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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.993.53805" ID-GBIF-Dataset="153d057d-c190-4db9-aa8b-4da3cb04ae64" ID-PMC="PMC7683496" ID-Pensoft-Pub="1313-2970-993-47" ID-Pensoft-UUID="2D7B2EB8CB285CBCA90A27064C6B1F17" ID-PubMed="33262675" ID-ZooBank="C40989DB80634C9FA481E7AA82CA924B" ModsDocID="1313-2970-993-47" checkinTime="1605571755153" checkinUser="pensoft" docAuthor="Ruan, Yongying, Douglas, Hume B., Qiu, Lu, Chen, Xiaoqin &amp; Jiang, Shihong" docDate="2020" docId="F997E33FA32F5490997E2AD787B38A54" docLanguage="en" docName="ZooKeys 993: 47-120" docOrigin="ZooKeys 993" docSource="http://dx.doi.org/10.3897/zookeys.993.53805" docTitle="Phorocardius yunnanensis Ruan &amp; Douglas 2020, sp. nov." docType="treatment" docUuid="66DD489B-D095-4CCE-86A0-40676B3F1DA3" docUuidSource="ZooBank" docVersion="4" id="2D7B2EB8CB285CBCA90A27064C6B1F17" lastPageNumber="47" masterDocId="2D7B2EB8CB285CBCA90A27064C6B1F17" masterDocTitle="Revision of Chinese Phorocardius species (Coleoptera, Elateridae, Cardiophorinae)" masterLastPageNumber="120" masterPageNumber="47" pageNumber="47" updateTime="1668169908327" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Chinese Phorocardius species (Coleoptera, Elateridae, Cardiophorinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ruan, Yongying</mods:namePart>
<mods:affiliation>School of Applied Chemistry and Biological Technology, Shenzhen Polytechnic, Shenzhen, Guangdong 518055, China</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5025-5592</mods:nameIdentifier>
<mods:nameIdentifier type="email">yongyingruan@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Douglas, Hume B.</mods:namePart>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Ave., Ottawa, Ontario, K 1 A 0 C 6, Canada</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-1722-7554</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Qiu, Lu</mods:namePart>
<mods:affiliation>Institute of Entomology, College of Plant Protection, Southwest University, Beibei, Chongqing 400716, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Xiaoqin</mods:namePart>
<mods:affiliation>School of Applied Chemistry and Biological Technology, Shenzhen Polytechnic, Shenzhen, Guangdong 518055, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Shihong</mods:namePart>
<mods:affiliation>School of Applied Chemistry and Biological Technology, Shenzhen Polytechnic, Shenzhen, Guangdong 518055, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>993</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>47</mods:start>
<mods:end>120</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.993.53805</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.993.53805</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-993-47</mods:identifier>
<mods:identifier type="ZooBank">C40989DB80634C9FA481E7AA82CA924B</mods:identifier>
<mods:identifier type="Pensoft-UUID">2D7B2EB8CB285CBCA90A27064C6B1F17</mods:identifier>
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<treatment ID-GBIF-Taxon="169091756" LSID="urn:lsid:zoobank.org:act:66DD489B-D095-4CCE-86A0-40676B3F1DA3" httpUri="http://treatment.plazi.org/id/F997E33FA32F5490997E2AD787B38A54" lastPageNumber="47" pageId="0" pageNumber="47">
<subSubSection pageId="0" pageNumber="47" type="nomenclature">
<paragraph pageId="0" pageNumber="47">
10.
<taxonomicName LSID="F997E33F-A32F-5490-997E-2AD787B38A54" authority="Ruan &amp; Douglas" authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis" status="sp. nov.">Phorocardius yunnanensis Ruan &amp; Douglas</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="47">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A holotype, male, dorsal view B paratype, female, dorsal view C paratype, female (Yunnan, Xi-shuan-ban-na, Meng-zhe), ventral view D aedeagus of holotype, dorsal view E aedeagus of holotype, ventral view, arrow indicating apex of paramere F aedeagus of holotype, lateral view G female abdominal sternite VIII, dorsal view (paratype) H ovipositor of paratype, dorsal view I distal (upper side) and proximal sclerites of bursa copulatrix (paratype)." figureDoi="10.3897/zookeys.993.53805.figure19" httpUri="https://binary.pensoft.net/fig/475452" pageId="0" pageNumber="47">Figs 19</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A pronotum and scutellum of holotype, dorsal view B posterior edge of pronotum, left side, dorsal view, arrows showing sublateral incisions C head of holotype, frontal view D apex of last abdominal ventrite (ventrite V) of male and female, ventral view (paratypes), arrow showing apical lateral incisions E paratype, ventral view of pro- and mesothorax; indicating shapes of prosternal process, mesepisternum (upper, green arrow on antero-mesal angle) and projections on posterior edge of mesosternum (lower, red arrow)." figureDoi="10.3897/zookeys.993.53805.figure20" httpUri="https://binary.pensoft.net/fig/475453" pageId="0" pageNumber="47">, 20</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Shape of supraantennal carina of Chinese Phorocardius species. Arrows indicating supraantennal carina forked near eye. A P. alterlineatus Ruan &amp; Douglas, sp. nov. B P. flavistriolatus Ruan &amp; Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan &amp; Douglas, sp. nov. G P. rufiposterus Ruan &amp; Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan &amp; Douglas, sp. nov. J P. zhiweii Ruan, Douglas &amp; Qiu, sp. nov. K head, frontal-lateral view, arrows indicating forked supraantennal carina." figureDoi="10.3897/zookeys.993.53805.figure23" httpUri="https://binary.pensoft.net/fig/475457" pageId="0" pageNumber="47">, 23I</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Lateral view of prosternal process and mesosternum of Chinese Phorocardius species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i. e., on anteroventral angle of mesosternal fossa according to Douglas 2003). A P. alterlineatus Ruan &amp; Douglas, sp. nov. B P. flavistriolatus Ruan &amp; Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan &amp; Douglas, sp. nov. G P. rufiposterus Ruan &amp; Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan &amp; Douglas, sp. nov. J P. zhiweii Ruan, Douglas &amp; Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47">, 24I</figureCitation>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Female abdominal ventrite V in Chinese Phorocardius species (ventral view). A P. alterlineatus Ruan &amp; Douglas, sp. nov. B P. flavistriolatus Ruan &amp; Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus F P. rufiposterus Ruan &amp; Douglas, sp. nov. G P. unguicularis H P. yunnanensis Ruan &amp; Douglas, sp. nov, arrows showing incisions near apex." figureDoi="10.3897/zookeys.993.53805.figure26" httpUri="https://binary.pensoft.net/fig/475460" pageId="0" pageNumber="47">, 26H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="type locality">
<paragraph pageId="0" pageNumber="47">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="47">Yunnan Prov., Xi-shuang-ban-na, Meng-a (alt. 1050-1080 m).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="etymology">
<paragraph pageId="0" pageNumber="47">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="47">The name of this species refers to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="distribution">
<paragraph pageId="0" pageNumber="47">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="47">China (currently endemic to Yunnan).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="differential diagnosis">
<paragraph pageId="0" pageNumber="47">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="47">Body length greater than 7.0 mm; integument black and shiny (non-metallic), elytron without yellow stripes, appendages yellow-brown. Prothorax: procoxal cavities narrowly open; prosternal process not strongly narrowed posterad to ventral apex in ventral view, with apex truncate to slightly rounded. Pterothorax: scutellar shield with posterior apex pointed. Tarsal claw with ventral apex not smaller than dorsal apex. Male genitalia: paramere acute in ventral view with small, acute preapical lateral expansion, without apical mesal callus. Female: apex of last abdominal ventrite (ventrite V) truncate to slightly convex, bent dorsad, each side with an incision.</paragraph>
<paragraph pageId="0" pageNumber="47">This species is distinctive for having the dorsum entirely black and shiny and legs entirely yellow-brown (except for brown-black coxae).</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
Ruan &amp; Douglas, sp. nov. is close to
<taxonomicName lsidName="P. vicinus" pageId="0" pageNumber="47" rank="species" species="vicinus">
<emphasis italics="true" pageId="0" pageNumber="47">P. vicinus</emphasis>
</taxonomicName>
in brown-black body color, but distinguishable by the following. In
<taxonomicName lsidName="P. yunnanensis" pageId="0" pageNumber="47" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">P. yunnanensis</emphasis>
</taxonomicName>
Ruan &amp; Douglas, sp. nov., pronotum longer with length of pronotum to elytra ratio 0.37-0.40 (excluding posterior angle) or ca. 0.43 (including posterior angle); antennae, palpi of mouthparts and legs yellow-brown to brown; and proximal sclerites of bursa copulatrix with basal edge concave. According to Kollar (1848) and type material (NHMW),
<taxonomicName lsidName="P. vicinus" pageId="0" pageNumber="47" rank="species" species="vicinus">
<emphasis italics="true" pageId="0" pageNumber="47">P. vicinus</emphasis>
</taxonomicName>
has length of pronotum to elytra ratio only 0.33; appendages brown-black to black; and proximal sclerites of bursa copulatrix not concave at basal edge.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="description">
<paragraph pageId="0" pageNumber="47">Description.</paragraph>
<paragraph pageId="0" pageNumber="47">(based on all type specimens (25♂, 24♀)) Dorsum black and shiny, venter brown-black with last 2-3 ventrites yellow-brown. Antennae brown to yellow-brown, with first two antennomeres slightly lighter in color. Legs entirely yellow-brown (except coxae brown-black). Surface of body with yellow pubescence.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="description">
<paragraph pageId="0" pageNumber="47">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="47">(based on all type specimens) Male body length 7.3-8.6 mm, width 2.6-3.3 mm. Female body length 7.3-10.4 mm, width 2.8-3.8 mm. Body length to width ratio 2.6-2.9. Pronotal width to length ratio 1.1-1.2. Pronotal width to body width ratio 0.84-0.91. Elytral length to pronotal length ratio 2.5-2.7; elytron length to width ratio 3.6-3.9.</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">
<emphasis italics="true" pageId="0" pageNumber="47">Head</emphasis>
.
</emphasis>
Frons and vertex with interspaces between punctures 1-2
<normalizedToken originalValue="×">x</normalizedToken>
average puncture diameter; punctures slightly sparser at centre of vertex. Frontal carina in frontal view straight at middle, curved dorsally at sides. Distance between eyes to width of eye ratio 3.4-4.1. Antenna with apex reaching to or slightly reaching beyond posterior angle of pronotum in male, not reaching to posterior angle in female. Antenna length to body length ratio, in male 0.34-0.39; in female 0.32-0.36. Proportions of antennomere lengths (male): 100 (scape); 57-65; 73-79; 78-80; 77-84; 78-89; 74-89; 77-89; 75-79; 73-83; 105-116.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">
<emphasis italics="true" pageId="0" pageNumber="47">Prothorax</emphasis>
.
</emphasis>
Pronotum in dorsal view: sides evenly convex from anterior edge to constriction near posterior fifth, widest near mid-length; posterior angles with lateral sides almost straight, not bulged; surface with small punctures, interspaces between punctures 1.5-3
<normalizedToken originalValue="×">x</normalizedToken>
average puncture diameter. In ventral view, ventral surface of prosternal process with sides carinate and gradually narrow from anterior to posterior end, with apex rounded. In lateral view, prosternal process with ventral surface curved slightly dorsad, posterior end concave (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Lateral view of prosternal process and mesosternum of Chinese Phorocardius species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i. e., on anteroventral angle of mesosternal fossa according to Douglas 2003). A P. alterlineatus Ruan &amp; Douglas, sp. nov. B P. flavistriolatus Ruan &amp; Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan &amp; Douglas, sp. nov. G P. rufiposterus Ruan &amp; Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan &amp; Douglas, sp. nov. J P. zhiweii Ruan, Douglas &amp; Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47">24I</figureCitation>
, upper arrow). Procoxal cavities narrowly open.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Pterothorax</emphasis>
(Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A pronotum and scutellum of holotype, dorsal view B posterior edge of pronotum, left side, dorsal view, arrows showing sublateral incisions C head of holotype, frontal view D apex of last abdominal ventrite (ventrite V) of male and female, ventral view (paratypes), arrow showing apical lateral incisions E paratype, ventral view of pro- and mesothorax; indicating shapes of prosternal process, mesepisternum (upper, green arrow on antero-mesal angle) and projections on posterior edge of mesosternum (lower, red arrow)." figureDoi="10.3897/zookeys.993.53805.figure20" httpUri="https://binary.pensoft.net/fig/475453" pageId="0" pageNumber="47">20E</figureCitation>
,
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Lateral view of prosternal process and mesosternum of Chinese Phorocardius species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i. e., on anteroventral angle of mesosternal fossa according to Douglas 2003). A P. alterlineatus Ruan &amp; Douglas, sp. nov. B P. flavistriolatus Ruan &amp; Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan &amp; Douglas, sp. nov. G P. rufiposterus Ruan &amp; Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan &amp; Douglas, sp. nov. J P. zhiweii Ruan, Douglas &amp; Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47">24I</figureCitation>
). Mesepisternum in ventral view with antero-mesal angle broadly rounded mesad of a notch, facing antero-mesally (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A pronotum and scutellum of holotype, dorsal view B posterior edge of pronotum, left side, dorsal view, arrows showing sublateral incisions C head of holotype, frontal view D apex of last abdominal ventrite (ventrite V) of male and female, ventral view (paratypes), arrow showing apical lateral incisions E paratype, ventral view of pro- and mesothorax; indicating shapes of prosternal process, mesepisternum (upper, green arrow on antero-mesal angle) and projections on posterior edge of mesosternum (lower, red arrow)." figureDoi="10.3897/zookeys.993.53805.figure20" httpUri="https://binary.pensoft.net/fig/475453" pageId="0" pageNumber="47">20E</figureCitation>
, upper (green) arrow); Projections on posterior edge of mesosternum: in ventral view weak (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A pronotum and scutellum of holotype, dorsal view B posterior edge of pronotum, left side, dorsal view, arrows showing sublateral incisions C head of holotype, frontal view D apex of last abdominal ventrite (ventrite V) of male and female, ventral view (paratypes), arrow showing apical lateral incisions E paratype, ventral view of pro- and mesothorax; indicating shapes of prosternal process, mesepisternum (upper, green arrow on antero-mesal angle) and projections on posterior edge of mesosternum (lower, red arrow)." figureDoi="10.3897/zookeys.993.53805.figure20" httpUri="https://binary.pensoft.net/fig/475453" pageId="0" pageNumber="47">20E</figureCitation>
, lower (red) arrow); in lateral view weak to absent, not produced anteriorly (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Lateral view of prosternal process and mesosternum of Chinese Phorocardius species. Upper green arrows indicating shape of posterior end of prosternal process; lower red arrows indicating projections (absent or present) on posterior edge of mesosternum (i. e., on anteroventral angle of mesosternal fossa according to Douglas 2003). A P. alterlineatus Ruan &amp; Douglas, sp. nov. B P. flavistriolatus Ruan &amp; Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus. F P. minutus Ruan &amp; Douglas, sp. nov. G P. rufiposterus Ruan &amp; Douglas, sp. nov. H P. unguicularis. I P. yunnanensis Ruan &amp; Douglas, sp. nov. J P. zhiweii Ruan, Douglas &amp; Qiu, sp. nov." figureDoi="10.3897/zookeys.993.53805.figure24" httpUri="https://binary.pensoft.net/fig/475458" pageId="0" pageNumber="47">24I</figureCitation>
, lower (red) arrow). Scutellar shield: width to length ratio 0.7-0.93; anterolateral edges slightly sinuate; posterior edge pointed. Elytra: upper edge of epipleura with minute serrations.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Legs</emphasis>
. Length ratio of metatarsomeres I-V (excluding claws): 100; 73-79; 65-79; 45-54; 138-154. Claw with ventral apex almost as large as dorsal apex.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Abdomen</emphasis>
. Lateral edges of visible abdominal ventrites I-V with minute serrations.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Male genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A holotype, male, dorsal view B paratype, female, dorsal view C paratype, female (Yunnan, Xi-shuan-ban-na, Meng-zhe), ventral view D aedeagus of holotype, dorsal view E aedeagus of holotype, ventral view, arrow indicating apex of paramere F aedeagus of holotype, lateral view G female abdominal sternite VIII, dorsal view (paratype) H ovipositor of paratype, dorsal view I distal (upper side) and proximal sclerites of bursa copulatrix (paratype)." figureDoi="10.3897/zookeys.993.53805.figure19" httpUri="https://binary.pensoft.net/fig/475452" pageId="0" pageNumber="47">19D-F</figureCitation>
). Robust in dorsal view, slender in lateral view. Median lobe in ventral view narrowing from base to basal third, parallel-sided and slender from basal third to apex, apex broadly rounded to truncate. Median lobe in lateral view evenly curved ventrad from base to apex, apex rounded and dilated. Paramere in dorsal view: wide from base to mid-length, gradually narrowed beyond mid-length, apex slanted; preapical lateral expansion small and sharp, facing laterally; apical mesal callus absent; width 2-2.5
<normalizedToken originalValue="×">x</normalizedToken>
median lobe width (measured at mid-length of paramere and median lobe respectively). Paramere in lateral view straight from base to mid-length, curved ventrad from mid-length to apex; apex without sharp hook-shaped preapical ventral expansion.
</paragraph>
<caption doi="10.3897/zookeys.993.53805.figure19" httpUri="https://binary.pensoft.net/fig/475452" pageId="0" pageNumber="47" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 19.</emphasis>
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
Ruan &amp; Douglas, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
holotype, male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
paratype, female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
paratype, female (Yunnan, Xi-shuan-ban-na, Meng-zhe), ventral view
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
aedeagus of holotype, dorsal view
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
aedeagus of holotype, ventral view, arrow indicating apex of paramere
<emphasis bold="true" pageId="0" pageNumber="47">F</emphasis>
aedeagus of holotype, lateral view
<emphasis bold="true" pageId="0" pageNumber="47">G</emphasis>
female abdominal sternite VIII, dorsal view (paratype)
<emphasis bold="true" pageId="0" pageNumber="47">H</emphasis>
ovipositor of paratype, dorsal view
<emphasis bold="true" pageId="0" pageNumber="47">I</emphasis>
distal (upper side) and proximal sclerites of bursa copulatrix (paratype).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Female</emphasis>
. Body length slightly larger than male (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A holotype, male, dorsal view B paratype, female, dorsal view C paratype, female (Yunnan, Xi-shuan-ban-na, Meng-zhe), ventral view D aedeagus of holotype, dorsal view E aedeagus of holotype, ventral view, arrow indicating apex of paramere F aedeagus of holotype, lateral view G female abdominal sternite VIII, dorsal view (paratype) H ovipositor of paratype, dorsal view I distal (upper side) and proximal sclerites of bursa copulatrix (paratype)." figureDoi="10.3897/zookeys.993.53805.figure19" httpUri="https://binary.pensoft.net/fig/475452" pageId="0" pageNumber="47">19B, C</figureCitation>
), apex of abdominal ventrite V in ventral view truncate to slightly convex, bent dorsad, each side with an incision (Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A pronotum and scutellum of holotype, dorsal view B posterior edge of pronotum, left side, dorsal view, arrows showing sublateral incisions C head of holotype, frontal view D apex of last abdominal ventrite (ventrite V) of male and female, ventral view (paratypes), arrow showing apical lateral incisions E paratype, ventral view of pro- and mesothorax; indicating shapes of prosternal process, mesepisternum (upper, green arrow on antero-mesal angle) and projections on posterior edge of mesosternum (lower, red arrow)." figureDoi="10.3897/zookeys.993.53805.figure20" httpUri="https://binary.pensoft.net/fig/475453" pageId="0" pageNumber="47">20D</figureCitation>
,
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Female abdominal ventrite V in Chinese Phorocardius species (ventral view). A P. alterlineatus Ruan &amp; Douglas, sp. nov. B P. flavistriolatus Ruan &amp; Douglas, sp. nov. C P. florentini. D P. magnus. E P. manuleatus F P. rufiposterus Ruan &amp; Douglas, sp. nov. G P. unguicularis H P. yunnanensis Ruan &amp; Douglas, sp. nov, arrows showing incisions near apex." figureDoi="10.3897/zookeys.993.53805.figure26" httpUri="https://binary.pensoft.net/fig/475460" pageId="0" pageNumber="47">26H</figureCitation>
, indicated by blue arrow) (in male, apex of abdominal ventrite V not bent dorsad, each side weakly concave, see Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Phorocardius yunnanensis Ruan &amp; Douglas, sp. nov. A pronotum and scutellum of holotype, dorsal view B posterior edge of pronotum, left side, dorsal view, arrows showing sublateral incisions C head of holotype, frontal view D apex of last abdominal ventrite (ventrite V) of male and female, ventral view (paratypes), arrow showing apical lateral incisions E paratype, ventral view of pro- and mesothorax; indicating shapes of prosternal process, mesepisternum (upper, green arrow on antero-mesal angle) and projections on posterior edge of mesosternum (lower, red arrow)." figureDoi="10.3897/zookeys.993.53805.figure20" httpUri="https://binary.pensoft.net/fig/475453" pageId="0" pageNumber="47">20D</figureCitation>
). Bursa copulatrix with proximal sclerites large, semi-spherical, base with deep concavity, apex narrowed and angulate; with many spines on internal surface: each with 9-11 large ones on mesal edge, 10-12 smaller ones on disc.
</paragraph>
<caption doi="10.3897/zookeys.993.53805.figure20" httpUri="https://binary.pensoft.net/fig/475453" pageId="0" pageNumber="47" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 20.</emphasis>
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
Ruan &amp; Douglas, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
pronotum and scutellum of holotype, dorsal view
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
posterior edge of pronotum, left side, dorsal view, arrows showing sublateral incisions
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
head of holotype, frontal view
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
apex of last abdominal ventrite (ventrite V) of male and female, ventral view (paratypes), arrow showing apical lateral incisions
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
paratype, ventral view of pro- and mesothorax; indicating shapes of prosternal process, mesepisternum (upper, green arrow on antero-mesal angle) and projections on posterior edge of mesosternum (lower, red arrow).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="type material">
<paragraph pageId="0" pageNumber="47">Type material.</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Holotype</emphasis>
. ♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-a (勐阿), 1050-1080 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.VI.6, leg. Fuji Pu [in Chinese]; 3) Holotype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Paratypes</emphasis>
(34♂42♀). 1♂1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-a (勐阿), 1050-1080 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.VI.9-10, leg. Shuyong Wang [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-a (勐阿), 1000 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.V.19, leg. Fuji Pu [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 2♂3♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-zhe (勐遮), 1200 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.VI.15, leg. Fuji Pu [in Chinese]; 3)
<taxonomicName baseAuthorityName="Miwa" baseAuthorityYear="1927" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yanagiharae" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yanagiharae">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yanagiharae</emphasis>
</taxonomicName>
, det. Siqin Ge; 4) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 5♂6♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-zhe (勐遮), 1200 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.VI.15, leg. Shuyong Wang [in Chinese]; 3)
<taxonomicName baseAuthorityName="Miwa" baseAuthorityYear="1927" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yanagiharae" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yanagiharae">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yanagiharae</emphasis>
</taxonomicName>
(Miwa), det. Shihong Jiang, 1998; 4) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-zhe (勐遮), 1200 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.IV.14, leg. Shuyong Wang [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-zhe (勐遮), 1200 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.VI.17, leg. Zhi-zi Chen [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-zhe (勐遮), 870 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.VII.7, leg. Fuji-Pu [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Meng-zhe (勐遮), Nan-nuo-shan, 1100 m, 1957.IV.28, leg. Fuji Pu, light trap [in Chinese]; 2) [same information as label 1, in Russian]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-hun (勐混), 1200 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.V.28, leg. Shuyong Wang [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-hun (勐混), 1000-1200 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.V.21, leg. Leyi Zheng [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 2♂1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-hun (勐混), 1200-1400 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.V.17-24, leg. Xuwu Meng [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Meng-hun (勐混), 1200 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.V.17, leg. Xuwu Meng [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Xi-shuang-ban-na, Da-meng-long (大勐龙), 650 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.VI.9, leg. Shuyong Wang [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Jing-dong (景东), Dong-jia-fen, 1250 m, 1956.V.27, leg. Zha-gu-liang-ye-fu [in Chinese]; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Jing-dong (景东), 1170 m, 1956.VI.30, leg. Zha-gu-liang-ye-fu, light trap [in Chinese]; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Jing-dong (景东), Dong-jia-fen, 1250 m, 1956.VI.29, leg. Krizhanovsky [in Chinese]; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 2♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Jing-dong (景东), Dong-jia-fen, 1250 m, 1956.VI.3, leg. Krizhanovsky [in Chinese]; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Bao-shan (保山) to Yong-ping (永平), 1955.V.28, leg. Le Wu [in Chinese]; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, east of Bao-shan (保山), Lang-cang river 1200 m, 1955.V.28, leg. Bu-xi-ke [in Chinese]; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 2♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Si-mao (思茅), 1380 m, Chinese Academy of Sciences [in Chinese]; 2) 1958.VI.7, leg. Shuyong Wang [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 2♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Si-mao (思茅), Pu-wen, 950-1200 m, 1957.V.11, leg. Guangji Hong &amp; Zhirang Meng [in Chinese]; 2) [same information as label 1, in Russian]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 3♂12♀ (SZPT, ex. LQCC), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Pu-er, Si-mao (思茅), 1400 m, 9-11.V.2018, Meizihu Park, leg. Jianyue Qiu &amp; Hao xu, Shenzhen Polytechnic [in Chinese]; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 6♂3♀ (SZPT), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Lin-cang, Yun-xian County, Man-wang township (漫湾镇), light trap, VI-VIII, 2018, leg. Zichun Xiong, Shenzhen Polytechnic [in Chinese]; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂2♀ (SZPT), labels: 1) Nabang Town,
<normalizedToken originalValue="那邦镇">那邦镇</normalizedToken>
, Yingjiang County,
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, 2018-IV-3, 252 m, Lu Qiu Leg.; 2) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 4♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Cang-yuan (沧源), 750-790 m, Chinese Academy of Sciences [in Chinese]; 2) 1980.V.19-22, leg. Jinwen Shang [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Cang-yuan (沧源), ban-lao, 1100 m, Chinese Academy of Sciences [in Chinese]; 2) 1980.V.18, leg. Hongxing Li [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♂ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Lu-xi (潞西), 1250 m, Chinese Academy of Sciences [in Chinese]; 2) 1980.V.18, leg. Jinwen Shang [in Chinese]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019. • 1♀ (IZCAS), labels: 1)
<emphasis bold="true" pageId="0" pageNumber="47">Yunnan</emphasis>
, Rui-li (瑞丽), 1400 m, 1956.VI.6, leg. Benshou Zhou [in Chinese]; 2) [same information as label 1, in Russian]; 3) Paratype
<taxonomicName authorityName="Ruan &amp; Douglas" authorityYear="2020" class="Insecta" family="Elateridae" genus="Phorocardius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phorocardius yunnanensis" order="Coleoptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">Phorocardius yunnanensis</emphasis>
</taxonomicName>
sp. nov. Des. Ruan &amp; Douglas, 2019.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="remarks">
<paragraph pageId="0" pageNumber="47">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="47">Based on specimen information, this species is currently only known from mountainous southwest Yunnan, China. It inhabits mainly middle elevations (ca. 650-1400 m). Southwest Yunnan is rainy and humid, subtropical to tropical with subtropical evergreen broad-leaf forest or tropical rain forest. Specimens were collected by sweep-netting and light traps, indicating both diurnal and nocturnal activity.</paragraph>
</subSubSection>
</treatment>
</document>