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<document id="C0AC3DDF7755BC4DB40C8D14C5B4F92F" ID-CLB-Dataset="99512" ID-DOI="10.5252/geodiversitas2022v44a23" ID-GBIF-Dataset="1818f0c9-dbee-4fa4-b69a-ad6d3712913b" ID-ISSN="1638-9395" ID-Zenodo-Dep="6819412" ID-ZooBank="urn:lsid:zoobank.org:pub:999E7A6A-3781-4746-8519-62E1010D3880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657550474273" checkinUser="felipe" docAuthor="Klopfstein, Seraina" docDate="2022" docId="FC1A686EFFB5FFED8F0245FAFB780A84" docLanguage="en" docName="Geodiversitas.44.23.645-664.pdf.imf" docOrigin="Geodiversitas 44 (23)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Crusopimpla rettigi Klopfstein 2022, n. sp." docType="treatment" docUuid="CEF582AD-D348-4DF2-9C66-05C6E3E80885" docUuidSource="ZooBank" docVersion="8" lastPageNumber="653" masterDocId="00231016FFBFFFE78E394169FF940D6F" masterDocTitle="High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)" masterLastPageNumber="664" masterPageNumber="645" pageNumber="653" updateTime="1699358479615" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="95298AA8E61879A2F2FC681BFD41A7C7">High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)</mods:title>
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<mods:namePart id="1E90518516D1E5A66613CDBD92B70D44">Klopfstein, Seraina</mods:namePart>
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<mods:title id="7AEDB70DEDE83BB7A36C61F23D4A9835">Geodiversitas</mods:title>
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<mods:part id="2966A518A513C1532CE8B65463E19DFA">
<mods:date id="FFE99312FA924F176F2A1D0E89572DF8">2022</mods:date>
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<mods:number id="36BB4D908A63F2100BAD87751285B728">2022-07-07</mods:number>
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<mods:identifier id="883AC9C6C2CD513E74DE54162DDCF5E7" type="DOI">10.5252/geodiversitas2022v44a23</mods:identifier>
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<treatment id="FC1A686EFFB5FFED8F0245FAFB780A84" ID-DOI="http://doi.org/10.5281/zenodo.6857802" ID-GBIF-Taxon="197821804" ID-Zenodo-Dep="6857802" LSID="urn:lsid:zoobank.org:act:CEF582AD-D348-4DF2-9C66-05C6E3E80885" httpUri="http://treatment.plazi.org/id/FC1A686EFFB5FFED8F0245FAFB780A84" lastPageNumber="653" pageId="10" pageNumber="653">
<subSubSection id="3CA98AF3FFB5FFED8F0245FAFDC509C1" box="[315,593,1171,1198]" pageId="10" pageNumber="653" type="nomenclature">
<paragraph id="740CD978FFB5FFED8F0245FAFDC509C1" blockId="10.[315,593,1171,1198]" box="[315,593,1171,1198]" pageId="10" pageNumber="653">
<heading id="2F446E14FFB5FFED8F0245FAFDC509C1" box="[315,593,1171,1198]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="653" reason="2">
<taxonomicName id="B3B3A2FBFFB5FFED8F0245FAFD9B09C1" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[315,527,1171,1198]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="653" phylum="Arthropoda" rank="species" species="rettigi" status="sp. nov.">
<emphasis id="46C7056AFFB5FFED8F0245FAFD9B09C1" bold="true" box="[315,527,1171,1198]" italics="true" pageId="10" pageNumber="653">Crusopimpla rettigi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFB5FFED8C2F45FDFDC509C1" box="[534,593,1172,1198]" pageId="10" pageNumber="653" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB5FFED8FA745DDFC9C0868" pageId="10" pageNumber="653" type="description">
<paragraph id="740CD978FFB5FFED8FA745DDFE7909A1" blockId="10.[414,493,1204,1231]" box="[414,493,1204,1231]" pageId="10" pageNumber="653">
(
<figureCitation id="EC88C5FDFFB5FFED8F9E45DDFE7009A0" box="[423,484,1204,1231]" captionStart="FIG" captionStartId="10.[132,143,802,819]" captionTargetBox="[132,1447,215,759]" captionTargetId="figure-581@10.[699,983,487,760]" captionTargetPageId="10" captionText="FIG. 5. — Holotype of Crusopimpla rettigi n. sp. (FUR-11209), photograph (A) and interpretative drawing (B), where dotted lines represent uncertain and/or interpolated interpretations. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6819424" httpUri="https://zenodo.org/record/6819424/files/figure.png" pageId="10" pageNumber="653">Fig. 5</figureCitation>
)
</paragraph>
<paragraph id="740CD978FFB5FFED8EBD459FFC9C0868" blockId="10.[132,776,1270,1287]" box="[132,776,1270,1287]" pageId="10" pageNumber="653">
<uri id="0022D57AFFB5FFED8EBD459FFC9C0868" box="[132,776,1270,1287]" pageId="10" pageNumber="653">
urn:lsid:zoobank.org:act:
<uuid id="0015E3ADFFB5FFED8F65459FFC9C0868" box="[348,776,1270,1287]" pageId="10" pageNumber="653">CEF582AD-D348-4DF2-9C66-05C6E3E80885</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB5FFED8EBD4451FC9C0805" pageId="10" pageNumber="653" type="materials_examined">
<paragraph id="740CD978FFB5FFED8EBD4451FC9C0805" blockId="10.[132,776,1335,1386]" pageId="10" pageNumber="653">
TYPE MATERIAL. —
<materialsCitation id="C4DBD325FFB5FFED8F05445EFEAB0805" ID-GBIF-Occurrence="3863921311" collectionCode="FUR" collectorName="E. Rettig" country="Denmark" location="MorsO Kommune" municipality="Jutland" pageId="10" pageNumber="653" specimenCode="FUR-11209" specimenCount="1" specimenCount-female="1" stateProvince="Limfjord" typeStatus="holotype">
<typeStatus id="AB0867DAFFB5FFED8F05445EFE020820" box="[316,406,1335,1359]" pageId="10" pageNumber="653" type="holotype">Holotype</typeStatus>
<specimenCount id="62B512F1FFB5FFED8FA3445EFE4F0820" box="[410,475,1335,1359]" pageId="10" pageNumber="653" type="female">female</specimenCount>
:
<specimenCode id="24157103FFB5FFED8FDB4451FDC3083F" box="[482,599,1336,1360]" pageId="10" pageNumber="653">FUR-11209</specimenCode>
(part and counterpart; leg.
<collectorName id="D946BCAEFFB5FFED8EE7443BFEAC0805" box="[222,312,1362,1386]" pageId="10" pageNumber="653">E. Rettig</collectorName>
)
</materialsCitation>
.
<materialsCitation id="C4DBD325FFB5FFED8F7E443BFD2C0805" ID-GBIF-Occurrence="3863921314" box="[327,696,1362,1386]" collectionCode="FUR" country="Denmark" location="MorsO Kommune" municipality="Jutland" pageId="10" pageNumber="653" specimenCode="FUR-11219" specimenCount="1" specimenCount-female="1" stateProvince="Limfjord" typeStatus="paratype">
<typeStatus id="AB0867DAFFB5FFED8F7E443BFE370805" box="[327,419,1362,1386]" pageId="10" pageNumber="653" type="paratype">Paratypes</typeStatus>
(both
<specimenCount id="62B512F1FFB5FFED8FDB443BFDBB0805" box="[482,559,1362,1386]" pageId="10" pageNumber="653" type="female">females</specimenCount>
):
<specimenCode id="24157103FFB5FFED8C07443BFD2C0805" box="[574,696,1362,1386]" pageId="10" pageNumber="653">FUR-11219</specimenCode>
</materialsCitation>
<materialsCitation id="C4DBD325FFB5FFED8C80443BFC9C0805" ID-GBIF-Occurrence="3863921302" box="[697,776,1362,1386]" collectionCode="FUR" country="Denmark" location="Svalklit" municipality="Jutland" pageId="10" pageNumber="653" specimenCode="FUR-13074" specimenCount="1" specimenCount-female="1" stateProvince="Limfjord" typeStatus="paratype">
,
<specimenCode id="24157103FFB5FFED8CF8443BFC900805" box="[705,772,1362,1386]" pageId="10" pageNumber="653">13074</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB5FFED8EBD44EBFD3F08DB" pageId="10" pageNumber="653" type="etymology">
<paragraph id="740CD978FFB5FFED8EBD44EBFD3F08DB" blockId="10.[131,775,1409,1460]" pageId="10" pageNumber="653">
ETYMOLOGY. — This species is dedicated to Mr Erwin Rettig, who collected the
<specimenCount id="62B512F1FFB5FFED8F3C44F5FE0B08DB" box="[261,415,1436,1460]" pageId="10" pageNumber="653" type="generic">three specimens</specimenCount>
included in the
<typeStatus id="AB0867DAFFB5FFED8C7944F5FDFE08DB" box="[576,618,1436,1460]" pageId="10" pageNumber="653">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB5FFED8EBD44A5FD120B77" pageId="10" pageNumber="653" type="materials_examined">
<paragraph id="740CD978FFB5FFED8EBD44A5FD120B77" blockId="10.[132,777,1484,1561]" pageId="10" pageNumber="653">
TYPE HORIZON AND LOCALITY. —
<collectingCountry id="0CA499E8FFB5FFED8FD144A5FDDF088B" box="[488,587,1484,1508]" name="Denmark" pageId="10" pageNumber="653">Denmark</collectingCountry>
,
<collectingMunicipality id="94684302FFB5FFED8C6E44A5FD37088B" box="[599,675,1484,1508]" pageId="10" pageNumber="653">Jutland</collectingMunicipality>
,
<collectingRegion id="B677179AFFB5FFED8C8944A5FC9D088B" box="[688,777,1484,1508]" pageId="10" pageNumber="653">Limfjord</collectingRegion>
region,
<location id="716C8FA3FFB5FFED8EED448CFE190891" LSID="urn:lsid:plazi:treatment:FC1A686EFFB5FFED8F0245FAFB780A84:716C8FA3FFB5FFED8EED448CFE190891" box="[212,397,1509,1534]" country="Denmark" municipality="Jutland" name="MorsO Kommune" pageId="10" pageNumber="653" stateProvince="Limfjord">MorsØ Kommune</location>
,
<location id="716C8FA3FFB5FFED8FA3448FFDFA0891" LSID="urn:lsid:plazi:treatment:FC1A686EFFB5FFED8F0245FAFB780A84:716C8FA3FFB5FFED8FA3448FFDFA0891" box="[410,622,1510,1534]" country="Denmark" municipality="Jutland" name="Klinten ved Klitgard" pageId="10" pageNumber="653" stateProvince="Limfjord">Klinten ved Klitgård</location>
(holotype and FUR-11219), Svalklit (FUR-13074). Cement stone.
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB5FFED8EBD4758FE6B0A6E" pageId="10" pageNumber="653" type="diagnosis">
<paragraph id="740CD978FFB5FFED8EBD4758FEBD0B12" blockId="10.[132,775,1584,1793]" pageId="10" pageNumber="653">
DIAGNOSIS. — Taxonomic placement: this species is very similar to
<taxonomicName id="B3B3A2FBFFB5FFED8E984722FEF30B0C" authorityName="Klopfstein" authorityYear="2022" box="[161,359,1611,1635]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="653" phylum="Arthropoda" rank="species" species="violina">
<emphasis id="46C7056AFFB5FFED8E984722FEF30B0C" box="[161,359,1611,1635]" italics="true" pageId="10" pageNumber="653">Crusopimpla violina</emphasis>
</taxonomicName>
see reasoning for subfamily and genus placement there.
</paragraph>
<paragraph id="740CD978FFB5FFED8EBD4716FE6B0A6E" blockId="10.[132,775,1584,1793]" pageId="10" pageNumber="653">
Species diagnosis: together with
<taxonomicName id="B3B3A2FBFFB5FFED8F864716FDEA0BF8" authorityName="Klopfstein" authorityYear="2022" box="[447,638,1663,1687]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="653" phylum="Arthropoda" rank="species" species="violina">
<emphasis id="46C7056AFFB5FFED8F864716FDEA0BF8" box="[447,638,1663,1687]" italics="true" pageId="10" pageNumber="653">Crusopimpla violina</emphasis>
</taxonomicName>
,
<taxonomicName id="B3B3A2FBFFB5FFED8CB247E9FD4B0BF8" authorityName="Klopfstein" authorityYear="2022" box="[651,735,1664,1687]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="653" phylum="Arthropoda" rank="species" species="rettigi">
<emphasis id="46C7056AFFB5FFED8CB247E9FD4B0BF8" box="[651,735,1664,1687]" italics="true" pageId="10" pageNumber="653">C. rettigi</emphasis>
</taxonomicName>
can be distinguished from other species of the genus by their colour pattern. From
<emphasis id="46C7056AFFB5FFED8ECF47DCFEC80BA3" box="[246,348,1716,1740]" italics="true" pageId="10" pageNumber="653">
<taxonomicName id="B3B3A2FBFFB5FFED8ECF47DCFECC0BA3" authorityName="Klopfstein" authorityYear="2022" box="[246,344,1716,1740]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="653" phylum="Arthropoda" rank="species" species="violina">C. violina</taxonomicName>
,
</emphasis>
it can be distinguished by its much stouter tergites and the entirely orange antennae, while they are at least partly dark brown in the other species.
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFB5FFED8EBD465BFB780A84" pageId="10" pageNumber="653" type="description">
<paragraph id="740CD978FFB5FFED8EBD465BFE810A23" blockId="10.[130,776,1842,2028]" box="[132,277,1842,1868]" pageId="10" pageNumber="653">DESCRIPTION</paragraph>
<paragraph id="740CD978FFB5FFED8EBD463BFE960A04" blockId="10.[130,776,1842,2028]" box="[132,258,1874,1899]" pageId="10" pageNumber="653">
<emphasis id="46C7056AFFB5FFED8EBD463BFE960A04" box="[132,258,1874,1899]" italics="true" pageId="10" pageNumber="653">Preservation</emphasis>
</paragraph>
<paragraph id="740CD978FFB5FFED8EBD4618FAD00941" blockId="10.[130,776,1842,2028]" lastBlockId="10.[811,1457,917,1071]" pageId="10" pageNumber="653">
<typeStatus id="AB0867DAFFB5FFED8EBD4618FF730AE4" box="[132,231,1905,1931]" pageId="10" pageNumber="653" type="holotype">Holotype</typeStatus>
and
<typeStatus id="AB0867DAFFB5FFED8F22461BFEE00AE3" box="[283,372,1906,1932]" pageId="10" pageNumber="653" type="paratype">paratype</typeStatus>
FUR-
<quantity id="B34B749DFFB5FFED8F834618FD8C0AE4" box="[442,536,1905,1931]" metricMagnitude="2" metricUnit="m" metricValue="2.8496259999999998" pageId="10" pageNumber="653" unit="in" value="11219.0">11219 in</quantity>
dorsal,
<typeStatus id="AB0867DAFFB5FFED8C53461BFD570AE3" box="[618,707,1906,1932]" pageId="10" pageNumber="653" type="paratype">paratype</typeStatus>
FUR-
<quantity id="B34B749DFFB5FFED8EBD46F8FF4A0AC4" box="[132,222,1937,1963]" metricMagnitude="2" metricUnit="m" metricValue="3.320796" pageId="10" pageNumber="653" unit="in" value="13074.0">13074 in</quantity>
lateral view.Head in rather well preserved,in FUR-13074 including mandibles.Antennae nearly complete in
<typeStatus id="AB0867DAFFB5FFED8C4C46D8FD440AA4" box="[629,720,1969,1995]" pageId="10" pageNumber="653" type="holotype">holotype</typeStatus>
, only base preserved in
<typeStatus id="AB0867DAFFB5FFED8F0446BBFE370A83" box="[317,419,2002,2028]" pageId="10" pageNumber="653" type="paratype">paratypes</typeStatus>
. Mesosoma rather well preserved, including part of propodeal carination; fore wings in
<typeStatus id="AB0867DAFFB5FFED8B6C42FCFA250EC0" box="[1365,1457,917,943]" pageId="10" pageNumber="653" type="holotype">holotype</typeStatus>
complete, in
<typeStatus id="AB0867DAFFB5FFED8D8F42DCFB8C0EA0" box="[950,1048,949,975]" pageId="10" pageNumber="653" type="paratype">paratypes</typeStatus>
partial, hind wings indicated but with carination only weakly preserved, a bit better in FUR-11219. Metasoma in
<typeStatus id="AB0867DAFFB5FFED8D83429DFB820961" box="[954,1046,1012,1038]" pageId="10" pageNumber="653" type="holotype">holotype</typeStatus>
complete, in FUR-11219 missing most of ovipositor and in FUR-13074 from
<specimenCode id="24157103FFB5FFED8A87457CFB4B0940" box="[1214,1247,1045,1071]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="653">T7</specimenCode>
onwards.
</paragraph>
<paragraph id="740CD978FFB5FFED8D14453DFB9C0862" blockId="10.[811,1456,1108,1293]" pageId="10" pageNumber="653">
<emphasis id="46C7056AFFB5FFED8D14453DFBAB0901" bold="true" box="[813,1087,1108,1134]" pageId="10" pageNumber="653">Body 8.6-10.5 (10.5) mm</emphasis>
. Head and mesosoma black or dark brown, wing venation dark brown in forewing, with light base of pterostigma, and light brownish in hind wings. Antennae entirely orange (
<typeStatus id="AB0867DAFFB5FFED8DE545DDFBD509A0" box="[988,1089,1204,1231]" pageId="10" pageNumber="653" type="holotype">holotype</typeStatus>
).
<specimenCode id="24157103FFB5FFED8A6A45DDFBE009A1" box="[1107,1140,1204,1230]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="653">T1</specimenCode>
dark at base, orange at apex, remaining tergites entirely orange, legs orange; ovipositor sheaths dark brown.
</paragraph>
<paragraph id="740CD978FFB5FFED8D14445AFB1E08C2" blockId="10.[812,1455,1331,1454]" pageId="10" pageNumber="653">
<emphasis id="46C7056AFFB5FFED8D14445AFCE40822" bold="true" box="[813,880,1331,1357]" pageId="10" pageNumber="653">Head.</emphasis>
Transverse, wider than long; mandible with two teeth (FUR-13074), the lower a bit shorter; antennae with maybe 24-30 segments, median flagellomeres only slightly longer than wide, apical ones quadrate.
</paragraph>
<paragraph id="740CD978FFB5FFED8D1444BAFB790A43" blockId="10.[811,1458,1491,1836]" pageId="10" pageNumber="653">
<emphasis id="46C7056AFFB5FFED8D1444BAFC3E0882" bold="true" box="[813,938,1491,1517]" pageId="10" pageNumber="653">Mesosoma.</emphasis>
Rather stout; notauli only visible anteriorly, rest unclear; propodeum distinctly shortened, with some faint traces of carinae. Fore wing 6.1-8.25 (8.25) mm; areolet closed, quadrate, with uneven sides, 4-M very short, 2r-m about as long as 3r-m; 2m-cu with two bullae, bent outwards; pterostigma 2.4-2.6 × longer than wide; 1cu-a meeting M + Cu a little distad from 1-M; 3-Cu clearly longer than 2cu-a; radial cell 2.5-3.0 × longer than wide. Hind wing with 1-Rs about 1.9 × longer than 1rs-m, 1-Cu about 1.1 × as long as cu-a. Hind legs not well preserved, but rather stout, hind femur maybe 3.0-3.2 × longer than wide.
</paragraph>
<paragraph id="740CD978FFB5FFED8D14463BFB780A84" blockId="10.[811,1457,1873,2028]" pageId="10" pageNumber="653">
<emphasis id="46C7056AFFB5FFED8D14463BFC3D0A03" bold="true" box="[813,937,1874,1900]" pageId="10" pageNumber="653">Metasoma.</emphasis>
With T1 about 0.7-0.85 (0.85) × longer than wide, with strong median longitudinal carina converging on basal half, then parallel and diverging towards the posterior end; T2 0.4-0.45 (0.45) × as long as wide. Ovipositor sheaths 0.30 × as long as fore wing, rather robust.
</paragraph>
</subSubSection>
</treatment>
</document>