198 lines
26 KiB
XML
198 lines
26 KiB
XML
<document id="691B777F0F2B0C7BAE403EE4C848B449" ID-DOI="http://dx.doi.org/10.3897/zookeys.764.24581" ID-GBIF-Dataset="a620743e-5023-417f-b5a9-00a281b10f6c" ID-PMC="PMC5997757" ID-Pensoft-Pub="1313-2970-764-111" ID-PubMed="29899676" ID-ZooBank="0983E67CC321498ABD95A2AF1038A6EE" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-764-111" ModsDocOrigin="ZooKeys 764" ModsDocTitle="Taxonomic revision proves Trachusapubescens (Morawitz, 1872) sensu lato to be a complex of allopatric and sympatric species in South-Eastern Europe and Western Asia (Hymenoptera, Apoidea, Anthidiini)" checkinTime="1528216734164" checkinUser="pensoft" docAuthor="Kasparek, Max" docDate="2018" docId="F41C3B316870B20C67392D43C9EF2D6F" docLanguage="en" docName="ZooKeys 764: 111-144" docOrigin="ZooKeys 764" docSource="http://dx.doi.org/10.3897/zookeys.764.24581" docTitle="Trachusa pubescens Morawitz 1872" docType="treatment" docVersion="6" lastPageNumber="111" masterDocId="7A49D127FFBF5569B868FF95FFBFFFF4" masterDocTitle="Taxonomic revision proves Trachusapubescens (Morawitz, 1872) sensu lato to be a complex of allopatric and sympatric species in South-Eastern Europe and Western Asia (Hymenoptera, Apoidea, Anthidiini)" masterLastPageNumber="144" masterPageNumber="111" pageNumber="111" updateTime="1701315234354" updateUser="plazi">
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<mods:title id="A8DC2922FCC0FE66531F67BD724C2E3D">Taxonomic revision proves Trachusapubescens (Morawitz, 1872) sensu lato to be a complex of allopatric and sympatric species in South-Eastern Europe and Western Asia (Hymenoptera, Apoidea, Anthidiini)</mods:title>
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<mods:namePart id="F593290EF1CE4D3403B3E51C918BB137">Kasparek, Max</mods:namePart>
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<mods:date id="4607DC416813F626A85E236BFBECCB2E">2018</mods:date>
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<mods:number id="98E014185447167CB3ADC94849D5EE0F">764</mods:number>
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<mods:classification id="7DD9D27BA86E359ECD1DE13D16DFADC0">journal article</mods:classification>
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<mods:identifier id="95612C7F7E1737727B1DB034ED7D289E" type="DOI">http://dx.doi.org/10.3897/zookeys.764.24581</mods:identifier>
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<treatment id="F41C3B316870B20C67392D43C9EF2D6F" ID-GBIF-Taxon="145906848" LSID="urn:lsid:plazi:treatment:F41C3B316870B20C67392D43C9EF2D6F" httpUri="http://treatment.plazi.org/id/F41C3B316870B20C67392D43C9EF2D6F" lastPageNumber="111" pageId="0" pageNumber="111">
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<subSubSection id="48D63D973EAB2ACC7E220E54A96F11BC" pageId="0" pageNumber="111" type="nomenclature">
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<paragraph id="30C46AC3CD6776881A4AFFE392C0C887" pageId="0" pageNumber="111">
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<taxonomicName id="C3F068A5053F6A4CD7216DD114A8E0C1" ID-CoL="57PMR" authority="Morawitz, 1872" authorityName="Morawitz" authorityYear="1872" class="Insecta" family="Megachilidae" genus="Trachusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachusa pubescens" order="Hymenoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="pubescens">Trachusa pubescens (Morawitz, 1872)</taxonomicName>
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Figs 11, 12, 14, 15
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</paragraph>
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</subSubSection>
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<subSubSection id="D031B811B71A166ECD610FA28B529B12" pageId="0" pageNumber="111" type="reference_group">
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<paragraph id="101C341CDEABDD6655E4FDC94AD46DC3" pageId="0" pageNumber="111">
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<taxonomicName id="8E788F0AFF046C3A9C2B7D072131DFF7" class="Insecta" family="Megachilidae" genus="Anthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthidium pubescens" order="Hymenoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="pubescens">Anthidium pubescens</taxonomicName>
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<bibRefCitation id="10F17F8953FD41B7333F05E3A4B69F01" author="Morawitz, F" journalOrPublisher="Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva)" pageId="0" pageNumber="111" pagination="45 - 63" title="Neue suedrussische Bienen." volume="9" year="1872">Morawitz 1872</bibRefCitation>
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: 59-60 [Note: While the date of publication is given in many publications as 1873, it has now been accepted as 27.vi.1872. See
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<bibRefCitation id="A9EDDE69D522A195456AC7B9FBDC9B1A" author="Pesenko, YA" journalOrPublisher="Denisia" pageId="0" pageNumber="111" pagination="1 - 616" title="Annotated bibliography of Russian and Soviet publications on the bees (Hymenoptera: Apoidea; excluding Apis mellifera): 1771 - 2002." volume="11" year="2003">Pesenko and Astafurova (2003)</bibRefCitation>
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].
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</paragraph>
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<paragraph id="F12AEF8E20A1EBC0B185CDFA042DEBD6" pageId="0" pageNumber="111">
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<taxonomicName id="13E51E13E30BF3A17305A0C1AF282521" class="Insecta" family="Megachilidae" genus="Anthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthidium pubescens" order="Hymenoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="pubescens">Anthidium pubescens</taxonomicName>
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<bibRefCitation id="8546471EA0D4DF55C329C1C00EB6F74B" author="Morawitz, F" journalOrPublisher="Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva)" pageId="0" pageNumber="111" pagination="129 - 189" title="Die Bienen Daghestans." volume="10" year="1874">Morawitz (1874)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="479516CF4A2605B694D66A6B8171CC50" pageId="0" pageNumber="111" type="lectotype">
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<paragraph id="D04779E41B377F33E70C457138992C3F" pageId="0" pageNumber="111">Lectotype.</paragraph>
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<paragraph id="96B85EC2256CE59456A71BF07D5015D3" pageId="0" pageNumber="111">Male, Derbent, Dagestan, Russia (ZISP). Photographs provided by Yu. Astafurova.</paragraph>
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<paragraph id="34FC8EAF06D0C922589787C187EA620B" pageId="0" pageNumber="111">
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The species was described on the basis of two males collected in "Hab. in Caucaso, Derbent" (photographs of original labels in
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<bibRefCitation id="FF7F1F1321A425CB7F7630FC128E47A7" author="Proshchalykin, MYu" journalOrPublisher="Far Eastern Entomologist" pageId="0" pageNumber="111" pagination="1 - 34" title="The species-group names of bees (Hymenoptera: Apoidea, Apiformes) described from Crimea, North Caucasus, European part of Russia and Ural. Part II. Families Andrenidae and Megachilidae." volume="328" year="2017">Proshchalykinet al. 2017</bibRefCitation>
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). There are two males in ZISP from this locality, which corresponds to the original description of
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<bibRefCitation id="04D001871A529F4589B9BECA1ABC0236" author="Morawitz, F" journalOrPublisher="Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva)" pageId="0" pageNumber="111" pagination="45 - 63" title="Neue suedrussische Bienen." volume="9" year="1872">Morawitz (1872</bibRefCitation>
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,
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<bibRefCitation id="C680EBA2FCCA4D31A98FD47052BD8598" author="Morawitz, F" journalOrPublisher="Horae Societatis Entomologicae Rossicae (Trudy Russkago entomologicheskago obshchestva)" pageId="0" pageNumber="111" pagination="129 - 189" title="Die Bienen Daghestans." volume="10" year="1874">1874</bibRefCitation>
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). One of these males was designated as lectotype by Proshchalykin and Astafurova in 2016 (
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<bibRefCitation id="3C340F8CAA3B204EB75BB3A803ED670D" author="Proshchalykin, MYu" journalOrPublisher="Far Eastern Entomologist" pageId="0" pageNumber="111" pagination="1 - 34" title="The species-group names of bees (Hymenoptera: Apoidea, Apiformes) described from Crimea, North Caucasus, European part of Russia and Ural. Part II. Families Andrenidae and Megachilidae." volume="328" year="2017">Proshchalykin et al. 2017</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="F1D45E52703829A03FD95D3A62F164C9" pageId="0" pageNumber="111" type="materials_examined">
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<paragraph id="51DD4D06EA394CAC9EE348D296CD3509" pageId="0" pageNumber="111">Material examined.</paragraph>
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<paragraph id="6CDC55978E8825A897DA0D28F3DE85AC" pageId="0" pageNumber="111">
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Females have been assigned to
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<taxonomicName id="CF07AC79878A2CBE1CB691612BBC554D" lsidName="T. pubescens" pageId="0" pageNumber="111" rank="species" species="pubescens">T. pubescens</taxonomicName>
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on the basis of the location of the collection area (as long as no other species of the species group are known to occur in this region: Crimea, southern Greece, northern Iran) or because they were collected together with males which were positively identified as
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<taxonomicName id="0F7170F8B6C4FF84B33B2FED749F8968" lsidName="T. pubescens" pageId="0" pageNumber="111" rank="species" species="pubescens">T. pubescens</taxonomicName>
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(Turkey). GREECE: 2♀, 6♂, Alt-Korinth (ancient Corinth), Peloponnes, 03.06.1964, M. Schwarz leg. (6ex. cMS, 2ex. cMAV); 1♀, same location, 21.05.1964, 1♀, same location, 23.05.1962, M. Schwarz leg. (cMS); 1♀, same location, 01.06.1974, M. Schwarz leg. (cMAV); 1♂, Alt-Korinth (ancient Corinth), Solomos, Peloponnes, 24.05.1964, M. Schwarz leg. (cMS); 1♂, Athens, 02.06.1962, H. Hunder leg. (cMS); 2♀, Corinth, Peloponnes, 24.05.1962, M. Schwarz leg. (1 ex. cMAV, 1 ex. cMS); 1♀, Chelmos, Kalavrita, Peloponnes, 31.05.1962, H. Hamann leg. (cMS); 2♂, Chalkis (Euboea), V.1926, Holtz leg. (ZMB); 1♂, Hellas Alt-Korinth, 21.06.1996, W. Arens leg. (cWA); 1♂, ibid., 07.06.1997, W. Arens leg. (cWA). IRAN: 2♂, Elburs, Polour 22 km N Ab Ali, 13.-14.7.1965, A. G. Soika & G. A. Mavromoustakis leg. (OLL); 3♀, ibid., 11.07.1965, A. G. Soika & G. A. Mavromoustakis leg. (OLL); 1♀, Tehran, 5.-8.5.1972, H. Bytinski-Salz leg. (OLL); 3♂, Elburs, Ilekah road 4 km above Pol-e Zanguleh, 2450 m, 14.7.1967, Baker leg. (SEMC, Baker collection); 1♀, 1♂, Kakan, Yasouj, 9.07.2009 (cAM). MACEDONIA (Former Yugoslav Republic): 1♀, 1♂, Prilep, 01.06.1965, K. Warncke leg. (OLL). RUSSIA: 1♂ (paralectotype), Derbent, Dagestan (ZISP). Photographs provided by Yu. Astafurova. TURKEY: 1♀, 3♂, Turkey ("Asia Minor"), 1890 (OLL, ZMB); 1♀, 1♂, Ankara, 16.06.1934, A. Seitz leg., "Anthidium laticeps F. Mor. det. J. D. Alfken 1934" (ZMB); 1♂, ibdid., 21.06.1934, A. Seitz leg., "Archianthidium maximum Friese Pasteels det. 1967" (SMF); 1♂, ibid., 24.06.1934, A. Seitz leg., "A. pubescens Pasteels det. 1976" (SMF); 1♂, ibid., 25.06.1934, A. Seitz leg., "A. laticeps / Archianthidium pubescens J. Pasteels det. 1976" (SMF); 3♂, Ankara: 40 km W of Ayas, 26.06.1998, J. Halada leg. (cMS); 5♂, Isparta:
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<normalizedToken id="E6DD1713245C2B53487446C74A647806" originalValue="Karakuş">Karakus</normalizedToken>
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<normalizedToken id="D004E64035801B0CCCA809F2EAA3E0FD" originalValue="Dağı">Dagi</normalizedToken>
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Centr. (
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<geoCoordinate id="BCFF2B73A1313046CA75468D2220723B" direction="north" orientation="latitude" precision="925" value="38.25">38°15'N</geoCoordinate>
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,
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<geoCoordinate id="31C26CE0F673D8957281C7395A598211" direction="east" orientation="longitude" precision="925" value="30.65">30°39'E</geoCoordinate>
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), 1460 m, 11.07.2006, J. Halada leg. (cMS); 2♂, Denizli: 35 km SSE Denizli (
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<geoCoordinate id="7EB0FF9150616A9AB8118001CF329102" direction="north" orientation="latitude" precision="925" value="37.616665">37°37'N</geoCoordinate>
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,
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<geoCoordinate id="07216B8F50A5C8839FCCB3D6189E7847" direction="east" orientation="longitude" precision="925" value="29.283333">29°17'E</geoCoordinate>
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) 970 m, 05.07.2006, J. Halada leg.; 1♀, 2♂, 28 km SSE
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<normalizedToken id="064193374119B66457ACA0D6E3C7DEEE" originalValue="Kütahya">Kuetahya</normalizedToken>
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(
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<geoCoordinate id="8E743EE63445DF6635FC68F0E491100B" direction="north" orientation="latitude" precision="925" value="39.216667">39°13'N</geoCoordinate>
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,
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<geoCoordinate id="7DEB4D84E12AB8B68DD92C7BA324FEC8" direction="east" orientation="longitude" precision="925" value="30.133333">30°08'E</geoCoordinate>
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) 1110 m, 12.07.2006, J. Halada leg. (cMS); 3♀, 4♂, Tatvan, Van
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<normalizedToken id="BDD3CA10E8E8B8E7FF6B62D65E2662B4" originalValue="Gölü">Goelue</normalizedToken>
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, 01.07.2000, M. Halada leg. (cMS); 2♀, 3♂, Bitlis, Nemrut
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<normalizedToken id="075587E54134DC3D89E3F9F07F47CA81" originalValue="Dağı">Dagi</normalizedToken>
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(2000 m); 28.07.1986, I. Blank leg. (OLL); 2♀, 1♂, Siirt: 10 km S, 23.-24.6.1985, M. Schwarz leg. (cMS); 2♀, 1♂, Adiyaman:
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<normalizedToken id="9395DF15C013D7B92DF05A3EF333C6CD" originalValue="Gölbaşı">Goelbasi</normalizedToken>
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, 21.06.1985, M. Schwarz leg. (cMS); 1♂,
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<normalizedToken id="5D93883FD112B4F75AB7DAB6C2BFB0DC" originalValue="Hakkâri">Hakkari</normalizedToken>
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: 16 km SE
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<normalizedToken id="0B9FF910EE4DC27E76BE483877A6BBBD" originalValue="Yüksekova">Yueksekova</normalizedToken>
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(1700 m), 28.06.1985, M. Schwarz leg. (cMS); 1♂,
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<normalizedToken id="189BF9CA5FE0AD61C79E0BD8CF95A8CE" originalValue="Hakkâri">Hakkari</normalizedToken>
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: Sat Mountain S
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<normalizedToken id="1668F29202C08AE7F35CBD6D7F17E8C4" originalValue="Varegös">Varegoes</normalizedToken>
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(2000 m), 06.08.1982, K. Warncke leg. (OLL). TURKMENISTAN: 1♂, West Kopet Dagh [
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<normalizedToken id="9A6D6985B7FADB36CC69BB69C55F5241" originalValue="Köpetdag">Koepetdag</normalizedToken>
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], Syunt Mts., 21.06.1953, O. Kryzhanovskiy leg. (ZISP). Photographs provided by Yu. Astafurova. UKRAINE: 1♂, Crimea, 03.08.1937 (SIZK); 1♂, Crimea (exact location illegible), before 1908 (SIZK); 1♂, Crimea (
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<normalizedToken id="A54B6A8B5D52CE075947A0F1D1FFD6DA" originalValue="“Tauria”">"Tauria"</normalizedToken>
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; exact location illegible), 22.06.1914, V. Pliginski leg., (cMAV); 2♂, Crimea (exact location illegible), without date (cMAV); 1♂, Crimea (
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<normalizedToken id="96AED1F7704B525248C9B3363D7EDFB5" originalValue="“Tauria”">"Tauria"</normalizedToken>
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), Sevastopol, 26.06.1911, V. Pliginski leg. (cMAV); 1♂, Crimea (
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<normalizedToken id="FE49B2DFBC8BBA0B44009F1915D53A9F" originalValue="“Tauria”">"Tauria"</normalizedToken>
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), 21.06.1909, V. Pliginski leg. (cMAV); 1♂, Crimea, Savastopol, 26.06.1986 (SIZK); 1♂, ibid., 08.07.1912, V. Pliginski leg. (cMAV); 1♂, Karadag, Crimea, 01.07.1919 (OLL); 1♀, ibid., 17.06.1923 (OLL); 1♂, ibid., 23.06.1925, Kistjakovsky leg. (SIZK); 1♂, ibid., 22.06.1929, J. Paramonum leg. (SIZK); 1♀, 1♂, Crimea, Feodossija, 17.06.1995, C. Ivanov leg. (SIZK).
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</paragraph>
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</subSubSection>
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<subSubSection id="5A491673D334D723F12E5D5F8C86607A" pageId="0" pageNumber="111" type="materials_examined">
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<paragraph id="F7EEA08AFD9903B92206D646B512C33D" pageId="0" pageNumber="111">Other material.</paragraph>
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<paragraph id="9F3070A4D0BBC034531566286862EE47" pageId="0" pageNumber="111">
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TURKEY: The following females from Turkey are attributed to this species as "Trachusa aff. pubescens": 1♀, Ankara, 22.06.1973, K. Warncke leg. (OLL); 1♀, Ankara:
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<normalizedToken id="5FDBD639F3526880DA5F6A25302A6D22" originalValue="Kızılcahamam">Kizilcahamam</normalizedToken>
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; 18.06.1985, M. Schwarz leg. (cMS); 1♀, Ankara: 10 km S Ankara, 05.06.1988, K. Warncke leg. (OLL); 1♀, Konya: 10 km S Karaman, 19.06.1985, M. Schwarz leg. (cMS); 1♀, Konya: Sille; 08.06.1972, J. Heinrich leg. (SMF); 1♀,
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<normalizedToken id="836269475E760406EED846CFD6FF72CD" originalValue="Akşehir">Aksehir</normalizedToken>
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, 07.1934, (OLL), "A. interruptum Pasteels det. 1967 / A. pubescens det. Warncke"; 2♀,
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<normalizedToken id="4CFD1F38D7054D792F73E85D97FCDE66" originalValue="Akşehir">Aksehir</normalizedToken>
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(
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<normalizedToken id="987D188277CA07675B04E24A2E7078B0" originalValue="“Ak-Chehir“">"Ak-Chehir"</normalizedToken>
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), 1900, Korb leg. (OLL); 3♀,
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<normalizedToken id="17B9D1E60E84E46EB3CB9E7F9C54B279" originalValue="Hakkâri">Hakkari</normalizedToken>
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: 19 km S
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<normalizedToken id="E5C90FF821DB572CD67EA1A21760AD0A" originalValue="Beyetüşşebap">Beyetuessebap</normalizedToken>
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(1200 m), 26.06.1985, M. Schwarz leg. (cMS); 1♀, Malatya: Erkenek 60 km SW Malatya (1300 m), 26.06.2000, M. Halada leg. (cMS); 1♀,
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<normalizedToken id="09BD4AAF92E83524816B5D7952D1EE76" originalValue="Hakkâri">Hakkari</normalizedToken>
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: 16 km SE
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<normalizedToken id="5341BEEE624F88A6B80BCFB4C88E12E2" originalValue="Yüksekova">Yueksekova</normalizedToken>
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(1700 m), 28.06.1985, M. Schwarz leg. (cMS); 1♀,
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<normalizedToken id="F16940D62DBCEBBC4148B609E33DA353" originalValue="Hakkâri">Hakkari</normalizedToken>
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,
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<normalizedToken id="94A687E146C5B518715730860A2F7B01" originalValue="Suvari-Halil-Paß">Suvari-Halil-Pass</normalizedToken>
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<normalizedToken id="2CD3EDB7842D80077045EB32D040642A" originalValue="östl">oestl</normalizedToken>
|
||
.
|
||
<normalizedToken id="9471D8D02E8FD86B6E1C27EBC689D56D" originalValue="Beytüşşebap">Beytuessebap</normalizedToken>
|
||
(2300 m), 03.08.1982, K. Warncke leg. (OLL).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE01048E310B4ED361D5BFA195E2B4DE" pageId="0" pageNumber="111" type="differential diagnosis">
|
||
<paragraph id="D4B89E860E894738A15822020C997278" pageId="0" pageNumber="111">Differential diagnosis.</paragraph>
|
||
<paragraph id="0AA17E937491C0D9C74FCD6D387C1A09" pageId="0" pageNumber="111">The male is characterised by the following morphological features: Apical margin of clypeus straight or only slightly curved inwards and shallowly crenulated, usually without individually discernible tubercles; lateral projections of T6 subacute, apex of median projection emarginate (variable between a shallowly emarginate and a narrow V-shaped incision); punctation of T6 dense and coarse; median projection of T7 broadened toward apex. In some specimens in which the median emargination of T6 is inconspicuous or absent, the median projection of T7 is widened. For identification at least one of these two characters should hold true. Margins of yellow maculations on abdominal terga normally irregularly ragged.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB967A854FF417C66B6F73923AD05E0" pageId="0" pageNumber="111" type="variability">
|
||
<paragraph id="DC88E982AFBD78AFA91ED7907C96B107" pageId="0" pageNumber="111">Variability.</paragraph>
|
||
<paragraph id="21222850F7B4114937A2A95CA1F0C31F" pageId="0" pageNumber="111">Four OUs were distinguished: Greece (including one from FYR of Macedonia), Crimea, Anatolia, and northern Iran. Members of the Greek and Crimean OUs are characterised by yellow maculation in the genal area and a dark vertex. The genal maculation usually extends from the top of the eye to the lower end of the eye but is sometimes irregular (margins ragged or maculations broken). Only one specimen from the Crimea has yellow on the vertex. In the Anatolian OU, however, the yellow maculation of the genal area usually extends up to the middle of the vertex and merges, or at least the lateral bands are contiguous. There is, however, some variation within populations and, for example, specimens with merged maculation and with widely separated maculations can be found together. The colour pattern of northern Iranian specimens is in general close to the Greek and Crimean OUs but one specimen with subcontiguous bands is available.</paragraph>
|
||
<paragraph id="744A1B20D62221EC847CB576F5A817CA" pageId="0" pageNumber="111">The yellow colouration on T2 in the Crimean, Greek and northern Iranian OUs is confined to lateral bands far apart from each other, whereas in the Anatolian OU the gap between the lateral bands is narrower and two small yellow spots are normally situated between them.</paragraph>
|
||
<paragraph id="13BC7A09AD520590E63FC20EF7ABAA92" pageId="0" pageNumber="111">Specimens of the northern Iranian OU are smaller than those in the other populations. Despite a small sample size (N = 5), the Iranian specimens proved to be significantly smaller than Anatolian specimens in 22 of 27 morphological features (at least p <0.05; t-test). Anatolian specimens in turn do not differ in size from Crimean and Greek specimens.</paragraph>
|
||
<paragraph id="0696D5237BB8A50A35B67F6AE993905A" pageId="0" pageNumber="111">
|
||
A Discriminant Analysis (DA) was carried out to find out whether the OUs attributed to
|
||
<taxonomicName id="B2EB8D3700777AA1DF66CAD5FC3C674B" lsidName="T. pubescens" pageId="0" pageNumber="111" rank="species" species="pubescens">T. pubescens</taxonomicName>
|
||
s. str. are different from each other in morphometric characters. Fig. 4 shows that the specimens from Greece, Crimea, and Anatolia form three distinctive clusters. In their morphometric characters northern Iranian specimens are between the Greek and Anatolian specimens but their separation is less clear. Nevertheless, in a confusion matrix, which summarises the reclassification of the observations and enables us to see quickly the percentage of well-classified observations, all specimens (100%) of all four groups were correctly classified.
|
||
</paragraph>
|
||
<paragraph id="1E9842459FA0B3731865FBDC1C34F52E" pageId="0" pageNumber="111">
|
||
Altogether, the populations of
|
||
<taxonomicName id="4F379C27A1F3A656B742E423049826ED" lsidName="T. pubescens" pageId="0" pageNumber="111" rank="species" species="pubescens">T. pubescens</taxonomicName>
|
||
s. str. from Greece, the Crimea, Anatolia, and northern Iran can be separated based on a set of morphological features. Northern Iranian specimens are smaller than all others. The colouration pattern of the vertex and gena enables most Anatolian specimens to be distinguished from the others, but due to variation within populations this feature is not regarded here as being of taxonomic significance. The pattern of the yellow colouration on the first three terga is on average different in the Anatolian population from the other OUs. All these features could justify giving the OUs subspecies rank. However, the holotype of
|
||
<taxonomicName id="48E944F730AA8FF35B698193C66E899C" lsidName="T. pubescens" pageId="0" pageNumber="111" rank="species" species="pubescens">T. pubescens</taxonomicName>
|
||
could not be examined and, although the description is detailed enough to allow unambiguous species attribution within the
|
||
<taxonomicName id="5A8767ACD58BFF88B312B55E707C26B3" lsidName="T. pubescens" pageId="0" pageNumber="111" rank="species" species="pubescens">T. pubescens</taxonomicName>
|
||
species group, it is not detailed enough to decide whether the colouration and/or morphometric features agree with one of the OUs described here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2435DAB1118FE0B9B07941608D88B76B" pageId="0" pageNumber="111" type="flower preference">
|
||
<paragraph id="58B15C1BD18FA15C4B0209AA8A9E9343" pageId="0" pageNumber="111">Flower preference.</paragraph>
|
||
<paragraph id="810E06D44D5CA74D4C32090360094603" pageId="0" pageNumber="111">No information available.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CDE50CADC62198C327FDFE1C13A0036D" pageId="0" pageNumber="111" type="distribution">
|
||
<paragraph id="26D96370A808A752A5B6F6D513827D79" pageId="0" pageNumber="111">Distribution.</paragraph>
|
||
<paragraph id="F3AD305F92C76AC20E1C96FDC0B1CC8F" pageId="0" pageNumber="111">
|
||
The distribution area extends from Crimea and southern Greece over Anatolia and the Caucasus to the Kopet Dagh Mountain in Turkmenistan. In the south, the distribution area extends into the Zagross Mountains of Iran. Possible hybridisation with
|
||
<taxonomicName id="9B805074D68C60E73003D05716971946" lsidName="T. balcanica" pageId="0" pageNumber="111" rank="species" species="balcanica">T. balcanica</taxonomicName>
|
||
sp. n. takes place in the F.Y.R. Macedonia.
|
||
</paragraph>
|
||
<paragraph id="B28EA0200F2AE63EDED3564F34C994BA" pageId="0" pageNumber="111">
|
||
<bibRefCitation id="91FFA9090070129B20BEA0A59B9AAF34" author="Mocsary, A" journalOrPublisher="Magyar Tudomanyos Akademia, Mathematikai es Termeszettudomanyi Koezlemenyek" pageId="0" pageNumber="111" pagination="1 - 70" title="Ujabb Adatok Temesmegye Hartyaroepű Faunajahoz." volume="16" year="1879">
|
||
<normalizedToken id="BA054BD7A96913B375B3418DB405F8A8" originalValue="Mocsáry">Mocsary</normalizedToken>
|
||
(1879
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="5793A68BD863E9D0F471BD82EAB551F1" author="Mocsary, A" journalOrPublisher="Termeszetrajzi Fuezetek Kiadija a Magyar Nemzeti Muezeum [Naturhistorische Hefte herausgegeben vom Ungarischen National-Museum]" pageId="0" pageNumber="111" pagination="241 - 278" title="Species generis Anthidium Fabr. regionis Palaearcticae." volume="8" year="1884">1884</bibRefCitation>
|
||
) describes material from Hungary with an emarginate T6. While this character alone does not allow unambiguous species identification, it may be interpreted as an indication that the distribution of
|
||
<taxonomicName id="0C66CA4CFD5251EEC17E9FA3C9247608" lsidName="T. pubescens" pageId="0" pageNumber="111" rank="species" species="pubescens">T. pubescens</taxonomicName>
|
||
may extend further north than indicated by the records presented in this paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |