C7056AFFB6FFEE8D14477AFCE60B42" bold="true" box="[813,882,1555,1581]" pageId="9" pageNumber="652">Head.</emphasis>
Transverse, wider than long; only base of antenna preserved, with scape rather short and wide.
</paragraph>
<paragraph id="740CD978FFB6FFEE8D14471BFA300A83" blockId="9.[811,1457,1650,2028]" pageId="9" pageNumber="652">
<emphasis id="46C7056AFFB6FFEE8D14471BFC3C0BE3" bold="true" box="[813,936,1650,1676]" pageId="9" pageNumber="652">Mesosoma.</emphasis>
Rather stout; notauli unclear, maybe indicated at front of mesoscutum; propodeum distinctly shortened, with nearly circular spiracle, pleural, lateral longitudinal and at least partial median longitudinal carinae present, both transverse carinae evident at least medially. Fore wing
<quantity id="B34B749DFFB6FFEE8B3A479BFAC90A63" box="[1283,1373,1778,1804]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="652" unit="mm" value="3.8">3.8 mm</quantity>
; areolet closed, quadrate, with uneven sides, 4-M very short, 2r-m about as long as 3r-m; 2m-cu not well preserved, only very little bent outwards; pterostigma 2.7 × longer than wide; 1cu-a meeting M + Cu clearly distad from 1-M; 3-Cu clearly longer than 2cu-a; radial cell 2.4 × longer than wide.Hind wing with 1-Rs about 1.5 × longer than 1rs-m, 1-Cu maybe 1.5 × as long as cu-a. Hind legs not well preserved, but rather stout.
</paragraph>
<caption id="20CC89F0FFB5FFED8EBD424BFE4F0E25" ID-DOI="http://doi.org/10.5281/zenodo.6819424" ID-Zenodo-Dep="6819424" httpUri="https://zenodo.org/record/6819424/files/figure.png" pageId="10" pageNumber="653" startId="10.[132,143,802,819]" targetBox="[132,1447,215,759]" targetPageId="10">
<paragraph id="740CD978FFB5FFED8EBD424BFE4F0E25" blockId="10.[132,1455,802,842]" pageId="10" pageNumber="653">
FIG. 5. — Holotype of
<taxonomicName id="B3B3A2FBFFB5FFED8F00424BFE430E5C" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[313,471,802,819]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="653" phylum="Arthropoda" rank="species" species="rettigi" status="sp. nov.">
<emphasis id="46C7056AFFB5FFED8F00424BFE430E5C" box="[313,471,802,819]" italics="true" pageId="10" pageNumber="653">Crusopimpla rettigi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFB5FFED8FE5424BFD9F0E5C" box="[476,523,802,819]" pageId="10" pageNumber="653" rank="species">n. sp.</taxonomicNameLabel>
(FUR-11209), photograph (
<emphasis id="46C7056AFFB5FFED8CD4424BFD6E0E5C" bold="true" box="[749,762,802,819]" pageId="10" pageNumber="653">A</emphasis>
) and interpretative drawing (
<emphasis id="46C7056AFFB5FFED8DDE424BFC600E5C" bold="true" box="[999,1012,802,819]" pageId="10" pageNumber="653">B</emphasis>
), where dotted lines represent uncertain and/or interpolated interpretations. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="740CD978FFB5FFED8EBD42FCFC910921" blockId="10.[130,775,917,1103]" pageId="10" pageNumber="653">
<emphasis id="46C7056AFFB5FFED8EBD42FCFF6E0EC0" bold="true" box="[132,250,917,943]" pageId="10" pageNumber="653">Metasoma.</emphasis>
With T1 showing an expansion just after its base in lateral view and an indication of what is probably the spiracle at about 0.25 its length; median longitudinal carina not well visible but maybe indicated;T2 probably shorter than wide (but lateral view), following tergites clearly transverse. Ovipositor sheaths at least 0.34 × as long as fore wing (tip not preserved),rather robust.
</paragraph>
</subSubSection>
</treatment>
</document>