treatments-xml/data/F4/2D/21/F42D2116FFF7910EFF6E220227EB81AD.xml

<document id="4E3DA1A634A5524CABDF9A38A88A7B22" ID-CLB-Dataset="24083" ID-DOI="10.11646/zootaxa.4732.1.3" ID-GBIF-Dataset="a1d73d3e-e33e-4a19-aab8-55876eb1408b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3662541" ID-ZooBank="F4F9A5CF-63B3-4557-BB12-E3914664AC8F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581432325729" checkinUser="plazi" docAuthor="Trýzna, Miloš &amp; Baňař, Petr" docDate="2020" docId="F42D2116FFF7910EFF6E220227EB81AD" docLanguage="en" docName="zootaxa.4732.1.3.pdf" docOrigin="Zootaxa 4732 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holophloeus loebli Tryzna &amp; Banar 2020, sp. nov." docType="treatment" docUuid="E111F11B-6AB0-4C8F-8E11-698119E4BA93" docUuidSource="ZooBank" docVersion="5" lastPageNumber="87" masterDocId="0814596EFFF49106FFF927012276860C" masterDocTitle="A new species of Holophloeus Jordan (Coleoptera: Anthribidae) from eastern Madagascar with ecological notes on it and H. tuberosus (Fairmaire, 1897)" masterLastPageNumber="98" masterPageNumber="79" pageNumber="82" updateTime="1698781830297" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="73AE0A5468F3D958E9C7A5A18EBD07A0">A new species of Holophloeus Jordan (Coleoptera: Anthribidae) from eastern Madagascar with ecological notes on it and H. tuberosus (Fairmaire, 1897)</mods:title>
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<mods:namePart id="0D2EA92B50E22F9F25EA1A483BECD48A">Trýzna, Miloš</mods:namePart>
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<mods:namePart id="ACCD5B114BC6871CB7C53A20D0DB54F2">Baňař, Petr</mods:namePart>
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<mods:date id="C6A5F7B9C0C34CF5DC6E484E1FAA24F2">2020</mods:date>
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<treatment id="F42D2116FFF7910EFF6E220227EB81AD" ID-DOI="http://doi.org/10.5281/zenodo.3663731" ID-GBIF-Taxon="162362126" ID-Zenodo-Dep="3663731" LSID="urn:lsid:zoobank.org:act:E111F11B-6AB0-4C8F-8E11-698119E4BA93" httpUri="http://treatment.plazi.org/id/F42D2116FFF7910EFF6E220227EB81AD" lastPageId="8" lastPageNumber="87" pageId="3" pageNumber="82">
<subSubSection id="349EC38BFFF79105FF6E220220C08312" box="[151,694,1283,1310]" pageId="3" pageNumber="82" type="nomenclature">
<paragraph id="7C3B9000FFF79105FF6E220220C08312" blockId="3.[151,694,1283,1346]" box="[151,694,1283,1310]" pageId="3" pageNumber="82">
<heading id="2773276CFFF79105FF6E220220C08312" bold="true" box="[151,694,1283,1310]" fontSize="11" level="1" pageId="3" pageNumber="82" reason="1">
<emphasis id="4EF04C12FFF79105FF6E220220C08312" bold="true" box="[151,694,1283,1310]" pageId="3" pageNumber="82">
<taxonomicName id="BB84EB83FFF79105FF6E220220278312" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" authorityYear="2020" box="[151,593,1283,1310]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="82" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="4EF04C12FFF79105FF6E2202230A8312" bold="true" box="[151,380,1283,1310]" italics="true" pageId="3" pageNumber="82">Holophloeus loebli</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="55C3F169FFF79105FDA0220520C08312" box="[601,694,1284,1310]" pageId="3" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="349EC38BFFF79105FF6E222920C0834E" box="[151,694,1320,1346]" pageId="3" pageNumber="82" type="description">
<paragraph id="7C3B9000FFF79105FF6E222920C0834E" blockId="3.[151,694,1283,1346]" box="[151,694,1320,1346]" pageId="3" pageNumber="82">
(
<figureCitation id="E4BF8C85FFF79105FF6622292299834E" box="[159,239,1320,1346]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="3" pageNumber="82">Figs. 4</figureCitation>
, 
<figureCitation id="E4BF8C85FFF79105FF0222292343834E" box="[251,309,1320,1346]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="3" pageNumber="82">6–10</figureCitation>
, 
<figureCitation id="E4BF8C85FFF79105FEB8222923FF834E" box="[321,393,1320,1346]" captionStart="FIGURES 16–19. 16, 18" captionStartId="6.[151,264,1600,1625]" captionTargetBox="[151,1436,237,1575]" captionTargetId="figure@6.[151,1436,237,1575]" captionTargetPageId="6" captionText="FIGURES 16–19. 16, 18, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 17, 19, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 16–17, pronotum, dorsal view; 18–19, elytra, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="3" pageNumber="82">16, 18</figureCitation>
, 
<figureCitation id="E4BF8C85FFF79105FE6C222923A8834E" box="[405,478,1320,1346]" captionStart="FIGURES 20–23. 20, 22" captionStartId="7.[151,264,1545,1570]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure@7.[151,1436,181,1520]" captionTargetPageId="7" captionText="FIGURES 20–23. 20, 22, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 21, 23, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 20–21, head, ventral view; 22–23, prosternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662552" httpUri="https://zenodo.org/record/3662552/files/figure.png" pageId="3" pageNumber="82">20, 22</figureCitation>
, 
<figureCitation id="E4BF8C85FFF79105FE102229202D834E" box="[489,603,1320,1346]" captionStart="FIGURES 24–29. 24, 26, 28" captionStartId="9.[151,264,1945,1970]" captionTargetBox="[178,1402,181,1920]" captionTargetId="figure@9.[178,1409,181,1920]" captionTargetPageId="9" captionText="FIGURES 24–29. 24, 26, 28, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 25, 27, 29, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 24–25, mesosternum and metasternum, dorsal view; 26–27, abdominal ventrites I–V, dorsal view; 28–29, abdominal ventrite V, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662555" httpUri="https://zenodo.org/record/3662555/files/figure.png" pageId="3" pageNumber="82">24, 26, 28</figureCitation>
, 
<figureCitation id="E4BF8C85FFF79105FD9E222920D9834E" box="[615,687,1320,1346]" captionStart-0="FIGURES 34–35" captionStart-1="FIGURES 36–37" captionStart-2="FIGURES 38–39" captionStart-3="FIGURES 40–41" captionStart-4="FIGURE 42" captionStartId-0="12.[151,264,1958,1983]" captionStartId-1="13.[151,264,1956,1981]" captionStartId-2="14.[151,264,1963,1988]" captionStartId-3="15.[151,264,1928,1953]" captionStartId-4="16.[151,260,1535,1562]" captionTargetBox-0="[164,1423,181,1932]" captionTargetBox-1="[164,1423,181,1930]" captionTargetBox-2="[164,1423,181,1937]" captionTargetBox-3="[151,1436,181,1903]" captionTargetId-0="figure@12.[164,1423,181,1932]" captionTargetId-1="figure@13.[164,1423,181,1930]" captionTargetId-2="figure@14.[164,1423,181,1937]" captionTargetId-3="figure@15.[151,1436,181,1903]" captionTargetId-4="figure@16.[347,1240,179,1510]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="16" captionText-0="FIGURES 34–35. Collecting site of Holophloeus loebli Trýzna &amp; Baňař sp. nov. in Andasibe National Park, Analamazaotra forest, 2016." captionText-1="FIGURES 36–37. Collecting site of Holophloeus loebli Trýzna &amp; Baňař sp. nov.; 36, Andasibe National Park, Analamazaotra forest, 2016; 37, type locality in Mantadia National Park, Eulophia circuit, 2017." captionText-2="FIGURES 38–39. Type locality of Holophloeus loebli Trýzna &amp; Baňař sp. nov. in Mantadia National Park, Eulophia circuit, 2017; 38, primary forest with single huge torso of lying trunk; 39, detail of cover of bark with specimens." captionText-3="FIGURES 40–41. Type locality of Holophloeus loebli Trýzna &amp; Baňař sp. nov. in Mantadia National Park, Eulophia circuit, 2017, detail with specimen and exit hole." captionText-4="FIGURE 42. Distribution of Holophloeus species in Madagascar." figureDoi-0="http://doi.org/10.5281/zenodo.3662561" figureDoi-1="http://doi.org/10.5281/zenodo.3662563" figureDoi-2="http://doi.org/10.5281/zenodo.3662566" figureDoi-3="http://doi.org/10.5281/zenodo.3662568" figureDoi-4="http://doi.org/10.5281/zenodo.3690719" httpUri-0="https://zenodo.org/record/3662561/files/figure.png" httpUri-1="https://zenodo.org/record/3662563/files/figure.png" httpUri-2="https://zenodo.org/record/3662566/files/figure.png" httpUri-3="https://zenodo.org/record/3662568/files/figure.png" httpUri-4="https://zenodo.org/record/3690719/files/figure.png" pageId="3" pageNumber="82">34–42</figureCitation>
)
</paragraph>
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<paragraph id="7C3B9000FFF79105FF6E227122C283A2" blockId="3.[151,1437,1391,2030]" pageId="3" pageNumber="82">
<materialsCitation id="CCEC9A5DFFF79105FF6E227122C283A2" ID-GBIF-Occurrence="2572159694" country="Madagascar" elevation="993" latitude="-18.80161" location="Mantadia National Park" longLatPrecision="1" longitude="48.429142" municipality="East" pageId="3" pageNumber="82" specimenCount="993" stateProvince="Toamasina" typeStatus="holotype">
<emphasis id="4EF04C12FFF79105FF6E227123438386" bold="true" box="[151,309,1391,1418]" pageId="3" pageNumber="82">
<typeStatus id="A33F2EA2FFF79105FF6E227122A48386" box="[151,210,1392,1418]" pageId="3" pageNumber="82">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="9C5F0A7AFFF79105FEC22271231D8386" box="[315,363,1392,1418]" pageId="3" pageNumber="82">East</collectingMunicipality>
<collectingCountry id="0493D090FFF79105FE882271238C8386" box="[369,506,1392,1418]" name="Madagascar" pageId="3" pageNumber="82">Madagascar</collectingCountry>
, 
<collectingRegion id="BE405EE2FFF79105FDFD2271209E8386" box="[516,744,1392,1418]" country="Madagascar" name="Toamasina" pageId="3" pageNumber="82">Toamasina province</collectingRegion>
, 
<location id="795BC6DBFFF79105FD0A227121898386" LSID="urn:lsid:plazi:treatment:F42D2116FFF7910EFF6E220227EB81AD:795BC6DBFFF79105FD0A227121898386" box="[755,1023,1392,1418]" country="Madagascar" latitude="-18.80161" longLatPrecision="1" longitude="48.429142" municipality="East" name="Mantadia National Park" pageId="3" pageNumber="82" stateProvince="Toamasina">Mantadia National Park</location>
, 
<geoCoordinate id="19B0F6C7FFF79105FBF3227126C58386" box="[1034,1203,1391,1418]" degrees="18" direction="south" minutes="48" orientation="latitude" pageId="3" pageNumber="82" precision="1" seconds="05.8" value="-18.80161">S 18°48´05.8´´</geoCoordinate>
, 
<geoCoordinate id="19B0F6C7FFF79105FB44227127118386" box="[1213,1383,1391,1418]" degrees="48" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="82" precision="1" seconds="44.9" value="48.429142">E 48°25´44.9´´</geoCoordinate>
, 
<specimenCount id="6A825B89FFF79105FA8B227122C283A2" pageId="3" pageNumber="82" type="generic">
<quantity id="BB7C3DE5FFF79105FA8B227122C683A2" metricMagnitude="2" metricUnit="m" metricValue="9.93" pageId="3" pageNumber="82" unit="m" value="993.0">
<elevation id="F7A97733FFF79105FA8B227122C683A2" metricMagnitude="2" metricUnit="m" metricValue="9.93" pageId="3" pageNumber="82" unit="m" value="993.0">993 m</elevation>
</quantity>
.
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</paragraph>
<paragraph id="7C3B9000FFF79105FF3E22B9266983DE" blockId="3.[151,1437,1391,2030]" box="[199,1055,1463,1490]" pageId="3" pageNumber="82">
<emphasis id="4EF04C12FFF79105FF3E22B9230383DE" bold="true" box="[199,373,1463,1490]" pageId="3" pageNumber="82">Type material.</emphasis>
<materialsCitation id="CCEC9A5DFFF79105FE8522B9266983DE" ID-GBIF-Occurrence="2572159688" box="[380,1055,1463,1490]" country="Madagascar" location="TOAMASINA PROVINCE" pageId="3" pageNumber="82" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<emphasis id="4EF04C12FFF79105FE8522B921D183C3" bold="true" box="[380,935,1463,1490]" pageId="3" pageNumber="82">
<typeStatus id="A33F2EA2FFF79105FE8522B9239F83DE" box="[380,489,1464,1490]" pageId="3" pageNumber="82" type="holotype">Holotype</typeStatus>
(male): 
<collectingCountry id="0493D090FFF79105FDB522B6216883DE" box="[588,798,1463,1490]" name="Madagascar" pageId="3" pageNumber="82">MADAGASCAR</collectingCountry>
: 
<collectingRegion id="BE405EE2FFF79105FCD322B921D183C3" box="[810,935,1464,1490]" country="Madagascar" name="Toamasina" pageId="3" pageNumber="82">TOAMASINA</collectingRegion>
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PROVINCE
<emphasis id="4EF04C12FFF79105FBEF22B9266983DE" bold="true" box="[1046,1055,1464,1490]" pageId="3" pageNumber="82">:</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="7C3B9000FFF79102FF3E22DA26DF874F" blockId="3.[151,1437,1391,2030]" lastBlockId="4.[151,1437,152,647]" lastPageId="4" lastPageNumber="83" pageId="3" pageNumber="82">
<emphasis id="4EF04C12FFF79105FF3E22DA229283FA" bold="true" box="[199,228,1499,1526]" pageId="3" pageNumber="82">‘E</emphasis>
<materialsCitation id="CCEC9A5DFFF79105FF0922DA23918032" ID-GBIF-Occurrence="2572159697" collectionCode="BSNPC, MTDC" country="Madagascar" latitude="-18.80161" location="TOAMASINA PROVINCE" longLatPrecision="1" longitude="48.429142" pageId="3" pageNumber="82" specimenCount="88" specimenCount-female="48" specimenCount-male="40" stateProvince="Toamasina" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FF0922DA264483FF" bold="true" box="[240,1074,1499,1526]" pageId="3" pageNumber="82">
<collectingCountry id="0493D090FFF79105FF0922DA23CB83FA" box="[240,445,1499,1526]" name="Madagascar" pageId="3" pageNumber="82">MADAGASCAR</collectingCountry>
, 
<date id="083AB6C0FFF79105FE2822DD238283FA" box="[465,500,1500,1526]" pageId="3" pageNumber="82" value="2016-12-30">30.</date>
XII. / 2016, MANTADIA N. P., 993 M, “EULOPHIA
</emphasis>
CIRCUIT 
<emphasis id="4EF04C12FFF79105FB6A22DA20508016" bold="true" pageId="3" pageNumber="82">
“, / 
<geoCoordinate id="19B0F6C7FFF79105FB3022DA270183FA" box="[1225,1399,1499,1526]" degrees="18" direction="south" minutes="48" orientation="latitude" pageId="3" pageNumber="82" precision="1" seconds="05.8" value="-18.80161">S 18°48´05.8´´</geoCoordinate>
, 
<geoCoordinate id="19B0F6C7FFF79105FA7322DD235F8016" degrees="48" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="82" precision="1" seconds="44.9" value="48.429142">E 48°25´44.9´´</geoCoordinate>
, / MILOŠ TRÝZNA LEG.
</emphasis>
’ [p] (
<collectionCode id="1A9508C5FFF79105FD9E210120B08016" box="[615,710,1536,1562]" pageId="3" pageNumber="82">BSNPC</collectionCode>
). 
<emphasis id="4EF04C12FFF79105FD2022FE21C48016" bold="true" box="[729,946,1535,1562]" pageId="3" pageNumber="82">
<typeStatus id="A33F2EA2FFF79105FD2022FE21488016" box="[729,830,1535,1562]" pageId="3" pageNumber="82" type="allotype">Allotype</typeStatus>
(female):
</emphasis>
the same data as holotype (
<collectionCode id="1A9508C5FFF79105FB17210127338016" box="[1262,1349,1536,1562]" pageId="3" pageNumber="82">MTDC</collectionCode>
). 
<emphasis id="4EF04C12FFF79105FAA0210123918032" bold="true" pageId="3" pageNumber="82">
<typeStatus id="A33F2EA2FFF79105FAA0210122A38032" pageId="3" pageNumber="82" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="6A825B89FFF79105FF1C212223398032" box="[229,335,1571,1598]" pageId="3" pageNumber="82" type="male">40 males</specimenCount>
, 
<specimenCount id="6A825B89FFF79105FEA5212523A98032" box="[348,479,1571,1598]" pageId="3" pageNumber="82" type="female">48 females</specimenCount>
)
</emphasis>
</materialsCitation>
<emphasis id="4EF04C12FFF79105FE1E2122239A8032" bold="true" box="[487,492,1571,1598]" pageId="3" pageNumber="82">:</emphasis>
<materialsCitation id="CCEC9A5DFFF79105FE0A21252134806E" ID-GBIF-Occurrence="2572159700" collectionCode="MTDC, MMBC, BMNH, CMNC, MNHN, MTD, NMPC, ZSMC" country="Madagascar" location="TOAMASINA PROVINCE" pageId="3" pageNumber="82" specimenCount="43" specimenCount-female="20" specimenCount-male="23" stateProvince="Toamasina" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FE0A212520998032" bold="true" box="[499,751,1571,1598]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FE0A2125202A8032" box="[499,604,1572,1598]" pageId="3" pageNumber="82" type="male">23 males</specimenCount>
, 
<specimenCount id="6A825B89FFF79105FD902125209D8032" box="[617,747,1571,1598]" pageId="3" pageNumber="82" type="female">20 females</specimenCount>
:
</emphasis>
the same data as holotype (
<collectionCode id="1A9508C5FFF79105FBDF2125260D8032" box="[1062,1147,1572,1598]" pageId="3" pageNumber="82">MTDC</collectionCode>
, 3 pairs in 
<collectionCode id="1A9508C5FFF79105FB0F212527268032" box="[1270,1360,1572,1598]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="82">MMBC</collectionCode>
, 1 pair each in 
<collectionCode id="1A9508C5FFF79105FF1621492330806E" box="[239,326,1608,1634]" pageId="3" pageNumber="82">BMNH</collectionCode>
, 
<collectionCode id="1A9508C5FFF79105FEA8214923D0806E" box="[337,422,1608,1634]" pageId="3" pageNumber="82">CMNC</collectionCode>
, 
<collectionCode id="1A9508C5FFF79105FE4B2149207C806E" box="[434,522,1608,1634]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="82">MNHN</collectionCode>
, 
<collectionCode id="1A9508C5FFF79105FDEF21492021806E" box="[534,599,1608,1634]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="3" pageNumber="82">MTD</collectionCode>
, 
<collectionCode id="1A9508C5FFF79105FD9B214920C7806E" box="[610,689,1608,1634]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="82">NMPC</collectionCode>
and 
<collectionCode id="1A9508C5FFF79105FD102149214C806E" box="[745,826,1608,1634]" country="Germany" httpUri="http://grbio.org/cool/a6ms-py1z" name="Zoologische Staatssammlung" pageId="3" pageNumber="82">ZSMC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="CCEC9A5DFFF79105FCB7214923EA808A" ID-GBIF-Occurrence="2572159691" collectingDate="2017-01-17" collectingDateMax="2017-01-22" collectingDateMin="2017-01-17" collectionCode="MTDC" country="Madagascar" location="TOAMASINA PROVINCE" pageId="3" pageNumber="82" specimenCount="29" specimenCount-female="18" specimenCount-male="11" stateProvince="Toamasina" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FCB72149263F806E" bold="true" box="[846,1097,1607,1634]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FCB7214921C3806E" box="[846,949,1608,1634]" pageId="3" pageNumber="82" type="male">11 males</specimenCount>
, 
<specimenCount id="6A825B89FFF79105FC3B21492632806E" box="[962,1092,1607,1634]" pageId="3" pageNumber="82" type="female">18 females</specimenCount>
:
</emphasis>
the same data as holotype but 
<date id="083AB6C0FFF79105FF6E216D2358808A" box="[151,302,1644,1670]" pageId="3" pageNumber="82" value="2017-01-17" valueMax="2017-01-22" valueMin="2017-01-17">
<collectingDate id="187E4F28FFF79105FF6E216D2358808A" box="[151,302,1644,1670]" pageId="3" pageNumber="82" value="2017-01-17" valueMax="2017-01-22" valueMin="2017-01-17">17.-22.i.2017</collectingDate>
</date>
(
<collectionCode id="1A9508C5FFF79105FEC4216D23E2808A" box="[317,404,1644,1670]" pageId="3" pageNumber="82">MTDC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="CCEC9A5DFFF79105FE51216D21A380A6" ID-GBIF-Occurrence="2572159692" collectingDate="2016-02-05" collectionCode="MTDC" collectorName="Milos Tryzna &amp; M. Tryzna" country="Madagascar" county="Indri" elevation="955" latitude="-18.938" location="Andasibe" longLatPrecision="1" longitude="48.419197" municipality="Analamazaotra" pageId="3" pageNumber="82" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FE51216D2057808A" bold="true" box="[424,545,1643,1670]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FE51216D206A808A" box="[424,540,1643,1670]" pageId="3" pageNumber="82" type="female">2 females</specimenCount>
:
</emphasis>
‘E MADAGASCAR, 
<date id="083AB6C0FFF79105FCE6216A21BF808A" box="[799,969,1643,1670]" pageId="3" pageNumber="82" value="2007-02-05" valueMax="2007-02-13" valueMin="2007-02-05">5.-13.ii. / 2007</date>
, 
<location id="795BC6DBFFF79105FC2A216D264B808A" LSID="urn:lsid:plazi:treatment:F42D2116FFF7910EFF6E220227EB81AD:795BC6DBFFF79105FC2A216D264B808A" box="[979,1085,1644,1670]" country="Madagascar" county="Indri" latitude="-18.938" longLatPrecision="1" longitude="48.419197" municipality="Analamazaotra" name="Andasibe" pageId="3" pageNumber="82">Andasibe</location>
N.P., 
<quantity id="BB7C3DE5FFF79105FB78216A26BD808A" box="[1153,1227,1643,1670]" metricMagnitude="2" metricUnit="m" metricValue="9.55" pageId="3" pageNumber="82" unit="m" value="955.0">
<elevation id="F7A97733FFF79105FB78216A26BD808A" box="[1153,1227,1643,1670]" metricMagnitude="2" metricUnit="m" metricValue="9.55" pageId="3" pageNumber="82" unit="m" value="955.0">955 m</elevation>
</quantity>
, / “Analamazaotra forest”, / 
<geoCoordinate id="19B0F6C7FFF79105FF06219123D180A6" box="[255,423,1679,1706]" degrees="18" direction="south" minutes="56" orientation="latitude" pageId="3" pageNumber="82" precision="1" seconds="45.0" value="-18.945833">S 18°56´45.0´´</geoCoordinate>
, 
<geoCoordinate id="19B0F6C7FFF79105FE4B2191202D80A6" box="[434,603,1679,1706]" degrees="48" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="82" precision="1" seconds="08.0" value="48.41889">E 48°25´08.0´´</geoCoordinate>
, / 
<collectorName id="D171F5D6FFF79105FD8C2191217C80A6" box="[629,778,1680,1706]" pageId="3" pageNumber="82">Miloš Trýzna</collectorName>
leg.’ [p] (
<collectionCode id="1A9508C5FFF79105FC8F219121BB80A6" box="[886,973,1680,1706]" pageId="3" pageNumber="82">MTDC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="CCEC9A5DFFF79105FC262191232880FE" ID-GBIF-Occurrence="2572159701" collectingDate="2016-02-05" collectionCode="MTDC" collectorName="Milos Tryzna &amp; M. Tryzna" country="Madagascar" county="Indri" elevation="955" latitude="-18.938" location="Andasibe" longLatPrecision="1" longitude="48.419197" municipality="Analamazaotra" pageId="3" pageNumber="82" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FC262191264080A6" bold="true" box="[991,1078,1680,1706]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FC262191264480A6" box="[991,1074,1680,1706]" pageId="3" pageNumber="82" type="male">1 male</specimenCount>
:
</emphasis>
‘E MADAGASCAR, 
<date id="083AB6C0FFF79105FACB218E27E180A6" box="[1330,1431,1679,1706]" pageId="3" pageNumber="82" value="2016-02-05">
<collectingDate id="187E4F28FFF79105FACB218E27E180A6" box="[1330,1431,1679,1706]" pageId="3" pageNumber="82" value="2016-02-05">5.ii.2016</collectingDate>
</date>
, / Andasibe N.P., “ 
<collectingMunicipality id="9C5F0A7AFFF79105FE9B21B5207980C2" box="[354,527,1716,1742]" pageId="3" pageNumber="82">Analamazaotra</collectingMunicipality>
/ forest”, circuit 
<collectingCounty id="955AE88CFFF79105FD2E21B5217B80C2" box="[727,781,1716,1742]" pageId="3" pageNumber="82">Indri</collectingCounty>
1, / 
<quantity id="BB7C3DE5FFF79105FCBF21B521E780C2" box="[838,913,1716,1742]" metricMagnitude="2" metricUnit="m" metricValue="9.79" pageId="3" pageNumber="82" unit="m" value="979.0">
<elevation id="F7A97733FFF79105FCBF21B521E780C2" box="[838,913,1716,1742]" metricMagnitude="2" metricUnit="m" metricValue="9.79" pageId="3" pageNumber="82" unit="m" value="979.0">979 m</elevation>
</quantity>
, 
<geoCoordinate id="19B0F6C7FFF79105FC6621B5263D80C2" box="[927,1099,1715,1742]" degrees="18" direction="south" minutes="56" orientation="latitude" pageId="3" pageNumber="82" precision="1" seconds="16.8" value="-18.938">S 18°56´16.8´´</geoCoordinate>
, / 
<geoCoordinate id="19B0F6C7FFF79105FB9321B5276180C2" box="[1130,1303,1715,1742]" degrees="48" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="82" precision="1" seconds="09.1" value="48.419197">E 48°25´09.1´´</geoCoordinate>
, 
<collectorName id="D171F5D6FFF79105FADC21B527EA80C2" box="[1317,1436,1716,1742]" pageId="3" pageNumber="82">M. Trýzna</collectorName>
leg.’ [p] (
<collectionCode id="1A9508C5FFF79105FF0621D9232080FE" box="[255,342,1752,1778]" pageId="3" pageNumber="82">MTDC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="CCEC9A5DFFF79105FE9021D9263D811A" ID-GBIF-Occurrence="2572159698" collectingDate="2019-01-22" collectingDateMax="2019-01-29" collectingDateMin="2019-01-22" collectionCode="MTDC" collectorName="Andasibe N. P. &amp; M. Tryzna" country="Madagascar" elevation="947" latitude="-18.94461" location="Analamazaotra" longLatPrecision="1" longitude="48.424084" municipality="Aventure" pageId="3" pageNumber="82" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FE9021D923A180FE" bold="true" box="[361,471,1751,1778]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FE9021D923A580FE" box="[361,467,1751,1778]" pageId="3" pageNumber="82" type="female">1 female</specimenCount>
:
</emphasis>
‘MADAGASCAR, 
<quantity id="BB7C3DE5FFF79105FD4421D9217180FE" box="[701,775,1752,1778]" metricMagnitude="2" metricUnit="m" metricValue="9.47" pageId="3" pageNumber="82" unit="m" value="947.0">
<elevation id="F7A97733FFF79105FD4421D9217180FE" box="[701,775,1752,1778]" metricMagnitude="2" metricUnit="m" metricValue="9.47" pageId="3" pageNumber="82" unit="m" value="947.0">947 m</elevation>
</quantity>
, / 
<collectorName id="D171F5D6FFF79105FCD921D921B480FE" box="[800,962,1752,1778]" pageId="3" pageNumber="82">Andasibe N.P.</collectorName>
, 
<date id="083AB6C0FFF79105FC3721D9261180FE" box="[974,1127,1752,1778]" pageId="3" pageNumber="82" value="2019-01-22" valueMax="2019-01-29" valueMin="2019-01-22">
<collectingDate id="187E4F28FFF79105FC3721D9261180FE" box="[974,1127,1752,1778]" pageId="3" pageNumber="82" value="2019-01-22" valueMax="2019-01-29" valueMin="2019-01-22">22.-29.i.2019</collectingDate>
</date>
, / 
<location id="795BC6DBFFF79105FB7921D9275A80FE" LSID="urn:lsid:plazi:treatment:F42D2116FFF7910EFF6E220227EB81AD:795BC6DBFFF79105FB7921D9275A80FE" box="[1152,1324,1752,1778]" country="Madagascar" latitude="-18.94461" longLatPrecision="1" longitude="48.424084" municipality="Aventure" name="Analamazaotra" pageId="3" pageNumber="82">Analamazaotra</location>
, 
<collectingMunicipality id="9C5F0A7AFFF79105FACF21D927EA80FE" box="[1334,1436,1752,1778]" pageId="3" pageNumber="82">Aventure</collectingMunicipality>
circ., / 
<geoCoordinate id="19B0F6C7FFF79105FF1D21FD23F8811A" box="[228,398,1787,1814]" degrees="18" direction="south" minutes="56" orientation="latitude" pageId="3" pageNumber="82" precision="1" seconds="40.6" value="-18.94461">S 18°56´40.6´´</geoCoordinate>
, 
<geoCoordinate id="19B0F6C7FFF79105FE6021FD2035811A" box="[409,579,1787,1814]" degrees="48" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="82" precision="1" seconds="26.7" value="48.424084">E 48°25´26.7´´</geoCoordinate>
, / primary forest, 
<collectorName id="D171F5D6FFF79105FCF321FD2108811A" box="[778,894,1788,1814]" pageId="3" pageNumber="82">M. Trýzna</collectorName>
leg.’ [p] (
<collectionCode id="1A9508C5FFF79105FC1521FD2635811A" box="[1004,1091,1788,1814]" pageId="3" pageNumber="82">MTDC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="CCEC9A5DFFF79105FBAF21FD2288818E" ID-GBIF-Occurrence="2572159689" collectingDate="1997-12-10" collectingDateMax="1997-12-18" collectingDateMin="1997-12-10" collectionCode="MTDC" collectorName="Pacholatko &amp; Sic &amp; Fairm. &amp; Frieser" country="Madagascar" determinerName="R. Frieser" pageId="3" pageNumber="82" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FBAF21FD26B5811A" bold="true" box="[1110,1219,1787,1814]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FBAF21FD26C9811A" box="[1110,1215,1787,1814]" pageId="3" pageNumber="82" type="female">1 female</specimenCount>
:
</emphasis>
‘MADAGASKAR c. / MORAMANGA env. / 
<date id="083AB6C0FFF79105FE282021200B8136" box="[465,637,1824,1850]" pageId="3" pageNumber="82" value="1997-12-10" valueMax="1997-12-18" valueMin="1997-12-10">
<collectingDate id="187E4F28FFF79105FE282021200B8136" box="[465,637,1824,1850]" pageId="3" pageNumber="82" value="1997-12-10" valueMax="1997-12-18" valueMin="1997-12-10">10.-18.xii.1997</collectingDate>
</date>
/ P. 
<collectorName id="D171F5D6FFF79105FD4A202121598136" box="[691,815,1824,1850]" pageId="3" pageNumber="82">Pacholátko</collectorName>
lgt.’ [p] // ‘ 
<taxonomicName id="BB84EB83FFF79105FC4D2021263E8136" authorityName="Jordan" authorityYear="1928" box="[948,1096,1824,1850]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="82" phylum="Arthropoda" rank="genus">Holophloeus</taxonomicName>
/ tuberculosus (
<collectorName id="D171F5D6FFF79105FAFB2021275A8137" box="[1282,1324,1824,1851]" pageId="3" pageNumber="82">Sic</collectorName>
!) / 
<collectorName id="D171F5D6FFF79105FAAB202127EB8136" box="[1362,1437,1824,1850]" pageId="3" pageNumber="82">Fairm.</collectorName>
[h] /det. 
<determinerName id="89E77CC4FFF79105FF032045231C8152" box="[250,362,1860,1886]" pageId="3" pageNumber="82">R. Frieser</determinerName>
[p] 2007 [h]’ // ‘CUM TYPO / COMPARATUM’ [red printed, a typical label used by 
<collectorName id="D171F5D6FFF79105FABC204227E08152" box="[1349,1430,1859,1886]" pageId="3" pageNumber="82">Frieser</collectorName>
] (
<collectionCode id="1A9508C5FFF79105FF6620692280818E" box="[159,246,1896,1922]" pageId="3" pageNumber="82">MTDC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="CCEC9A5DFFF79105FEF32069205881C6" ID-GBIF-Occurrence="2572159690" collectionCode="MNHN, MTDC" collectorName="Madagascar &amp; Antsianaka &amp; Lac Alaotra &amp; Perrot Freres &amp; Semestre &amp; Museum Paris &amp; Coll. &amp; R. Oberthur" country="Madagascar" pageId="3" pageNumber="82" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FEF320692397818E" bold="true" box="[266,481,1895,1922]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FEF32069232D818E" box="[266,347,1896,1922]" pageId="3" pageNumber="82" type="male">1 male</specimenCount>
, 
<specimenCount id="6A825B89FFF79105FE91206923AB818E" box="[360,477,1895,1922]" pageId="3" pageNumber="82" type="female">2 females</specimenCount>
:
</emphasis>
‘ 
<collectorName id="D171F5D6FFF79105FE142069200F818E" box="[493,633,1896,1922]" pageId="3" pageNumber="82">Madagascar</collectorName>
/ 
<collectorName id="D171F5D6FFF79105FD762069217B818E" box="[655,781,1896,1922]" pageId="3" pageNumber="82">Antsianaka</collectorName>
[= forest 
<collectorName id="D171F5D6FFF79105FC852069218C818E" box="[892,1018,1896,1922]" pageId="3" pageNumber="82">Antsianaka</collectorName>
near 
<collectorName id="D171F5D6FFF79105FBC0206926B6818E" box="[1081,1216,1895,1922]" pageId="3" pageNumber="82">Lac Alaotra</collectorName>
] / 
<collectorName id="D171F5D6FFF79105FB2720692706818E" box="[1246,1392,1896,1922]" pageId="3" pageNumber="82">Perrot Frères</collectorName>
/ 2 
<superScript id="8BF13D48FFF79105FA6C206627EA8179" attach="none" box="[1429,1436,1895,1909]" fontSize="6" pageId="3" pageNumber="82">e</superScript>
<collectorName id="D171F5D6FFF79105FF6E208D228881AA" box="[151,254,1932,1958]" pageId="3" pageNumber="82">Semestre</collectorName>
1893 [p]’ // ‘ 
<collectorName id="D171F5D6FFF79105FE72208D204681AA" box="[395,560,1932,1958]" pageId="3" pageNumber="82">Muséum Paris</collectorName>
/ ex. 
<collectorName id="D171F5D6FFF79105FD95208D20D281AA" box="[620,676,1932,1958]" pageId="3" pageNumber="82">Coll.</collectorName>
/ 
<collectorName id="D171F5D6FFF79105FD41208D214B81AA" box="[696,829,1932,1958]" pageId="3" pageNumber="82">R. Oberthür</collectorName>
/ 1952’ [p, typical blue acquisition label of 
<collectionCode id="1A9508C5FFF79105FAD9208A270F81AA" box="[1312,1401,1931,1958]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="82">MNHN</collectionCode>
] (1 pair in 
<collectionCode id="1A9508C5FFF79105FF1E20B1234981C6" box="[231,319,1968,1994]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="82">MNHN</collectionCode>
, 
<specimenCount id="6A825B89FFF79105FEB220B123DA81C6" box="[331,428,1968,1994]" pageId="3" pageNumber="82" type="female">1 female</specimenCount>
in 
<collectionCode id="1A9508C5FFF79105FE2920B1205081C6" box="[464,550,1968,1994]" pageId="3" pageNumber="82">MTDC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="CCEC9A5DFFF79105FDC020B121AC81E2" ID-GBIF-Occurrence="2572159699" collectionCode="MNHN" collectorName="Madagascar &amp; Fenerive &amp; E. Perrot &amp; Museum Paris &amp; Coll. &amp; Oberthur" country="Madagascar" pageId="3" pageNumber="82" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<emphasis id="4EF04C12FFF79105FDC020B120D181C6" bold="true" box="[569,679,1967,1994]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FDC020B120D481C6" box="[569,674,1967,1994]" pageId="3" pageNumber="82" type="female">1 female</specimenCount>
:
</emphasis>
‘ 
<collectorName id="D171F5D6FFF79105FD4B20B1214881C6" box="[690,830,1968,1994]" pageId="3" pageNumber="82">Madagascar</collectorName>
/ 
<collectorName id="D171F5D6FFF79105FCAC20B121C181C6" box="[853,951,1968,1994]" pageId="3" pageNumber="82">Fénerive</collectorName>
/ 
<collectorName id="D171F5D6FFF79105FC3420B1264681C6" box="[973,1072,1968,1994]" pageId="3" pageNumber="82">E. Perrot</collectorName>
[p] // 
<collectorName id="D171F5D6FFF79105FB8F20B1276181C6" box="[1142,1303,1968,1994]" pageId="3" pageNumber="82">Muséum Paris</collectorName>
/ ex. 
<collectorName id="D171F5D6FFF79105FAAF20B127F881C6" box="[1366,1422,1968,1994]" pageId="3" pageNumber="82">Coll.</collectorName>
/ R. 
<collectorName id="D171F5D6FFF79105FF4120D5236B81E2" box="[184,285,2004,2030]" pageId="3" pageNumber="82">Oberthür</collectorName>
/ 1952’ [p, typical blue acquisition label of 
<collectionCode id="1A9508C5FFF79105FCF120D2211781E2" box="[776,865,2003,2030]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="82">MNHN</collectionCode>
] (
<collectionCode id="1A9508C5FFF79105FC8F20D521A481E2" box="[886,978,2004,2030]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="82">MNHN</collectionCode>
)
</materialsCitation>
; 
<emphasis id="4EF04C12FFF79105FC1C20D5263E81E2" bold="true" box="[997,1096,2004,2030]" pageId="3" pageNumber="82">
<specimenCount id="6A825B89FFF79105FC1C20D5263581E2" box="[997,1091,2004,2030]" pageId="3" pageNumber="82" type="male">2 males</specimenCount>
:
</emphasis>
‘Madagascar / Fénerive / Per- rot Frères, 1 
<superScript id="8BF13D48FFF09102FED92799235B86AA" attach="left" box="[288,301,152,166]" fontSize="6" pageId="4" pageNumber="83">er</superScript>
Trim. 1896’ [p] // ‘Muséum Paris / ex. Coll. / R. Oberthür / 1952’ [p, typical blue acquisition label of 
<collectionCode id="1A9508C5FFF09102FF6E27BD228686DB" box="[151,240,188,215]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="83">MNHN</collectionCode>
] (
<specimenCount id="6A825B89FFF09102FEF027BC234886DB" box="[265,318,189,215]" pageId="4" pageNumber="83" type="generic">1 ex.</specimenCount>
in 
<collectionCode id="1A9508C5FFF09102FE9A27BC23CD86DB" box="[355,443,189,215]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="83">MNHN</collectionCode>
, 
<specimenCount id="6A825B89FFF09102FE3E27BC238886DB" box="[455,510,189,215]" pageId="4" pageNumber="83" type="generic">1 ex.</specimenCount>
in 
<collectionCode id="1A9508C5FFF09102FDDA27BC200F86DB" box="[547,633,189,215]" pageId="4" pageNumber="83">MTDC</collectionCode>
)
<materialsCitation id="CCEC9A5DFFF09102FD7827BC26BA86F7" ID-GBIF-Occurrence="2572159695" collectingDate="2004-12-15" collectingDateMax="2004-12-25" collectingDateMin="2004-12-15" collectionCode="MTDC" collectorName="R. Andreeva &amp; I. Andreev" country="Madagascar" location="Ranomafana" pageId="4" pageNumber="83" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
; 
<emphasis id="4EF04C12FFF09102FD7427BC21D386D8" bold="true" box="[653,933,188,215]" pageId="4" pageNumber="83">
<specimenCount id="6A825B89FFF09102FD7427BC208086DB" box="[653,758,188,215]" pageId="4" pageNumber="83" type="female">1 female</specimenCount>
: 
<collectingRegion id="BE405EE2FFF09102FCFB27BC21D386D8" box="[770,933,189,215]" country="Madagascar" name="Fianarantsoa" pageId="4" pageNumber="83">FIANARANTSOA</collectingRegion>
</emphasis>
PROVINCE
<emphasis id="4EF04C12FFF09102FBEC27BC266886DB" bold="true" box="[1045,1054,189,215]" pageId="4" pageNumber="83">:</emphasis>
‘ 
<collectingCountry id="0493D090FFF09102FBD327BC268E86DB" box="[1066,1272,189,215]" name="Madagascar" pageId="4" pageNumber="83">MADAGASKAR</collectingCountry>
/ 
<location id="795BC6DBFFF09102FAF627BC27EB86DB" LSID="urn:lsid:plazi:treatment:F42D2116FFF7910EFF6E220227EB81AD:795BC6DBFFF09102FAF627BC27EB86DB" box="[1295,1437,189,215]" country="Madagascar" name="Ranomafana" pageId="4" pageNumber="83" stateProvince="Fianarantsoa">Ranomafana</location>
res. [= Ranomafana National Park] / 
<date id="083AB6C0FFF09102FDD527E1209286F7" box="[556,740,224,251]" pageId="4" pageNumber="83" value="2004-12-15" valueMax="2004-12-25" valueMin="2004-12-15">
<collectingDate id="187E4F28FFF09102FDD527E1209286F7" box="[556,740,224,251]" pageId="4" pageNumber="83" value="2004-12-15" valueMax="2004-12-25" valueMin="2004-12-15">15.-25. xii. 2004</collectingDate>
</date>
/ 
<collectorName id="D171F5D6FFF09102FD0E27E021F386F7" box="[759,901,225,251]" pageId="4" pageNumber="83">R. Andreeva</collectorName>
, 
<collectorName id="D171F5D6FFF09102FC7727E0267586F7" box="[910,1027,225,251]" pageId="4" pageNumber="83">I. Andreev</collectorName>
leg.’ [p] (
<collectionCode id="1A9508C5FFF09102FB9427E026B286F7" box="[1133,1220,225,251]" pageId="4" pageNumber="83">MTDC</collectionCode>
)
</materialsCitation>
<materialsCitation id="CCEC9A5DFFF09102FB3527E020E3874F" ID-GBIF-Occurrence="2572159696" collectingDate="2019-01-08" collectingDateMax="2019-01-12" collectingDateMin="2019-01-08" collectionCode="MTDC" collectorName="Ranomafana N. P. &amp; M. Tryzna" country="Madagascar" elevation="1000" latitude="-21.263111" location="Talatakely" longLatPrecision="1" longitude="47.422417" pageId="4" pageNumber="83" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
; 
<emphasis id="4EF04C12FFF09102FB2F27E027EA86F7" bold="true" box="[1238,1436,224,251]" pageId="4" pageNumber="83">
<specimenCount id="6A825B89FFF09102FB2F27E0275386F7" box="[1238,1317,225,251]" pageId="4" pageNumber="83" type="male">1 male</specimenCount>
, 
<specimenCount id="6A825B89FFF09102FAC927E027EE86F7" box="[1328,1432,224,251]" pageId="4" pageNumber="83" type="female">1 female</specimenCount>
:
</emphasis>
‘ 
<collectingCountry id="0493D090FFF09102FF622604231C8713" box="[155,362,261,287]" name="Madagascar" pageId="4" pageNumber="83">MADAGASCAR</collectingCountry>
, 
<quantity id="BB7C3DE5FFF09102FE8F260423BD8713" box="[374,459,261,287]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="83" unit="m" value="1000.0">
<elevation id="F7A97733FFF09102FE8F260423BD8713" box="[374,459,261,287]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="83" unit="m" value="1000.0">1000 m</elevation>
</quantity>
/ 
<collectorName id="D171F5D6FFF09102FE1B260420DE8713" box="[482,680,261,287]" pageId="4" pageNumber="83">Ranomafana N.P.</collectorName>
, circuit / 
<location id="795BC6DBFFF09102FCE8260421F08713" LSID="urn:lsid:plazi:treatment:F42D2116FFF7910EFF6E220227EB81AD:795BC6DBFFF09102FCE8260421F08713" box="[785,902,261,287]" country="Madagascar" latitude="-21.263111" longLatPrecision="1" longitude="47.422417" name="Talatakely" pageId="4" pageNumber="83">Talatakely</location>
, 
<date id="083AB6C0FFF09102FC6B2604266A8713" box="[914,1052,261,287]" pageId="4" pageNumber="83" value="2019-01-08" valueMax="2019-01-12" valueMin="2019-01-08">
<collectingDate id="187E4F28FFF09102FC6B2604266A8713" box="[914,1052,261,287]" pageId="4" pageNumber="83" value="2019-01-08" valueMax="2019-01-12" valueMin="2019-01-08">8.-12.i.2019</collectingDate>
</date>
, / 
<geoCoordinate id="19B0F6C7FFF09102FBCE260426978713" box="[1079,1249,260,287]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="4" pageNumber="83" precision="1" seconds="47.2" value="-21.263111">S 21°15´47.2´´</geoCoordinate>
, 
<geoCoordinate id="19B0F6C7FFF09102FB14260427E18713" box="[1261,1431,260,287]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="4" pageNumber="83" precision="1" seconds="20.7" value="47.422417">E 47°25´20.7´´</geoCoordinate>
, / primary forest, 
<collectorName id="D171F5D6FFF09102FEAA262823BE874F" box="[339,456,297,323]" pageId="4" pageNumber="83">M. Trýzna</collectorName>
leg.’ [p] (
<collectionCode id="1A9508C5FFF09102FDCF262820FB874F" box="[566,653,297,323]" pageId="4" pageNumber="83">MTDC</collectionCode>
)
</materialsCitation>
<materialsCitation id="CCEC9A5DFFF09102FD6C262826D3874F" ID-GBIF-Occurrence="2572159693" box="[661,1189,296,323]" collectionCode="MNHN" country="Madagascar" pageId="4" pageNumber="83" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
; 
<emphasis id="4EF04C12FFF09102FD592628211D874F" bold="true" box="[672,875,296,323]" pageId="4" pageNumber="83">
<specimenCount id="6A825B89FFF09102FD5926282086874F" box="[672,752,297,323]" pageId="4" pageNumber="83" type="male">1 male</specimenCount>
, 
<specimenCount id="6A825B89FFF09102FD0426282110874F" box="[765,870,296,323]" pageId="4" pageNumber="83" type="female">1 female</specimenCount>
:
</emphasis>
‘Madagascar’ [p] (
<collectionCode id="1A9508C5FFF09102FBB8262826EB874F" box="[1089,1181,297,323]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="83">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="7C3B9000FFF09102FF3E264C23D38787" blockId="4.[151,1437,152,647]" pageId="4" pageNumber="83">
Red label [p]: ‘ 
<typeStatus id="A33F2EA2FFF09102FE91264C2077876B" box="[360,513,333,359]" pageId="4" pageNumber="83" type="holotype">HOLOTYPE</typeStatus>
/ 
<typeStatus id="A33F2EA2FFF09102FDEA264C20D2876B" box="[531,676,333,359]" pageId="4" pageNumber="83" type="allotype">ALLOTYPE</typeStatus>
/ 
<typeStatus id="A33F2EA2FFF09102FD4F264C2137876B" box="[694,833,333,359]" pageId="4" pageNumber="83" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="BB84EB83FFF09102FCAD264D219E876B" authorityName="Jordan" authorityYear="1928" box="[852,1000,332,359]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="4EF04C12FFF09102FCAD264D219E876B" bold="true" box="[852,1000,332,359]" italics="true" pageId="4" pageNumber="83">Holophloeus</emphasis>
</taxonomicName>
/ 
<emphasis id="4EF04C12FFF09102FC02264D26EF876B" bold="true" box="[1019,1177,332,359]" pageId="4" pageNumber="83">
<taxonomicName id="BB84EB83FFF09102FC02264D264D876B" authority="Trýzna &amp; Baňař, 2020" authorityName="Trýzna &amp; Baňař" authorityYear="2020" box="[1019,1083,332,359]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="83" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="4EF04C12FFF09102FC02264D264D876B" bold="true" box="[1019,1083,332,359]" italics="true" pageId="4" pageNumber="83">loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="55C3F169FFF09102FBB8264C26EF876B" box="[1089,1177,333,359]" pageId="4" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/ M. Trýzna &amp; P. Baňař det., 2018 (resp. 2019)’.
</paragraph>
</subSubSection>
<subSubSection id="349EC38BFFF0910EFF3E269520F485C7" lastPageId="8" lastPageNumber="87" pageId="4" pageNumber="83" type="description">
<paragraph id="7C3B9000FFF09102FF3E26952125848B" blockId="4.[151,1437,152,647]" pageId="4" pageNumber="83">
<emphasis id="4EF04C12FFF09102FF3E269520AA87A3" bold="true" box="[199,732,404,431]" pageId="4" pageNumber="83">
Description. Male 
<typeStatus id="A33F2EA2FFF09102FE582694207E87A3" box="[417,520,405,431]" pageId="4" pageNumber="83" type="holotype">holotype</typeStatus>
(female 
<typeStatus id="A33F2EA2FFF09102FD94269520B987A3" box="[621,719,404,431]" pageId="4" pageNumber="83" type="allotype">allotype</typeStatus>
).
</emphasis>
<emphasis id="4EF04C12FFF09102FD1B269721F287A3" box="[738,900,406,431]" italics="true" pageId="4" pageNumber="83">Measurements</emphasis>
(in mm): Total body length—8.82 (8.00). Head: total length—1.96 (1.80), length of rostrum—1.29 (1.12), maximum width of rostrum—1.04 (1.12), length of eye— 0.67 (0.68), maximum width across eyes—1.49 (1.52), minimum distance between eyes—0.56 (0.60). Antenna: length of segments: II—0.33 (0.28), III—0.71 (0.45), IV—0.69 (0.45), V—0.62 (0.35), VI—0.67 (0.28), VII—0.53 (0.25), VIII—0.44 (0.20), IX—0.33 (0.28), X—0.22 (0.18), XI—0.38 (0.28), width of segment IX—0.28 (0.25). Pronotum: maximum length—2.09 (1.88), maximum width—2.73 (2.72), minimum width—1.33 (1.44). Elytra: maximum length—4.45 (4.08), maximum width—3.31 (3.32).
</paragraph>
<caption id="28FBC088FFF09102FF6E20B7205781FF" ID-DOI="http://doi.org/10.5281/zenodo.3662545" ID-Zenodo-Dep="3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="4" pageNumber="83" startId="4.[151,264,1974,1999]" targetBox="[193,1318,681,1949]" targetPageId="4">
<paragraph id="7C3B9000FFF09102FF6E20B7205781FF" blockId="4.[151,1436,1974,2035]" pageId="4" pageNumber="83">
<emphasis id="4EF04C12FFF09102FF6E20B7232581C3" bold="true" box="[151,339,1974,1999]" pageId="4" pageNumber="83">FIGURES 4–5. 4,</emphasis>
<taxonomicName id="BB84EB83FFF09102FEA320B620A281C3" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" authorityYear="2020" box="[346,724,1974,1999]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="83" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="4EF04C12FFF09102FEA320B6206D81C3" box="[346,539,1975,1999]" italics="true" pageId="4" pageNumber="83">Holophloeus loebli</emphasis>
<emphasis id="4EF04C12FFF09102FDD820B720A281C3" bold="true" box="[545,724,1974,1999]" pageId="4" pageNumber="83">Trýzna &amp; Baňař</emphasis>
</taxonomicName>
<emphasis id="4EF04C12FFF09102FD2320B6214781C3" bold="true" box="[730,817,1975,1999]" pageId="4" pageNumber="83">
<taxonomicNameLabel id="55C3F169FFF09102FD2320B6215A81C3" box="[730,812,1975,1999]" pageId="4" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype, male; 
<emphasis id="4EF04C12FFF09102FC2220B6219881C3" bold="true" box="[987,1006,1975,1999]" pageId="4" pageNumber="83">5,</emphasis>
<taxonomicName id="BB84EB83FFF09102FC0D20B627EF81C3" authority="(Fairmaire, 1897)" authorityName="in Mt." baseAuthorityName="Fairmaire" baseAuthorityYear="1897" box="[1012,1433,1974,1999]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="83" phylum="Arthropoda" rank="species" species="tuberosus">
<emphasis id="4EF04C12FFF09102FC0D20B626A881C3" box="[1012,1246,1975,1999]" italics="true" pageId="4" pageNumber="83">Holophloeus tuberosus</emphasis>
(
<bibRefCitation id="1815EDF1FFF09102FB1320B727E781C3" author="Fairmaire, L. M. H." box="[1258,1425,1974,1999]" pageId="4" pageNumber="83" pagination="164 - 204" refId="ref7170" refString="Fairmaire, L. M. H. (1897) Materiaux pour la faune coleopterique de la region malgache. 3 e note (suite). Annales de la Societe Entomologique de Belgique, 41, 164 - 204." type="journal article" year="1897">Fairmaire, 1897</bibRefCitation>
)
</taxonomicName>
, male from Mt. ďAmbre National Park.
</paragraph>
</caption>
<caption id="28FBC088FFF19103FF6E207621C981F7" ID-DOI="http://doi.org/10.5281/zenodo.3662547" ID-Zenodo-Dep="3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="5" pageNumber="84" startId="5.[151,264,1911,1936]" targetBox="[197,1388,197,1883]" targetPageId="5">
<paragraph id="7C3B9000FFF19103FF6E207621C981F7" blockId="5.[151,1436,1911,2044]" pageId="5" pageNumber="84">
<emphasis id="4EF04C12FFF19103FF6E2076233F819C" bold="true" box="[151,329,1911,1936]" pageId="5" pageNumber="84">FIGURES 6–15.</emphasis>
Male terminalia. 
<emphasis id="4EF04C12FFF19103FDFB2076204D819C" bold="true" box="[514,571,1911,1936]" pageId="5" pageNumber="84">6–10,</emphasis>
<taxonomicName id="BB84EB83FFF19103FDBA207621B3819C" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" authorityYear="2020" box="[579,965,1911,1936]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="4EF04C12FFF19103FDBA207621718183" box="[579,775,1911,1935]" italics="true" pageId="5" pageNumber="84">Holophloeus loebli</emphasis>
<emphasis id="4EF04C12FFF19103FCF7207621B3819C" bold="true" box="[782,965,1911,1936]" pageId="5" pageNumber="84">Trýzna &amp; Baňař</emphasis>
</taxonomicName>
<emphasis id="4EF04C12FFF19103FC3520792653819C" bold="true" box="[972,1061,1912,1936]" pageId="5" pageNumber="84">
<taxonomicNameLabel id="55C3F169FFF19103FC3520792657819C" box="[972,1057,1912,1936]" pageId="5" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype male; 
<emphasis id="4EF04C12FFF19103FB3720762764819C" bold="true" box="[1230,1298,1911,1936]" pageId="5" pageNumber="84">11–15,</emphasis>
<taxonomicName id="BB84EB83FFF19103FAE2207623C381B8" authority="(Fairmaire, 1897)" authorityName="in Mt." baseAuthorityName="Fairmaire" baseAuthorityYear="1897" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="tuberosus">
<emphasis id="4EF04C12FFF19103FAE22076228F81BF" italics="true" pageId="5" pageNumber="84">Holophloeus tuberosus</emphasis>
(
<bibRefCitation id="1815EDF1FFF19103FEFF209A23D881B8" author="Fairmaire, L. M. H." box="[262,430,1947,1972]" pageId="5" pageNumber="84" pagination="164 - 204" refId="ref7170" refString="Fairmaire, L. M. H. (1897) Materiaux pour la faune coleopterique de la region malgache. 3 e note (suite). Annales de la Societe Entomologique de Belgique, 41, 164 - 204." type="journal article" year="1897">Fairmaire, 1897</bibRefCitation>
)
</taxonomicName>
, male from Mt. ďAmbre National Park; 
<emphasis id="4EF04C12FFF19103FCAB209A21FC81B8" bold="true" box="[850,906,1947,1972]" pageId="5" pageNumber="84">6, 11,</emphasis>
tegmen, dorsal view; 
<emphasis id="4EF04C12FFF19103FB94209D26D081B8" bold="true" box="[1133,1190,1947,1972]" pageId="5" pageNumber="84">7, 12,</emphasis>
aedeagus, ventral view; 
<emphasis id="4EF04C12FFF19103FF6E20BE22A781D4" bold="true" box="[151,209,1983,2008]" pageId="5" pageNumber="84">8, 13,</emphasis>
inner sac of aedeagus, latero-ventral view; 
<emphasis id="4EF04C12FFF19103FD6A20BE20BB81D4" bold="true" box="[659,717,1983,2008]" pageId="5" pageNumber="84">9, 14,</emphasis>
segment VIII and sternite IX, ventral view; 
<emphasis id="4EF04C12FFF19103FB6E20BE26AB81D4" bold="true" box="[1175,1245,1983,2008]" pageId="5" pageNumber="84">10, 15,</emphasis>
apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm.
</paragraph>
</caption>
<paragraph id="7C3B9000FFF29100FF3E2799275B86DB" blockId="6.[151,1436,152,215]" pageId="6" pageNumber="85">
<emphasis id="4EF04C12FFF29100FF3E2799233E86BF" bold="true" box="[199,328,152,179]" pageId="6" pageNumber="85">Coloration</emphasis>
of the cuticle black, only tarsomeres III–IV in all legs dark brown, claws slightly lighter. Proximal and distal part of tarsomere I and proximal part of tarsomere II covered with dense whitish hairs (
<figureCitation id="E4BF8C85FFF29100FB2227BC275686DB" box="[1243,1312,189,215]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="6" pageNumber="85">Fig. 4</figureCitation>
).
</paragraph>
<caption id="28FBC088FFF29100FF6E214127778071" ID-DOI="http://doi.org/10.5281/zenodo.3662550" ID-Zenodo-Dep="3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="6" pageNumber="85" startId="6.[151,264,1600,1625]" targetBox="[151,1436,237,1575]" targetPageId="6">
<paragraph id="7C3B9000FFF29100FF6E214127778071" blockId="6.[151,1436,1600,1661]" pageId="6" pageNumber="85">
<emphasis id="4EF04C12FFF29100FF6E214123E98055" bold="true" box="[151,415,1600,1625]" pageId="6" pageNumber="85">FIGURES 16–19. 16, 18,</emphasis>
<taxonomicName id="BB84EB83FFF29100FE5F214121568055" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" authorityYear="2020" box="[422,800,1600,1625]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="4EF04C12FFF29100FE5F2141201E8054" box="[422,616,1600,1624]" italics="true" pageId="6" pageNumber="85">Holophloeus loebli</emphasis>
<emphasis id="4EF04C12FFF29100FD97214121568055" bold="true" box="[622,800,1600,1625]" pageId="6" pageNumber="85">Trýzna &amp; Baňař</emphasis>
</taxonomicName>
<emphasis id="4EF04C12FFF29100FCDF2140210B8055" bold="true" box="[806,893,1601,1625]" pageId="6" pageNumber="85">
<taxonomicNameLabel id="55C3F169FFF29100FCDF2140210F8055" box="[806,889,1601,1625]" pageId="6" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype, male; 
<emphasis id="4EF04C12FFF29100FBD12141261B8055" bold="true" box="[1064,1133,1600,1625]" pageId="6" pageNumber="85">17, 19,</emphasis>
<taxonomicName id="BB84EB83FFF29100FB8A2141236D8070" authority="(Fairmaire, 1897)" authorityName="in Mt." baseAuthorityName="Fairmaire" baseAuthorityYear="1897" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="tuberosus">
<emphasis id="4EF04C12FFF29100FB8A2141272B8054" box="[1139,1373,1600,1624]" italics="true" pageId="6" pageNumber="85">Holophloeus tuberosus</emphasis>
(
<bibRefCitation id="1815EDF1FFF29100FA93214123628070" author="Fairmaire, L. M. H." pageId="6" pageNumber="85" pagination="164 - 204" refId="ref7170" refString="Fairmaire, L. M. H. (1897) Materiaux pour la faune coleopterique de la region malgache. 3 e note (suite). Annales de la Societe Entomologique de Belgique, 41, 164 - 204." type="journal article" year="1897">Fairmaire, 1897</bibRefCitation>
)
</taxonomicName>
, male from Mt. ďAmbre National Park; 
<emphasis id="4EF04C12FFF29100FD4C2165208D8071" bold="true" box="[693,763,1636,1661]" pageId="6" pageNumber="85">16–17,</emphasis>
pronotum, dorsal view; 
<emphasis id="4EF04C12FFF29100FC0A2165264F8071" bold="true" box="[1011,1081,1636,1661]" pageId="6" pageNumber="85">18–19,</emphasis>
elytra, dorsal view.
</paragraph>
</caption>
<paragraph id="7C3B9000FFF29100FF3E21AB22A181E9" blockId="6.[151,1437,1706,2021]" pageId="6" pageNumber="85">
<emphasis id="4EF04C12FFF29100FF3E21AB234C80C9" bold="true" box="[199,314,1706,1733]" pageId="6" pageNumber="85">Vestiture.</emphasis>
<emphasis id="4EF04C12FFF29100FEB821AA230880C9" box="[321,382,1707,1733]" italics="true" pageId="6" pageNumber="85">Head</emphasis>
naked, 
<emphasis id="4EF04C12FFF29100FE2121AD203280C9" box="[472,580,1708,1733]" italics="true" pageId="6" pageNumber="85">pronotum</emphasis>
covered with very fine sparse indistinct appressed blackish setae, on lateral sides of pronotum and on prosternum setae whitish. 
<emphasis id="4EF04C12FFF29100FD0E21CE214B80E5" box="[759,829,1743,1769]" italics="true" pageId="6" pageNumber="85">Elytra</emphasis>
, including all tubercles covered with very short fine sparse appressed blackish setae. 
<emphasis id="4EF04C12FFF29100FDFE21F220078101" box="[519,625,1779,1805]" italics="true" pageId="6" pageNumber="85">Antennae</emphasis>
with short fine appressed setae, antennomeres VIII–XI in addition with several long hairs on distal part. All 
<emphasis id="4EF04C12FFF29100FDCC20162014813D" box="[565,610,1815,1841]" italics="true" pageId="6" pageNumber="85">legs</emphasis>
covered with fine setae, all femora with indistinct light sparse appressed setae particularly in distal part, all tibiae with dense black setae, appressed on external part and distinctly subdecumbent in internal part. All tarsomeres covered with decumbent dense black setae, proximal and distal part of tarsomere I and proximal part of tarsomere II also covered with dense whitish decumbent hairs. Venter of thorax and abdominal sternites with very short sparse whitish pubescence. 
<emphasis id="4EF04C12FFF29100FC7C20A6218681CD" box="[901,1008,1959,1985]" italics="true" pageId="6" pageNumber="85">Pygidium</emphasis>
covered with very fine appressed dark setae.
</paragraph>
<caption id="28FBC088FFF39101FF6E21082766804A" ID-DOI="http://doi.org/10.5281/zenodo.3662552" ID-Zenodo-Dep="3662552" httpUri="https://zenodo.org/record/3662552/files/figure.png" pageId="7" pageNumber="86" startId="7.[151,264,1545,1570]" targetBox="[151,1436,181,1520]" targetPageId="7">
<paragraph id="7C3B9000FFF39101FF6E21082766804A" blockId="7.[151,1436,1545,1606]" pageId="7" pageNumber="86">
<emphasis id="4EF04C12FFF39101FF6E210823D6802E" bold="true" box="[151,416,1545,1570]" pageId="7" pageNumber="86">FIGURES 20–23. 20, 22,</emphasis>
<taxonomicName id="BB84EB83FFF39101FE5F210B2156802E" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" authorityYear="2020" box="[422,800,1545,1570]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="4EF04C12FFF39101FE5F210B201E802E" box="[422,616,1546,1570]" italics="true" pageId="7" pageNumber="86">Holophloeus loebli</emphasis>
<emphasis id="4EF04C12FFF39101FD9721082156802E" bold="true" box="[622,800,1545,1570]" pageId="7" pageNumber="86">Trýzna &amp; Baňař</emphasis>
</taxonomicName>
<emphasis id="4EF04C12FFF39101FCDF210B210B802E" bold="true" box="[806,893,1546,1570]" pageId="7" pageNumber="86">
<taxonomicNameLabel id="55C3F169FFF39101FCDF210B210F802E" box="[806,889,1546,1570]" pageId="7" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holotype, male; 
<emphasis id="4EF04C12FFF39101FBD12108261B802E" bold="true" box="[1064,1133,1545,1570]" pageId="7" pageNumber="86">21, 23,</emphasis>
<taxonomicName id="BB84EB83FFF39101FB8A210B236D804A" authority="(Fairmaire, 1897)" authorityName="in Mt." baseAuthorityName="Fairmaire" baseAuthorityYear="1897" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="tuberosus">
<emphasis id="4EF04C12FFF39101FB8A210B272B802E" box="[1139,1373,1546,1570]" italics="true" pageId="7" pageNumber="86">Holophloeus tuberosus</emphasis>
(
<bibRefCitation id="1815EDF1FFF39101FA9321082362804A" author="Fairmaire, L. M. H." pageId="7" pageNumber="86" pagination="164 - 204" refId="ref7170" refString="Fairmaire, L. M. H. (1897) Materiaux pour la faune coleopterique de la region malgache. 3 e note (suite). Annales de la Societe Entomologique de Belgique, 41, 164 - 204." type="journal article" year="1897">Fairmaire, 1897</bibRefCitation>
)
</taxonomicName>
, male from Mt. ďAmbre National Park; 
<emphasis id="4EF04C12FFF39101FD4C212C208D804A" bold="true" box="[693,763,1581,1606]" pageId="7" pageNumber="86">20–21,</emphasis>
head, ventral view; 
<emphasis id="4EF04C12FFF39101FC30212C2679804A" bold="true" box="[969,1039,1581,1606]" pageId="7" pageNumber="86">22–23,</emphasis>
prosternum, dorsal view.
</paragraph>
</caption>
<paragraph id="7C3B9000FFF3910EFF3E217220148417" blockId="7.[151,1443,1651,2039]" lastBlockId="8.[151,1437,153,1295]" lastPageId="8" lastPageNumber="87" pageId="7" pageNumber="86">
<emphasis id="4EF04C12FFF39101FF3E217223378082" bold="true" box="[199,321,1651,1678]" pageId="7" pageNumber="86">Structure.</emphasis>
<emphasis id="4EF04C12FFF39101FEBE217523F28082" box="[327,388,1652,1678]" italics="true" pageId="7" pageNumber="86">Head</emphasis>
relatively long, rostrum broad (
<figureCitation id="E4BF8C85FFF39101FD1E217521408082" box="[743,822,1652,1678]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="7" pageNumber="86">Figs. 4</figureCitation>
, 
<figureCitation id="E4BF8C85FFF39101FCBB217521178082" box="[834,865,1652,1678]" captionStart="FIGURES 20–23. 20, 22" captionStartId="7.[151,264,1545,1570]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure@7.[151,1436,181,1520]" captionTargetPageId="7" captionText="FIGURES 20–23. 20, 22, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 21, 23, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 20–21, head, ventral view; 22–23, prosternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662552" httpUri="https://zenodo.org/record/3662552/files/figure.png" pageId="7" pageNumber="86">20</figureCitation>
), flat in middle, with indistinct depression between antennal scrobes, with distinct longitudinal carina in central part of rostrum. Dorsal part of rostrum and frons very strongly sculptured, this sculpturation also on lateral sides of rostrum. Ratio of rostrum length to maximum width 
<quantity id="BB7C3DE5FFF39101FF6E21E1229280F6" box="[151,228,1760,1786]" metricMagnitude="-2" metricUnit="m" metricValue="3.1496" pageId="7" pageNumber="86" unit="in" value="1.24">1.24 in</quantity>
male, 1.00 in female. Ventral part of head only with fine and sparse punctation (
<figureCitation id="E4BF8C85FFF39101FBAC21E126DE80F6" box="[1109,1192,1760,1786]" captionStart="FIGURES 20–23. 20, 22" captionStartId="7.[151,264,1545,1570]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure@7.[151,1436,181,1520]" captionTargetPageId="7" captionText="FIGURES 20–23. 20, 22, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 21, 23, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 20–21, head, ventral view; 22–23, prosternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662552" httpUri="https://zenodo.org/record/3662552/files/figure.png" pageId="7" pageNumber="86">Fig. 20</figureCitation>
). Eyes not emarginate anteriorly, dorsal ocular index 
<quantity id="BB7C3DE5FFF39101FE10200520408112" box="[489,566,1796,1822]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="7" pageNumber="86" unit="in" value="1.2">1.20 in</quantity>
male, 
<quantity id="BB7C3DE5FFF39101FD87200520BD8112" box="[638,715,1796,1822]" metricMagnitude="-2" metricUnit="m" metricValue="3.302" pageId="7" pageNumber="86" unit="in" value="1.3">1.30 in</quantity>
female. Ratio of maximum width across eyes to maximum width of rostrum 
<quantity id="BB7C3DE5FFF39101FEEA20292316814E" box="[275,352,1832,1858]" metricMagnitude="-2" metricUnit="m" metricValue="3.6322" pageId="7" pageNumber="86" unit="in" value="1.43">1.43 in</quantity>
male, 
<quantity id="BB7C3DE5FFF39101FE522029238E814E" box="[427,504,1832,1858]" metricMagnitude="-2" metricUnit="m" metricValue="3.4544" pageId="7" pageNumber="86" unit="in" value="1.36">1.36 in</quantity>
female. 
<emphasis id="4EF04C12FFF39101FDA1202820B4814F" box="[600,706,1833,1859]" italics="true" pageId="7" pageNumber="86">Antennae</emphasis>
(
<figureCitation id="E4BF8C85FFF39101FD2820292162814E" box="[721,788,1832,1858]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="7" pageNumber="86">Fig. 4</figureCitation>
) of male distinctly extended to posterior margin of pronotum, antennae of females shorter, reaching approximately posterior margin of pronotum. Scape and pedicel thick, funicle thin, antennomeres IX–XI not forming compact club but slack and gradually extended in both sexes. Antennomere IX only slightly longer than wider in both sexes, X as long as wide in male, slightly transverse in female, XI slightly longer than wider in male, as long as wide in female. 
<emphasis id="4EF04C12FFF39101FC8220B8219C81DE" box="[891,1002,1977,2002]" italics="true" pageId="7" pageNumber="86">Pronotum</emphasis>
transverse, ratio of its length to maximum width 
<quantity id="BB7C3DE5FFF39101FEE620DD231B81FA" box="[287,365,2012,2038]" metricMagnitude="-2" metricUnit="m" metricValue="1.9558" pageId="7" pageNumber="86" unit="in" value="0.77">0.77 in</quantity>
male, 
<quantity id="BB7C3DE5FFF39101FE4020DD207E81FA" box="[441,520,2012,2038]" metricMagnitude="-2" metricUnit="m" metricValue="1.7526" pageId="7" pageNumber="86" unit="in" value="0.69">0.69 in</quantity>
female, widest at distinct dorsal transverse carina, from which strongly narrowed anteriad and posteriad (
<figureCitation id="E4BF8C85FFFC910EFE5A2798238E86B8" box="[419,504,153,180]" captionStart="FIGURES 16–19. 16, 18" captionStartId="6.[151,264,1600,1625]" captionTargetBox="[151,1436,237,1575]" captionTargetId="figure@6.[151,1436,237,1575]" captionTargetPageId="6" captionText="FIGURES 16–19. 16, 18, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 17, 19, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 16–17, pronotum, dorsal view; 18–19, elytra, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="8" pageNumber="87">Fig. 16</figureCitation>
). Dorsal antebasal transverse carina with arch in middle, here shortly interrupted, continuously-undulate to sides and interrupted before lateral lobe. Lateral carina of pronotum short, in contact with dorsal transverse carina forming a distinct lobe. Basal longitudinal carinula of pronotum absent. Pronotal disc with one well-defined tubercle, surrounding area with shallow depressions. 
<emphasis id="4EF04C12FFFC910EFC41260421888713" box="[952,1022,261,287]" italics="true" pageId="8" pageNumber="87">Elytra</emphasis>
broadly suboval to rectangular (
<figureCitation id="E4BF8C85FFFC910EFA96260422C3874F" captionStart="FIGURES 16–19. 16, 18" captionStartId="6.[151,264,1600,1625]" captionTargetBox="[151,1436,237,1575]" captionTargetId="figure@6.[151,1436,237,1575]" captionTargetPageId="6" captionText="FIGURES 16–19. 16, 18, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 17, 19, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 16–17, pronotum, dorsal view; 18–19, elytra, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="8" pageNumber="87">Fig. 18</figureCitation>
), narrowed evenly in distal fifth of their length, ratio of the maximum length to maximum width of elytra 
<quantity id="BB7C3DE5FFFC910EFAB6262827EA874F" box="[1359,1436,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="3.4036" pageId="8" pageNumber="87" unit="in" value="1.34">1.34 in</quantity>
male, 
<quantity id="BB7C3DE5FFFC910EFF25264C235C876B" box="[220,298,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="3.1242" pageId="8" pageNumber="87" unit="in" value="1.23">1.23 in</quantity>
female. Elytra with numerous well-defined tubercles on even-numbered elytral intervals: second elytral interval with three well-defined protruded subbasal, median and postmedian and one small preapical tubercles, fourth elytral interval with two well-defined antemedian and postmedian and one small preapical tubercles, sixth and eighth elytral intervals with numerous small tubercles (
<figureCitation id="E4BF8C85FFFC910EFCD326B8210F87DF" box="[810,889,441,467]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="8" pageNumber="87">Figs. 4</figureCitation>
, 
<figureCitation id="E4BF8C85FFFC910EFC7D26B821D587DF" box="[900,931,441,467]" captionStart="FIGURES 16–19. 16, 18" captionStartId="6.[151,264,1600,1625]" captionTargetBox="[151,1436,237,1575]" captionTargetId="figure@6.[151,1436,237,1575]" captionTargetPageId="6" captionText="FIGURES 16–19. 16, 18, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 17, 19, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 16–17, pronotum, dorsal view; 18–19, elytra, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="8" pageNumber="87">18</figureCitation>
). 
<emphasis id="4EF04C12FFFC910EFC4E26B8265787DF" box="[951,1057,441,467]" italics="true" pageId="8" pageNumber="87">Abdomen</emphasis>
shorter than broad. Ventrites II–V with prominent margins on proximal part in male (
<figureCitation id="E4BF8C85FFFC910EFD3F26DC213B87FB" box="[710,845,477,503]" captionStart="FIGURES 24–29. 24, 26, 28" captionStartId="9.[151,264,1945,1970]" captionTargetBox="[178,1402,181,1920]" captionTargetId="figure@9.[178,1409,181,1920]" captionTargetPageId="9" captionText="FIGURES 24–29. 24, 26, 28, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 25, 27, 29, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 24–25, mesosternum and metasternum, dorsal view; 26–27, abdominal ventrites I–V, dorsal view; 28–29, abdominal ventrite V, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662555" httpUri="https://zenodo.org/record/3662555/files/figure.png" pageId="8" pageNumber="87">Figs. 26, 28</figureCitation>
). 
<emphasis id="4EF04C12FFFC910EFCA626DC21BC87FB" box="[863,970,477,503]" italics="true" pageId="8" pageNumber="87">Pygidium</emphasis>
longer than wide in male, wider than long in female, slightly narrowing posteriorly.
</paragraph>
<paragraph id="7C3B9000FFFC910EFF3E25242086851B" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="87">
<emphasis id="4EF04C12FFFC910EFF3E252420D18433" box="[199,679,549,575]" italics="true" pageId="8" pageNumber="87">Male genitalia and surrounding structures</emphasis>
. Tegmen robust (
<figureCitation id="E4BF8C85FFFC910EFC94252421C5844C" box="[877,947,549,576]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 6</figureCitation>
), tegminal plate wide, basal piece strongly, tegminal apodeme weakly sclerotized. Apex of tegmen conspicuously broadened. Aedeagus long and robust (
<figureCitation id="E4BF8C85FFFC910EFA96254822D1848B" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 7</figureCitation>
), its inner sac (
<figureCitation id="E4BF8C85FFFC910EFEB5256C23FB848B" box="[332,397,621,647]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 8</figureCitation>
) with several symmetrical and asymmetrical densely serrated structures. Pedon sub-rectangular in shape, with short and strongly sclerotized apical projection. Tectum with very long process, longer than apex of pedon (
<figureCitation id="E4BF8C85FFFC910EFEF425B423FC84C3" box="[269,394,693,719]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Figs. 7, 10</figureCitation>
). Tergite VIII strongly sclerotized, with two antero-lateral projections, sternite VIII (
<figureCitation id="E4BF8C85FFFC910EFAB625B427E284C3" box="[1359,1428,693,719]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 9</figureCitation>
) sub-divided into two kidney-shaped hemisternites. Sternite IX (= spiculum gastrale) with strongly sclerotized arms touching sternite VIII on its posterior margin (
<figureCitation id="E4BF8C85FFFC910EFD6425FC2094851B" box="[669,738,765,791]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="7C3B9000FFFC910EFF3E242120F485C7" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="87">
<emphasis id="4EF04C12FFFC910EFF3E242123CC8537" bold="true" box="[199,442,800,827]" pageId="8" pageNumber="87">Sexual dimorphism.</emphasis>
<emphasis id="4EF04C12FFFC910EFE3A242020668537" box="[451,528,801,827]" italics="true" pageId="8" pageNumber="87">Males:</emphasis>
body comparatively more slender than in females, rostrum slightly longer and more slender. Antennae extended distinctly beyond posterior margin of pronotum, antennomere X as long as wide. Abdominal ventrites flat in middle, ventrites II–V with prominent margins on proximal part. 
<emphasis id="4EF04C12FFFC910EFB4A2468276D858F" box="[1203,1307,873,899]" italics="true" pageId="8" pageNumber="87">Females:</emphasis>
body more robust than in male. Antennae shorter, approximately reaching posterior margin of pronotum, antennomere X slightly transverse. Abdominal ventrites rounded.
</paragraph>
</subSubSection>
<subSubSection id="349EC38BFFFC910EFF3E24D426FE82AF" pageId="8" pageNumber="87" type="etymology">
<paragraph id="7C3B9000FFFC910EFF3E24D42047821F" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="87">
<emphasis id="4EF04C12FFFC910EFF3E24D4233885E3" bold="true" box="[199,334,981,1007]" pageId="8" pageNumber="87">Etymology.</emphasis>
Patronym, dedicated to our dear colleague and friend Ivan Löbl (
<collectingRegion id="BE405EE2FFFC910EFBDB24D4260B85E3" box="[1058,1149,981,1007]" country="Switzerland" name="Geneve" pageId="8" pageNumber="87">Geneva</collectingRegion>
), eminent specialist in the field of the taxonomy of 
<taxonomicName id="BB84EB83FFFC910EFE5724F8205B821F" authorityName="Linnaeus" authorityYear="1758" box="[430,557,1017,1043]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="8" pageNumber="87" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
.
</paragraph>
<paragraph id="7C3B9000FFFC910EFF3E231D26FE82AF" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="87">
<emphasis id="4EF04C12FFFC910EFF3E231D238F823B" bold="true" box="[199,505,1052,1079]" pageId="8" pageNumber="87">Collecting circumstances.</emphasis>
The 
<typeStatus id="A33F2EA2FFFC910EFDCC231C20EC823B" box="[565,666,1053,1079]" pageId="8" pageNumber="87" type="holotype">holotype</typeStatus>
, 
<typeStatus id="A33F2EA2FFFC910EFD5F231C2176823B" box="[678,768,1053,1079]" pageId="8" pageNumber="87" type="allotype">allotype</typeStatus>
and most of the 
<typeStatus id="A33F2EA2FFFC910EFC3B231C265B823B" box="[962,1069,1053,1079]" pageId="8" pageNumber="87" type="paratype">paratypes</typeStatus>
were collected in tropical moist primary forest at an elevation of 993 metres on the lower side of a fallen tree trunk. The rotten trunk had a diameter about 
<quantity id="BB7C3DE5FFFC910EFF27236523538273" box="[222,293,1124,1151]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="87" unit="cm" value="50.0">50 cm</quantity>
and was located in a distinctly shaded area. Microhabitat of the type locality is on 
<figureCitation id="E4BF8C85FFFC910EFB192364271F8273" box="[1248,1385,1125,1151]" captionStart-0="FIGURES 36–37" captionStart-1="FIGURES 38–39" captionStart-2="FIGURES 40–41" captionStartId-0="13.[151,264,1956,1981]" captionStartId-1="14.[151,264,1963,1988]" captionStartId-2="15.[151,264,1928,1953]" captionTargetBox-0="[164,1423,181,1930]" captionTargetBox-1="[164,1423,181,1937]" captionTargetBox-2="[151,1436,181,1903]" captionTargetId-0="figure@13.[164,1423,181,1930]" captionTargetId-1="figure@14.[164,1423,181,1937]" captionTargetId-2="figure@15.[151,1436,181,1903]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURES 36–37. Collecting site of Holophloeus loebli Trýzna &amp; Baňař sp. nov.; 36, Andasibe National Park, Analamazaotra forest, 2016; 37, type locality in Mantadia National Park, Eulophia circuit, 2017." captionText-1="FIGURES 38–39. Type locality of Holophloeus loebli Trýzna &amp; Baňař sp. nov. in Mantadia National Park, Eulophia circuit, 2017; 38, primary forest with single huge torso of lying trunk; 39, detail of cover of bark with specimens." captionText-2="FIGURES 40–41. Type locality of Holophloeus loebli Trýzna &amp; Baňař sp. nov. in Mantadia National Park, Eulophia circuit, 2017, detail with specimen and exit hole." figureDoi-0="http://doi.org/10.5281/zenodo.3662563" figureDoi-1="http://doi.org/10.5281/zenodo.3662566" figureDoi-2="http://doi.org/10.5281/zenodo.3662568" httpUri-0="https://zenodo.org/record/3662563/files/figure.png" httpUri-1="https://zenodo.org/record/3662566/files/figure.png" httpUri-2="https://zenodo.org/record/3662568/files/figure.png" pageId="8" pageNumber="87">Figs. 37–41</figureCitation>
. For details, see section on Ecological notes on Madagascan species of the genus 
<emphasis id="4EF04C12FFFC910EFC08238826FE82AF" box="[1009,1160,1161,1187]" italics="true" pageId="8" pageNumber="87">
<taxonomicName id="BB84EB83FFFC910EFC08238826F282AF" authorityName="Jordan" authorityYear="1928" box="[1009,1156,1161,1187]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="genus">Holophloeus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="349EC38BFFFC910EFF3E23AD270582CB" box="[199,1395,1196,1223]" pageId="8" pageNumber="87" type="distribution">
<paragraph id="7C3B9000FFFC910EFF3E23AD270582CB" blockId="8.[151,1437,153,1295]" box="[199,1395,1196,1223]" pageId="8" pageNumber="87">
<emphasis id="4EF04C12FFFC910EFF3E23AD231782CB" bold="true" box="[199,353,1196,1223]" pageId="8" pageNumber="87">Distribution.</emphasis>
All specimens were collected in east 
<collectingCountry id="0493D090FFFC910EFCFE23AC21E682CB" box="[775,912,1197,1223]" name="Madagascar" pageId="8" pageNumber="87">Madagascar</collectingCountry>
, in 
<collectingRegion id="BE405EE2FFFC910EFC4E23AC264682CB" box="[951,1072,1197,1223]" country="Madagascar" name="Toamasina" pageId="8" pageNumber="87">Toamasina</collectingRegion>
and 
<collectingRegion id="BE405EE2FFFC910EFB9F23AC268382CB" box="[1126,1269,1197,1223]" country="Madagascar" name="Fianarantsoa" pageId="8" pageNumber="87">Fianarantsoa</collectingRegion>
Provinces.
</paragraph>
</subSubSection>
<subSubSection id="349EC38BFFFC910EFF3E23D127EB81AD" pageId="8" pageNumber="87" type="diagnosis">
<paragraph id="7C3B9000FFFC910EFF3E23D1277F8303" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="87">
<emphasis id="4EF04C12FFFC910EFF3E23D123A782E7" bold="true" box="[199,465,1232,1259]" pageId="8" pageNumber="87">Differential diagnosis.</emphasis>
<taxonomicName id="BB84EB83FFFC910EFE2523D0210D82E7" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" authorityYear="2020" box="[476,891,1233,1259]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="4EF04C12FFFC910EFE2523D020C382E7" box="[476,693,1233,1259]" italics="true" pageId="8" pageNumber="87">Holophloeus loebli</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<emphasis id="4EF04C12FFFC910EFC7F23D0219582E7" bold="true" box="[902,995,1233,1259]" pageId="8" pageNumber="87">
<taxonomicNameLabel id="55C3F169FFFC910EFC7F23D0219582E7" box="[902,995,1233,1259]" pageId="8" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="BB84EB83FFFC910EFB7A23D322AF8303" authority="(Fairmaire, 1897)" authorityName="in Mt." baseAuthorityName="Fairmaire" baseAuthorityYear="1897" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="tuberosus">
<emphasis id="4EF04C12FFFC910EFB7A23D3276382E7" box="[1155,1301,1233,1259]" italics="true" pageId="8" pageNumber="87">H. tuberosus</emphasis>
(
<bibRefCitation id="1815EDF1FFFC910EFADE23D022A68303" author="Fairmaire, L. M. H." pageId="8" pageNumber="87" pagination="164 - 204" refId="ref7170" refString="Fairmaire, L. M. H. (1897) Materiaux pour la faune coleopterique de la region malgache. 3 e note (suite). Annales de la Societe Entomologique de Belgique, 41, 164 - 204." type="journal article" year="1897">Fairmaire, 1897</bibRefCitation>
)
</taxonomicName>
from which it can be distinguished by the morphological characters given in the following key:
</paragraph>
<paragraph id="7C3B9000FFFC910EFF6E225F27EA8077" blockId="8.[151,1437,1373,1953]" pageId="8" pageNumber="87">
(1) Specimens more robust, ratio of body length to maximum width 
<quantity id="BB7C3DE5FFFC910EFCBC225C21F1837F" box="[837,903,1373,1395]" metricMagnitude="-2" metricUnit="m" metricValue="6.7564" pageId="8" pageNumber="87" unit="in" value="2.66">2.66 in</quantity>
male, 
<quantity id="BB7C3DE5FFFC910EFC3F225F267E837F" box="[966,1032,1373,1395]" metricMagnitude="-2" metricUnit="m" metricValue="6.1214" pageId="8" pageNumber="87" unit="in" value="2.41">2.41 in</quantity>
female (
<figureCitation id="E4BF8C85FFFC910EFBA3225C26E58378" box="[1114,1171,1373,1396]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="8" pageNumber="87">Fig. 4</figureCitation>
). Pronotum transverse (
<figureCitation id="E4BF8C85FFFC910EFA8C225C228A839D" captionStart="FIGURES 16–19. 16, 18" captionStartId="6.[151,264,1600,1625]" captionTargetBox="[151,1436,237,1575]" captionTargetId="figure@6.[151,1436,237,1575]" captionTargetPageId="6" captionText="FIGURES 16–19. 16, 18, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 17, 19, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 16–17, pronotum, dorsal view; 18–19, elytra, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="8" pageNumber="87">Fig. 16</figureCitation>
), ratio of its length to width at carina 
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male, 
<quantity id="BB7C3DE5FFFC910EFD10227A215A839D" box="[745,812,1403,1425]" metricMagnitude="-2" metricUnit="m" metricValue="1.7526" pageId="8" pageNumber="87" unit="in" value="0.69">0.69 in</quantity>
female. Elytra suboval (
<figureCitation id="E4BF8C85FFFC910EFBE1227A2629839D" box="[1048,1119,1403,1425]" captionStart="FIGURES 16–19. 16, 18" captionStartId="6.[151,264,1600,1625]" captionTargetBox="[151,1436,237,1575]" captionTargetId="figure@6.[151,1436,237,1575]" captionTargetPageId="6" captionText="FIGURES 16–19. 16, 18, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 17, 19, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 16–17, pronotum, dorsal view; 18–19, elytra, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="8" pageNumber="87">Fig. 18</figureCitation>
), ratio of the maximum length to maximum width of elytra 
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male, 
<quantity id="BB7C3DE5FFFC910EFDA4229920E983A2" box="[605,671,1432,1454]" metricMagnitude="-2" metricUnit="m" metricValue="3.1242" pageId="8" pageNumber="87" unit="in" value="1.23">1.23 in</quantity>
female. Elytral tubercles robust, more prominent, in cross-section rather spherical. Subbasal, median and postmedian tubercles on the second elytral interval very prominent (see also from lateral view). Pronotal and elytral tubercles entirely black. Lateral carina of pronotum reaching one third of pronotal length. Prosternum and mesosternum with deep sparse microsculpture (
<figureCitation id="E4BF8C85FFFC910EFD4822F121758009" box="[689,771,1520,1542]" captionStart="FIGURES 20–23. 20, 22" captionStartId="7.[151,264,1545,1570]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure@7.[151,1436,181,1520]" captionTargetPageId="7" captionText="FIGURES 20–23. 20, 22, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 21, 23, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 20–21, head, ventral view; 22–23, prosternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662552" httpUri="https://zenodo.org/record/3662552/files/figure.png" pageId="8" pageNumber="87">Figs. 22</figureCitation>
, 
<figureCitation id="E4BF8C85FFFC910EFCF722F1215F800A" box="[782,809,1520,1542]" captionStart="FIGURES 24–29. 24, 26, 28" captionStartId="9.[151,264,1945,1970]" captionTargetBox="[178,1402,181,1920]" captionTargetId="figure@9.[178,1409,181,1920]" captionTargetPageId="9" captionText="FIGURES 24–29. 24, 26, 28, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 25, 27, 29, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 24–25, mesosternum and metasternum, dorsal view; 26–27, abdominal ventrites I–V, dorsal view; 28–29, abdominal ventrite V, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662555" httpUri="https://zenodo.org/record/3662555/files/figure.png" pageId="8" pageNumber="87">24</figureCitation>
). Male abdominal ventrite V short and transverse (
<figureCitation id="E4BF8C85FFFC910EFAE022F127E7800A" box="[1305,1425,1520,1542]" captionStart="FIGURES 24–29. 24, 26, 28" captionStartId="9.[151,264,1945,1970]" captionTargetBox="[178,1402,181,1920]" captionTargetId="figure@9.[178,1409,181,1920]" captionTargetPageId="9" captionText="FIGURES 24–29. 24, 26, 28, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 25, 27, 29, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 24–25, mesosternum and metasternum, dorsal view; 26–27, abdominal ventrites I–V, dorsal view; 28–29, abdominal ventrite V, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662555" httpUri="https://zenodo.org/record/3662555/files/figure.png" pageId="8" pageNumber="87">Figs. 26, 28</figureCitation>
), ratio of its length to width 0.40, and distinctly wider than width of pygidium in apical view. Antennomere IX in male shorter, only 1.19x longer than wide (
<figureCitation id="E4BF8C85FFFC910EFDFA212A2049804D" box="[515,575,1579,1601]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="8" pageNumber="87">Fig. 4</figureCitation>
). 
<emphasis id="4EF04C12FFFC910EFDAA212A2081804C" box="[595,759,1579,1600]" italics="true" pageId="8" pageNumber="87">Male terminalia:</emphasis>
tegminal plate wider and robust, its apex broadened (
<figureCitation id="E4BF8C85FFFC910EFAFF212A2734804D" box="[1286,1346,1579,1601]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 6</figureCitation>
). Tectum with very long process, longer than apex of pedon (
<figureCitation id="E4BF8C85FFFC910EFD2C214921498052" box="[725,831,1608,1630]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Figs. 7, 10</figureCitation>
). Sternite IX (= spiculum gastrale) touching sternite VIII at its posterior margin (
<figureCitation id="E4BF8C85FFFC910EFE77216423BE8070" box="[398,456,1637,1660]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 9</figureCitation>
) .................................................... 
<emphasis id="4EF04C12FFFC910EFBBC216727EA8077" bold="true" box="[1093,1436,1637,1659]" pageId="8" pageNumber="87">
<taxonomicName id="BB84EB83FFFC910EFBBC2167273C8077" authority="Tryzna &amp; Banar" authorityName="Tryzna &amp; Banar" authorityYear="2020" box="[1093,1354,1637,1659]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="4EF04C12FFFC910EFBBC216726ED8077" bold="true" box="[1093,1179,1637,1659]" italics="true" pageId="8" pageNumber="87">H. loebli</emphasis>
Trýzna &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="55C3F169FFFC910EFAA9216727EA8077" box="[1360,1436,1638,1659]" pageId="8" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="7C3B9000FFFC910EFF6E218227EB81AD" blockId="8.[151,1437,1373,1953]" pageId="8" pageNumber="87">
(2) Specimens more slender, ratio of body length to maximum width 
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male, 
<quantity id="BB7C3DE5FFFC910EFC2A218226608095" box="[979,1046,1667,1689]" metricMagnitude="-2" metricUnit="m" metricValue="6.858" pageId="8" pageNumber="87" unit="in" value="2.7">2.70 in</quantity>
female (
<figureCitation id="E4BF8C85FFFC910EFB91218226D58095" box="[1128,1187,1667,1689]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="8" pageNumber="87">Fig. 5</figureCitation>
). Pronotum slightly transverse (
<figureCitation id="E4BF8C85FFFC910EFED821A1231180BA" box="[289,359,1696,1718]" captionStart="FIGURES 16–19. 16, 18" captionStartId="6.[151,264,1600,1625]" captionTargetBox="[151,1436,237,1575]" captionTargetId="figure@6.[151,1436,237,1575]" captionTargetPageId="6" captionText="FIGURES 16–19. 16, 18, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 17, 19, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 16–17, pronotum, dorsal view; 18–19, elytra, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="8" pageNumber="87">Fig. 17</figureCitation>
), ratio of its length to width at carina 
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male, 
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female. Elytra suboval (
<figureCitation id="E4BF8C85FFFC910EFB8F21A126CB80BA" box="[1142,1213,1696,1718]" captionStart="FIGURES 16–19. 16, 18" captionStartId="6.[151,264,1600,1625]" captionTargetBox="[151,1436,237,1575]" captionTargetId="figure@6.[151,1436,237,1575]" captionTargetPageId="6" captionText="FIGURES 16–19. 16, 18, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 17, 19, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 16–17, pronotum, dorsal view; 18–19, elytra, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662550" httpUri="https://zenodo.org/record/3662550/files/figure.png" pageId="8" pageNumber="87">Fig. 19</figureCitation>
), ratio of the maximum length to maximum width of elytra 
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male, 
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female. Elytral tubercles smaller, less prominent, in cross-section suboval. Subbasal, median and postmedian tubercles on the second elytral interval not prominent (see also from lateral view). Pronotal and elytral tubercles each with apex brownish. Lateral carina of pronotum very short to almost lacking. Prosternum and mesosternum with shallow dense microsculpture (
<figureCitation id="E4BF8C85FFFC910EFD0A201421338127" box="[755,837,1813,1835]" captionStart="FIGURES 20–23. 20, 22" captionStartId="7.[151,264,1545,1570]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure@7.[151,1436,181,1520]" captionTargetPageId="7" captionText="FIGURES 20–23. 20, 22, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 21, 23, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 20–21, head, ventral view; 22–23, prosternum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662552" httpUri="https://zenodo.org/record/3662552/files/figure.png" pageId="8" pageNumber="87">Figs. 23</figureCitation>
, 
<figureCitation id="E4BF8C85FFFC910EFCA92014211D8127" box="[848,875,1813,1835]" captionStart="FIGURES 24–29. 24, 26, 28" captionStartId="9.[151,264,1945,1970]" captionTargetBox="[178,1402,181,1920]" captionTargetId="figure@9.[178,1409,181,1920]" captionTargetPageId="9" captionText="FIGURES 24–29. 24, 26, 28, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 25, 27, 29, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 24–25, mesosternum and metasternum, dorsal view; 26–27, abdominal ventrites I–V, dorsal view; 28–29, abdominal ventrite V, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662555" httpUri="https://zenodo.org/record/3662555/files/figure.png" pageId="8" pageNumber="87">25</figureCitation>
). Male abdominal ventrite V weakly transverse (
<figureCitation id="E4BF8C85FFFC910EFABE2014228B8145" captionStart="FIGURES 24–29. 24, 26, 28" captionStartId="9.[151,264,1945,1970]" captionTargetBox="[178,1402,181,1920]" captionTargetId="figure@9.[178,1409,181,1920]" captionTargetPageId="9" captionText="FIGURES 24–29. 24, 26, 28, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 25, 27, 29, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 24–25, mesosternum and metasternum, dorsal view; 26–27, abdominal ventrites I–V, dorsal view; 28–29, abdominal ventrite V, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3662555" httpUri="https://zenodo.org/record/3662555/files/figure.png" pageId="8" pageNumber="87">Figs. 27, 29</figureCitation>
), ratio of its length to width 0.60, and slender, only slightly wider than width of pygidium in apical view. Antennomere IX in male longer, 1.44x longer than wide (
<figureCitation id="E4BF8C85FFFC910EFDB3205120F3816A" box="[586,645,1872,1894]" captionStart="FIGURES 4–5. 4" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[193,1318,681,1949]" captionTargetId="figure@4.[180,1406,669,1949]" captionTargetPageId="4" captionText="FIGURES 4–5. 4, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype, male; 5, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park." figureDoi="http://doi.org/10.5281/zenodo.3662545" httpUri="https://zenodo.org/record/3662545/files/figure.png" pageId="8" pageNumber="87">Fig. 5</figureCitation>
). 
<emphasis id="4EF04C12FFFC910EFD612051214D8169" box="[664,827,1872,1893]" italics="true" pageId="8" pageNumber="87">Male terminalia:</emphasis>
tegminal plate narrower, its apex thin, not broadened (
<figureCitation id="E4BF8C85FFFC910EFAB3205127E6816A" box="[1354,1424,1872,1894]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 11</figureCitation>
). Tectum strongly reduced, without process (
<figureCitation id="E4BF8C85FFFC910EFD79206C2082818F" box="[640,756,1901,1923]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Figs. 12, 15</figureCitation>
). Sternite IX (= spiculum gastrale) touching sternite VIII in the middle of its length (
<figureCitation id="E4BF8C85FFFC910EFE9B208A23DE81AD" box="[354,424,1931,1953]" captionStart="FIGURES 6–15" captionStartId="5.[151,264,1911,1936]" captionTargetBox="[197,1388,197,1883]" captionTargetId="figure@5.[189,1399,181,1885]" captionTargetPageId="5" captionText="FIGURES 6–15. Male terminalia. 6–10, Holophloeus loebli Trýzna &amp; Baňař sp. nov., holotype male; 11–15, Holophloeus tuberosus (Fairmaire, 1897), male from Mt. ďAmbre National Park; 6, 11, tegmen, dorsal view; 7, 12, aedeagus, ventral view; 8, 13, inner sac of aedeagus, latero-ventral view; 9, 14, segment VIII and sternite IX, ventral view; 10, 15, apex of aedeagus, dorsal view. Red arrows indicate important details (see Key). Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3662547" httpUri="https://zenodo.org/record/3662547/files/figure.png" pageId="8" pageNumber="87">Fig. 14</figureCitation>
)......................................................... 
<taxonomicName id="BB84EB83FFFC910EFB9D208A27EB81AD" authority="(Fairmaire, 1897)" authorityName="in Mt." baseAuthorityName="Fairmaire" baseAuthorityYear="1897" box="[1124,1437,1931,1953]" class="Insecta" family="Anthribidae" genus="Holophloeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="tuberosus">
<emphasis id="4EF04C12FFFC910EFB9D208A27EB81AD" bold="true" box="[1124,1437,1931,1953]" pageId="8" pageNumber="87">
<emphasis id="4EF04C12FFFC910EFB9D208A269581AD" bold="true" box="[1124,1251,1931,1953]" italics="true" pageId="8" pageNumber="87">H. tuberosus</emphasis>
(
<bibRefCitation id="1815EDF1FFFC910EFB16208A27E381AD" author="Fairmaire, L. M. H." box="[1263,1429,1931,1953]" pageId="8" pageNumber="87" pagination="164 - 204" refId="ref7170" refString="Fairmaire, L. M. H. (1897) Materiaux pour la faune coleopterique de la region malgache. 3 e note (suite). Annales de la Societe Entomologique de Belgique, 41, 164 - 204." type="journal article" year="1897">Fairmaire, 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>