280 lines
34 KiB
XML
280 lines
34 KiB
XML
<document id="FDA13193FFD24608F42D27C6C7EA3A57" ID-CLB-Dataset="54359" ID-DOI="10.11646/zootaxa.5274.1.1" ID-GBIF-Dataset="95f4a28d-f058-4cb0-9ad3-bf8652c3ed04" ID-ISSN="1175-5326" ID-Zenodo-Dep="7888156" ID-ZooBank="F60E9DF7-6E56-449E-B6D8-4069D4F9D1D0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1682928899127" checkinUser="plazi" docAuthor="Seifert, Bernhard" docDate="2023" docId="F8463F1440619C7FFF1961E6FF70AFB4" docLanguage="en" docName="zootaxa.5274.1.1.pdf" docOrigin="Zootaxa 5274 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5274.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cardiocondyla elegans var. ulianini Emery 1889" docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="047F476C407E9C5FFF8E6709FF89A80D" masterDocTitle="A revision of the Palaearctic species of the ant genus Cardiocondyla Emery 1869 (Hymenoptera: Formicidae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="32" updateTime="1699493006202" updateUser="plazi">
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<mods:title id="28F27260B781F9CA8815963A02941E8B">A revision of the Palaearctic species of the ant genus Cardiocondyla Emery 1869 (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="3A1952E95D2266A5BE648CC4B89FD765">Seifert, Bernhard</mods:namePart>
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<mods:title id="71B5F2374F45CD9A1ADA8A7C722D2553">Zootaxa</mods:title>
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<mods:part id="F2FD96EE8D53E4AA40E7CB74A9C417ED">
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<mods:date id="FAD1D053D24B32AF6679FBF2247248DF">2023</mods:date>
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<mods:number id="D04F755FD0B965EC933A456878EB8A18">2023-04-28</mods:number>
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<mods:number id="FE5456134AF0F941570999B8A20339D7">5274</mods:number>
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<mods:url id="974300E88835062A816D145D9F3B52D1">http://dx.doi.org/10.11646/zootaxa.5274.1.1</mods:url>
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<mods:classification id="68942B6C417F215300CA82615E5AA513">journal article</mods:classification>
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<mods:identifier id="D84B7D89A13D3095428BE60EDEF96646" type="DOI">10.11646/zootaxa.5274.1.1</mods:identifier>
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<mods:identifier id="AECA79DADD6BB34FE15A495A419C995E" type="GBIF-Dataset">95f4a28d-f058-4cb0-9ad3-bf8652c3ed04</mods:identifier>
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<mods:identifier id="0B68604FAA76D1217CFB0375591797BC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="F8463F1440619C7FFF1961E6FF70AFB4" ID-DOI="http://doi.org/10.5281/zenodo.7893244" ID-GBIF-Taxon="209580619" ID-Zenodo-Dep="7893244" LSID="urn:lsid:plazi:treatment:F8463F1440619C7FFF1961E6FF70AFB4" httpUri="http://treatment.plazi.org/id/F8463F1440619C7FFF1961E6FF70AFB4" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<subSubSection id="38F5DD8940619C40FF1961E6FC12AF04" box="[151,923,1774,1802]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="70508E0240619C40FF1961E6FC12AF04" blockId="31.[151,1437,1774,1982]" box="[151,923,1774,1802]" pageId="31" pageNumber="32">
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<heading id="2B18396E40619C40FF1961E6FC12AF04" box="[151,923,1774,1802]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="5">
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<taxonomicName id="B7EFF58140619C40FF1961E6FD33AF04" authority="Emery 1889" authorityName="Emery" authorityYear="1889" box="[151,698,1775,1802]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="variety" species="elegans" variety="ulianini">
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<emphasis id="429B521040619C40FF1961E6FE12AF04" box="[151,411,1775,1801]" italics="true" pageId="31" pageNumber="32">Cardiocondyla elegans</emphasis>
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var.
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<emphasis id="429B521040619C40FE5C61E6FDA0AF04" box="[466,553,1775,1801]" italics="true" pageId="31" pageNumber="32">ulianini</emphasis>
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Emery 1889
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</taxonomicName>
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[
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<typeStatus id="AF5430A040619C40FD4961F9FD73AF07" box="[711,762,1776,1802]" pageId="31" pageNumber="32">type</typeStatus>
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investigation]
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="38F5DD8940619C7FFF19601AFD6EAAA4" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="70508E0240619C40FF19601AFC0CAFB0" blockId="31.[151,1437,1774,1982]" pageId="31" pageNumber="32">
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<materialsCitation id="C087845F40619C40FF19601AFC08AFB0" collectionCode="MCSN" country="Turkmenistan" county="Investigated" location="This" municipality="The" pageId="31" pageNumber="32" specimenCount="1" specimenCount-worker="1" typeStatus="lectotype">
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<location id="7530D8D940619C40FF19601AFF40AF20" LSID="urn:lsid:plazi:treatment:F8463F1440619C7FFF1961E6FF70AFB4:7530D8D940619C40FF19601AFF40AF20" box="[151,201,1811,1837]" country="Turkmenistan" county="Investigated" municipality="The" name="This" pageId="31" pageNumber="32">This</location>
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taxon has been described from material of the
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<location id="7530D8D940619C40FD59601AFC5AAF20" LSID="urn:lsid:plazi:treatment:F8463F1440619C7FFF1961E6FF70AFB4:7530D8D940619C40FD59601AFC5AAF20" box="[727,979,1811,1837]" country="Turkmenistan" county="Investigated" municipality="The" name="Fedchenko Expedition" pageId="31" pageNumber="32">Fedchenko Expedition</location>
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to “Turkestan”.
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<collectingMunicipality id="9034147840619C40FB02601AFB31AF20" box="[1164,1208,1811,1837]" pageId="31" pageNumber="32">The</collectingMunicipality>
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geographic position of the type locality is unknown and should be somewhere in
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<collectingCountry id="08F8CE9240619C40FCC7603EFC6EAF5C" box="[841,999,1847,1873]" name="Turkmenistan" pageId="31" pageNumber="32">Turkmenistan</collectingCountry>
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,
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<collectingCountry id="08F8CE9240619C40FC7C603EFBF9AF5C" box="[1010,1136,1847,1873]" name="Uzbekistan" pageId="31" pageNumber="32">Uzbekistan</collectingCountry>
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or
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<collectingCountry id="08F8CE9240619C40FB19603EFA80AF5C" box="[1175,1289,1847,1873]" name="Tajikistan" pageId="31" pageNumber="32">Tajikistan</collectingCountry>
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.
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<collectingCounty id="9931F68E40619C40FA9D603EFA15AF5C" box="[1299,1436,1847,1873]" pageId="31" pageNumber="32">Investigated</collectingCounty>
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were
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<typeStatus id="AF5430A040619C40FF596052FEB7AF78" box="[215,318,1883,1909]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
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and
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<typeStatus id="AF5430A040619C40FEF76052FD99AF78" box="[377,528,1883,1909]" pageId="31" pageNumber="32" type="paralectotype">paralectotype</typeStatus>
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worker from
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<collectionCode id="16FE16C740619C40FD3C6052FC88AF78" box="[690,769,1883,1909]" country="Italy" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale, Verona" pageId="31" pageNumber="32">MCSN</collectionCode>
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Genova, labelled “Turkestan Fedtschenko, mus. Moscou” and “
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<taxonomicName id="B7EFF58140619C40FF416076FD19AF94" authority="Em." authorityName="Emery" authorityYear="1889" box="[207,656,1919,1946]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="variety" species="elegans" variety="ulianini">Cardiocondyla elegans var. ulianini Em.</taxonomicName>
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” and
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<specimenCount id="66E9458B40619C40FD406089FCC2AF94" box="[718,843,1919,1946]" pageId="31" pageNumber="32" type="worker">one worker</specimenCount>
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from MHN Genève, designated by Forel as “Cotype” and labelled “
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<taxonomicName id="B7EFF58140619C40FEBE60AAFCF1AFB0" authority="Em., Turkestan" authorityName="Em., Turkestan" box="[304,888,1955,1982]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="variety" species="elegans" variety="ulianini">Cardiocondyla elegans var. ulianini Em., Turkestan</taxonomicName>
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”
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="70508E02405E9C7FFF19679FFBB7A8F4" blockId="32.[151,1437,150,1977]" pageId="32" pageNumber="33">
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<emphasis id="429B5210405E9C7FFF19679FFE2BA8BD" bold="true" box="[151,418,150,176]" pageId="32" pageNumber="33">All material examined</emphasis>
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.
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<materialsCitation id="C087845F405E9C7FFE3E679EFB22A8DB" country="Afghanistan" location="For" municipality="This" pageId="32" pageNumber="33" specimenCount="44" specimenCount-worker="44">
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Numeric phenotypical data were available in 23 samples with
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<specimenCount id="66E9458B405E9C7FFBE0679EFB66A8BC" box="[1134,1263,151,177]" pageId="32" pageNumber="33" type="worker">44 workers</specimenCount>
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.
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<location id="7530D8D9405E9C7FFB74679EFAA8A8BC" LSID="urn:lsid:plazi:treatment:F8463F1440619C7FFF1961E6FF70AFB4:7530D8D9405E9C7FFB74679EFAA8A8BC" box="[1274,1313,151,177]" country="Afghanistan" municipality="This" name="For" pageId="32" pageNumber="33">For</location>
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details see supplementary information SI1, SI2.
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<collectingMunicipality id="90341478405E9C7FFDB367B2FDE6A8D8" box="[573,623,187,213]" pageId="32" pageNumber="33">This</collectingMunicipality>
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material originated from
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<collectingCountry id="08F8CE92405E9C7FFC1D67B2FB95A8D8" box="[915,1052,187,213]" name="Afghanistan" pageId="32" pageNumber="33">Afghanistan</collectingCountry>
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(3 samples)
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</materialsCitation>
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, Caucasus (3),
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<collectingCountry id="08F8CE92405E9C7FFAD767B2FA15A8D8" box="[1369,1436,187,213]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
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(1),
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<materialsCitation id="C087845F405E9C7FFF4867D6FEFBA8F4" box="[198,370,223,249]" country="Kazakhstan" isTailEnumeration="true" location="For" municipality="This" pageId="32" pageNumber="33" specimenCount="1">
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<collectingCountry id="08F8CE92405E9C7FFF4867D6FEC0A8F4" box="[198,329,223,249]" name="Kazakhstan" pageId="32" pageNumber="33">Kazakhstan</collectingCountry>
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(8)
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</materialsCitation>
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, Kyrghistan (3), allegedly
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<collectingCountry id="08F8CE92405E9C7FFD1367D6FCA6A8F4" box="[669,815,223,249]" name="Saudi Arabia" pageId="32" pageNumber="33">Saudi Arabia</collectingCountry>
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(1) and the
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<collectingCountry id="08F8CE92405E9C7FFC3867D6FB99A8F4" box="[950,1040,223,249]" name="Ukraine" pageId="32" pageNumber="33">Ukraine</collectingCountry>
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(3).
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</paragraph>
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<paragraph id="70508E02405E9C7FFF19660BFD6EAAA4" blockId="32.[151,1437,150,1977]" pageId="32" pageNumber="33">
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<emphasis id="429B5210405E9C7FFF19660BFEF8A910" bold="true" box="[151,369,258,285]" pageId="32" pageNumber="33">Geographic range</emphasis>
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. Continuously distributed across the steppe and semidesert zone from the S
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<collectingCountry id="08F8CE92405E9C7FFB63660AFACEA910" box="[1261,1351,259,285]" name="Ukraine" pageId="32" pageNumber="33">Ukraine</collectingCountry>
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(
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<geoCoordinate id="15DBE8C5405E9C7FFAD4660AFA1DA910" box="[1370,1428,259,285]" degrees="32" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="55555" value="32.0">32°E</geoCoordinate>
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) eastwards to N Xinjang (
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<geoCoordinate id="15DBE8C5405E9C7FFE23662EFE60A94C" box="[429,489,295,321]" degrees="88" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="55555" value="88.0">88°E</geoCoordinate>
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). The northern range border is demarcated by
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<geoCoordinate id="15DBE8C5405E9C7FFC62662FFBB5A94C" box="[1004,1084,294,321]" degrees="46.5" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5555" value="46.5">46.5°N</geoCoordinate>
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in the
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<collectingCountry id="08F8CE92405E9C7FFB06662EFB6BA94C" box="[1160,1250,295,321]" name="Ukraine" pageId="32" pageNumber="33">Ukraine</collectingCountry>
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and
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<geoCoordinate id="15DBE8C5405E9C7FFA96662EFAE1A94C" box="[1304,1384,295,321]" degrees="46.7" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5555" value="46.7">46.7°N</geoCoordinate>
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in E
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<collectingCountry id="08F8CE92405E9C7FFF196642FE93A968" box="[151,282,331,357]" name="Kazakhstan" pageId="32" pageNumber="33">Kazakhstan</collectingCountry>
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and the southern border by
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<geoCoordinate id="15DBE8C5405E9C7FFDC16642FD02A968" box="[591,651,331,357]" degrees="34" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="55555" value="34.0">34°N</geoCoordinate>
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in
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<collectingCountry id="08F8CE92405E9C7FFD256642FCB1A968" box="[683,824,331,357]" name="Afghanistan" pageId="32" pageNumber="33">Afghanistan</collectingCountry>
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. The altitudinal range extends from
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<quantity id="B71723E7405E9C7FFB496643FB76A968" box="[1223,1279,330,357]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="32" pageNumber="33" unit="m" value="25.0">25 m</quantity>
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below zero in the Caspian region to
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<quantity id="B71723E7405E9C7FFE036666FE6BA984" box="[397,482,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="32" pageNumber="33" unit="m" value="1800.0">1800 m</quantity>
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in
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<collectingCountry id="08F8CE92405E9C7FFD8B6666FD1BA984" box="[517,658,367,393]" name="Afghanistan" pageId="32" pageNumber="33">Afghanistan</collectingCountry>
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. A morphologically typical sample, centrally placed in the
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<taxonomicName id="B7EFF581405E9C7FFAAA6666FA15A984" authorityName="Emery" authorityYear="1889" box="[1316,1436,367,393]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ulianini">
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<emphasis id="429B5210405E9C7FFAAA6666FA15A984" box="[1316,1436,367,393]" italics="true" pageId="32" pageNumber="33">C. ulianini</emphasis>
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</taxonomicName>
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cluster by PCA considering the 16 standard characters, allegedly from Hofuf /
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<collectingCountry id="08F8CE92405E9C7FFBBC669AFB4EA9A0" box="[1074,1223,403,429]" name="Saudi Arabia" pageId="32" pageNumber="33">Saudi Arabia</collectingCountry>
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(
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<geoCoordinate id="15DBE8C5405E9C7FFB54669BFAA6A9A0" box="[1242,1327,402,429]" degrees="25.4" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="5555" value="25.4">25.4°N</geoCoordinate>
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,
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<geoCoordinate id="15DBE8C5405E9C7FFAB1669AFA06A9A3" box="[1343,1423,403,430]" degrees="49.6" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="5555" value="49.6">49.6°E</geoCoordinate>
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), appears extremely isolated. The most likely explanation is confusion of labels after side-by-side stereomicroscopic comparison of unmounted specimens belonging to different samples as it was a frequent investigation practice by Cedric Collingwood—the donor of several samples in the SMN Görlitz collection.This putting of specimens in wrong tubes repeatedly lead to dramatic zoogeographic confusion: a male of the exclusively Himalayan
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<taxonomicName id="B7EFF581405E9C7FFB7A652AFA14AA31" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[1268,1437,547,572]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="crinitus">
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<emphasis id="429B5210405E9C7FFB7A652AFA14AA31" box="[1268,1437,547,572]" italics="true" pageId="32" pageNumber="33">Lasius crinitus</emphasis>
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||
</taxonomicName>
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was “transferred” by Collingwood to
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<collectingCountry id="08F8CE92405E9C7FFDC0654EFD18AA6C" box="[590,657,583,609]" name="Spain" pageId="32" pageNumber="33">Spain</collectingCountry>
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, a worker of the exclusively Australasian
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<taxonomicName id="B7EFF581405E9C7FFBF0654EFEA8AA88" authority="Seifert 2003" authorityName="Seifert" authorityYear="2003" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="paranuda">
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<emphasis id="429B5210405E9C7FFBF0654EFA15AA6C" box="[1150,1436,583,609]" italics="true" pageId="32" pageNumber="33">Cardiocondyla paranuda</emphasis>
|
||
<bibRefCitation id="147EF3F3405E9C7FFF196562FEA8AA88" author="Seifert, B." box="[151,289,619,645]" pageId="32" pageNumber="33" pagination="203 - 338" refId="ref43774" refString="Seifert, B. (2003) The ant genus Cardiocondyla (Insecta: Hymenoptera: Formicidae) - a taxonomic revision of the C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi, and C. minutior species groups. Annalen des Naturhistorischen Museums, Wien, Serie B, 104, 203 - 338." type="journal article" year="2003">Seifert 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
to the Sahara desert (see
|
||
<bibRefCitation id="147EF3F3405E9C7FFDCC6562FC84AA88" author="Seifert, B. & Okita, I. & Heinze, J." box="[578,781,619,645]" pageId="32" pageNumber="33" pagination="324 - 356" refId="ref44079" refString="Seifert, B., Okita, I. & Heinze, J. (2017) A taxonomic revision of the Cardiocondyla nuda group (Hymenoptera: Formicidae). Zootaxa, 4290 (2), 324 - 356. https: // doi. org / 10.11646 / zootaxa. 4290.2.4" type="journal article" year="2017">
|
||
Seifert
|
||
<emphasis id="429B5210405E9C7FFD1D6562FD45AA88" box="[659,716,619,645]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
) or a worker of
|
||
<taxonomicName id="B7EFF581405E9C7FFC4A6562FB56AA88" authorityName="Forel" authorityYear="1892" box="[964,1247,619,645]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bulgarica">
|
||
<emphasis id="429B5210405E9C7FFC4A6562FB56AA88" box="[964,1247,619,645]" italics="true" pageId="32" pageNumber="33">Cardiocondyla bulgarica</emphasis>
|
||
</taxonomicName>
|
||
, restricted to the Balkans and
|
||
<collectingCountry id="08F8CE92405E9C7FFEA86586FE20AAA4" box="[294,425,655,681]" name="Turkey" pageId="32" pageNumber="33">Asia Minor</collectingCountry>
|
||
, to inner
|
||
<collectingCountry id="08F8CE92405E9C7FFD9E6586FDEAAAA4" box="[528,611,655,681]" name="Tunisia" pageId="32" pageNumber="33">Tunisia</collectingCountry>
|
||
(see p. 37).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="38F5DD89405E9C7FFF1965BBFB8DAE78" pageId="32" pageNumber="33" type="diagnosis">
|
||
<paragraph id="70508E02405E9C7FFF1965BBFCCAADE8" blockId="32.[151,1437,150,1977]" pageId="32" pageNumber="33">
|
||
<emphasis id="429B5210405E9C7FFF1965BBFE83AAC1" bold="true" box="[151,266,690,716]" pageId="32" pageNumber="33">Diagnosis</emphasis>
|
||
: --Worker (
|
||
<tableCitation id="3D6DBBB9405E9C7FFE1E65BAFE53AAC0" box="[400,474,691,717]" captionStart="TABLE 2" captionStartId="39.[152,244,151,175]" captionTargetPageId="39" captionText="TABLE 2. Measurements of workers of the Cardiocondyla ulianini group. Explanation of data arrangement as in Tab. 1" httpUri="http://table.plazi.org/id/2490DE8A40599C78FF16679EFA15A8BD" pageId="32" pageNumber="33" tableUuid="2490DE8A40599C78FF16679EFA15A8BD">Tab. 2</tableCitation>
|
||
,
|
||
<figureCitation id="E8D49287405E9C7FFE6965BAFDE6AAC0" box="[487,623,690,717]" captionStart="FIGURES 22–25" captionStartId="13.[151,264,1034,1058]" captionTargetBox="[241,1347,181,1010]" captionTargetId="figure-15@13.[241,1347,181,1010]" captionTargetPageId="13" captionText="FIGURES 22–25. Cardiocondyla ulianini; Fig. 22: head in dorsal view; Fig. 23: lateral view; Fig. 24: dorsal view; Fig. 25: head surface between inner eye margin and paramedian vertex. Kyrgistan: 42.44.41N, 75.49.50E, 2000.07.30-91." figureDoi="http://doi.org/10.5281/zenodo.7888212" httpUri="https://zenodo.org/record/7888212/files/figure.png" pageId="32" pageNumber="33">Figs. 22–25</figureCitation>
|
||
, key). Medium-sized, CS 522 µm. Head moderately elongated, CL/CW 1.149. Postocular distance rather low, PoOc/CL 0.411. Scape moderately long, SL/CS 0.810. Eye rather large, EYE/CS 0.251. Occipital margin suggestively concave. Frons rather broad (FRS/CS 0.257), frontal carinae slightly converging immediately caudal of FRS level (FL/FR 1.060). Dorsal profile of promesonotum and of propodeum convex with a well-developed metanotal depression (MGr/CS 3.44 %). Propodeal spines rather short, more triangular than spiny, SP/CS 0.120), their axis in profile deviating by about 45° from longitudinal axis of mesosoma, their bases approached (SPBA/CS 0.236). Petiole rather narrower and much higher than wide (PeW/CS 0.290, PeH/ CS 0.336), its node in dorsal aspect slightly longer than wide; in profile with a moderately long peduncle that is about 1.25fold as long as high and with a moderately steep anterior slope of the node (about 60° relative to ventral profile). Postpetiole twice as wide as high (PpW/CS 0.564, PpH/CS 0.279), in dorsal view heard-shaped, with a concave anterior margin and convex sides; postpetiolar sternite with the anteromedian portion significantly more bulging than its anteroparamedian portion; in lateral view this anteromedian bulge forms a small, obtusely-angled, rounded corner and changes into the helcium with a distinct angle. Clypeus smooth and shiny, with only suggested of microrugulae. Vertex in overall impression shiny, completely without carinulae or microrugae, very weak and fine rugulae are restricted to frontal laminae and frontolateral head (genae). Vertex with numerous foveolae the diameter of which is smaller than the width of the brilliantly shining interspaces (dFOV 13.8), the interspaces with scattered very fine stickman-like fragments of a microreticulum; outer margin of foveolae shallow, not very clearly demarcated and internal foveolar surface often with longitudinal carinulae (
|
||
<figureCitation id="E8D49287405E9C7FFB8E621EFBDEAD3C" box="[1024,1111,1302,1329]" captionStart="FIGURES 22–25" captionStartId="13.[151,264,1034,1058]" captionTargetBox="[241,1347,181,1010]" captionTargetId="figure-15@13.[241,1347,181,1010]" captionTargetPageId="13" captionText="FIGURES 22–25. Cardiocondyla ulianini; Fig. 22: head in dorsal view; Fig. 23: lateral view; Fig. 24: dorsal view; Fig. 25: head surface between inner eye margin and paramedian vertex. Kyrgistan: 42.44.41N, 75.49.50E, 2000.07.30-91." figureDoi="http://doi.org/10.5281/zenodo.7888212" httpUri="https://zenodo.org/record/7888212/files/figure.png" pageId="32" pageNumber="33">Fig. 25</figureCitation>
|
||
). Dorsal promesonotum and propodeum glabrous. Promesonotum with scattered and shallow foveolae of 10–12 µm diameter. Lateral meso- and metapleurae rather shiny and delicately longitudinally striate-reticulate, contrasting the glabrous parts of mesonotum and pronotum. Dorsum of waist glabrous except for a very delicate microreticulum. First gaster tergite glabrous. Pubescence on whole body rather short and dilute, PLG/CS 5.89 %, sqPDG 4.89. Color of head, mesosoma, and gaster varying from pale yellowish brown to blackish brown.
|
||
</paragraph>
|
||
<paragraph id="70508E02405E9C7FFF1962E7FB8DAE78" blockId="32.[151,1437,150,1977]" pageId="32" pageNumber="33">
|
||
<emphasis id="429B5210405E9C7FFF1962E7FD2CAE04" bold="true" box="[151,677,1518,1545]" pageId="32" pageNumber="33">Taxonomic comments and clustering results</emphasis>
|
||
.
|
||
<taxonomicName id="B7EFF581405E9C7FFD3D62E6FC30AE04" authorityName="Emery" authorityYear="1889" box="[691,953,1519,1545]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ulianini">
|
||
<emphasis id="429B5210405E9C7FFD3D62E6FC30AE04" box="[691,953,1519,1545]" italics="true" pageId="32" pageNumber="33">Cardiocondyla ulianini</emphasis>
|
||
</taxonomicName>
|
||
is a characteristic species that differs from members of the four species of the
|
||
<taxonomicName id="B7EFF581405E9C7FFDA2611AFD2DAE20" baseAuthorityName="Seifert" baseAuthorityYear="2018" box="[556,676,1555,1581]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="429B5210405E9C7FFDA2611AFD2DAE20" box="[556,676,1555,1581]" italics="true" pageId="32" pageNumber="33">C. elegans</emphasis>
|
||
</taxonomicName>
|
||
group as well as from
|
||
<emphasis id="429B5210405E9C7FFC20611AFB58AE21" box="[942,1233,1555,1581]" italics="true" pageId="32" pageNumber="33">
|
||
<taxonomicName id="B7EFF581405E9C7FFC20611AFBC9AE20" authorityName="Forel" authorityYear="1892" box="[942,1088,1555,1581]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bulgarica">C. bulgarica</taxonomicName>
|
||
,
|
||
<taxonomicName id="B7EFF581405E9C7FFBC2611AFB58AE21" authorityName="Seifert" authorityYear="2003" box="[1100,1233,1555,1581]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="persiana">C. persiana</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="B7EFF581405E9C7FFA9F611AFA15AE20" authorityName=", Forel" authorityYear="1913" box="[1297,1436,1555,1581]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="sahlbergi">
|
||
<emphasis id="429B5210405E9C7FFA9F611AFA15AE20" box="[1297,1436,1555,1581]" italics="true" pageId="32" pageNumber="33">C. sahlbergi</emphasis>
|
||
</taxonomicName>
|
||
by the structure of vertex foveolae and several morphometric differences (
|
||
<tableCitation id="3D6DBBB9405E9C7FFC7D613EFBC1AE5C" box="[1011,1096,1591,1617]" captionStart="TABLE 1" captionStartId="31.[152,244,151,175]" captionTargetPageId="31" captionText="TABLE 1. Measurements of workers of the Cardiocondyla elegans group. Data are given in the sequence arithmetic mean ± standard deviation [minimum, maximum]. For characters with fewer data than indicated in the head of the columns, the number of measurements is given after the square brackets." httpUri="http://table.plazi.org/id/2490DE8A40619C40FF16679EFC98A8FA" pageId="32" pageNumber="33" tableUuid="2490DE8A40619C40FF16679EFC98A8FA">Tabs. 1</tableCitation>
|
||
and
|
||
<tableCitation id="3D6DBBB9405E9C7FFB0A613EFB1CAE5C" box="[1156,1173,1591,1617]" captionStart="TABLE 2" captionStartId="39.[152,244,151,175]" captionTargetPageId="39" captionText="TABLE 2. Measurements of workers of the Cardiocondyla ulianini group. Explanation of data arrangement as in Tab. 1" httpUri="http://table.plazi.org/id/2490DE8A40599C78FF16679EFA15A8BD" pageId="32" pageNumber="33" tableUuid="2490DE8A40599C78FF16679EFA15A8BD">2</tableCitation>
|
||
). For differences to
|
||
<taxonomicName id="B7EFF581405E9C7FFA0D613EFF7EAE78" authorityName="Seifert" authorityYear="2003" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="littoralis">
|
||
<emphasis id="429B5210405E9C7FFA0D613EFF7EAE78" italics="true" pageId="32" pageNumber="33">C. littoralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B7EFF581405E9C7FFE886152FE1AAE79" authority="Seifert, 2023" authorityName="Seifert" authorityYear="2023" box="[262,403,1627,1653]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="caspiense" status="sp. nov.">
|
||
<emphasis id="429B5210405E9C7FFE886152FE1AAE79" box="[262,403,1627,1653]" italics="true" pageId="32" pageNumber="33">C. caspiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="429B5210405E9C7FFE146152FE50AE78" bold="true" box="[410,473,1627,1653]" pageId="32" pageNumber="33">
|
||
<taxonomicNameLabel id="59A8EF6B405E9C7FFE146152FE50AE78" box="[410,473,1627,1653]" pageId="32" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="B7EFF581405E9C7FFE696152FD09AE78" authorityName="Seifert" authorityYear="2003" box="[487,640,1627,1653]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="gallilaeica">
|
||
<emphasis id="429B5210405E9C7FFE696152FD09AE78" box="[487,640,1627,1653]" italics="true" pageId="32" pageNumber="33">C. gallilaeica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B7EFF581405E9C7FFD386152FCB0AE78" authorityName="Seifert" authorityYear="2003" box="[694,825,1627,1653]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="israelica">
|
||
<emphasis id="429B5210405E9C7FFD386152FCB0AE78" box="[694,825,1627,1653]" italics="true" pageId="32" pageNumber="33">C. israelica</emphasis>
|
||
</taxonomicName>
|
||
see these species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="38F5DD89405E9C7FFF196177FF70AFB4" pageId="32" pageNumber="33" type="biology_ecology">
|
||
<paragraph id="70508E02405E9C7FFF196177FF70AFB4" blockId="32.[151,1437,150,1977]" pageId="32" pageNumber="33">
|
||
<emphasis id="429B5210405E9C7FFF196177FF7EAE95" bold="true" box="[151,247,1662,1688]" pageId="32" pageNumber="33">Biology.</emphasis>
|
||
Habitats observed in E Dagestan, Kyrghistan and
|
||
<collectingCountry id="08F8CE92405E9C7FFCA96176FC23AE94" box="[807,938,1663,1689]" name="Kazakhstan" pageId="32" pageNumber="33">Kazakhstan</collectingCountry>
|
||
were in three cases sandy river terraces with very sparse dry steppe vegetation and once a flat, salty and periodically inundated loess soil at a lake margin. Nests had one or two simple entrance holes. Two excavated nests contained two and four fully sclerotized ergatoid males with shear-shaped mandibles. The presence of more than
|
||
<specimenCount id="66E9458B405E9C7FFC9461E2FC0BAF08" box="[794,898,1771,1797]" pageId="32" pageNumber="33" type="adult">one adult</specimenCount>
|
||
male in these nests and the fact that only one of these
|
||
<specimenCount id="66E9458B405E9C7FFF596006FEB6AF24" box="[215,319,1807,1833]" pageId="32" pageNumber="33" type="male">six males</specimenCount>
|
||
showed an injury (an amputated tibia) indicates that males of
|
||
<taxonomicName id="B7EFF581405E9C7FFC7E6006FB7DAF24" authorityName="Emery" authorityYear="1889" box="[1008,1268,1807,1833]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ulianini">
|
||
<emphasis id="429B5210405E9C7FFC7E6006FB7DAF24" box="[1008,1268,1807,1833]" italics="true" pageId="32" pageNumber="33">Cardiocondyla ulianini</emphasis>
|
||
</taxonomicName>
|
||
tend to coexist and avoid fighting at least in adult stage. Alate gynes were observed in SE
|
||
<collectingCountry id="08F8CE92405E9C7FFC54603AFBD4AF40" box="[986,1117,1843,1869]" name="Kazakhstan" pageId="32" pageNumber="33">Kazakhstan</collectingCountry>
|
||
<date id="0451A8C2405E9C7FFBEA603AFA83AF40" box="[1124,1290,1843,1869]" pageId="32" pageNumber="33" value="2001-08-07">7 August 2001</date>
|
||
.
|
||
<bibRefCitation id="147EF3F3405E9C7FFA9B603AFEF7AF7C" author="Marikovsky, P. I. & Yakushkin, V. T" pageId="32" pageNumber="33" pagination="57 - 61" refId="ref43370" refString="Marikovsky, P. I. & Yakushkin, V. T, (1974) Muravyei Cardiocondyla ulianini Em., 1889 i sistematicheskoye polosheniye " parasiticheskovo muravya Xenometra ". Isvestiya Akademii Nauk Kazakhskoy SSR, seria biologicheskaya, 3, 57 - 61." type="journal article" year="1974">Marikovsky & Yakushkin (1974)</bibRefCitation>
|
||
described the biology of supposedly
|
||
<taxonomicName id="B7EFF581405E9C7FFCA6605EFBA7AF7C" authorityName="Emery" authorityYear="1889" box="[808,1070,1879,1905]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ulianini">
|
||
<emphasis id="429B5210405E9C7FFCA6605EFBA7AF7C" box="[808,1070,1879,1905]" italics="true" pageId="32" pageNumber="33">Cardiocondyla ulianini</emphasis>
|
||
</taxonomicName>
|
||
from SE
|
||
<collectingCountry id="08F8CE92405E9C7FFB10605EFAADAF7C" box="[1182,1316,1879,1905]" name="Kazakhstan" pageId="32" pageNumber="33">Kazakhstan</collectingCountry>
|
||
. However, a possible confusion with sympatric
|
||
<taxonomicName id="B7EFF581405E9C7FFDB46072FD34AF98" authorityName="Seifert" authorityYear="2003" box="[570,701,1915,1941]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="littoralis">
|
||
<emphasis id="429B5210405E9C7FFDB46072FD34AF98" box="[570,701,1915,1941]" italics="true" pageId="32" pageNumber="33">C. littoralis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="B7EFF581405E9C7FFD696072FCFFAF99" authority="Seifert, 2023" authorityName="Seifert" authorityYear="2023" box="[743,886,1915,1941]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="caspiense" status="sp. nov.">
|
||
<emphasis id="429B5210405E9C7FFD696072FCFFAF99" box="[743,886,1915,1941]" italics="true" pageId="32" pageNumber="33">C. caspiense</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="429B5210405E9C7FFCF16072FC49AF98" bold="true" box="[895,960,1915,1941]" pageId="32" pageNumber="33">
|
||
<taxonomicNameLabel id="59A8EF6B405E9C7FFCF16072FC49AF98" box="[895,960,1915,1941]" pageId="32" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
makes the interpretation of their findings difficult.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |