169 lines
19 KiB
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169 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.463.8499" ID-PMC="PMC4294300" ID-Pensoft-Pub="1313-2970-463-21" ID-Pensoft-UUID="3E5CFFC6FFC91628FFF6E355951DFFF8" ID-PubMed="25589861" ID-Zenodo-Dep="578654" ID-ZooBank="71C1AE92B0CE4D26B1E5076E480180AD" ModsDocID="1313-2970-463-21" checkinTime="1451244962410" checkinUser="pensoft" docAuthor="Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. & Titarenko, Andrey Y." docDate="2014" docId="F374DCCD1AA3511AB7B1CFB44FB5BFA6" docLanguage="en" docName="ZooKeys 463: 21-56" docOrigin="ZooKeys 463" docPubDate="2014-12-12" docSource="http://dx.doi.org/10.3897/zookeys.463.8499" docTitle="Carabus (Archiplectes) besleticus subsp. mtsaranus Kurnakov 1972" docType="treatment" docVersion="3" id="3E5CFFC6FFC91628FFF6E355951DFFF8" lastPageNumber="29" masterDocId="3E5CFFC6FFC91628FFF6E355951DFFF8" masterDocTitle="Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="56" masterPageNumber="21" pageNumber="27" updateTime="1668159733340" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Solodovnikov, Igor A.</mods:namePart>
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<mods:affiliation>Vitebsk State P. M. Masherov University, Moskovskyi prospect, 33, Vitebsk 210038, Belarus</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zamotajlov, Alexandr S.</mods:namePart>
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<mods:affiliation>Kuban State Agrarian University, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">a_zamotajlov@mail.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fominykh, Dmitriy D.</mods:namePart>
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<mods:affiliation>Russian Entomological Society, Kuban Branch, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Titarenko, Andrey Y.</mods:namePart>
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<mods:affiliation>Publicly Traded Company " Morpho Absoloni ", Bolotnikovskaya St. 5 / 3, Moscow 117556, Russia</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-12-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>463</mods:number>
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<mods:extent unit="page">
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<mods:start>21</mods:start>
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<mods:end>56</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.463.8499</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.463.8499</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-463-21</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578654</mods:identifier>
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<treatment ID-GBIF-Taxon="182234815" LSID="urn:lsid:plazi:treatment:F374DCCD1AA3511AB7B1CFB44FB5BFA6" httpUri="http://treatment.plazi.org/id/F374DCCD1AA3511AB7B1CFB44FB5BFA6" lastPageId="8" lastPageNumber="29" pageId="6" pageNumber="27">
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<subSubSection pageId="6" pageNumber="27" type="nomenclature">
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<paragraph pageId="6" pageNumber="27">
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<taxonomicName LSID="F374DCCD-1AA3-511A-B7B1-CFB44FB5BFA6" authority="Kurnakov, 1972" authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus Kurnakov, 1972</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="6" pageNumber="27">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="6" pageNumber="27">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="6" pageNumber="27">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="6" pageNumber="27">, 4</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="6" pageNumber="27">, 7</figureCitation>
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<figureCitation captionStart="Figures 16–21" captionStartId="F9" captionText="Figures 16 - 21. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) besleticus besleticus and Carabus (Archiplectes) besleticus mtsaranus. 16 - 18 Carabus (Archiplectes) besleticus besleticus 19 - 21 Carabus (Archiplectes) besleticus mtsaranus." pageId="6" pageNumber="27">, 19-21</figureCitation>
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<figureCitation captionStart="Figures 50–57" captionStartId="F14" captionText="Figures 50 - 57. Dorsal habitus, Carabus (Archiplectes) besleticus mtsaranus, Abkhazia, Zashirbara Mt. Range, valley of River Mtsara." pageId="6" pageNumber="27">, 50-57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="28" type="reference_group">
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<paragraph pageId="7" pageNumber="28">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Neoplectes) satyrus subsp. mtsaranus" order="Coleoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Neoplectes" subSpecies="mtsaranus">
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<pageBreakToken pageId="7" pageNumber="28" start="start">Carabus</pageBreakToken>
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(Neoplectes) satyrus mtsaranus
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</taxonomicName>
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Kurnakov, 1972: 114 (
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<normalizedToken originalValue="“Chaîne">"Chaine</normalizedToken>
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Bzybienne
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<normalizedToken originalValue="près">pres</normalizedToken>
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du village de Mtsara").
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</paragraph>
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<paragraph pageId="7" pageNumber="28">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus subsp. mtsaranus" order="Coleoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) satyrus mtsaranus</taxonomicName>
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:
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<bibRefCitation author="Gottwald, J" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="23" pageNumber="44" pagination="278 - 314" refId="B5" refString="Gottwald, J, 1985. Revision der Untergattung Archiplectes Gottwald (Plectes auct.) der Gattung Carabus (Coleoptera, Carabidae). Acta Entomologica Bohemoslovaca 824: 278 - 314" title="Revision der Untergattung Archiplectes Gottwald (Plectes auct.) der Gattung Carabus (Coleoptera, Carabidae)." volume="824" year="1985">Gottwald 1985</bibRefCitation>
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: 310;
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="Catalogue of Palaearctic Coleoptera" pageId="23" pageNumber="44" pagination="118 - 201" refId="B2" refString="Bousquet, Y, Brezina, B, Davies, A, Farkac, J, Smetana, A, 2003. Tribe Carabini Latreille, 1802. Catalogue of Palaearctic Coleoptera Vol. 1: 118 - 201" title="Tribe Carabini Latreille, 1802." volume="Vol. 1" year="2003">Bousquet et al. 2003</bibRefCitation>
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: 132;
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<bibRefCitation author="Retezar, I" journalOrPublisher="Mesterprint Kft., Budapest" pageId="23" pageNumber="44" refId="B8" refString="Retezar, I, 2008. The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics. Mesterprint Kft., Budapest" title="The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics." year="2008">
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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2008
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</bibRefCitation>
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: 41.
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</paragraph>
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<paragraph pageId="7" pageNumber="28">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) satyrus subsp. mtsaranus" order="Coleoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Tribax" subSpecies="mtsaranus">Carabus (Tribax) satyrus mtsaranus</taxonomicName>
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:
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<bibRefCitation author="Deuve, T" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="23" pageNumber="44" refId="B3" refString="Deuve, T, 2004. Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae). Pensoft Publishers, Sofia-Moscow" title="Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae)." year="2004">Deuve 2004</bibRefCitation>
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: 274.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="28" type="comparative material examined">
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<paragraph pageId="7" pageNumber="28">Comparative material examined.</paragraph>
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<paragraph pageId="7" pageNumber="28">134 specimens were examined (31 specimens measured, 18 male genitalia preparations studied): 12 males, 17 females Abkhazia, Gudauta Distr., valley of River Aapsta, 700 m, V-VII.1993, leg. A. Zamotajlov, F. Miroshnikov (cZAM, cFDD); 49 males, 56 females, Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova, V. Kotsur, N. Pichugin (cSIA, cKVM, cPSM, cPNYu, cFDD).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="28" type="description">
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<paragraph pageId="7" pageNumber="28">Description.</paragraph>
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<paragraph pageId="7" pageNumber="28">Large form, males 35.3-37.1 (35.8) mm, females 37.5-41.3 (39.3) mm long, robust. Underside black, dorsum normally with bright metallic lustre, dark blue, violet, black-violet, black, rather seldom green, bronze, crimson, females matte. Body massive.</paragraph>
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<paragraph pageId="7" pageNumber="28">
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Head not inflated. Pronotum of variable shape, from subcordate to cordate, transverse. PW/PL = 1.14-1.48 (1.35) in males and 1.26-1.56 (1.41) in females, with base narrower than in the nominotypical subspecies, PW/PB = 1.15-1.26 (1.20) in males and 1.13-1.32 (1.23) in females, hind angles strongly protruding backwards and sidewards, seldom only backwards, pointed apically. Median groove distinct, disk transversally or irregularly rugose, with rugosity gradually strengthening towards median groove. Elytra strongly elongate to ovate, more elongate in males, with quite inconspicuous depression in the middle in females and seldom in males. EL/EW = 1.65-1.85 (1.75) in males and 1.73-1.98 (1.80) in females. Elytral sculpture identical in males and females, forming coarse and precise series of short links. Table
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<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric characteristic of males (n = 16) and females (n = 31) of Carabus (Archiplectes) satyrus and males (n = 83) and females (n = 91) of Carabus (Archiplectes) besleticus subspecies." httpUri="http://table.plazi.org/id/E3AA304075ED326595338A0A005BF64B" pageId="7" pageNumber="28" tableUuid="E3AA304075ED326595338A0A005BF64B">2</tableCitation>
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lists the main morphometric measurements of the studied populations.
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</paragraph>
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<paragraph pageId="7" pageNumber="28">
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The shape of the male genitalia is practically identical to that of the other
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
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taxa. The endophallus differs from the one in
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<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
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, mainly in the shape of the dorsal appendix, which is more elongate and possesses a more extended form, characteristic for populations dwelling in an interfluve of r. Kealasur and Bzyb.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="29" pageId="7" pageNumber="28" type="differential diagnosis">
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<paragraph pageId="7" pageNumber="28">Differential diagnosis.</paragraph>
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<paragraph lastPageId="8" lastPageNumber="29" pageId="7" pageNumber="28">
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Resembles
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<taxonomicName authority="besleticus" authorityName="besleticus Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. besleticus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="besleticus">
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<pageBreakToken pageId="8" pageNumber="29" start="start">Carabus</pageBreakToken>
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(Archiplectes) besleticus besleticus
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</taxonomicName>
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from which it differs in a more elongate and less ovate body shape, the different form of the hind angles of the pronotum, and in having a more coarse elytral sculpture with precise series of rather short links. The color of dorsum is usually more dull and darker than in the nominotypical and other subspecies. For the morphometric characters of this subspecies see Figures
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="8" pageNumber="29">1</figureCitation>
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-
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="8" pageNumber="29">4</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="29" type="distribution">
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<paragraph pageId="8" pageNumber="29">Distribution.</paragraph>
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<paragraph pageId="8" pageNumber="29">Populates southern slopes of Bzybian Mt. Range at an altitude of 400-1500 m in an interfluve of r. Gumista - Western Gumista and Khipsta.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="29" type="habitat">
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<paragraph pageId="8" pageNumber="29">Habitat.</paragraph>
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<paragraph pageId="8" pageNumber="29">
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Prefers beech and beech-chestnut forest sites, sometimes admixed with box, rich in ground litter at karstic landforms (sometimes even populating also quite sharp and steep ones). This subspecies also occurs at alpestrine and alpine meadows. Activity of imago, depending upon the altitude of the habitation, proceeds from April to August.
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<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) apollo subsp. tenebricosus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="apollo" subGenus="Archiplectes" subSpecies="tenebricosus">Carabus (Archiplectes) apollo tenebricosus</taxonomicName>
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Kurnakov, 1962,
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<taxonomicName authorityName="Mandl" authorityYear="1955" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Microplectes) argonautarum subsp. reischitzi" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="argonautarum" subGenus="Microplectes" subSpecies="reischitzi">Carabus (Microplectes) argonautarum reischitzi</taxonomicName>
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Mandl, 1955,
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<taxonomicName authority="apschuanus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) apschuanus subsp. apschuanus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="apschuanus" subGenus="Tribax" subSpecies="apschuanus">Carabus (Tribax) apschuanus apschuanus</taxonomicName>
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Rost, 1893,
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) constantinovi subsp. otcharensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="constantinovi" subGenus="Tribax" subSpecies="otcharensis">Carabus (Tribax) constantinovi otcharensis</taxonomicName>
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Kurnakov, 1970.
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<taxonomicName authorityName="Motschulsky" authorityYear="1840" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Megodontus) septemcarinatus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="septemcarinatus" subGenus="Megodontus">Carabus (Megodontus) septemcarinatus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Sphodristocarabus) armeniacus subsp. dvorschaki" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="armeniacus" subGenus="Sphodristocarabus" subSpecies="dvorschaki">Carabus (Sphodristocarabus) armeniacus dvorschaki</taxonomicName>
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Mandl, 1975, and
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procerus) caucasicus subsp. colchicus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subGenus="Procerus" subSpecies="colchicus">Carabus (Procerus) caucasicus colchicus</taxonomicName>
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occur together with this subspecies.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |