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83 lines
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<document id="6F0979C3357118088B860ED0C6DB4A70" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629618962334" checkinUser="felipe" docAuthor="Fend, En. V." docDate="2009" docId="F44687BC04376266FF70EFADFD26FC56" docLanguage="en" docName="zt02077p030.pdf" docOrigin="Zootaxa 2077" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Kincaidiana Altman 1936" docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="087FFFC404356264FFE7EE0FFF82FFAA" masterDocTitle="An evaluation of the genus Kincaidiana Altman, 1936, with the designation of Altmanella n. gen. (Annelida, Clitellata, Lumbriculidae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="3" updateTime="1699084233931" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3555EC36694456C71BA57C48C19AB243">An evaluation of the genus Kincaidiana Altman, 1936, with the designation of Altmanella n. gen. (Annelida, Clitellata, Lumbriculidae)</mods:title>
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<mods:namePart id="4E43E7B123F38C677B1610DA2F808FB2">Fend, En. V.</mods:namePart>
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<mods:date id="7D146F603046701B5D8CCCEE668C6830">2009</mods:date>
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<mods:number id="DABDB3667F77912ACF0C88CBECB386B3">2077</mods:number>
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<treatment id="F44687BC04376266FF70EFADFD26FC56" ID-DOI="http://doi.org/10.5281/zenodo.5333277" ID-Zenodo-Dep="5333277" LSID="urn:lsid:plazi:treatment:F44687BC04376266FF70EFADFD26FC56" httpUri="http://treatment.plazi.org/id/F44687BC04376266FF70EFADFD26FC56" lastPageNumber="3" pageId="2" pageNumber="3">
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<subSubSection id="34F5652104376266FF70EFADFE5BFE16" box="[151,473,418,444]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="7C5036AA04376266FF70EFADFE5BFE16" blockId="2.[151,473,418,444]" box="[151,473,418,444]" pageId="2" pageNumber="3">
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<heading id="271881C604376266FF70EFADFE5BFE16" bold="true" box="[151,473,418,444]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<taxonomicName id="BBEF4D2904376266FF70EFADFE5BFE16" ID-CoL="84PF6" authority="Altman, 1936" authorityName="Altman" authorityYear="1936" box="[151,473,418,444]" class="Clitellata" family="Lumbriculidae" genus="Kincaidiana" kingdom="Animalia" order="Lumbriculida" pageId="2" pageNumber="3" phylum="Annelida" rank="genus">
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<emphasis id="4E9BEAB804376266FF70EFADFE5BFE16" bold="true" box="[151,473,418,444]" pageId="2" pageNumber="3">
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<emphasis id="4E9BEAB804376266FF70EFADFEA9FE16" bold="true" box="[151,299,418,444]" italics="true" pageId="2" pageNumber="3">Kincaidiana</emphasis>
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<bibRefCitation id="187E4B5B04376266FED5EFADFE5BFE16" author="Altman" box="[306,473,418,444]" firstAuthor="Altman" pageId="2" pageNumber="3" pagination="1 - 137" refId="ref15043" refString="Altman L. C. (1936) Oligochaeta of Washington. University of Washington Publications in Biology, 4, 1 - 137." type="journal article" year="1936">Altman, 1936</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="34F5652104376266FF70EFE4FD26FC56" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="7C5036AA04376266FF70EFE4FD26FC56" blockId="2.[151,1436,491,1020]" pageId="2" pageNumber="3">
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<emphasis id="4E9BEAB804376266FF70EFE4FE1BFDAF" bold="true" box="[151,409,491,517]" pageId="2" pageNumber="3">Emended diagnosis</emphasis>
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(modified from
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<bibRefCitation id="187E4B5B04376266FD89EFE3FC93FDAC" author="Cook" box="[622,785,492,518]" firstAuthor="Cook" pageId="2" pageNumber="3" pagination="200 - 285" refId="ref15259" refString="Cook, D. G. (1971) Family Lumbriculidae. In: Brinkhurst R. O. & Jamieson B. G. M. Aquatic Oligochaeta of the World. Oliver and Boyd, Edinburgh, pp. 200 - 285." type="book chapter" year="1971">Cook [1971]</bibRefCitation>
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). Medium-sized to large worms with long, pseudosegmented proboscis (
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<figureCitation id="E4D42A2F04376266FE40EC1DFD91FD86" box="[423,531,530,556]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Figs. 1A</figureCitation>
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,
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<figureCitation id="E4D42A2F04376266FDC5EC1DFDEDFD86" box="[546,623,530,556]" captionStart="FIGURE 2" captionStartId="4.[151,257,1679,1703]" captionTargetBox="[154,1432,380,1658]" captionTargetId="figure-260@4.[151,1436,376,1664]" captionTargetPageId="4" captionText="FIGURE 2. Kincaidiana hexatheca. A–C, F and I–J from the Hoh River, Washington; D–E and G–H from the type series, Washington. A–B, whole worms, with A. freidris specimens (Owyhee River, Oregon) for size comparison. C, sagittal section in an anterior segment, showing transverse (circular) muscle layer of body wall. D, head of a dissected worm. E, spermatheca from a dissected worm. F, spermatheca from a sectioned worm, showing sperm lined up along ectal end of ampulla. G–H, male pores and atria of a dissected and a sectioned worm. I–J, sections of proximal and distal parts of vas deferens; proximal to the atrium, the vas has a thick muscle coat. Scale bars: A 10 mm, B 5 mm, C 50 µm, E–H 200 µm, I–J 50 µm." pageId="2" pageNumber="3">2A–B</figureCitation>
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). Chaetae bifid and enlarged in a few anterior segments (
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<figureCitation id="E4D42A2F04376266FACBEC1DFA08FD86" box="[1324,1418,530,556]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Fig. 1B</figureCitation>
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); simple-pointed posteriorly. No distinct pharyngeal pad (
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<figureCitation id="E4D42A2F04376266FCD0EC36FC1AFDF9" box="[823,920,569,595]" captionStart="FIGURE 2" captionStartId="4.[151,257,1679,1703]" captionTargetBox="[154,1432,380,1658]" captionTargetId="figure-260@4.[151,1436,376,1664]" captionTargetPageId="4" captionText="FIGURE 2. Kincaidiana hexatheca. A–C, F and I–J from the Hoh River, Washington; D–E and G–H from the type series, Washington. A–B, whole worms, with A. freidris specimens (Owyhee River, Oregon) for size comparison. C, sagittal section in an anterior segment, showing transverse (circular) muscle layer of body wall. D, head of a dissected worm. E, spermatheca from a dissected worm. F, spermatheca from a sectioned worm, showing sperm lined up along ectal end of ampulla. G–H, male pores and atria of a dissected and a sectioned worm. I–J, sections of proximal and distal parts of vas deferens; proximal to the atrium, the vas has a thick muscle coat. Scale bars: A 10 mm, B 5 mm, C 50 µm, E–H 200 µm, I–J 50 µm." pageId="2" pageNumber="3">Fig. 2D</figureCitation>
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). One or two pairs of lateral blood vessels, with many blind branches, in posterior segments (
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<figureCitation id="E4D42A2F04376266FD0AEC6FFCC9FDD0" box="[749,843,608,634]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Fig. 1C</figureCitation>
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). Nephridia begin on 11/12; postseptal expansion narrow and directed dorsally (
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<figureCitation id="E4D42A2F04376266FE1BEC89FDD8FD0A" box="[508,602,646,672]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Fig. 1D</figureCitation>
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). One pair of testes in IX, one pair of ovaries in X. Male pores on IX, near 9/10, usually paired and slightly inside chaetal lines (
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<figureCitation id="E4D42A2F04376266FCA0ECA2FC27FD6D" box="[839,933,685,711]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Fig. 1A</figureCitation>
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). Pendant penes within shallow sacs when retracted; everted sacs form low porophores (
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<figureCitation id="E4D42A2F04376266FD49ECDBFCBCFD44" box="[686,830,724,750]" captionStart="FIGURE 2" captionStartId="4.[151,257,1679,1703]" captionTargetBox="[154,1432,380,1658]" captionTargetId="figure-260@4.[151,1436,376,1664]" captionTargetPageId="4" captionText="FIGURE 2. Kincaidiana hexatheca. A–C, F and I–J from the Hoh River, Washington; D–E and G–H from the type series, Washington. A–B, whole worms, with A. freidris specimens (Owyhee River, Oregon) for size comparison. C, sagittal section in an anterior segment, showing transverse (circular) muscle layer of body wall. D, head of a dissected worm. E, spermatheca from a dissected worm. F, spermatheca from a sectioned worm, showing sperm lined up along ectal end of ampulla. G–H, male pores and atria of a dissected and a sectioned worm. I–J, sections of proximal and distal parts of vas deferens; proximal to the atrium, the vas has a thick muscle coat. Scale bars: A 10 mm, B 5 mm, C 50 µm, E–H 200 µm, I–J 50 µm." pageId="2" pageNumber="3">Figs. 2G–H</figureCitation>
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). Atria club-shaped to elongate-cylindrical, ental ¾ or more loosely covered with multicellular, pyriform prostate glands (
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<figureCitation id="E4D42A2F04376266FC21ECF5FBA8FCBE" box="[966,1066,762,788]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Figs. 1E</figureCitation>
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,
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<figureCitation id="E4D42A2F04376266FBD0ECF5FBDDFCBE" box="[1079,1119,762,788]" captionStart="FIGURE 2" captionStartId="4.[151,257,1679,1703]" captionTargetBox="[154,1432,380,1658]" captionTargetId="figure-260@4.[151,1436,376,1664]" captionTargetPageId="4" captionText="FIGURE 2. Kincaidiana hexatheca. A–C, F and I–J from the Hoh River, Washington; D–E and G–H from the type series, Washington. A–B, whole worms, with A. freidris specimens (Owyhee River, Oregon) for size comparison. C, sagittal section in an anterior segment, showing transverse (circular) muscle layer of body wall. D, head of a dissected worm. E, spermatheca from a dissected worm. F, spermatheca from a sectioned worm, showing sperm lined up along ectal end of ampulla. G–H, male pores and atria of a dissected and a sectioned worm. I–J, sections of proximal and distal parts of vas deferens; proximal to the atrium, the vas has a thick muscle coat. Scale bars: A 10 mm, B 5 mm, C 50 µm, E–H 200 µm, I–J 50 µm." pageId="2" pageNumber="3">2G</figureCitation>
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). One pair of male funnels, usually extending back one or more segments in sperm sac (
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<figureCitation id="E4D42A2F04376266FC69ED2EFC6DFC91" box="[910,1007,801,827]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Fig. 1E</figureCitation>
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). Vasa deferentia do not penetrate posterior septum (9/10); they join atria near-medially; muscle coat of atria extends for a short distance along vasa deferentia (
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<figureCitation id="E4D42A2F04376266FE87ED61FE51FC22" box="[352,467,878,904]" captionStart="FIGURE 2" captionStartId="4.[151,257,1679,1703]" captionTargetBox="[154,1432,380,1658]" captionTargetId="figure-260@4.[151,1436,376,1664]" captionTargetPageId="4" captionText="FIGURE 2. Kincaidiana hexatheca. A–C, F and I–J from the Hoh River, Washington; D–E and G–H from the type series, Washington. A–B, whole worms, with A. freidris specimens (Owyhee River, Oregon) for size comparison. C, sagittal section in an anterior segment, showing transverse (circular) muscle layer of body wall. D, head of a dissected worm. E, spermatheca from a dissected worm. F, spermatheca from a sectioned worm, showing sperm lined up along ectal end of ampulla. G–H, male pores and atria of a dissected and a sectioned worm. I–J, sections of proximal and distal parts of vas deferens; proximal to the atrium, the vas has a thick muscle coat. Scale bars: A 10 mm, B 5 mm, C 50 µm, E–H 200 µm, I–J 50 µm." pageId="2" pageNumber="3">Fig. 2I–J</figureCitation>
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). Female pores on chaetal lines at posterior septum of first post-atrial segment. Three pairs of spermathecae, in IX, X, and XI; pores slightly inside ventral chaetal lines, usually in line with or slightly anterior to chaetae (
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<figureCitation id="E4D42A2F04376266FE19EDB3FDD9FC7C" box="[510,603,956,982]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Fig. 1A</figureCitation>
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). Spermathecae with an elongate, thick duct and an elongate-tubular to elongate-pyriform ampulla (
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<figureCitation id="E4D42A2F04376266FE05EDEDFDC4FC56" box="[482,582,994,1020]" captionStart="FIGURE 1" captionStartId="3.[151,256,1641,1665]" captionTargetBox="[157,1430,187,1610]" captionTargetId="figure-182@3.[151,1436,187,1613]" captionTargetPageId="3" captionText="FIGURE 1. Kincaidiana hexatheca. A–D from the Hoh River, Washington; E from Inglenook Fen, California. A, an unmounted specimen in ventral view, showing male and spermathecal pores. B, representative chaetae: from top to bottom, a ventral chaeta in II, a dorsal pair in VIII, a ventral in VI, a ventral in X, and a ventral from mid-body. C, lateral blood vessels in two posterior segments. D, nephridium. E, reproductive organs. Scale bars: A 2mm, B 200 µm, C 1 mm, D 200 µm, E 300 µm." pageId="2" pageNumber="3">Figs. 1E</figureCitation>
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,
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<figureCitation id="E4D42A2F04376266FDB4EDEDFD1AFC56" box="[595,664,994,1020]" captionStart="FIGURE 2" captionStartId="4.[151,257,1679,1703]" captionTargetBox="[154,1432,380,1658]" captionTargetId="figure-260@4.[151,1436,376,1664]" captionTargetPageId="4" captionText="FIGURE 2. Kincaidiana hexatheca. A–C, F and I–J from the Hoh River, Washington; D–E and G–H from the type series, Washington. A–B, whole worms, with A. freidris specimens (Owyhee River, Oregon) for size comparison. C, sagittal section in an anterior segment, showing transverse (circular) muscle layer of body wall. D, head of a dissected worm. E, spermatheca from a dissected worm. F, spermatheca from a sectioned worm, showing sperm lined up along ectal end of ampulla. G–H, male pores and atria of a dissected and a sectioned worm. I–J, sections of proximal and distal parts of vas deferens; proximal to the atrium, the vas has a thick muscle coat. Scale bars: A 10 mm, B 5 mm, C 50 µm, E–H 200 µm, I–J 50 µm." pageId="2" pageNumber="3">2E–F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |