278 lines
24 KiB
XML
278 lines
24 KiB
XML
<document ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE008102AFFA6FE70DAFB9C836DBD" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.5:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="5" docTitle="Delopleurus krikkeni Frolov 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="149" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="148" updateTime="1682086239887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Frolov, Andrey V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>49</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>154</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/00222933.2014.909072</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3bad6762-1788-4f94-9cfd-1281b8619c55</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4002597</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4331159" ID-GBIF-Taxon="167368589" ID-Zenodo-Dep="4331159" LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD" httpUri="http://treatment.plazi.org/id/F91EE008102AFFA6FE70DAFB9C836DBD" lastPageId="20" lastPageNumber="149" pageId="19" pageNumber="148">
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<subSubSection box="[412,836,168,192]" pageId="19" pageNumber="148" type="nomenclature">
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<paragraph blockId="19.[412,836,168,224]" box="[412,836,168,192]" pageId="19" pageNumber="148">
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<heading box="[412,836,168,192]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="148" reason="2">
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<taxonomicName ID-CoL="34H59" authority="Frolov" authorityName="Frolov" authorityYear="2014" box="[412,733,168,192]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="148" phylum="Arthropoda" rank="species" species="krikkeni" status="sp. nov.">
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<emphasis bold="true" box="[412,645,168,192]" italics="true" pageId="19" pageNumber="148">Delopleurus krikkeni</emphasis>
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Frolov
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</taxonomicName>
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,
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<taxonomicNameLabel box="[746,836,169,192]" pageId="19" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[539,709,200,224]" pageId="19" pageNumber="148" type="description">
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<paragraph blockId="19.[412,836,168,224]" box="[539,709,200,224]" pageId="19" pageNumber="148">
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(
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<figureCitation box="[548,700,200,224]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6A–D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="148" type="diagnosis">
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<paragraph blockId="19.[144,1104,273,463]" box="[144,252,273,296]" pageId="19" pageNumber="148">
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<heading box="[144,252,273,296]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
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<emphasis box="[144,252,273,296]" italics="true" pageId="19" pageNumber="148">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="19.[144,1104,273,463]" pageId="19" pageNumber="148">
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This species can be easily separated from other
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<taxonomicName authorityName="Erichson" authorityYear="1847" box="[735,866,312,336]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="148" phylum="Arthropoda" rank="genus">
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<emphasis box="[735,866,312,336]" italics="true" pageId="19" pageNumber="148">Delopleurus</emphasis>
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</taxonomicName>
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species by sparsely punctate to impunctate disc and base of pronotum. Its pygidium is similar to that of
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<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[177,301,376,400]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="148" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
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<emphasis box="[177,301,376,400]" italics="true" pageId="19" pageNumber="148">D. fossatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[311,402,376,399]" pageId="19" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
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but differs in having convex basal border distinctly widened medially (
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<figureCitation box="[255,368,407,431]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6B</figureCitation>
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). From
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<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[461,583,408,432]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="148" phylum="Arthropoda" rank="species" species="fossatus" status="sp. nov.">
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<emphasis box="[461,583,408,432]" italics="true" pageId="19" pageNumber="148">D. fossatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[590,678,408,431]" pageId="19" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
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it also differs in having larger eyes (in dorsal view).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="148" type="description">
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<paragraph blockId="19.[144,1104,544,1053]" box="[144,271,544,567]" pageId="19" pageNumber="148">
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<heading box="[144,271,544,567]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
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<emphasis box="[144,271,544,567]" italics="true" pageId="19" pageNumber="148">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="19.[144,1104,544,1053]" box="[144,1099,583,607]" pageId="19" pageNumber="148">
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<emphasis box="[144,323,583,607]" italics="true" pageId="19" pageNumber="148">
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<typeStatus box="[144,249,583,607]" pageId="19" pageNumber="148">Holotype</typeStatus>
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, male.
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</emphasis>
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Body strongly convex, black, glabrous, length
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<quantity box="[863,949,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="148" unit="mm" value="4.6">4.6 mm</quantity>
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(
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<figureCitation box="[966,1083,583,607]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6A</figureCitation>
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).
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</paragraph>
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<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">Clypeus quadridentate. Two medial teeth acute, lateral ones right-angled. Clypeal surface coarsely punctate in anterior part and laterally, frons densely punctate (punctures separated by one to two puncture diameters).</paragraph>
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<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">Pronotum about two times wider than long. Anterior and lateral margins distinctly bordered, base feebly bordered laterally. Disc and base of pronotum impunctate, sides relatively densely punctate.</paragraph>
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<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">Elytra trapezoidal, as wide as long, shiny. Stria relatively deep, with punctures 1.5–2 times larger than striae width. Elytral intervals convex, with minute punctation.</paragraph>
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<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">Anterior tibiae with three outer teeth, without a small acute tooth between first outer tooth and apical spur. Anterior tibial spur bifurcated apically.</paragraph>
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<paragraph blockId="19.[144,1104,544,1053]" pageId="19" pageNumber="148">
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Pygidium with relatively strongly convex disc separate by deep circular fossa from border. The apical border wider medially and somewhat tapering laterally; apical border somewhat tuberculate in the middle (
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<figureCitation box="[652,766,997,1021]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6B</figureCitation>
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).
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</paragraph>
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<paragraph blockId="19.[144,1104,544,1053]" box="[186,862,1029,1053]" pageId="19" pageNumber="148">
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Parameres in lateral view with small processes (
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<figureCitation box="[730,846,1029,1053]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6C</figureCitation>
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).
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</paragraph>
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<paragraph blockId="19.[144,1103,1109,1165]" pageId="19" pageNumber="148">
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<emphasis box="[144,319,1109,1133]" italics="true" pageId="19" pageNumber="148">
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<typeStatus box="[144,248,1110,1133]" pageId="19" pageNumber="148">Paratype</typeStatus>
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, male.
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</emphasis>
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Differs from the
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<typeStatus box="[514,612,1109,1133]" pageId="19" pageNumber="148">holotype</typeStatus>
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in being slightly larger (length
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<quantity box="[961,1047,1109,1133]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="19" pageNumber="148" unit="mm" value="4.8">4.8 mm</quantity>
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) and having only anteromedial part of pronotal disc impunctate.
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</paragraph>
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<paragraph blockId="19.[144,346,1221,1245]" box="[144,346,1221,1245]" pageId="19" pageNumber="148">
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<heading box="[144,346,1221,1245]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
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<emphasis box="[144,224,1221,1245]" italics="true" pageId="19" pageNumber="148">Female</emphasis>
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unknown.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="148" type="distribution">
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<paragraph blockId="19.[144,1104,1325,1548]" box="[144,276,1325,1349]" pageId="19" pageNumber="148">
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||
<heading box="[144,276,1325,1349]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
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||
<emphasis box="[144,276,1325,1349]" italics="true" pageId="19" pageNumber="148">Distribution</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="19.[144,1104,1325,1548]" pageId="19" pageNumber="148">
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<collectingCounty box="[144,188,1364,1388]" pageId="19" pageNumber="148">The</collectingCounty>
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locality where the
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<typeStatus box="[428,526,1364,1388]" pageId="19" pageNumber="148">holotype</typeStatus>
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was collected is uncertain (Adrian Davis, pers. comm.).
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<collectingMunicipality box="[246,270,1397,1420]" pageId="19" pageNumber="148">In</collectingMunicipality>
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1896, ‘
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<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FE8ADF279F926A98" box="[358,468,1396,1420]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="Samburu'" pageId="19" pageNumber="148" stateProvince="Turkana">Samburu’</location>
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might refer to a large area to the northwest of
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<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FBEDDF269A096A98" box="[1025,1103,1397,1420]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="Mount" pageId="19" pageNumber="148" stateProvince="Turkana">Mount</location>
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<collectingCountry box="[144,218,1429,1453]" name="Kenya" pageId="19" pageNumber="148">Kenya</collectingCountry>
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and to the south of
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<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FE2CDFC79FBF6AB8" box="[448,505,1428,1452]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="Lake" pageId="19" pageNumber="148" stateProvince="Turkana">Lake</location>
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<collectingRegion box="[513,614,1428,1452]" country="Kenya" name="Turkana" pageId="19" pageNumber="148">Turkana</collectingRegion>
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, that was occupied by the
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<collectingRegion box="[912,1015,1428,1452]" country="Kenya" name="Samburu" pageId="19" pageNumber="148">Samburu</collectingRegion>
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people.
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<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FF7CDFE79EFF6ADF" box="[144,185,1460,1483]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="But" pageId="19" pageNumber="148" stateProvince="Turkana">But</location>
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it might also be the railway station (
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<geoCoordinate box="[596,676,1460,1484]" degrees="3" direction="south" minutes="46" orientation="latitude" pageId="19" pageNumber="148" precision="925" value="-3.7666667">
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3°46
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<emphasis bold="true" box="[647,653,1460,1484]" italics="true" pageId="19" pageNumber="148">ʹ</emphasis>
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S
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</geoCoordinate>
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,
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<geoCoordinate box="[688,787,1460,1484]" pageId="19" pageNumber="148">
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39°16
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<emphasis bold="true" box="[752,762,1460,1484]" italics="true" pageId="19" pageNumber="148">ʺ</emphasis>
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E
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</geoCoordinate>
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) on the railroad Mombasa–Nairobi. This latter locality is some
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<quantity box="[562,647,1492,1516]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="148" unit="km" value="100.0">100 km</quantity>
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east-northeast of the locality where the
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||
<typeStatus box="[144,243,1524,1548]" pageId="19" pageNumber="148">paratype</typeStatus>
|
||
was collected,
|
||
<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102AFFA1FE4DDFA79CFB691F" box="[417,701,1524,1548]" country="Kenya" county="The" latitude="-3.7666667" municipality="In" name="Mkomazi Game Reserve" pageId="19" pageNumber="148" stateProvince="Turkana">Mkomazi Game Reserve</location>
|
||
(
|
||
<figureCitation box="[719,837,1524,1548]" captionStart="Figure 6" captionStartId="8.[212,279,738,760]" captionTargetBox="[214,1170,168,697]" captionTargetId="figure@8.[213,1170,168,706]" captionTargetPageId="8" captionText="Figure 6. Delopleurus krikkeni sp. nov., holotype. (A) habitus; (B) pygidium; (C) aedeagus in lateral view and internal sac; (D) locality map." figureDoi="http://doi.org/10.5281/zenodo.4002609" httpUri="https://zenodo.org/record/4002609/files/figure.png" pageId="19" pageNumber="148">Figure 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[144,828,1628,1691]" pageId="19" pageNumber="148" type="etymology">
|
||
<paragraph blockId="19.[144,828,1628,1691]" box="[144,265,1628,1652]" pageId="19" pageNumber="148">
|
||
<heading box="[144,265,1628,1652]" fontSize="10" level="3" pageId="19" pageNumber="148" reason="5">
|
||
<emphasis box="[144,265,1628,1652]" italics="true" pageId="19" pageNumber="148">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="19.[144,828,1628,1691]" box="[144,828,1667,1691]" pageId="19" pageNumber="148">The species is named after Jan Krikken (Naturalis, Leiden).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="20" pageNumber="149" type="materials_examined">
|
||
<paragraph blockId="20.[212,1172,168,328]" box="[212,368,168,192]" pageId="20" pageNumber="149">
|
||
<heading box="[212,368,168,192]" fontSize="10" level="3" pageId="20" pageNumber="149" reason="5">
|
||
<emphasis box="[212,368,168,192]" italics="true" pageId="20" pageNumber="149">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="20.[212,1172,168,328]" pageId="20" pageNumber="149">
|
||
<materialsCitation ID-GBIF-Occurrence="2849122352" collectingDateMax="1836-11-20" collectingDateMin="1896-10-31" collectionCode="BMNH" collectorName="B. E. Africa & C. S. Bretton" country="Kenya" location="Samburu" pageId="20" pageNumber="149" specimenCount="1" stateProvince="Samburu" typeStatus="holotype">
|
||
<typeStatus box="[212,321,208,232]" pageId="20" pageNumber="149">Holotype</typeStatus>
|
||
,
|
||
<specimenCount box="[333,387,208,232]" pageId="20" pageNumber="149" type="male">male</specimenCount>
|
||
with labels ‘
|
||
<collectingRegion box="[533,636,208,232]" country="Kenya" name="Samburu" pageId="20" pageNumber="149">Samburu</collectingRegion>
|
||
[
|
||
<collectingCountry box="[651,755,208,232]" name="Kenya" pageId="20" pageNumber="149">KENYA</collectingCountry>
|
||
]
|
||
<date box="[770,934,208,232]" pageId="20" pageNumber="149">30.X to 20.XI.</date>
|
||
[18]96’, ‘
|
||
<collectorName box="[1034,1171,208,232]" pageId="20" pageNumber="149">B. E. Africa</collectorName>
|
||
<collectorName box="[212,362,240,264]" pageId="20" pageNumber="149">C. S. Bretton</collectorName>
|
||
98-12’, ‘diff. from the others Krikken 1975’ (
|
||
<collectionCode box="[882,974,240,264]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="20" pageNumber="149">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="2849122306" collectingDate="1995-04-07" collectingDateMax="1995-04-08" collectingDateMin="1995-04-07" collectionCode="BMNH" collectorName="Mkomazi GR & J. G. Davies" country="Tanzania" location="Ibaya Camp" pageId="20" pageNumber="149" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus box="[994,1097,241,265]" pageId="20" pageNumber="149">Paratype</typeStatus>
|
||
, male:
|
||
<collectingCountry box="[212,367,272,296]" name="Tanzania" pageId="20" pageNumber="149">TANZANIA</collectingCountry>
|
||
:
|
||
<collectorName box="[383,551,272,296]" pageId="20" pageNumber="149">Mkomazi GR</collectorName>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:F91EE008102AFFA6FE70DAFB9C836DBD:746807C5102DFFA6FDD5DB439C896E3C" box="[569,719,272,296]" country="Tanzania" name="Ibaya Camp" pageId="20" pageNumber="149">Ibaya Camp</location>
|
||
, unburnt grassland,
|
||
<collectingMethod box="[967,1107,272,296]" pageId="20" pageNumber="149">pitfall traps</collectingMethod>
|
||
,
|
||
<date pageId="20" pageNumber="149" value="1995-04-07" valueMax="1995-04-08" valueMin="1995-04-07">
|
||
<collectingDate pageId="20" pageNumber="149" value="1995-04-07" valueMax="1995-04-08" valueMin="1995-04-07">7–8. IV.1995</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[317,451,304,328]" pageId="20" pageNumber="149">J.G. Davies</collectorName>
|
||
leg. (
|
||
<collectionCode box="[516,609,304,328]" pageId="20" pageNumber="149">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="20" pageNumber="149" type="discussion">
|
||
<paragraph blockId="20.[212,1172,394,681]" box="[212,310,394,418]" pageId="20" pageNumber="149">
|
||
<heading box="[212,310,394,418]" fontSize="10" level="3" pageId="20" pageNumber="149" reason="5">
|
||
<emphasis box="[212,310,394,418]" italics="true" pageId="20" pageNumber="149">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="20.[212,1172,394,681]" pageId="20" pageNumber="149">
|
||
Two new species are described in this work from the Eastern Arc Region,
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov" authorityYear="2014" box="[1046,1172,434,458]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="krikkeni" status="sp. nov.">
|
||
<emphasis box="[1046,1066,435,458]" italics="true" pageId="20" pageNumber="149">D</emphasis>
|
||
.
|
||
<emphasis box="[1081,1172,434,458]" italics="true" pageId="20" pageNumber="149">krikkeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[212,302,467,490]" pageId="20" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[363,494,467,490]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
|
||
<emphasis box="[363,383,467,490]" italics="true" pageId="20" pageNumber="149">D</emphasis>
|
||
.
|
||
<emphasis box="[400,494,467,490]" italics="true" pageId="20" pageNumber="149">naviauxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[503,594,467,490]" pageId="20" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
|
||
The former species is known from two males and the latter from two females only. I believe that these specimens are not conspecific because the male specimens lack a characteristic small tooth apicad of the first major outer tooth on the anterior tibae, which is found in the
|
||
<taxonomicName authorityName="Frolov et Cambefort" authorityYear="2014" box="[866,998,562,586]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="naviauxi">
|
||
<emphasis box="[866,886,563,586]" italics="true" pageId="20" pageNumber="149">D</emphasis>
|
||
.
|
||
<emphasis box="[904,998,562,585]" italics="true" pageId="20" pageNumber="149">naviauxi</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus box="[1008,1110,561,585]" pageId="20" pageNumber="149">holotype</typeStatus>
|
||
. Sexdependent modifications of the legs are common in the
|
||
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[835,983,593,617]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="153" phylum="Arthropoda" rank="subFamily" subFamily="Scarabaeinae">Scarabaeinae</taxonomicName>
|
||
but found in the males only. I think that this additional tibial tooth is sex-independent and should be present in the males of
|
||
<taxonomicName authority="Frolov, 2014" authorityName="Frolov et Cambefort" authorityYear="2014" box="[480,610,658,681]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="149" phylum="Arthropoda" rank="species" species="naviauxi" status="sp. nov.">
|
||
<emphasis box="[480,500,658,681]" italics="true" pageId="20" pageNumber="149">D</emphasis>
|
||
.
|
||
<emphasis box="[516,610,658,681]" italics="true" pageId="20" pageNumber="149">naviauxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[618,709,658,681]" pageId="20" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |