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<document id="8B36C95A76B0FE31A8ED792F389B4805" ID-CLB-Dataset="99512" ID-DOI="10.5252/geodiversitas2022v44a23" ID-GBIF-Dataset="1818f0c9-dbee-4fa4-b69a-ad6d3712913b" ID-ISSN="1638-9395" ID-Zenodo-Dep="6819412" ID-ZooBank="urn:lsid:zoobank.org:pub:999E7A6A-3781-4746-8519-62E1010D3880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657550474273" checkinUser="felipe" docAuthor="Klopfstein, Seraina" docDate="2022" docId="FC1A686EFFAEFFF58F2545DDFE890A84" docLanguage="en" docName="Geodiversitas.44.23.645-664.pdf.imf" docOrigin="Geodiversitas 44 (23)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Xanthopimpla crescendae Klopfstein 2022, n. sp." docType="treatment" docUuid="343B9824-38A4-45C6-8730-E0B51D695735" docUuidSource="ZooBank" docVersion="9" lastPageNumber="661" masterDocId="00231016FFBFFFE78E394169FF940D6F" masterDocTitle="High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)" masterLastPageNumber="664" masterPageNumber="645" pageNumber="660" updateTime="1699358479615" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="551BC6C7F73489D9A7F20E8B333DCCCC">High diversity of pimpline parasitoid wasps (Hymenoptera, Ichneumonidae, Pimplinae) from the lowermost Eocene Fur Formation (Denmark)</mods:title>
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<mods:namePart id="0E6776AB97499B098BE659A14225373E">Klopfstein, Seraina</mods:namePart>
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<mods:title id="8558EA967AFE8E7A0C2AB90700961BE6">Geodiversitas</mods:title>
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<mods:date id="B0C5FC2313A2D9C01B5DC8A04DD6F906">2022</mods:date>
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<mods:number id="A0116945AEC113999BF106902E64BF68">2022-07-07</mods:number>
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<mods:identifier id="4E51E0F9ECA46C568873D91BA8D612E9" type="DOI">10.5252/geodiversitas2022v44a23</mods:identifier>
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<treatment id="FC1A686EFFAEFFF58F2545DDFE890A84" ID-DOI="http://doi.org/10.5281/zenodo.6857746" ID-GBIF-Taxon="197821813" ID-Zenodo-Dep="6857746" LSID="urn:lsid:zoobank.org:act:343B9824-38A4-45C6-8730-E0B51D695735" httpUri="http://treatment.plazi.org/id/FC1A686EFFAEFFF58F2545DDFE890A84" lastPageId="18" lastPageNumber="661" pageId="17" pageNumber="660">
<subSubSection id="3CA98AF3FFAEFFF68F2545DDFDE409A0" box="[284,624,1204,1231]" pageId="17" pageNumber="660" type="nomenclature">
<paragraph id="740CD978FFAEFFF68F2545DDFDE409A0" blockId="17.[284,624,1204,1231]" box="[284,624,1204,1231]" pageId="17" pageNumber="660">
<heading id="2F446E14FFAEFFF68F2545DDFDE409A0" box="[284,624,1204,1231]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="660" reason="2">
<taxonomicName id="B3B3A2FBFFAEFFF68F2545DDFDBA09A0" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[284,558,1204,1231]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="660" phylum="Arthropoda" rank="species" species="crescendae" status="sp. nov.">
<emphasis id="46C7056AFFAEFFF68F2545DDFDBA09A0" bold="true" box="[284,558,1204,1231]" italics="true" pageId="17" pageNumber="660">Xanthopimpla crescendae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFAEFFF68C0C45DCFDE409A0" box="[565,624,1205,1231]" pageId="17" pageNumber="660" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAEFFF68FAE45BDFC920847" pageId="17" pageNumber="660" type="description">
<paragraph id="740CD978FFAEFFF68FAE45BDFE600980" blockId="17.[407,500,1236,1263]" box="[407,500,1236,1263]" pageId="17" pageNumber="660">
(
<figureCitation id="EC88C5FDFFAEFFF68F9945BDFE7F0981" box="[416,491,1236,1262]" captionStart="FIG" captionStartId="17.[132,143,823,840]" captionTargetBox="[132,1439,215,780]" captionTargetId="figure-631@17.[698,1267,213,781]" captionTargetPageId="17" captionText="FIG. 11. — Holotype of Xanthopimpla crescendae n. sp. (FUR-11214), photograph (A) with details and interpretative drawing (B), where dotted lines represent uncertain and/or interpolated interpretations. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6819436" httpUri="https://zenodo.org/record/6819436/files/figure.png" pageId="17" pageNumber="660">Fig. 11</figureCitation>
)
</paragraph>
<paragraph id="740CD978FFAEFFF68EBD447EFC920847" blockId="17.[132,774,1303,1320]" box="[132,774,1303,1320]" pageId="17" pageNumber="660">
<uri id="0022D57AFFAEFFF68EBD447EFC920847" box="[132,774,1303,1320]" pageId="17" pageNumber="660">
urn:lsid:zoobank.org:act:
<uuid id="0015E3ADFFAEFFF68F5B447EFC920847" box="[354,774,1303,1320]" pageId="17" pageNumber="660">343B9824-38A4-45C6-8730-E0B51D695735</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAEFFF68EBD4434FE8F08C6" pageId="17" pageNumber="660" type="materials_examined">
<paragraph id="740CD978FFAEFFF68EBD4434FE8F08C6" blockId="17.[131,775,1372,1449]" pageId="17" pageNumber="660">
TYPE MATERIAL. —
<materialsCitation id="C4DBD325FFAEFFF68F7E4435FECC08E0" ID-GBIF-Occurrence="3863921326" collectionCode="FUR" collectorName="E. Rettig" country="Denmark" location="MorsO Kommune" municipality="Jutland" pageId="17" pageNumber="660" specimenCode="FUR-11214" specimenCount="1" specimenCount-male="1" stateProvince="Limfjord" typeStatus="holotype">
<typeStatus id="AB0867DAFFAEFFF68F7E4435FE30081B" box="[327,420,1372,1396]" pageId="17" pageNumber="660" type="holotype">Holotype</typeStatus>
<specimenCode id="24157103FFAEFFF68F934435FDB6081B" box="[426,546,1372,1396]" pageId="17" pageNumber="660">FUR-11214</specimenCode>
(
<specimenCount id="62B512F1FFAEFFF68C094435FDF6081B" box="[560,610,1372,1396]" pageId="17" pageNumber="660" type="male">male</specimenCount>
?, part and counterpart; leg.
<collectorName id="D946BCAEFFAEFFF68ECF441EFED908E0" box="[246,333,1399,1423]" pageId="17" pageNumber="660">E. Rettig</collectorName>
),
</materialsCitation>
<materialsCitation id="C4DBD325FFAEFFF68F65441EFD1808E0" ID-GBIF-Occurrence="3863921304" box="[348,652,1399,1423]" collectionCode="FUR" country="Denmark" location="Svalklit" municipality="Jutland" pageId="17" pageNumber="660" specimenCode="FUR-13065" specimenCount="1" specimenCount-female="1" stateProvince="Limfjord" typeStatus="paratype">
<typeStatus id="AB0867DAFFAEFFF68F65441EFE2208E0" box="[348,438,1399,1423]" pageId="17" pageNumber="660" type="paratype">paratypes</typeStatus>
<specimenCode id="24157103FFAEFFF68F82441EFDA608E0" box="[443,562,1399,1423]" pageId="17" pageNumber="660">FUR-13065</specimenCode>
(
<specimenCount id="62B512F1FFAEFFF68C04441EFD1608E0" box="[573,642,1399,1423]" pageId="17" pageNumber="660" type="female">female</specimenCount>
),
</materialsCitation>
<materialsCitation id="C4DBD325FFAEFFF68CA9441EFE8F08C6" ID-GBIF-Occurrence="3863921316" collectionCode="FUR" country="Denmark" location="Skive, Ostklinten" municipality="Jutland" pageId="17" pageNumber="660" specimenCode="FUR-10918" specimenCount="1" stateProvince="Limfjord">
<specimenCode id="24157103FFAEFFF68CA9441EFC9308E0" box="[656,775,1399,1423]" pageId="17" pageNumber="660">FUR-10918</specimenCode>
(
<specimenCount id="62B512F1FFAEFFF68EB044F8FF3C08C6" box="[137,168,1425,1449]" pageId="17" pageNumber="660">sex</specimenCount>
unknown).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAEFFF68EBD44AFFC920897" pageId="17" pageNumber="660" type="etymology">
<paragraph id="740CD978FFAEFFF68EBD44AFFC920897" blockId="17.[130,776,1477,1528]" pageId="17" pageNumber="660">ETYMOLOGY. — This wasp is dedicated to my dear friend Tabia Stoffel, nickname “Crescendo”, for her unwavering support over the years.</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAEFFF68EBD477DFE730BC1" pageId="17" pageNumber="660" type="materials_examined">
<paragraph id="740CD978FFAEFFF68EBD477DFC920B0F" blockId="17.[130,776,1555,1632]" pageId="17" pageNumber="660">
TYPE HORIZON AND LOCALITY. —
<collectingCountry id="0CA499E8FFAEFFF68F84477AFD8D0B44" box="[445,537,1555,1579]" name="Denmark" pageId="17" pageNumber="660">Denmark</collectingCountry>
,
<collectingMunicipality id="94684302FFAEFFF68C18477AFDF30B44" box="[545,615,1555,1579]" pageId="17" pageNumber="660">Jutland</collectingMunicipality>
,
<collectingRegion id="B677179AFFAEFFF68C56477AFD560B44" box="[623,706,1555,1579]" pageId="17" pageNumber="660">Limfjord</collectingRegion>
region,
<location id="716C8FA3FFAEFFF68EBB4745FEA70B2A" LSID="urn:lsid:plazi:treatment:FC1A686EFFAEFFF58F2545DDFE890A84:716C8FA3FFAEFFF68EBB4745FEA70B2A" box="[130,307,1580,1605]" country="Denmark" municipality="Jutland" name="MorsO Kommune" pageId="17" pageNumber="660" stateProvince="Limfjord">MorsØ Kommune</location>
, Klinten ved Klitgård (holotype); Svalklit (FUR- 13065). Skive Kommune, Østklinten (FUR-10918).Cement stone.
</paragraph>
<paragraph id="740CD978FFAEFFF68EBD4714FE730BC1" blockId="17.[131,776,1660,1711]" pageId="17" pageNumber="660">
MATERIAL EXAMINED. — Tentatively identified as the same species:
<materialsCitation id="C4DBD325FFAEFFF68EBA47FFFE730BC1" ID-GBIF-Occurrence="3863921306" box="[131,487,1686,1711]" collectionCode="FUR" pageId="17" pageNumber="660" specimenCode="FUR-13068, 16642, 17232, 17235" specimenCount="1">
<specimenCode id="24157103FFAEFFF68EBA47FFFF6A0BC1" box="[131,254,1686,1710]" pageId="17" pageNumber="660">FUR-13068</specimenCode>
,
<specimenCode id="24157103FFAEFFF68F3147FFFEDE0BC1" box="[264,330,1686,1710]" pageId="17" pageNumber="660">16642</specimenCode>
,
<specimenCode id="24157103FFAEFFF68F6D47FEFE020BC0" box="[340,406,1687,1711]" pageId="17" pageNumber="660">17232</specimenCode>
,
<specimenCode id="24157103FFAEFFF68F9947FFFE760BC1" box="[416,482,1686,1710]" pageId="17" pageNumber="660">17235</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAEFFF68EBD47A2FAD709A0" pageId="17" pageNumber="660" type="diagnosis">
<paragraph id="740CD978FFAEFFF68EBD47A2FAD709A0" blockId="17.[131,777,1738,2028]" lastBlockId="17.[812,1457,916,1231]" pageId="17" pageNumber="660">
DIAGNOSIS. — Taxonomic placement: the areolet shape, outwardsbowed 2m-cu in the forewing, as well as the short and stout T1 and robust, protruding ovipositor clearly indicate
<taxonomicName id="B3B3A2FBFFAEFFF68C154796FD040A78" box="[556,656,1791,1815]" pageId="17" pageNumber="647" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
. But most of all, there is clear evidence for placing this species in
<taxonomicName id="B3B3A2FBFFAEFFF68C434673FC940A5D" authorityName="Saussure" authorityYear="1892" box="[634,768,1818,1842]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="660" phylum="Arthropoda" rank="genus">
<emphasis id="46C7056AFFAEFFF68C434673FC940A5D" box="[634,768,1818,1842]" italics="true" pageId="17" pageNumber="660">Xanthopimpla</emphasis>
</taxonomicName>
: narrow (twisted) mandibles (visible in
<typeStatus id="AB0867DAFFAEFFF68FC5465DFDC20A23" box="[508,598,1844,1868]" pageId="17" pageNumber="660" type="holotype">holotype</typeStatus>
), yellow or orange ground colouration with paired, dark markings on several tergites, stout legs, extensive carination of the propodeum, hind wing with 1-Cu shorter than cu-a (nervellus broken above middle), large claws. Species diagnosis: this species stands out within the genus by its almost or entirely dark mesoscutum and broad light mark at the base of the pterostigma, a combination not seen in any extant nor in the three known fossil species of the genus.
<taxonomicName id="B3B3A2FBFFAEFFF68AE042FDFC2B0EA8" authority="Khalaim, 2008" authorityName="Khalaim" authorityYear="2008" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="660" phylum="Arthropoda" rank="species" species="biamosa">
<emphasis id="46C7056AFFAEFFF68AE042FDFA3A0EC3" box="[1241,1454,916,940]" italics="true" pageId="17" pageNumber="660">Xanthopimpla biamosa</emphasis>
<bibRefCitation id="1022A489FFAEFFF68D1442C6FC2B0EA8" author="KHALAIM A. I." box="[813,959,943,967]" pageId="17" pageNumber="660" pagination="101 - 112" refId="ref16048" refString="KHALAIM A. I. 2008. - Fossil ichneumon wasps (Hymenoptera: Ichneumonidae) from Biamo (Russia), Oligocene. Alavesia 2: 101 - 112." type="journal article" year="2008">Khalaim, 2008</bibRefCitation>
</taxonomicName>
from the Oligocene of Biamo (
<collectingCountry id="0CA499E8FFAEFFF68AD242C6FAB80EA8" box="[1259,1324,943,967]" name="Russia" pageId="17" pageNumber="660">Russia</collectingCountry>
) furthermore has much shorter and stouter hind femora and two lateral dark marks on tergite 1, instead of its base being dark.
<taxonomicName id="B3B3A2FBFFAEFFF68B09428DFB900979" authority="Spasojevic et al., 2018" authorityName="Spasojevic" authorityYear="2018" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="660" phylum="Arthropoda" rank="species" species="messelensis">
<emphasis id="46C7056AFFAEFFF68B09428DFA250E93" box="[1328,1457,996,1020]" italics="true" pageId="17" pageNumber="660">X. messelensis</emphasis>
Spasojevic
<emphasis id="46C7056AFFAEFFF68DAF4296FC520979" box="[918,966,1022,1046]" italics="true" pageId="17" pageNumber="660">et al.</emphasis>
, 2018
</taxonomicName>
from the mid Eocene deposit in Messel has a very narrow, triangular areolet and dark basal bands instead of clearly disconnected spots on tergites 2-3. And
<taxonomicName id="B3B3A2FBFFAEFFF68B32455DFB3B0909" authority="Spasojevic, Wedmann &amp; Klopfstein, 2018" authorityName="Spasojevic, Wedmann &amp; Klopfstein" authorityYear="2018" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="660" phylum="Arthropoda" rank="species" species="praeclara">
<emphasis id="46C7056AFFAEFFF68B32455DFAE90924" box="[1291,1405,1075,1099]" italics="true" pageId="17" pageNumber="660">X. praeclara</emphasis>
<bibRefCitation id="1022A489FFAEFFF68BBC455AFB3B0909" author="SPASOJEVIC T. &amp; WEDMANN S. &amp; KLOPFSTEIN S." pageId="17" pageNumber="660" pagination="0197477" refId="ref17435" refString="SPASOJEVIC T., WEDMANN S. &amp; KLOPFSTEIN S. 2018 b. - Seven remarkable new fossil species of parasitoid wasps (Hymenoptera, Ichneumonidae) from the Eocene Messel Pit. PLoS One 13: e 0197477. https: // doi. org / 10.1371 / journal. pone. 0197477" type="journal article" year="2018">Spasojevic, Wedmann &amp; Klopfstein, 2018</bibRefCitation>
</taxonomicName>
, also from Messel, can be distinguished by the entirely yellow tergite 2. Among the here-in described Fur fossil species,
<taxonomicName id="B3B3A2FBFFAEFFF68A0445EAFB5209F5" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[1085,1222,1155,1178]" class="Insecta" family="Ichneumonidae" genus="Theronia" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="660" phylum="Arthropoda" rank="species" species="nigriscutum" status="sp. nov.">
<emphasis id="46C7056AFFAEFFF68A0445EAFB5209F5" box="[1085,1222,1155,1178]" italics="true" pageId="17" pageNumber="660">T. nigriscutum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFAEFFF68AF545EAFA9609F5" box="[1228,1282,1155,1178]" pageId="17" pageNumber="660" rank="species">n. sp.</taxonomicNameLabel>
is most similar to
<taxonomicName id="B3B3A2FBFFAEFFF68D1445F7FC0609DA" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[813,914,1181,1205]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="660" phylum="Arthropoda" rank="species" species="ciboisae" status="sp. nov.">
<emphasis id="46C7056AFFAEFFF68D1445F7FC0609DA" box="[813,914,1181,1205]" italics="true" pageId="17" pageNumber="660">X. ciboisae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFAEFFF68DA345F4FC4709DB" box="[922,979,1181,1204]" pageId="17" pageNumber="660" rank="species">n. sp.</taxonomicNameLabel>
, from which it can be distinguished by its dark mesoscutum, smaller size and more slender metasoma.
</paragraph>
</subSubSection>
<subSubSection id="3CA98AF3FFAEFFF58D14459DFE890A84" lastPageId="18" lastPageNumber="661" pageId="17" pageNumber="660" type="description">
<paragraph id="740CD978FFAEFFF68D14459DFC290861" blockId="17.[811,1457,1268,1613]" box="[813,957,1268,1294]" pageId="17" pageNumber="660">DESCRIPTION</paragraph>
<paragraph id="740CD978FFAEFFF68D14447DFC3F0842" blockId="17.[811,1457,1268,1613]" box="[813,939,1300,1325]" pageId="17" pageNumber="660">
<emphasis id="46C7056AFFAEFFF68D14447DFC3F0842" box="[813,939,1300,1325]" italics="true" pageId="17" pageNumber="660">Preservation</emphasis>
</paragraph>
<paragraph id="740CD978FFAEFFF68D14445AFB0F0B23" blockId="17.[811,1457,1268,1613]" pageId="17" pageNumber="660">
<typeStatus id="AB0867DAFFAEFFF68D14445AFC070822" box="[813,915,1331,1357]" pageId="17" pageNumber="660" type="holotype">Holotype</typeStatus>
preserved in dorsal view, with head including most of one and a partial second antenna, most of mesosoma, one nearly complete but somewhat distorted and one longitudinally folded forewing, two partial hind wings, partial hind legs, and metasoma except for its tip. Metasoma especially well-preserved, including traces of microsculpture on the tergites.
<typeStatus id="AB0867DAFFAEFFF68D55449AFC5C0B62" box="[876,968,1523,1549]" pageId="17" pageNumber="660" type="paratype">Paratype</typeStatus>
in dorsal view, head without antennae, most of mesosoma, partial fore wings, one nearly complete hind wing, metasoma except for its tip.
</paragraph>
<paragraph id="740CD978FFAEFFF68D14471BFB770A43" blockId="17.[811,1457,1650,1836]" pageId="17" pageNumber="660">
<emphasis id="46C7056AFFAEFFF68D14471BFBFF0BE3" bold="true" box="[813,1131,1650,1677]" pageId="17" pageNumber="660">
Body
<emphasis id="46C7056AFFAEFFF68D49471AFC160BE2" bold="true" box="[880,898,1651,1677]" italics="true" pageId="17" pageNumber="660">c.</emphasis>
7.8-9.3 (
<emphasis id="46C7056AFFAEFFF68DD7471AFB940BE2" bold="true" box="[1006,1024,1651,1677]" italics="true" pageId="17" pageNumber="660">c.</emphasis>
7.8) mm
</emphasis>
. Yellow to orange, with dark brown head, antennae uniformly orange. Dark on entire mesoscutum except maybe two lateral patches, basal part of propodeum, wing veins except basal half of pterostigma, basal two thirds of T1, paired spots on T2 to T4, on T5-6 these spots combining into a basal dark stripe.
</paragraph>
<paragraph id="740CD978FFAEFFF58D14463BFDF10901" blockId="17.[810,1455,1873,2028]" lastBlockId="18.[132,777,1012,1135]" lastPageId="18" lastPageNumber="661" pageId="17" pageNumber="660">
<emphasis id="46C7056AFFAEFFF68D14463BFCE40A03" bold="true" box="[813,880,1874,1900]" pageId="17" pageNumber="660">Head.</emphasis>
In
<typeStatus id="AB0867DAFFAEFFF68DAB4638FC650A04" box="[914,1009,1873,1899]" pageId="17" pageNumber="660" type="holotype">holotype</typeStatus>
turned upward to show from front, rather high with large compound eyes, these seemingly converging towards ventral (but this might be an artefact); with very narrow, darkened and bent mandibular tips than might indicate either a single tooth or strongly narrowed and twisted mandibles. Antennae incomplete, showing at least 23 clearly outlined flagellomeres (estimated:
<quantity id="B34B749DFFADFFF58FAF457DFE5B0941" box="[406,463,1044,1070]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="18" pageNumber="661" unit="in" value="26.0">26 in</quantity>
total), these in proximal half about twice as long as wide, after middle about 1.5 ×, and penultimate segments about as long as wide.
</paragraph>
<caption id="20CC89F0FFADFFF58EBD41B2FBBC0C31" ID-Table-UUID="20CC89F0FFADFFF58EBD41B2FBBC0C31" httpUri="http://table.plazi.org/id/20CC89F0FFADFFF58EBD41B2FBBC0C31" pageId="18" pageNumber="661" startId="18.[132,143,219,236]" targetBox="[134,1444,405,939]" targetIsTable="true" targetPageId="18">
<paragraph id="740CD978FFADFFF58EBD41B2FBBC0C31" blockId="18.[132,1457,219,350]" pageId="18" pageNumber="661">
TABLE 1. — Overview of characters supporting placement in
<taxonomicName id="B3B3A2FBFFADFFF58C4F41B2FD5F0D83" box="[630,715,219,236]" pageId="18" pageNumber="647" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
. Brackets indicate uncertain interpretation or visibility of only some aspect of a character.
<emphasis id="46C7056AFFADFFF58EBD419BFF1A0C6C" bold="true" box="[132,142,242,259]" pageId="18" pageNumber="661">1</emphasis>
, forewig with areolet triangular or oblique-quadrate;
<emphasis id="46C7056AFFADFFF58C73419BFDC00C6C" bold="true" box="[586,596,242,259]" pageId="18" pageNumber="661">2</emphasis>
, forewing vein 2m-cu bowed outwards, with two bullae (in brackets if bullae not visible);
<emphasis id="46C7056AFFADFFF58B79419BFADE0C6C" bold="true" box="[1344,1354,242,259]" pageId="18" pageNumber="661">3</emphasis>
, first tergite attached broadly to propodeum, parallel-sided or somewhat converging towards base; usually 1.0-2.0
<emphasis id="46C7056AFFADFFF58DDD4060FC7B0C76" bold="true" box="[996,1007,265,281]" pageId="18" pageNumber="661">×</emphasis>
longer than wide;
<emphasis id="46C7056AFFADFFF58AB54061FB020C76" bold="true" box="[1164,1174,264,281]" pageId="18" pageNumber="661">4</emphasis>
, spiracle of first tergite in front of middle; first sternite shorter than half the length of tergite;
<emphasis id="46C7056AFFADFFF58C5F4076FDE40C5F" bold="true" box="[614,624,287,304]" pageId="18" pageNumber="661">5</emphasis>
, ovipositor clearly protruding from metasomal apex, often long; if shorter than metasoma, then usually rather robust;
<emphasis id="46C7056AFFADFFF58F24405FFEB30C28" bold="true" box="[285,295,310,327]" pageId="18" pageNumber="661">6</emphasis>
, ventral valve of ovipositor with teeth, dorsal valve without a notch;
<emphasis id="46C7056AFFADFFF58D5C405FFCFB0C28" bold="true" box="[869,879,310,327]" pageId="18" pageNumber="661">7</emphasis>
, claws with basal lobe (present in many extant
<taxonomicName id="B3B3A2FBFFADFFF58B39405FFAC10C28" box="[1280,1365,310,327]" pageId="18" pageNumber="647" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
, especially
<taxonomicName id="B3B3A2FBFFADFFF58EBD4024FF470C31" box="[132,211,333,350]" pageId="18" pageNumber="662" rank="tribe" tribe="Ephialtini">Ephialtini</taxonomicName>
);
<emphasis id="46C7056AFFADFFF58EE64024FF7D0C31" bold="true" box="[223,233,333,350]" pageId="18" pageNumber="661">8</emphasis>
, unique character combination present that allows association with a particular genus in
<taxonomicName id="B3B3A2FBFFADFFF58DF64024FBB00C31" box="[975,1060,333,350]" pageId="18" pageNumber="647" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="740CD978FFADFFF58EBF4340FAA80EC4" pageId="18" pageNumber="661">
<table id="06B32BD8FFAD00188EBF40FCFA300EC4" box="[134,1444,405,939]" gridcols="10" gridrows="14" pageId="18" pageNumber="661">
<tr id="CA83DB3AFFAD00188EBF40FCFA300F52" box="[134,1444,405,573]" gridrow="0" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF40FCFE5A0F52" box="[134,462,405,573]" gridcol="0" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58EBF4340FF750F52" bold="true" box="[134,225,553,573]" pageId="18" pageNumber="661">Species</emphasis>
</th>
<th id="8952B246FFAD00188FE340FCFE630F52" box="[474,503,405,573]" gridcol="1" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58FE7408FFE600F56" bold="true" box="[478,500,486,569]" pageId="18" pageNumber="661">Areolet</emphasis>
<emphasis id="46C7056AFFADFFF58FE640B5FE7B0C8A" bold="true" box="[479,495,476,485]" pageId="18" pageNumber="661">1</emphasis>
</th>
<th id="8952B246FFAD00188C2940FCFDBE0F52" box="[528,554,405,573]" gridcol="2" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58C2A436EFDBD0C9A" bold="true" box="[531,553,440,568]" pageId="18" pageNumber="661">Vein 2 m-cu</emphasis>
<emphasis id="46C7056AFFADFFF58C2D40C7FDB00CD8" bold="true" box="[532,548,430,439]" pageId="18" pageNumber="661">2</emphasis>
</th>
<th id="8952B246FFAD00188C7140FCFDCA0F52" box="[584,606,405,573]" gridcol="3" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58C714081FDCA0C8E" bold="true" box="[584,606,468,569]" pageId="18" pageNumber="661">Tergite 1</emphasis>
<emphasis id="46C7056AFFADFFF58C7040A3FDCD0CBC" bold="true" box="[585,601,458,467]" pageId="18" pageNumber="661">3</emphasis>
</th>
<th id="8952B246FFAD00188C4440FCFD4E0F52" box="[637,730,405,573]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58C4440B2FD070F57" bold="true" box="[637,659,475,568]" pageId="18" pageNumber="661">Spiracle</emphasis>
<emphasis id="46C7056AFFADFFF58C4440BBFD190CB4" bold="true" box="[637,653,466,475]" pageId="18" pageNumber="661">4</emphasis>
<emphasis id="46C7056AFFADFFF58C9E40ABFD4E0F57" bold="true" box="[679,730,450,568]" pageId="18" pageNumber="661">Ovipositor shape</emphasis>
<emphasis id="46C7056AFFADFFF58CFC4083FD410C9C" bold="true" box="[709,725,490,499]" pageId="18" pageNumber="661">5</emphasis>
</th>
<th id="8952B246FFAD00188CCB40FCFC9C0F52" box="[754,776,405,573]" gridcol="6" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58CCB40ABFC9C0CD4" bold="true" box="[754,776,414,568]" pageId="18" pageNumber="661">Ovipositor tip</emphasis>
<emphasis id="46C7056AFFADFFF58CCA40FCFC970CF1" bold="true" box="[755,771,405,414]" pageId="18" pageNumber="661">6</emphasis>
</th>
<th id="8952B246FFAD00188D1240FCFCD20F52" box="[811,838,405,573]" gridcol="7" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58D164096FCD10F57" bold="true" box="[815,837,511,568]" pageId="18" pageNumber="661">Claw</emphasis>
<emphasis id="46C7056AFFADFFF58D16409FFCAB0C90" bold="true" box="[815,831,502,511]" pageId="18" pageNumber="661">7</emphasis>
</th>
<th id="8952B246FFAD00188D5A40FCFC020F52" box="[867,918,405,573]" gridcol="8" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58D5A409BFC020F56" bold="true" box="[867,918,443,569]" pageId="18" pageNumber="661">Genus diagnostics</emphasis>
<emphasis id="46C7056AFFADFFF58DB940D8FC040CD5" bold="true" box="[896,912,433,442]" pageId="18" pageNumber="661">8</emphasis>
</th>
<th id="8952B246FFAD00188D9940FCFA300F52" box="[928,1444,405,573]" gridcol="9" gridrow="0" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58D994343FC700F52" bold="true" box="[928,996,554,573]" pageId="18" pageNumber="661">Notes</emphasis>
</th>
</tr>
<tr id="CA83DB3AFFAD00188EBF4324FA300F0C" box="[134,1444,589,611]" gridrow="1" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF4324FE5A0F0C" box="[134,462,589,611]" gridcol="0" gridrow="1" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF4324FECC0F0C" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[134,344,589,611]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="collina" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58EBF4324FECC0F0C" box="[134,344,589,611]" italics="true" pageId="18" pageNumber="661">Crusopimpla collina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58F594324FE080F0C" box="[352,412,589,611]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE34324FE630F0C" box="[474,503,589,611]" gridcol="1" gridrow="1" pageId="18" pageNumber="661">++</td>
<td id="8952B246FFAD00188C294324FDBE0F0C" box="[528,554,589,611]" gridcol="2" gridrow="1" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C714324FDCA0F0C" box="[584,606,589,611]" gridcol="3" gridrow="1" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C444324FD1A0F0C" box="[637,654,589,611]" gridcol="4" gridrow="1" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188C8B4324FD4E0F0C" box="[690,730,589,611]" gridcol="5" gridrow="1" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188CCB4324FC9C0F0C" box="[754,776,589,611]" gridcol="6" gridrow="1" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188D124324FCD20F0C" box="[811,838,589,611]" gridcol="7" gridrow="1" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A4324FC020F0C" box="[867,918,589,611]" gridcol="8" gridrow="1" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D994324FA300F0C" box="[928,1444,589,611]" gridcol="9" gridrow="1" pageId="18" pageNumber="661">Paired bulges on T3-5 only rarely found outside</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF4303FA300FEF" box="[134,1444,618,640]" gridrow="2" pageId="18" pageNumber="661" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="8952B246FFAD00188D994303FA300FEF" box="[928,1444,618,640]" gridcol="9" gridrow="2" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58D8E4303FBB40FEF" box="[951,1056,618,640]" pageId="18" pageNumber="647" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF43ECFA300FF3" box="[134,1444,645,668]" gridrow="3" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF43ECFE5A0FF3" box="[134,462,645,668]" gridcol="0" gridrow="3" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF43EFFEE40FF3" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[134,368,646,668]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="elongata" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58EBF43EFFEE40FF3" box="[134,368,646,668]" italics="true" pageId="18" pageNumber="661">Crusopimpla elongata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58F4E43EFFE270FF3" box="[375,435,646,668]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE343ECFE630FF3" box="[474,503,645,668]" gridcol="1" gridrow="3" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C2943ECFDBE0FF3" box="[528,554,645,668]" gridcol="2" gridrow="3" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C7143ECFDCA0FF3" box="[584,606,645,668]" gridcol="3" gridrow="3" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C4443ECFD1A0FF3" box="[637,654,645,668]" gridcol="4" gridrow="3" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188C8B43ECFD4E0FF3" box="[690,730,645,668]" gridcol="5" gridrow="3" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188CCB43ECFC9C0FF3" box="[754,776,645,668]" gridcol="6" gridrow="3" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D1243ECFCD20FF3" box="[811,838,645,668]" gridcol="7" gridrow="3" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A43ECFC020FF3" box="[867,918,645,668]" gridcol="8" gridrow="3" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D9943ECFA300FF3" box="[928,1444,645,668]" gridcol="9" gridrow="3" pageId="18" pageNumber="661">Male only. Impressions on tergites not unique to</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF43CBFA300FD7" box="[134,1444,674,696]" gridrow="4" pageId="18" pageNumber="661" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="8952B246FFAD00188D9943CBFA300FD7" box="[928,1444,674,696]" gridcol="9" gridrow="4" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58D8E43CBFBB00FD7" box="[951,1060,674,696]" pageId="18" pageNumber="647" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
, but rare outside
</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF43D5FA300FBC" box="[134,1444,700,723]" gridrow="5" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF43D5FE5A0FBC" box="[134,462,700,723]" gridcol="0" gridrow="5" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF43D4FEC80FBC" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[134,348,701,723]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="minuta" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58EBF43D4FEC80FBC" box="[134,348,701,723]" italics="true" pageId="18" pageNumber="661">Crusopimpla minuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58F5D43D4FE340FBC" box="[356,416,701,723]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE343D5FE630FBC" box="[474,503,700,723]" gridcol="1" gridrow="5" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C2943D5FDBE0FBC" box="[528,554,700,723]" gridcol="2" gridrow="5" pageId="18" pageNumber="661">(+)</td>
<td id="8952B246FFAD00188C7143D5FDCA0FBC" box="[584,606,700,723]" gridcol="3" gridrow="5" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C4443D5FD1A0FBC" box="[637,654,700,723]" gridcol="4" gridrow="5" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C8B43D5FD4E0FBC" box="[690,730,700,723]" gridcol="5" gridrow="5" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188CCB43D5FC9C0FBC" box="[754,776,700,723]" gridcol="6" gridrow="5" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D1243D5FCD20FBC" box="[811,838,700,723]" gridcol="7" gridrow="5" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A43D5FC020FBC" box="[867,918,700,723]" gridcol="8" gridrow="5" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D9943D5FA300FBC" box="[928,1444,700,723]" gridcol="9" gridrow="5" pageId="18" pageNumber="661">2m-cu only slightly bowed outwards</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF43BEFA300F81" box="[134,1444,727,750]" gridrow="6" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF43BEFE5A0F81" box="[134,462,727,750]" gridcol="0" gridrow="6" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF43B1FEC40F81" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[134,336,728,750]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="rettigi" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58EBF43B1FEC40F81" box="[134,336,728,750]" italics="true" pageId="18" pageNumber="661">Crusopimpla rettigi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58F6143B1FE000F81" box="[344,404,728,750]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE343BEFE630F81" box="[474,503,727,750]" gridcol="1" gridrow="6" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C2943BEFDBE0F81" box="[528,554,727,750]" gridcol="2" gridrow="6" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C7143BEFDCA0F81" box="[584,606,727,750]" gridcol="3" gridrow="6" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C4443BEFD1A0F81" box="[637,654,727,750]" gridcol="4" gridrow="6" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188C8B43BEFD4E0F81" box="[690,730,727,750]" gridcol="5" gridrow="6" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188CCB43BEFC9C0F81" box="[754,776,727,750]" gridcol="6" gridrow="6" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D1243BEFCD20F81" box="[811,838,727,750]" gridcol="7" gridrow="6" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A43BEFC020F81" box="[867,918,727,750]" gridcol="8" gridrow="6" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D9943BEFA300F81" box="[928,1444,727,750]" gridcol="9" gridrow="6" pageId="18" pageNumber="661">–</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF439BFA300E66" box="[134,1444,754,777]" gridrow="7" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF439BFE5A0E66" box="[134,462,754,777]" gridcol="0" gridrow="7" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF439AFEC30E66" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[134,343,755,777]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="violina" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58EBF439AFEC30E66" box="[134,343,755,777]" italics="true" pageId="18" pageNumber="661">Crusopimpla violina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58F67439AFE0F0E66" box="[350,411,755,777]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE3439BFE630E66" box="[474,503,754,777]" gridcol="1" gridrow="7" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C29439BFDBE0E66" box="[528,554,754,777]" gridcol="2" gridrow="7" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C71439BFDCA0E66" box="[584,606,754,777]" gridcol="3" gridrow="7" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C44439BFD1A0E66" box="[637,654,754,777]" gridcol="4" gridrow="7" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188C8B439BFD4E0E66" box="[690,730,754,777]" gridcol="5" gridrow="7" pageId="18" pageNumber="661">(+)</td>
<td id="8952B246FFAD00188CCB439BFC9C0E66" box="[754,776,754,777]" gridcol="6" gridrow="7" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D12439BFCD20E66" box="[811,838,754,777]" gridcol="7" gridrow="7" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A439BFC020E66" box="[867,918,754,777]" gridcol="8" gridrow="7" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D99439BFA300E66" box="[928,1444,754,777]" gridcol="9" gridrow="7" pageId="18" pageNumber="661">Claws might show indication of basal lobe</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF4264FA300E4C" box="[134,1444,781,803]" gridrow="8" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF4264FE5A0E4C" box="[134,462,781,803]" gridcol="0" gridrow="8" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF4264FEE60E4C" authorityName="Klopfstein" authorityYear="2022" baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[134,370,781,803]" class="Insecta" family="Ichneumonidae" genus="Epitheronia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="stigmatica">
<emphasis id="46C7056AFFADFFF58EBF4264FEE60E4C" box="[134,370,781,803]" italics="true" pageId="18" pageNumber="661">Epitheronia stigmatica</emphasis>
</taxonomicName>
</th>
<td id="8952B246FFAD00188FE34264FE630E4C" box="[474,503,781,803]" gridcol="1" gridrow="8" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C294264FDBE0E4C" box="[528,554,781,803]" gridcol="2" gridrow="8" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C714264FDCA0E4C" box="[584,606,781,803]" gridcol="3" gridrow="8" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C444264FD1A0E4C" box="[637,654,781,803]" gridcol="4" gridrow="8" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C8B4264FD4E0E4C" box="[690,730,781,803]" gridcol="5" gridrow="8" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188CCB4264FC9C0E4C" box="[754,776,781,803]" gridcol="6" gridrow="8" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D124264FCD20E4C" box="[811,838,781,803]" gridcol="7" gridrow="8" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A4264FC020E4C" box="[867,918,781,803]" gridcol="8" gridrow="8" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D994264FA300E4C" box="[928,1444,781,803]" gridcol="9" gridrow="8" pageId="18" pageNumber="661">–</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF4243FA300E2F" box="[134,1444,810,832]" gridrow="9" pageId="18" pageNumber="661" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="8952B246FFAD00188EBF4243FE5A0E2F" box="[134,462,810,832]" gridcol="0" gridrow="9" pageId="18" pageNumber="661">
(
<bibRefCitation id="1022A489FFADFFF58E9B4243FEC10E2F" author="HENRIKSEN K. L." box="[162,341,810,832]" pageId="18" pageNumber="661" pagination="1 - 36" refId="ref15889" refString="HENRIKSEN K. L. 1922. - Eocene insects from Denmark. Danmarks geologiske Undersogelse 37: 1 - 36. https: // doi. org / 10.34194 / raekke 2. v 37.6823" type="journal article" year="1922">Henriksen, 1922</bibRefCitation>
), n. comb.
</th>
</tr>
<tr id="CA83DB3AFFAD00188EBF422DFA300E34" box="[134,1444,836,859]" gridrow="10" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF422DFE5A0E34" box="[134,462,836,859]" gridcol="0" gridrow="10" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF422CFF770E34" authorityName="Holmgren" authorityYear="1859" box="[134,227,837,859]" class="Insecta" family="Ichneumonidae" genus="Theronia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="genus">
<emphasis id="46C7056AFFADFFF58EBF422CFF770E34" box="[134,227,837,859]" italics="true" pageId="18" pageNumber="661">Theronia</emphasis>
</taxonomicName>
?
<taxonomicName id="B3B3A2FBFFADFFF58ECE422CFEDE0E34" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[247,330,837,859]" class="Insecta" family="Ichneumonidae" genus="Theronia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="furensis" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58ECE422CFEDE0E34" box="[247,330,837,859]" italics="true" pageId="18" pageNumber="661">furensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58F69422CFE190E34" box="[336,397,837,859]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE3422DFE630E34" box="[474,503,836,859]" gridcol="1" gridrow="10" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C29422DFDBE0E34" box="[528,554,836,859]" gridcol="2" gridrow="10" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C71422DFDCA0E34" box="[584,606,836,859]" gridcol="3" gridrow="10" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C44422DFD1A0E34" box="[637,654,836,859]" gridcol="4" gridrow="10" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188C8B422DFD4E0E34" box="[690,730,836,859]" gridcol="5" gridrow="10" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188CCB422DFC9C0E34" box="[754,776,836,859]" gridcol="6" gridrow="10" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D12422DFCD20E34" box="[811,838,836,859]" gridcol="7" gridrow="10" pageId="18" pageNumber="661">(+)</td>
<td id="8952B246FFAD00188D5A422DFC020E34" box="[867,918,836,859]" gridcol="8" gridrow="10" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D99422DFA300E34" box="[928,1444,836,859]" gridcol="9" gridrow="10" pageId="18" pageNumber="661">Claws enlarged</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF4236FA300E19" box="[134,1444,863,886]" gridrow="11" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF4236FE5A0E19" box="[134,462,863,886]" gridcol="0" gridrow="11" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF4209FF770E19" authorityName="Holmgren" authorityYear="1859" box="[134,227,864,886]" class="Insecta" family="Ichneumonidae" genus="Theronia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="genus">
<emphasis id="46C7056AFFADFFF58EBF4209FF770E19" box="[134,227,864,886]" italics="true" pageId="18" pageNumber="661">Theronia</emphasis>
</taxonomicName>
?
<taxonomicName id="B3B3A2FBFFADFFF58ECE4209FEE00E19" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[247,372,864,886]" class="Insecta" family="Ichneumonidae" genus="Theronia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="nigriscutum" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58ECE4209FEE00E19" box="[247,372,864,886]" italics="true" pageId="18" pageNumber="661">nigriscutum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58F434209FE230E19" box="[378,439,864,886]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE34236FE630E19" box="[474,503,863,886]" gridcol="1" gridrow="11" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C294236FDBE0E19" box="[528,554,863,886]" gridcol="2" gridrow="11" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C714236FDCA0E19" box="[584,606,863,886]" gridcol="3" gridrow="11" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C444236FD1A0E19" box="[637,654,863,886]" gridcol="4" gridrow="11" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188C8B4236FD4E0E19" box="[690,730,863,886]" gridcol="5" gridrow="11" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188CCB4236FC9C0E19" box="[754,776,863,886]" gridcol="6" gridrow="11" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D124236FCD20E19" box="[811,838,863,886]" gridcol="7" gridrow="11" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A4236FC020E19" box="[867,918,863,886]" gridcol="8" gridrow="11" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D994236FA300E19" box="[928,1444,863,886]" gridcol="9" gridrow="11" pageId="18" pageNumber="661">–</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF4213FA300EFF" box="[134,1444,890,912]" gridrow="12" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF4213FE5A0EFF" box="[134,462,890,912]" gridcol="0" gridrow="12" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF4213FEEC0EFF" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[134,376,890,912]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="ciboisae" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58EBF4213FEEC0EFF" box="[134,376,890,912]" italics="true" pageId="18" pageNumber="661">Xanthopimpla ciboisae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58F464213FE2F0EFF" box="[383,443,890,912]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE34213FE630EFF" box="[474,503,890,912]" gridcol="1" gridrow="12" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C294213FDBE0EFF" box="[528,554,890,912]" gridcol="2" gridrow="12" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C714213FDCA0EFF" box="[584,606,890,912]" gridcol="3" gridrow="12" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C444213FD1A0EFF" box="[637,654,890,912]" gridcol="4" gridrow="12" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188C8B4213FD4E0EFF" box="[690,730,890,912]" gridcol="5" gridrow="12" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188CCB4213FC9C0EFF" box="[754,776,890,912]" gridcol="6" gridrow="12" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D124213FCD20EFF" box="[811,838,890,912]" gridcol="7" gridrow="12" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A4213FC020EFF" box="[867,918,890,912]" gridcol="8" gridrow="12" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188D994213FA300EFF" box="[928,1444,890,912]" gridcol="9" gridrow="12" pageId="18" pageNumber="661">2m-cu with characteristic zig-zag for genus</td>
</tr>
<tr id="CA83DB3AFFAD00188EBF42FDFA300EC4" box="[134,1444,916,939]" gridrow="13" pageId="18" pageNumber="661">
<th id="8952B246FFAD00188EBF42FDFE5A0EC4" box="[134,462,916,939]" gridcol="0" gridrow="13" pageId="18" pageNumber="661">
<taxonomicName id="B3B3A2FBFFADFFF58EBF42FCFE1B0EC4" authority="Klopfstein, 2022" authorityName="Klopfstein" authorityYear="2022" box="[134,399,917,939]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="661" phylum="Arthropoda" rank="species" species="crescendae" status="sp. nov.">
<emphasis id="46C7056AFFADFFF58EBF42FCFE1B0EC4" box="[134,399,917,939]" italics="true" pageId="18" pageNumber="661">Xanthopimpla crescendae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5DF4B811FFADFFF58FAD42FCFE5A0EC4" box="[404,462,917,939]" pageId="18" pageNumber="661" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="8952B246FFAD00188FE342FDFE630EC4" box="[474,503,916,939]" gridcol="1" gridrow="13" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C2942FDFDBE0EC4" box="[528,554,916,939]" gridcol="2" gridrow="13" pageId="18" pageNumber="661">(+)</td>
<td id="8952B246FFAD00188C7142FDFDCA0EC4" box="[584,606,916,939]" gridcol="3" gridrow="13" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188C4442FDFD1A0EC4" box="[637,654,916,939]" gridcol="4" gridrow="13" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188C8B42FDFD4E0EC4" box="[690,730,916,939]" gridcol="5" gridrow="13" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188CCB42FDFC9C0EC4" box="[754,776,916,939]" gridcol="6" gridrow="13" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D1242FDFCD20EC4" box="[811,838,916,939]" gridcol="7" gridrow="13" pageId="18" pageNumber="661">–</td>
<td id="8952B246FFAD00188D5A42FDFC020EC4" box="[867,918,916,939]" gridcol="8" gridrow="13" pageId="18" pageNumber="661">+</td>
<td id="8952B246FFAD00188D9942FDFA300EC4" box="[928,1444,916,939]" gridcol="9" gridrow="13" pageId="18" pageNumber="661">2m-cu partially visible only in paratype</td>
</tr>
</table>
</paragraph>
<paragraph id="740CD978FFADFFF58EBD45FDFE920BE3" blockId="18.[130,777,1172,1676]" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58EBD45FDFE9709C1" bold="true" box="[132,259,1172,1198]" pageId="18" pageNumber="661">Mesosoma.</emphasis>
Rather slender, in
<typeStatus id="AB0867DAFFADFFF58FE045FDFDAD09C1" box="[473,569,1172,1198]" pageId="18" pageNumber="661" type="holotype">holotype</typeStatus>
distorted between mesoscutum and propodeum. Mesoscutum about as long as wide, with strongly converging but not meeting notauli. Propodeum with carination partially preserved (
<typeStatus id="AB0867DAFFADFFF58CAE459AFD680861" box="[663,764,1267,1294]" pageId="18" pageNumber="661" type="holotype">holotype</typeStatus>
), showing at least basal transverse, and lateral parts of apical transverse, and part of the median longitudinal carinae.Fore wing 5.5-6.3 (5.5) mm; areolet closed, quadrate, with uneven sides, 4-M very short, 2r-m a bit shorter than 3r-m; 2m-cu not well preserved in
<typeStatus id="AB0867DAFFADFFF58F5744FAFE5908C2" box="[366,461,1427,1453]" pageId="18" pageNumber="661" type="holotype">holotype</typeStatus>
but clearly bowing outwards in
<typeStatus id="AB0867DAFFADFFF58E9D44DAFE9008A2" box="[164,260,1459,1485]" pageId="18" pageNumber="661" type="paratype">paratype</typeStatus>
; pterostigma 3.7-3.9 (3.7) × longer than wide; 1cu-a meeting M + Cu opposite 1-M; 3-Cu a bit longer than 2cu-a; radial cell 3.2-3.3 × (3.3) longer than wide.Hind wing with 1-Rs about 1.8 × (
<typeStatus id="AB0867DAFFADFFF58FB0477AFE7D0B42" box="[393,489,1555,1581]" pageId="18" pageNumber="661" type="paratype">paratype</typeStatus>
) longer than 1rs-m, 1-Cu 0.5-0.7 × (0.5) as long as cu-a.Hind leg very short and stout, femur about 2.5 × longer than wide. Tarsal claw discernible, rather large.
</paragraph>
<paragraph id="740CD978FFADFFF58EBD47DBFE890A84" blockId="18.[130,777,1714,2028]" pageId="18" pageNumber="661">
<emphasis id="46C7056AFFADFFF58EBD47DBFF6F0BA3" bold="true" box="[132,251,1714,1740]" pageId="18" pageNumber="661">Metasoma.</emphasis>
With T1 about 1.1 × longer than wide, with strong median longitudinal carina converging on basal half, parallel posteriorly; T2 0.6 × as long as wide, basally with oblique grooves cutting off anterolateral corners, behind these with slowly converging grooves that cut off a roundish-trapezoid raised part medially bearing two confluent dark patches, this part with strong punctures visible. T3 with paired dark patches on basal half and pre-apical transverse groove indicated. T4-T7 with paired dark marks weakly indicated. Ovipositor not preserved.
</paragraph>
</subSubSection>
</treatment>
</document>
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