treatments-xml/data/F9/7F/17/F97F1711FFE93950E39F9FF72D45BFEA.xml
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<document id="F834EE62E67D31D09DDDBFED6F220485" ID-DOI="10.1111/zoj.12207" ID-ISSN="0024-4082" ID-Zenodo-Dep="5333281" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630173944707" checkinUser="felipe" docAuthor="Bragagnolo, Cibele, Hara, Marcos R. &amp; Pinto-Da-Rocha, Ricardo" docDate="2015" docId="F97F1711FFE93950E39F9FF72D45BFEA" docLanguage="en" docName="zoj12207.pdf" docOrigin="Zoological Journal of the Linnean Society 173 (2)" docSource="http://dx.doi.org/10.1111/zoj.12207" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Gonycranaus pluto Bragagnolo &amp; Hara &amp; Pinto-Da-Rocha 2015, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="317" masterDocId="05466F69FFFD3945E0259A282E28B935" masterDocTitle="A new family of Gonyleptoidea from South America (Opiliones, Laniatores)" masterLastPageNumber="319" masterPageNumber="296" pageNumber="316" updateTime="1699174272660" updateUser="plazi">
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<mods:title id="E414845679103B6A3863B1F67717936B">A new family of Gonyleptoidea from South America (Opiliones, Laniatores)</mods:title>
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<mods:namePart id="47CEFDEABFBAA3741773119CEB3DFDA0">Bragagnolo, Cibele</mods:namePart>
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<mods:namePart id="B21C02341EFD1C7DD783BD0DC9D386E8">Hara, Marcos R.</mods:namePart>
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<mods:namePart id="F5A9585EDA109647284B4964DCDC975D">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:title id="03E1C14BC4B41FD404D8F230C64413AC">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="409E44832172BFF078DE389E7074A4D9">2015</mods:date>
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<mods:number id="78E62D0670C64722C52C740E3C08B6FE">2015-01-29</mods:number>
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<mods:number id="C6D2CC045F5AA5D2E63C15C5A622C27B">173</mods:number>
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<treatment id="F97F1711FFE93950E39F9FF72D45BFEA" LSID="urn:lsid:plazi:treatment:F97F1711FFE93950E39F9FF72D45BFEA" httpUri="http://treatment.plazi.org/id/F97F1711FFE93950E39F9FF72D45BFEA" lastPageId="21" lastPageNumber="317" pageId="20" pageNumber="316">
<subSubSection id="39CCF58CFFE93951E39F9FF72B09BCC1" box="[954,1313,1503,1524]" pageId="20" pageNumber="316" type="nomenclature">
<paragraph id="7169A607FFE93951E39F9FF72B09BCC1" blockId="20.[954,1313,1503,1557]" box="[954,1313,1503,1524]" pageId="20" pageNumber="316">
<heading id="2A21116BFFE93951E39F9FF72B09BCC1" box="[954,1313,1503,1524]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="316" reason="4">
<taxonomicName id="B6D6DD84FFE93951E39F9FF72A96BCC6" ID-CoL="7WCWH" authority="Bragagnolo &amp; Hara &amp; Pinto-Da-Rocha, 2015" authorityName="Bragagnolo &amp; Hara &amp; Pinto-Da-Rocha" authorityYear="2015" box="[954,1214,1503,1524]" class="Arachnida" family="Gerdesiidae" genus="Gonycranaus" kingdom="Animalia" order="Opiliones" pageId="20" pageNumber="317" phylum="Arthropoda" rank="species" species="pluto" status="sp. nov.">GONYCRANAUS PLUTO</taxonomicName>
<emphasis id="43A27A15FFE93951E4E39FCA2B09BCC1" bold="true" box="[1222,1313,1503,1524]" pageId="20" pageNumber="316">
<taxonomicNameLabel id="5891C76EFFE93951E4E39FCA2B09BCC1" box="[1222,1313,1503,1524]" pageId="20" pageNumber="316" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39CCF58CFFE93951E43D9FD72DFFBFC6" pageId="20" pageNumber="316" type="description">
<paragraph id="7169A607FFE93951E43D9FD72AEBBF20" blockId="20.[954,1313,1503,1557]" box="[1048,1219,1535,1557]" pageId="20" pageNumber="316">
(
<figureCitation id="E9EDBA82FFE93951E4059FD72A45BF20" box="[1056,1133,1535,1557]" captionStart="Figure 6" captionStartId="9.[144,223,1779,1798]" captionTargetBox="[204,1364,198,1744]" captionTargetId="figure-0@9.[202,1364,194,1748]" captionTargetPageId="9" captionText="Figure 6. Gonycranaus pluto sp. nov. AF, male holotype: A, habitus, dorsal view; B, habitus, right lateral view; C, right trochanterpatella IV, dorsal view; D, right trochanterpatella IV, ventral view; E, right pedipalp, retrolateral view; F, right pedipalp, prolateral view. G, H, female paratype (MZSP 57197): G, habitus, dorsal view; H, habitus, right lateral view. C and D at same scale; E and F at same scale. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5333297" httpUri="https://zenodo.org/record/5333297/files/figure.png" pageId="20" pageNumber="316">FIGS 6</figureCitation>
;
<figureCitation id="E9EDBA82FFE93951E45F9FD72A93BF20" box="[1146,1211,1535,1557]" captionStart="Figure 9" captionStartId="12.[164,243,1832,1851]" captionTargetBox="[164,1444,198,1796]" captionTargetId="figure-0@12.[164,1444,193,1801]" captionTargetPageId="12" captionText="Figure 9. Distal part of penis. AC, Gonycranaus androgynus sp. nov. (holotype): A, dorsal view; B, left lateral view; C, ventral view. DF, Gonycranaus pluto sp. nov. (holotype): D, dorsal view; E, left lateral view; F, ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5333307" httpUri="https://zenodo.org/record/5333307/files/figure.png" pageId="20" pageNumber="316">9DF</figureCitation>
)
</paragraph>
<paragraph id="7169A607FFE93951E31F9C0D2D99BF0F" blockId="20.[826,1441,1573,1718]" box="[826,945,1573,1594]" pageId="20" pageNumber="316">
<emphasis id="43A27A15FFE93951E31F9C0D2D99BF0F" box="[826,945,1573,1594]" italics="true" pageId="20" pageNumber="316">Etymology</emphasis>
</paragraph>
<paragraph id="7169A607FFE93951E31F9C6C2DEBBF83" blockId="20.[826,1441,1573,1718]" pageId="20" pageNumber="316">After Pluto, one of the six Olympian gods and the god of the underworld previously named as Hades. In reference to the fact its species is restricted to cave environments.</paragraph>
<paragraph id="7169A607FFE93951E31F9CF72DFFBFC6" blockId="20.[826,1442,1758,1903]" box="[826,983,1758,1780]" pageId="20" pageNumber="316">
<emphasis id="43A27A15FFE93951E31F9CF72DFFBFC6" box="[826,983,1758,1780]" italics="true" pageId="20" pageNumber="316">
<typeStatus id="AE6D18A5FFE93951E31F9CF72D46BFC1" box="[826,878,1759,1780]" pageId="20" pageNumber="316">Type</typeStatus>
material
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="39CCF58CFFE93950E31F9CD52D45BFEA" lastPageId="21" lastPageNumber="317" pageId="20" pageNumber="316" type="materials_examined">
<paragraph id="7169A607FFE93950E31F9CD52CB6B840" blockId="20.[826,1442,1758,1903]" lastBlockId="21.[144,759,197,373]" lastPageId="21" lastPageNumber="317" pageId="20" pageNumber="316">
<materialsCitation id="C1BEAC5AFFE93951E31F9CD52A80BE65" collectingDate="2011-10-03" collectingDateMax="2011-10-06" collectingDateMin="2011-10-03" collectionCode="MZSP" collectorName="Gruta MP &amp; Bessi" country="Brazil" location="Morro do Pilar" pageId="20" pageNumber="316" specimenCode="MZSP 57196" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<collectingCountry id="09C1E697FFE93951E31F9CD52DB3BE27" box="[826,923,1789,1810]" name="Brazil" pageId="20" pageNumber="316">BRAZIL</collectingCountry>
.
<collectingRegion id="B31268E5FFE93951E3839CD52A68BE26" box="[934,1088,1789,1811]" country="Brazil" name="Minas Gerais" pageId="20" pageNumber="316">Minas Gerais</collectingRegion>
:
<location id="7409F0DCFFE93951E46E9CD52AD9BE26" LSID="urn:lsid:plazi:treatment:F97F1711FFE93950E39F9FF72D45BFEA:7409F0DCFFE93951E46E9CD52AD9BE26" box="[1099,1265,1789,1811]" country="Brazil" name="Morro do Pilar" pageId="20" pageNumber="316" stateProvince="Minas Gerais">Morro do Pilar</location>
,
<collectorName id="DC23C3D1FFE93951E4D99CD52B59BE27" box="[1276,1393,1789,1810]" pageId="20" pageNumber="316">Gruta MP</collectorName>
01B, 881337mN/667890mE SAD69,
<date id="056880C7FFE93951E4B59D332B0ABE04" box="[1168,1314,1819,1841]" pageId="20" pageNumber="316" value="2011-10-03" valueMax="2011-10-06" valueMin="2011-10-03">
<collectingDate id="152C792FFFE93951E4B59D332B0ABE04" box="[1168,1314,1819,1841]" pageId="20" pageNumber="316" value="2011-10-03" valueMax="2011-10-06" valueMin="2011-10-03">0306.X.2011</collectingDate>
</date>
,
<collectorName id="DC23C3D1FFE93951E5089D342B4FBE07" box="[1325,1383,1820,1842]" pageId="20" pageNumber="316">Bessi</collectorName>
<emphasis id="43A27A15FFE93951E5489D342BB3BE04" box="[1389,1435,1820,1841]" italics="true" pageId="20" pageNumber="316">et al</emphasis>
. leg,
<specimenCount id="67D06D8EFFE93951E34B9D122DAFBE65" box="[878,903,1850,1872]" pageId="20" pageNumber="316" type="male"></specimenCount>
<typeStatus id="AE6D18A5FFE93951E3B79D122DDBBE65" box="[914,1011,1850,1872]" pageId="20" pageNumber="316" type="holotype">holotype</typeStatus>
(
<specimenCode id="21700E7CFFE93951E4209D122A88BE65" box="[1029,1184,1850,1872]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="20" pageNumber="316" type="Museum">MZSP 57196</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="C1BEAC5AFFE93951E4939D132B05BE65" box="[1206,1325,1851,1872]" country="Brazil" location="Paratypes" pageId="20" pageNumber="316" specimenCount="1" stateProvince="Minas Gerais" typeStatus="paratype">
<typeStatus id="AE6D18A5FFE93951E4939D132B05BE65" box="[1206,1325,1851,1872]" pageId="20" pageNumber="316" type="paratype">
<location id="7409F0DCFFE93951E4939D132B05BE65" LSID="urn:lsid:plazi:treatment:F97F1711FFE93950E39F9FF72D45BFEA:7409F0DCFFE93951E4939D132B05BE65" box="[1206,1325,1851,1872]" country="Brazil" name="Paratypes" pageId="20" pageNumber="316" stateProvince="Minas Gerais">Paratypes</location>
</typeStatus>
</materialsCitation>
:
<materialsCitation id="C1BEAC5AFFE93950E51E9D122FB9B9CF" collectingDate="2012-02-28" collectionCode="MZSP" collectorName="Gruta MP &amp; Bessi" country="Brazil" lastPageId="21" lastPageNumber="317" location="Morro do Pilar" pageId="20" pageNumber="316" specimenCode="MZSP 57197" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="holotype">
<collectingCountry id="09C1E697FFE93951E51E9D122BB6BE7A" box="[1339,1438,1850,1871]" name="Brazil" pageId="20" pageNumber="316">BRAZIL</collectingCountry>
.
<collectingRegion id="B31268E5FFE93951E31F9D712DC6BE5A" box="[826,1006,1881,1903]" country="Brazil" name="Minas Gerais" pageId="20" pageNumber="316">Minas Gerais</collectingRegion>
:
<location id="7409F0DCFFE93951E4219D712AE5BE5A" LSID="urn:lsid:plazi:treatment:F97F1711FFE93950E39F9FF72D45BFEA:7409F0DCFFE93951E4219D712AE5BE5A" box="[1028,1229,1881,1903]" country="Brazil" name="Morro do Pilar" pageId="20" pageNumber="316" stateProvince="Minas Gerais">Morro do Pilar</location>
,
<collectorName id="DC23C3D1FFE93951E4C69D712B45BE5B" box="[1251,1389,1881,1902]" pageId="20" pageNumber="316">Gruta MP</collectorName>
01B, 7881337mN/667890mE SAD69,
<date id="056880C7FFE83950E1DB9AEE2C59B9EE" box="[510,625,198,219]" pageId="21" pageNumber="317" value="2012-02-28">
<collectingDate id="152C792FFFE83950E1DB9AEE2C59B9EE" box="[510,625,198,219]" pageId="21" pageNumber="317" value="2012-02-28">28.II.2012</collectingDate>
</date>
,
<collectorName id="DC23C3D1FFE83950E25B9AEE2C91B9E9" box="[638,697,198,220]" pageId="21" pageNumber="317">Bessi</collectorName>
<emphasis id="43A27A15FFE83950E2E59AEE2CD8B9EE" box="[704,752,198,219]" italics="true" pageId="21" pageNumber="317">et al</emphasis>
. leg.,
<specimenCount id="67D06D8EFFE83950E0E19ACD2EC4B9CF" box="[196,236,228,250]" pageId="21" pageNumber="317" type="female">1 ♀</specimenCount>
(
<specimenCode id="21700E7CFFE83950E0DF9ACD2FA2B9CF" box="[250,394,228,250]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57197</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C1BEAC5AFFE83950E1B89ACC2C86B9CF" box="[413,686,228,250]" collectingDate="2012-02-28" collectionCode="MZSP" collectorName="Gruta MP &amp; Bessi" country="Brazil" location="Morro do Pilar" pageId="21" pageNumber="317" specimenCode="MZSP 57192" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="holotype">
idem,
<specimenCount id="67D06D8EFFE83950E1C49ACD2C21B9CF" box="[481,521,228,250]" pageId="21" pageNumber="317" type="female">1 ♀</specimenCount>
(
<specimenCode id="21700E7CFFE83950E2329ACD2C8FB9CF" box="[535,679,228,250]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57192</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C1BEAC5AFFE83950E29F9ACC2F75B82C" collectingDate="2012-02-28" collectionCode="MZSP" collectorName="Gruta MP &amp; Bessi" country="Brazil" location="Morro do Pilar" pageId="21" pageNumber="317" specimenCode="MZSP 57242" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="holotype">
idem,
<specimenCount id="67D06D8EFFE83950E0B59B2B2E90B82C" box="[144,184,259,281]" pageId="21" pageNumber="317" type="female">1 ♀</specimenCount>
(
<specimenCode id="21700E7CFFE83950E0E39B2B2F7EB82C" box="[198,342,259,281]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57242</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="C1BEAC5AFFE83950E14C9B2B2C57B82C" box="[361,639,259,281]" collectingDate="2012-02-28" collectionCode="MZSP" collectorName="Gruta MP &amp; Bessi" country="Brazil" location="Morro do Pilar" pageId="21" pageNumber="317" specimenCode="MZSP 57243" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
idem,
<specimenCount id="67D06D8EFFE83950E1889B2B2FF2B82C" box="[429,474,259,281]" pageId="21" pageNumber="317" type="male">1 ♂</specimenCount>
(
<specimenCode id="21700E7CFFE83950E1C29B2B2C50B82C" box="[487,632,259,281]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57243</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C1BEAC5AFFE83950E2AF9B2B2F1FB802" collectingDate="2012-02-28" collectionCode="MZSP" collectorName="Gruta MP &amp; Bessi" country="Brazil" location="Morro do Pilar" pageId="21" pageNumber="317" specimenCode="MZSP 57244" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="holotype">
idem,
<specimenCount id="67D06D8EFFE83950E2EA9B2B2CDFB82C" box="[719,759,259,281]" pageId="21" pageNumber="317" type="female">1 ♀</specimenCount>
(
<specimenCode id="21700E7CFFE83950E0B29B0A2F07B802" box="[151,303,289,311]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57244</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C1BEAC5AFFE83950E1639B0A2C5AB802" box="[326,626,289,312]" collectingDate="2012-02-28" collectionCode="MZSP" collectorName="Gruta MP &amp; Bessi" country="Brazil" location="Morro do Pilar" pageId="21" pageNumber="317" specimenCode="MZSP 57245" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
idem,
<specimenCount id="67D06D8EFFE83950E1B79B0A2F97B802" box="[402,447,289,311]" pageId="21" pageNumber="317" type="male">1 ♂</specimenCount>
(
<specimenCode id="21700E7CFFE83950E1F79B0A2C42B802" box="[466,618,289,311]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57245</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C1BEAC5AFFE83950E2A79B0A2F19B863" collectingDate="2012-02-28" collectionCode="MZSP" collectorName="Gruta MP &amp; Bessi" country="Brazil" location="Morro do Pilar" pageId="21" pageNumber="317" specimenCode="MZSP 57246" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="holotype">
idem,
<specimenCount id="67D06D8EFFE83950E2E89B0A2CDFB802" box="[717,759,289,311]" pageId="21" pageNumber="317" type="female">1 ♀</specimenCount>
(
<specimenCode id="21700E7CFFE83950E0B29B682F03B863" box="[151,299,320,342]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57246</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C1BEAC5AFFE83950E1189B682F4FB840" collectingDate="2011-10-03" collectingDateMax="2011-10-06" collectingDateMin="2011-10-03" collectionCode="MZSP" collectorName="R. Bessi" country="Brazil" location="Morro do Pilar" pageId="21" pageNumber="317" specimenCode="MZSP 57247" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="holotype">
idem,
<date id="056880C7FFE83950E1A69B682C35B863" box="[387,541,320,342]" pageId="21" pageNumber="317" value="2011-10-03" valueMax="2011-10-06" valueMin="2011-10-03">
<collectingDate id="152C792FFFE83950E1A69B682C35B863" box="[387,541,320,342]" pageId="21" pageNumber="317" value="2011-10-03" valueMax="2011-10-06" valueMin="2011-10-03">0306.X.2011</collectingDate>
</date>
,
<collectorName id="DC23C3D1FFE83950E20D9B692CABB863" box="[552,643,320,342]" pageId="21" pageNumber="317">R. Bessi</collectorName>
<emphasis id="43A27A15FFE83950E2AF9B692C92B860" box="[650,698,320,342]" italics="true" pageId="21" pageNumber="317">et al</emphasis>
. leg.,
<specimenCount id="67D06D8EFFE83950E0B59B772E91B841" box="[144,185,350,372]" pageId="21" pageNumber="317" type="female">1 ♀</specimenCount>
(
<specimenCode id="21700E7CFFE83950E0EF9B772F48B840" box="[202,352,351,373]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57247</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C1BEAC5AFFE83950E1509B772CB2B840" box="[373,666,350,373]" collectingDate="2011-10-03" collectingDateMax="2011-10-06" collectingDateMin="2011-10-03" collectionCode="MZSP" collectorName="R. Bessi" country="Brazil" location="Morro do Pilar" pageId="21" pageNumber="317" specimenCode="MZSP 57248" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
idem,
<specimenCount id="67D06D8EFFE83950E19B9B772FC3B841" box="[446,491,350,372]" pageId="21" pageNumber="317" type="male">1 ♂</specimenCount>
(
<specimenCode id="21700E7CFFE83950E1D99B772CBAB840" box="[508,658,351,373]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP 57248</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="7169A607FFE83950E0B59BB52F28B887" blockId="21.[144,759,413,526]" box="[144,256,413,434]" pageId="21" pageNumber="317">
<emphasis id="43A27A15FFE83950E0B59BB52F28B887" box="[144,256,413,434]" italics="true" pageId="21" pageNumber="317">Diagnosis</emphasis>
</paragraph>
<paragraph id="7169A607FFE83950E0B59B932C80BB3B" blockId="21.[144,759,413,526]" pageId="21" pageNumber="317">
<taxonomicName id="B6D6DD84FFE83950E0B59B932F43B8E5" authorityName="Bragagnolo &amp; Hara &amp; Pinto-Da-Rocha" authorityYear="2015" box="[144,363,443,464]" class="Arachnida" family="Gerdesiidae" genus="Gonycranaus" kingdom="Animalia" order="Opiliones" pageId="21" pageNumber="317" phylum="Arthropoda" rank="species" species="pluto">
<emphasis id="43A27A15FFE83950E0B59B932F43B8E5" box="[144,363,443,464]" italics="true" pageId="21" pageNumber="317">Gonycranaus pluto</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="B6D6DD84FFE83950E2759B932CDEB8E5" authorityName="Bragagnolo &amp; Hara &amp; Pinto-Da-Rocha" authorityYear="2015" box="[592,758,443,464]" class="Arachnida" family="Gerdesiidae" genus="Gonycranaus" kingdom="Animalia" order="Opiliones" pageId="21" pageNumber="317" phylum="Arthropoda" rank="species" species="androgynus">
<emphasis id="43A27A15FFE83950E2759B932CDEB8E5" box="[592,758,443,464]" italics="true" pageId="21" pageNumber="317">G. androgynus</emphasis>
</taxonomicName>
and differs by legs IIV elongated, body coloration lightened, and absence of ventral process of penis.
</paragraph>
<paragraph id="7169A607FFE83950E0B5981E2F39BB7E" blockId="21.[144,759,566,1902]" box="[144,273,566,587]" pageId="21" pageNumber="317">
<emphasis id="43A27A15FFE83950E0B5981E2F39BB7E" box="[144,273,566,587]" italics="true" pageId="21" pageNumber="317">Description</emphasis>
</paragraph>
<paragraph id="7169A607FFE83950E0B5987C2FBABAC2" blockId="21.[144,759,566,1902]" pageId="21" pageNumber="317">
<emphasis id="43A27A15FFE83950E0B5987C2F6EBB5C" box="[144,326,596,617]" italics="true" pageId="21" pageNumber="317">
Male (
<typeStatus id="AE6D18A5FFE83950E0FC987C2F10BB5C" box="[217,312,596,617]" pageId="21" pageNumber="317" type="holotype">holotype</typeStatus>
):
</emphasis>
Dorsum (
<figureCitation id="E9EDBA82FFE83950E19E987C2C19BB5C" box="[443,561,595,618]" captionStart="Figure 6" captionStartId="9.[144,223,1779,1798]" captionTargetBox="[204,1364,198,1744]" captionTargetId="figure-0@9.[202,1364,194,1748]" captionTargetPageId="9" captionText="Figure 6. Gonycranaus pluto sp. nov. AF, male holotype: A, habitus, dorsal view; B, habitus, right lateral view; C, right trochanterpatella IV, dorsal view; D, right trochanterpatella IV, ventral view; E, right pedipalp, retrolateral view; F, right pedipalp, prolateral view. G, H, female paratype (MZSP 57197): G, habitus, dorsal view; H, habitus, right lateral view. C and D at same scale; E and F at same scale. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5333297" httpUri="https://zenodo.org/record/5333297/files/figure.png" pageId="21" pageNumber="317">Fig. 6A, B</figureCitation>
): Measurements: DSL 3.6; DSW 2.95; LI 15.2; LII 26.3; LIII 21.0; LIV 28.9. Leg IV/body length ratio: 8.02. Anterior margin of carapace with frontal hump covered by tubercles. Ocularium far from anterior margin of carapace, tuberculate, unarmed. Scutal area I entire; scutal areas IIV unarmed. Scutal groove II complete. Dorsal scutum outline type alpha, entirely tuberculate. One ozopore (anterior opening), covered by integumentary dome with slit-like opening obstructed by prolateral apophysis of coxa II (on lateral view). Posterior margin of dorsal scutum densely tuberculate, unarmed. Free tergites I III each with a row of tubercles. Anal operculum tuberculate, unarmed.
</paragraph>
<paragraph id="7169A607FFE83950E08D9E282F32BD01" blockId="21.[144,759,566,1902]" pageId="21" pageNumber="317">Venter. Coxa IIV, genital and free sternites tuberculate.</paragraph>
<paragraph id="7169A607FFE83950E08D9E152F43BD9A" blockId="21.[144,759,566,1902]" pageId="21" pageNumber="317">Chelicera. Enlarged. Segment I completely covered with pointed tubercles on bulla, segment II very enlarged with six teeth, segment III with six teeth (the basalmost largest).</paragraph>
<paragraph id="7169A607FFE83950E08D9E9F2CDFBF6E" blockId="21.[144,759,566,1902]" pageId="21" pageNumber="317">
Pedipalp (
<figureCitation id="E9EDBA82FFE83950E13E9E9F2FA4BDF8" box="[283,396,1207,1229]" captionStart="Figure 6" captionStartId="9.[144,223,1779,1798]" captionTargetBox="[204,1364,198,1744]" captionTargetId="figure-0@9.[202,1364,194,1748]" captionTargetPageId="9" captionText="Figure 6. Gonycranaus pluto sp. nov. AF, male holotype: A, habitus, dorsal view; B, habitus, right lateral view; C, right trochanterpatella IV, dorsal view; D, right trochanterpatella IV, ventral view; E, right pedipalp, retrolateral view; F, right pedipalp, prolateral view. G, H, female paratype (MZSP 57197): G, habitus, dorsal view; H, habitus, right lateral view. C and D at same scale; E and F at same scale. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5333297" httpUri="https://zenodo.org/record/5333297/files/figure.png" pageId="21" pageNumber="317">Fig. 6E, F</figureCitation>
). Coxa with one ventral pointed apophysis. Trochanter inflated, dorsally with four or five pointed tubercles, ventrally with one tubercle and one pointed apophysis. Femur dorsally with a median row of nine or ten pointed tubercles, one apical prolateral row of six tubercles, one apical retrolateral row of between three and five tubercles, and one prolateral apical pointed apophysis. Ventrally with a row of nine or ten high, enlarged pointed tubercles, the basalmost largest. Patella dorsolaterally with scattered tubercles, ventrally smooth. Tibia dorsolaterally with scattered tubercles; tibial setation, mesal IiiIiIi, ectal IiIiIi. Tarsus dorsolaterally smooth; tarsal setation, mesal iIiiIii, ectal iIiIiii. Tarsal claw as long as tarsus length.
</paragraph>
<paragraph id="7169A607FFE83950E08D9C4C2B02BB7F" blockId="21.[144,759,566,1902]" lastBlockId="21.[806,1422,198,831]" pageId="21" pageNumber="317">
Legs (
<figureCitation id="E9EDBA82FFE83950E0C99C4C2F77BF4C" box="[236,351,1636,1658]" captionStart="Figure 6" captionStartId="9.[144,223,1779,1798]" captionTargetBox="[204,1364,198,1744]" captionTargetId="figure-0@9.[202,1364,194,1748]" captionTargetPageId="9" captionText="Figure 6. Gonycranaus pluto sp. nov. AF, male holotype: A, habitus, dorsal view; B, habitus, right lateral view; C, right trochanterpatella IV, dorsal view; D, right trochanterpatella IV, ventral view; E, right pedipalp, retrolateral view; F, right pedipalp, prolateral view. G, H, female paratype (MZSP 57197): G, habitus, dorsal view; H, habitus, right lateral view. C and D at same scale; E and F at same scale. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5333297" httpUri="https://zenodo.org/record/5333297/files/figure.png" pageId="21" pageNumber="317">Fig. 6C, D</figureCitation>
). Coxa I with one prolateral and one retrolateral apophyses (retrolateral largest); coxa II and III with one prolateral and one retrolateral apophyses; coxa IV tuberculate, with one short bifid prolateral apical apophysis and one small apical retrolateral pointed tubercle (visible ventrally). Trochanters tibiae IIV tuberculate; trochanter IV with one retrolateral apical enlarged tubercle. Femur III densely tuberculate, ventrally with one retrolateral and one prolateral row of tubercles increasing in size apically. Femur IV straight and densely tuberculate, with one prodorsal and one retrodorsal apical enlarged, pointed tubercles; a row of retrolateral spiniform tubercles (increasing in size apically); ventrally with one retrolateral and one prolateral row of tubercles increasing in size apically, with one apical pointed apophysis. Patella III ventrally with one median enlarged tubercle and one apical pointed apophysis. Patella IV with four dorsoapical enlarged, pointed tubercles and ventrally with one apical pointed apophysis. Tibia IV with a retroventral row of tubercles increasing in size apically. Basitarsus I not inflated. Tarsal counts: 9(3), 18(3), 7, and 8.
</paragraph>
<paragraph id="7169A607FFE83950E31A987B2DF5BA15" blockId="21.[806,1422,198,831]" pageId="21" pageNumber="317">
Penis (
<figureCitation id="E9EDBA82FFE83950E3AC987B2DDEBB5C" box="[905,1014,595,617]" captionStart="Figure 9" captionStartId="12.[164,243,1832,1851]" captionTargetBox="[164,1444,198,1796]" captionTargetId="figure-0@12.[164,1444,193,1801]" captionTargetPageId="12" captionText="Figure 9. Distal part of penis. AC, Gonycranaus androgynus sp. nov. (holotype): A, dorsal view; B, left lateral view; C, ventral view. DF, Gonycranaus pluto sp. nov. (holotype): D, dorsal view; E, left lateral view; F, ventral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5333307" httpUri="https://zenodo.org/record/5333307/files/figure.png" pageId="21" pageNumber="317">Fig. 9DF</figureCitation>
). Ventral plate distal margin straight, its width a quarter of the basal portion, basally as wide as truncus apex; with three subapical pairs of unbranched setae, three basal pairs of unbranched long setae (longest set). Truncus apex slightly inflated. Glans sac subcylindrical. Stylus short, slightly curved. Without ventral process.
</paragraph>
<paragraph id="7169A607FFE83950E31A99012B45BA0A" blockId="21.[806,1422,198,831]" box="[831,1389,809,831]" pageId="21" pageNumber="317">Coloration. Background colour yellowish brown.</paragraph>
<paragraph id="7169A607FFE83950E30399482BAABADA" blockId="21.[806,1422,864,1008]" pageId="21" pageNumber="317">
<emphasis id="43A27A15FFE83950E30399482B33BA40" box="[806,1307,864,885]" italics="true" pageId="21" pageNumber="317">
Female (
<typeStatus id="AE6D18A5FFE83950E3AD99482DC3BA40" box="[904,1003,864,885]" pageId="21" pageNumber="317" type="paratype">paratype</typeStatus>
<collectionCode id="17C73EC2FFE83950E3D699482A13BA40" box="[1011,1083,864,885]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="317" type="Museum">MZSP</collectionCode>
57197;
<figureCitation id="E9EDBA82FFE83950E4B099482B25BA40" box="[1173,1293,864,885]" captionStart="Figure 6" captionStartId="9.[144,223,1779,1798]" captionTargetBox="[204,1364,198,1744]" captionTargetId="figure-0@9.[202,1364,194,1748]" captionTargetPageId="9" captionText="Figure 6. Gonycranaus pluto sp. nov. AF, male holotype: A, habitus, dorsal view; B, habitus, right lateral view; C, right trochanterpatella IV, dorsal view; D, right trochanterpatella IV, ventral view; E, right pedipalp, retrolateral view; F, right pedipalp, prolateral view. G, H, female paratype (MZSP 57197): G, habitus, dorsal view; H, habitus, right lateral view. C and D at same scale; E and F at same scale. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5333297" httpUri="https://zenodo.org/record/5333297/files/figure.png" pageId="21" pageNumber="317">Fig. 6G, H</figureCitation>
):
</emphasis>
Measurements: DSL 3.30; DSW 2.95; LI 16.6; LII 27.0; LIII 22.8; LIV 28.5. Leg IV/body length ratio: 8.6. Pedipalp: tibial setation, mesal IIiIi, ectal IiIiIi; tarsal setation, mesal iIiiI, ectal iIiIii. Tarsal counts: 8(3), 19(3), 7, and 8.
</paragraph>
<paragraph id="7169A607FFE83950E3039E392D58BDC9" blockId="21.[806,1422,1040,1277]" pageId="21" pageNumber="317">
<emphasis id="43A27A15FFE83950E3039E392A0DBD13" box="[806,1061,1040,1062]" italics="true" pageId="21" pageNumber="317">Variation in males (N</emphasis>
=
<emphasis id="43A27A15FFE83950E4679E392A76BD13" box="[1090,1118,1041,1062]" italics="true" pageId="21" pageNumber="317">3):</emphasis>
Measurements: DSL 3.3 3.5; DSW 2.802.85; LI 17.119.9; LII 30.531.0; LIII 21.722.3; LIV 28.929.4. Leg IV/body length ratio: 8.38.9. Pedipalp: femur dorsally with a median row of between nine and 11 pointed tubercles; tibial setation, mesal IiIiiI/IiIiIi, ectal IiIiIi/IiIIi; tarsal setation, mesal iIiiIii, ectal iIiIi. Tarsal counts: 910(3), 1820(3), 7, and 8.
</paragraph>
<paragraph id="7169A607FFE83950E3039F362AADBCFE" blockId="21.[806,1422,1309,1759]" pageId="21" pageNumber="317">
<emphasis id="43A27A15FFE83950E3039F362A06BC07" box="[806,1070,1309,1331]" italics="true" pageId="21" pageNumber="317">Variation in females (N</emphasis>
=
<emphasis id="43A27A15FFE83950E46F9F352A4EBC07" box="[1098,1126,1309,1330]" italics="true" pageId="21" pageNumber="317">2):</emphasis>
Measurements: DSL 2.9 3.0; DSW 2.62.65; LI 16.016.1; LII 27.033.5; LIII 21.5 21.9; LIV 28.529.95. Leg IV/body length ratio: 10.1. Pedipalp: tibial setation, mesal IiiIiI/IiIiIi, ectal IiIiIi; tarsal setation, mesal iIiiIii, ectal iIiIi. Tarsal segmentation: 89(3), 19(3), 7, and 8.
</paragraph>
<paragraph id="7169A607FFE83950E31A9FFD2D45BFEA" blockId="21.[806,1422,1309,1759]" pageId="21" pageNumber="317">
Notes.
<taxonomicName id="B6D6DD84FFE83950E3A89FFD2A48BCDC" authorityName="Bragagnolo &amp; Hara &amp; Pinto-Da-Rocha" authorityYear="2015" box="[909,1120,1492,1514]" class="Arachnida" family="Gerdesiidae" genus="Gonycranaus" kingdom="Animalia" order="Opiliones" pageId="21" pageNumber="317" phylum="Arthropoda" rank="species" species="pluto">
<emphasis id="43A27A15FFE83950E3A89FFD2A48BCDC" box="[909,1120,1492,1514]" italics="true" pageId="21" pageNumber="317">Gonycranaus pluto</emphasis>
</taxonomicName>
presents elongate appendages and depigmentation of the body and legs. It was only collected from dark areas of just one limestone cave (from the entrance to the deepest regions), despite thorough collecting efforts outside the entrance. These features are typical characteristics of Neotropical obligate cave-dwelling harvestmen (see for example
<bibRefCitation id="1547DBF6FFE83950E3039C832A5ABFF5" author="Perez-Gonzalez A &amp; Kury AB" box="[806,1138,1707,1729]" pageId="21" pageNumber="317" pagination="43 - 50" refId="ref13955" refString="Perez-Gonzalez A, Kury AB. 2002. A new remarkable troglomorphic gonyleptid from Brazil (Arachnida, Opiliones, Laniatores). Revista Iberica de Aracnologia 5: 43 - 50." type="journal article" year="2002">Pérez-Gonzalez &amp; Kury, 2002</bibRefCitation>
;
<bibRefCitation id="1547DBF6FFE83950E45B9C832D49BFEA" author="Hara MR &amp; Pinto-da-Rocha R" pageId="21" pageNumber="317" pagination="50 - 58" refId="ref13363" refString="Hara MR, Pinto-da-Rocha R. 2008. A new species of Brazilian troglobitic harvestman of the genus Iandumoema (Opiliones: Gonyleptidae). Zootaxa 1744: 50 - 58." type="journal article" year="2008">Hara &amp; Pinto-da-Rocha, 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>