432 lines
70 KiB
XML
432 lines
70 KiB
XML
<document id="4A7C3ADE9AAB98E8C2B3649717721DE1" ID-DOI="10.11646/zootaxa.3718.5.1" ID-GBIF-Dataset="d8493892-4f06-4e1a-b92a-f49f6adfb4e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="246828" ID-ZooBank="A34FF647-288B-4BD6-803C-0EADB27F2BC3" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460686746548" checkinUser="plazi" docAuthor="Bota-Sierra, Cornelio Andrés & Echeverri, Martha Isabel Wolff" docDate="2013" docId="E76CC555D006FFF3FF615F39F0BEF850" docLanguage="en" docName="zt03718p440.pdf" docOrigin="Zootaxa 3718 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesamphiagrion nataliae Bota-Sierra, sp. nov." docType="treatment" docVersion="8" lastPageNumber="430" masterDocId="1B55BD2DD01DFFEEFFF65845F35EFF8C" masterDocTitle="Taxonomic revision of Mesamphiagrion Kennedy, 1920 from Colombia (Odonata: Coenagrionidae), with the description of four new species" masterLastPageNumber="440" masterPageNumber="401" pageNumber="428" updateTime="1698351323746" updateUser="plazi">
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<mods:titleInfo id="059DF72EA658BAF730A3886C5A34F547">
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<mods:title id="227C4E6729681A3C9FBF1FA9E92C0E36">Taxonomic revision of Mesamphiagrion Kennedy, 1920 from Colombia (Odonata: Coenagrionidae), with the description of four new species</mods:title>
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<mods:name id="399D8F357D50406D0935186630B5210A" type="personal">
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<mods:namePart id="C8632C66888E9C5E91A69484629F439C">Bota-Sierra, Cornelio Andrés</mods:namePart>
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<mods:namePart id="D9A47626BDE021F86741FF947CF396EB">Echeverri, Martha Isabel Wolff</mods:namePart>
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<mods:typeOfResource id="1DF80294DB6AF7D972606BE257DDC2EA">text</mods:typeOfResource>
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<mods:title id="FAAE4BCFF2707980C8D285C640C1AC1F">Zootaxa</mods:title>
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<mods:date id="23BF0A7243D6FF35C2DD0C146A1E53A3">2013</mods:date>
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<mods:number id="73F3D3B65BB24529C79EECAF5E911D94">3718</mods:number>
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<treatment id="E76CC555D006FFF3FF615F39F0BEF850" ID-DOI="http://doi.org/10.5281/zenodo.5658567" ID-GBIF-Taxon="119580628" ID-Zenodo-Dep="5658567" LSID="urn:lsid:plazi:treatment:E76CC555D006FFF3FF615F39F0BEF850" httpUri="http://treatment.plazi.org/id/E76CC555D006FFF3FF615F39F0BEF850" lastPageId="29" lastPageNumber="430" pageId="27" pageNumber="428">
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<subSubSection id="27DF27C8D006FFF5FF615F39F648F857" pageId="27" pageNumber="428" type="nomenclature">
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<paragraph id="6F7A7443D006FFF5FF615F39F193F81A" blockId="27.[151,1436,1916,2011]" box="[151,717,1916,1942]" pageId="27" pageNumber="428">
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<heading id="3432C32FD006FFF5FF615F39F193F81A" bold="true" box="[151,717,1916,1942]" fontSize="11" level="1" pageId="27" pageNumber="428" reason="1">
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<emphasis id="5DB1A851D006FFF5FF615F39F193F81A" bold="true" box="[151,717,1916,1942]" pageId="27" pageNumber="428">
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<taxonomicName id="A8C50FC0D006FFF5FF615F39F139F81A" ID-CoL="9X3JL" authority="Bota-Sierra" authorityName="Bota-Sierra" box="[151,615,1916,1942]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="nataliae" status="sp. nov.">
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<emphasis id="5DB1A851D006FFF5FF615F39F290F81A" bold="true" box="[151,462,1916,1942]" italics="true" pageId="27" pageNumber="428">Mesamphiagrion nataliae</emphasis>
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Bota-Sierra
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</taxonomicName>
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<taxonomicNameLabel id="4682152AD006FFF5FD995F39F193F81A" box="[623,717,1916,1942]" pageId="27" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6F7A7443D006FFF5FF615FDBF648F857" blockId="27.[151,1436,1916,2011]" pageId="27" pageNumber="428">
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<figureCitation id="F7FE68C6D006FFF5FF615FDBF3BAF83B" box="[151,228,1950,1975]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="27" pageNumber="428">Figs. 9</figureCitation>
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a–e (diagnostic traits), 10a (habitus ♂), 10b (habitus androchrome ♀), 10c (habitus juvenile androchrome ♀), 10f (habitus juvenile ♂),
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(habitus gynochrome ♀), 13n (S7–
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<specimenCount id="79C3BFCAD006FFF5FC605F86F0ECF857" box="[918,946,1987,2011]" pageId="27" pageNumber="428" type="generic">10</specimenCount>
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♂), 14c, f (S7–10 ♀), 15 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="27DF27C8D001FFF2FF6158D2F03FFEE8" pageId="28" pageNumber="429" type="etymology">
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<paragraph id="6F7A7443D001FFF2FF6158D2F1D8FF59" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="429">
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<emphasis id="5DB1A851D001FFF2FF6158D2F240FF3C" bold="true" box="[151,286,151,176]" pageId="28" pageNumber="429">Etymology.</emphasis>
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Named
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<taxonomicName id="A8C50FC0D001FFF2FE8A58D2F286FF3C" box="[380,472,151,176]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="nataliae">
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<emphasis id="5DB1A851D001FFF2FE8A58D2F286FF3C" box="[380,472,151,176]" italics="true" pageId="28" pageNumber="429">nataliae</emphasis>
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</taxonomicName>
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(noun in the genitive case) after Dr. Natalia von Ellenrieder, whose work, advice, and unconditional help made this work possible.
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</paragraph>
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<paragraph id="6F7A7443D001FFF2FF31589AF03FFEE8" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="429">
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<emphasis id="5DB1A851D001FFF2FF31589AF2E0FF74" bold="true" box="[199,446,223,248]" pageId="28" pageNumber="429">Specimens examined</emphasis>
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(
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<specimenCount id="79C3BFCAD001FFF2FE3B58A5F137FF74" box="[461,617,223,248]" pageId="28" pageNumber="429" type="generic">24 specimens</specimenCount>
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: male
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<typeStatus id="B07ECAE1D001FFF2FD46589AF04BFF74" box="[688,789,223,248]" pageId="28" pageNumber="429" type="holotype">holotype</typeStatus>
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, allotype, and 22
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<typeStatus id="B07ECAE1D001FFF2FC2F58A5F717FF74" box="[985,1097,224,248]" pageId="28" pageNumber="429" type="paratype">paratypes</typeStatus>
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). Cundinamarca, Municipality Guasca:
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<specimenCount id="79C3BFCAD001FFF2FF0B5941F27EFE90" box="[253,288,260,284]" pageId="28" pageNumber="429" type="generic">9♂</specimenCount>
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and 3♀, Township El Salitre,
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<geoCoordinate id="0AF11284D001FFF2FD665941F181FE91" box="[656,735,260,285]" direction="north" orientation="latitude" pageId="28" pageNumber="429" precision="925" value="4.8333335">4°50'N</geoCoordinate>
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<geoCoordinate id="0AF11284D001FFF2FD1D5941F013FE91" box="[747,845,260,285]" direction="west" orientation="longitude" pageId="28" pageNumber="429" precision="925" value="-73.916664">73°55'W</geoCoordinate>
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, 2700M,
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<date id="1B7B5283D001FFF2FC375941F71AFE91" box="[961,1092,260,285]" pageId="28" pageNumber="429" value="2006-02-04">04/02/2006</date>
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, Leg: E. Realpe & L. Pérez, ANDES.
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<specimenCount id="79C3BFCAD001FFF2FEF4596DF27BFECC" box="[258,293,296,320]" pageId="28" pageNumber="429" type="generic">3♂</specimenCount>
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and 1♀, same data but CEUA.
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<typeStatus id="B07ECAE1D001FFF2FD705962F1AFFECC" box="[646,753,295,320]" pageId="28" pageNumber="429" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="79C3BFCAD001FFF2FD0A5962F041FECC" box="[764,799,295,320]" pageId="28" pageNumber="429" type="generic">5♂</specimenCount>
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and 2♀, Township Pueblo Viejo, Finca El Aji,
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<geoCoordinate id="0AF11284D001FFF2FAD9596DF6C2FECC" box="[1327,1436,296,320]" direction="north" orientation="latitude" pageId="28" pageNumber="429" precision="5" value="4.7934">4.7934°N</geoCoordinate>
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<geoCoordinate id="0AF11284D001FFF2FF615909F265FEE8" box="[151,315,332,356]" direction="west" orientation="longitude" pageId="28" pageNumber="429" precision="1" value="-73.923134">73.923131° W</geoCoordinate>
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,
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<quantity id="A83DD9A6D001FFF2FEB05909F2C9FEE9" box="[326,407,332,357]" metricMagnitude="3" metricUnit="m" metricValue="3.05" pageId="28" pageNumber="429" unit="m" value="3050.0">3050m</quantity>
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,
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<date id="1B7B5283D001FFF2FE545909F17AFEE8" box="[418,548,332,356]" pageId="28" pageNumber="429" value="2009-02-14">14/02/2009</date>
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, Leg: M. Sánchez, ANDES.
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</paragraph>
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</subSubSection>
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<subSubSection id="27DF27C8D001FFF3FF31592AF157FBF0" lastPageId="29" lastPageNumber="430" pageId="28" pageNumber="429" type="description">
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<paragraph id="6F7A7443D001FFF2FF31592AF163FD94" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="429">
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<emphasis id="5DB1A851D001FFF2FF31592AF17EFE04" bold="true" box="[199,544,367,392]" pageId="28" pageNumber="429">
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Description.
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<typeStatus id="B07ECAE1D001FFF2FEA9592AF290FE04" box="[351,462,367,392]" pageId="28" pageNumber="429" type="holotype">Holotype</typeStatus>
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. Head.
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</emphasis>
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Labium cream with light blue base. Base of mandible blue. Labrum blue with a basal small black spot and dorsolateral black margins. Gena blue. Anteclypeus blue. Postclypeus black. Frons black with pruinescence, light blue on area surrounding eye, gena, and clypeus, with medial transverse black stripe over clypeus, blue postocular spots reaching eye margin, and cream spot between ocelli and antenna. Antenna black. Back of head cream (
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<figureCitation id="F7FE68C6D001FFF2FE2659BAF17CFD94" box="[464,546,511,536]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="28" pageNumber="429">Fig. 10</figureCitation>
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a).
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</paragraph>
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<paragraph id="6F7A7443D001FFF2FF315A61F38EFC44" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="429">
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<emphasis id="5DB1A851D001FFF2FF315A61F278FDB1" bold="true" box="[199,294,548,573]" pageId="28" pageNumber="429">Thorax.</emphasis>
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Prothorax black, propleura blue becoming lighter to proximal portion. Medial lobe of posterior prothoracic lobe developed into a caudally projected squarish plate dorsally concave, lateral edges approximately half length of distal edge. Pterothorax black with blue antehumeral stripe, wide blue stripe covering mesepimeron and metepisternum, metepimeron light blue with pruinescence, and cream venter. Meta- and mesocoxae cream, procoxa blue. External side of femur black with small cream spot near base, medial side cream. Tibia external side black and medial side brown. Tarsi and claws brown (
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<figureCitation id="F7FE68C6D001FFF2FD035A92F016FD7C" box="[757,840,727,752]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="28" pageNumber="429">Fig. 10</figureCitation>
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a). Nine external metafemoral spurs, as long as space between them or shorter, gradually increasing in size toward apex. Six external metatibial spurs, as long as space between them or shorter, gradually decreasing in size toward apex. Tarsal claws with well developed supplementary tooth. Wings hyaline. Pt dark brown, ratio between distal and proximal sides about 1:1. CuP reaching CuPAA slightly distal to confluence of CuPAA with hind margin of wing. 14 Px in right and
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<quantity id="A83DD9A6D001FFF2FAF55B2DF662FC0C" box="[1283,1340,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="28" pageNumber="429" unit="in" value="12.0">12 in</quantity>
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left FW,
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<quantity id="A83DD9A6D001FFF2FF615BC9F391FC29" box="[151,207,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="28" pageNumber="429" unit="in" value="11.0">11 in</quantity>
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HW.
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<collectingCountry id="17D234D3D001FFF2FEF95BC9F26CFC28" box="[271,306,908,932]" name="Poland" pageId="28" pageNumber="429">RP</collectingCountry>
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2 branching between Px 4 and
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<quantity id="A83DD9A6D001FFF2FD7D5BC9F1EBFC29" box="[651,693,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="28" pageNumber="429" unit="in" value="5.0">5 in</quantity>
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left FW and under
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<quantity id="A83DD9A6D001FFF2FC795BC9F0E4FC29" box="[911,954,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="28" pageNumber="429" unit="in" value="5.0">5 in</quantity>
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right FW, between Px 4 and
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<quantity id="A83DD9A6D001FFF2FB095BC9F674FC29" box="[1279,1322,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="28" pageNumber="429" unit="in" value="5.0">5 in</quantity>
|
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HW (
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||
<figureCitation id="F7FE68C6D001FFF2FA985BC9F3EBFC44" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="28" pageNumber="429">Fig. 10</figureCitation>
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a).
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</paragraph>
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<paragraph id="6F7A7443D001FFF2FF315B91F1AAFA10" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="429">
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<emphasis id="5DB1A851D001FFF2FF315B91F21DFC61" bold="true" box="[199,323,980,1005]" pageId="28" pageNumber="429">Abdomen.</emphasis>
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Dorsum black and sides cream except: S1 distal margin and side blue. S2 side bluish. S4–7 with light blue ring on proximal margin. S7 with a dorsal blue spot on distal 9/10. S8–9 with blue dorsum. S10 with pale blue apical projections (
|
||
<figureCitation id="F7FE68C6D001FFF2FE555C59F2A1FBB8" box="[419,511,1052,1077]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="28" pageNumber="429">Figs. 10</figureCitation>
|
||
a, 12n). Genital ligula (
|
||
<figureCitation id="F7FE68C6D001FFF2FCF65C59F01CFBB9" box="[768,834,1052,1077]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="28" pageNumber="429">Fig. 9</figureCitation>
|
||
a–b) with apex concave, with pair of long lateroapical processes extending proximally and terminating in sharp point, also pair of short processes pointing distally about as long as 1/4 length of lateroapical processes and located basal to them, lateromedial lobes hemispherical in lateral view, not visible in ectal view, and inner medial process close to ligula’s flexure. Cercus (
|
||
<figureCitation id="F7FE68C6D001FFF2FB795CC2F78CFB2C" box="[1167,1234,1159,1184]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="28" pageNumber="429">Fig. 9</figureCitation>
|
||
c–d) external side black, medial side light blue, about 1/2 length of S10, with elongated dorsal process ending in small tooth, external ventroapical process pointed and shorter than dorsal process, inner ventroapical process extending beyond ventroapical external process and terminating in rounded tip which is most distal point of body in lateral view; in posterior view this process is curved outwards and covers the external ventroapical process. Basal process underdeveloped, extending to level of apex of paraproct and ending in blunt tip. Paraproct cream with dorsal process ending in black tip pointing distally, reaching about mid-length of cercus. Total length
|
||
<quantity id="A83DD9A6D001FFF2FB395D25F67CFAF4" box="[1231,1314,1376,1400]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="28" pageNumber="429" unit="mm" value="31.0">31 mm</quantity>
|
||
. Abdomen length
|
||
<quantity id="A83DD9A6D001FFF2FF125DC1F268FA10" box="[228,310,1412,1437]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="429" unit="mm" value="25.0">25 mm</quantity>
|
||
. FW length
|
||
<quantity id="A83DD9A6D001FFF2FE485DC1F14EFA10" box="[446,528,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="429" unit="mm" value="18.0">18 mm</quantity>
|
||
. HW length
|
||
<quantity id="A83DD9A6D001FFF2FD6B5DC1F1AEFA10" box="[669,752,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="28" pageNumber="429" unit="mm" value="17.0">17 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D001FFF2FF315DE2F60CF984" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="429">
|
||
<emphasis id="5DB1A851D001FFF2FF315DE2F145FA4C" bold="true" box="[199,539,1447,1472]" pageId="28" pageNumber="429">
|
||
Variation in male
|
||
<typeStatus id="B07ECAE1D001FFF2FE685DE2F148FA4C" box="[414,534,1447,1472]" pageId="28" pageNumber="429" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
As
|
||
<typeStatus id="B07ECAE1D001FFF2FDBF5DE2F1F5FA4C" box="[585,683,1447,1472]" pageId="28" pageNumber="429" type="holotype">holotype</typeStatus>
|
||
but seven to nine metafemoral external spurs. Six to eight external metatibial spurs. 12–14 Px in FW, 10–11Px in HW.
|
||
<collectingCountry id="17D234D3D001FFF2FD2A5D89F1A1FA68" box="[732,767,1484,1508]" name="Poland" pageId="28" pageNumber="429">RP</collectingCountry>
|
||
2 branching between Px 4 and
|
||
<quantity id="A83DD9A6D001FFF2FBAE5D89F7DAFA69" box="[1112,1156,1484,1509]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="28" pageNumber="429" unit="in" value="6.0">6 in</quantity>
|
||
FW, between Px 3 and
|
||
<quantity id="A83DD9A6D001FFF2FA785D89F3F3F984" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="28" pageNumber="429" unit="in" value="5.0">5 in</quantity>
|
||
HW. Total length
|
||
<quantity id="A83DD9A6D001FFF2FE8D5DB5F2A9F984" box="[379,503,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="28" pageNumber="429" unit="mm" value="30.0" valueMax="31.0" valueMin="29.0">29–31 mm</quantity>
|
||
. Abdomen length
|
||
<quantity id="A83DD9A6D001FFF2FD325DAAF01EF984" box="[708,832,1519,1544]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="28" pageNumber="429" unit="mm" value="24.0" valueMax="25.0" valueMin="23.0">23–25 mm</quantity>
|
||
. FW length
|
||
<quantity id="A83DD9A6D001FFF2FC3E5DB5F71AF984" box="[968,1092,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="28" pageNumber="429" unit="mm" value="17.5" valueMax="18.0" valueMin="17.0">17–18 mm</quantity>
|
||
. HW length
|
||
<quantity id="A83DD9A6D001FFF2FB275DAAF613F984" box="[1233,1357,1519,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="28" pageNumber="429" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D001FFF2FF315E51F2CAF9F9" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="429">
|
||
<emphasis id="5DB1A851D001FFF2FF315E51F153F9A1" bold="true" box="[199,525,1556,1581]" pageId="28" pageNumber="429">Variation in male juveniles</emphasis>
|
||
. Head and thorax, light blue instead of blue, brown instead of black (
|
||
<figureCitation id="F7FE68C6D001FFF2FAC65E51F6DDF9A1" box="[1328,1411,1556,1581]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="28" pageNumber="429">Fig. 10</figureCitation>
|
||
f). Coxae and legs cream. Pt brown. Dorsum of S1–3 red, but distal margin of S1 blue, distal margin of S2 black, and distal half of S3 black.
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D001FFF2FF315E3AF76AF889" blockId="28.[151,1436,151,2013]" pageId="28" pageNumber="429">
|
||
<emphasis id="5DB1A851D001FFF2FF315E3AF272F914" bold="true" box="[199,300,1663,1688]" pageId="28" pageNumber="429">Allotype</emphasis>
|
||
.
|
||
<emphasis id="5DB1A851D001FFF2FECB5E3AF2DAF914" bold="true" box="[317,388,1663,1688]" pageId="28" pageNumber="429">Head.</emphasis>
|
||
As
|
||
<typeStatus id="B07ECAE1D001FFF2FE405E3AF146F914" box="[438,536,1663,1688]" pageId="28" pageNumber="429" type="holotype">holotype</typeStatus>
|
||
but labium cream. Base of mandible brown. Labrum brown with small black midbasal spot and black dorsolateral margins. Postclypeus and anteclypeus brown, with pruinescence. Frons dark brown with light brown stripe crossing from eye to eye, proximal to vertex and surrounding occipital margin, blue postocular spots not reaching eye margin, approximately hemispherical (
|
||
<figureCitation id="F7FE68C6D001FFF2FC335EA9F749F889" box="[965,1047,1772,1797]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="28" pageNumber="429">Fig. 10</figureCitation>
|
||
b).
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D001FFF3FF315F4AF002FF74" blockId="28.[151,1436,151,2013]" lastBlockId="29.[151,1437,151,2013]" lastPageId="29" lastPageNumber="430" pageId="28" pageNumber="429">
|
||
<emphasis id="5DB1A851D001FFF2FF315F4AF278F8A4" bold="true" box="[199,294,1807,1832]" pageId="28" pageNumber="429">Thorax.</emphasis>
|
||
Anterior lobe of pronotum light blue. Medial lobe of posterior prothoracic lobe developed into caudally projected squarish plate concave dorsally, its lateral edge about half as long as distal edge (
|
||
<figureCitation id="F7FE68C6D001FFF2FACF5F71F6DEF8C0" box="[1337,1408,1844,1869]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="28" pageNumber="429">Fig. 9</figureCitation>
|
||
e). Mesepisternal plate brown approximately flat and triangular, carina extending from distal tip of mesostigmal plate to middorsal suture on each side (
|
||
<figureCitation id="F7FE68C6D001FFF2FDB45F39F1D0F818" box="[578,654,1916,1941]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="28" pageNumber="429">Fig. 9</figureCitation>
|
||
e). Pterothorax brown with pruinescence, middorsal stripe black, mesepisternum with light bluish brown stripe, wide light blue stripe on mesepimeron and distal metepisternum, metepisternum cream (
|
||
<figureCitation id="F7FE68C6D001FFF2FE6C5F81F2B3F851" box="[410,493,1988,2013]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="28" pageNumber="429">Fig. 10</figureCitation>
|
||
b). Coxa cream. External side of remainder of leg brown, medial side cream, femur with a small pale spot near apex. Seven external spurs on right metafemur and six on left. Six external spurs on right metatibia and seven on left. Pt brown. 12 Px in FW,
|
||
<quantity id="A83DD9A6D000FFF3FCEB58F9F008FF59" box="[797,854,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="29" pageNumber="430" unit="in" value="10.0">10 in</quantity>
|
||
HW.
|
||
<collectingCountry id="17D234D3D000FFF3FC6158F9F0E4FF58" box="[919,954,188,212]" name="Poland" pageId="29" pageNumber="430">RP</collectingCountry>
|
||
2 branching between Px 5 and
|
||
<quantity id="A83DD9A6D000FFF3FAE558F9F661FF59" box="[1299,1343,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="29" pageNumber="430" unit="in" value="6.0">6 in</quantity>
|
||
left FW and under
|
||
<quantity id="A83DD9A6D000FFF3FEFB589AF266FF74" box="[269,312,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="29" pageNumber="430" unit="in" value="6.0">6 in</quantity>
|
||
right FW, between Px 4 and
|
||
<quantity id="A83DD9A6D000FFF3FD8B589AF1F6FF74" box="[637,680,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="29" pageNumber="430" unit="in" value="5.0">5 in</quantity>
|
||
HW (
|
||
<figureCitation id="F7FE68C6D000FFF3FD1B589AF061FF74" box="[749,831,223,248]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="29" pageNumber="430">Fig. 10</figureCitation>
|
||
b).
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D000FFF3FF315941F7C9FD9B" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315941F21DFE91" bold="true" box="[199,323,260,285]" pageId="29" pageNumber="430">Abdomen.</emphasis>
|
||
S1–4 with pruinescence, dorsum black and sides greenish cream except: S1 distal margin blue. S4– 7 with incomplete light blue ring on proximal margin, S7 with a blue dorsal spot occupying about 2/3 of dorsum, located proximally and without reaching margins of S7, S8 with pair of dorsal rounded blue spots near proximal margin. S9 with thin pale blue middorsal line that does not reach margins of S9. S10 with medial triangular spot running from proximal edge to middle of segment (
|
||
<figureCitation id="F7FE68C6D000FFF3FD2B59D1F064FE21" box="[733,826,404,429]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="29" pageNumber="430">Figs. 10</figureCitation>
|
||
b, 13c). Vulvar spine cream with black tip. Ovipositor and paraproct cream, cercus brown. Subbasal plate of ovipositor triangular. Ovipositor distal apex reaching level of distal margin of S10. Cercus conical, as long as about half length of S10, its apex representing most distal point of body. Total length
|
||
<quantity id="A83DD9A6D000FFF3FE9E5A45F2E4FD9B" box="[360,442,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="430" unit="mm" value="30.0">30 mm</quantity>
|
||
. Abdomen length
|
||
<quantity id="A83DD9A6D000FFF3FD715A45F187FD9B" box="[647,729,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="29" pageNumber="430" unit="mm" value="23.0">23 mm</quantity>
|
||
. FW length
|
||
<quantity id="A83DD9A6D000FFF3FC945A45F0EAFD9B" box="[866,948,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="430" unit="mm" value="20.0">20 mm</quantity>
|
||
. HW length
|
||
<quantity id="A83DD9A6D000FFF3FBB75A45F7CDFD9B" box="[1089,1171,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="430" unit="mm" value="18.0">18 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D000FFF3FF315A61F761FD7C" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315A61F1C8FDB1" bold="true" box="[199,662,548,573]" pageId="29" pageNumber="430">
|
||
Variation in mature female
|
||
<typeStatus id="B07ECAE1D000FFF3FDEE5A61F1CEFDB1" box="[536,656,548,573]" pageId="29" pageNumber="430" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
As allotype but entire body can be covered in pruinescence.
|
||
<emphasis id="5DB1A851D000FFF3FAA35A61F6C2FDB1" bold="true" box="[1365,1436,548,573]" pageId="29" pageNumber="430">Head.</emphasis>
|
||
Base of mandible cream to greenish gray. Labrum cream to gray greenish with small black mediobasal indentation and dorsolateral black margins. Gena cream to gray. Anteclypeus gray and postclypeus brown along apical edge and gray along mediobasal margin. Antefrons dark brown to light green, postfrons with brown stripe from eye to eye proximal to vertex and surrounding it to occipital margin, or completely light brown with dark brown rings surrounding ocelli, postocular spots almost indiscernible, very light blue (
|
||
<figureCitation id="F7FE68C6D000FFF3FC265A92F77DFD7C" box="[976,1059,727,752]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="29" pageNumber="430">
|
||
Fig.
|
||
<quantity id="A83DD9A6D000FFF3FBF35A9DF77DFD7C" box="[1029,1059,728,752]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="29" pageNumber="430" unit="g" value="10.0">10</quantity>
|
||
</figureCitation>
|
||
g).
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D000FFF3FF315AB9F395FC28" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315AB9F278FC99" bold="true" box="[199,294,764,789]" pageId="29" pageNumber="430">Thorax.</emphasis>
|
||
Prothorax brown, anterior lobe of pronotum light blue, propleura blue becoming lighter proximally. Pterothorax light brown with black middorsal stripe, mesepisternal light stripe brown or blue, mesepimeron and distal metepisternum with light green stripe, metepisternum cream (
|
||
<figureCitation id="F7FE68C6D000FFF3FC575B01F0A8FCD0" box="[929,1014,836,861]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="29" pageNumber="430">
|
||
Fig.
|
||
<quantity id="A83DD9A6D000FFF3FC2E5B01F0A8FCD0" box="[984,1014,836,860]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="29" pageNumber="430" unit="g" value="10.0">10</quantity>
|
||
</figureCitation>
|
||
g). Six to eight external metafemoral spurs. Six to eight external metatibial spurs. 12–14 Px in FW,
|
||
<quantity id="A83DD9A6D000FFF3FCA65B2DF0ECFC0C" box="[848,946,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="29" pageNumber="430" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10–11 in</quantity>
|
||
HW.
|
||
<collectingCountry id="17D234D3D000FFF3FC025B2DF749FC0C" box="[1012,1047,872,896]" name="Poland" pageId="29" pageNumber="430">RP</collectingCountry>
|
||
2 branching between Px 3 and
|
||
<quantity id="A83DD9A6D000FFF3FA875B22F6C2FC0C" box="[1393,1436,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="29" pageNumber="430" unit="in" value="5.0">5 in</quantity>
|
||
HW.
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D000FFF3FF315BEAF7A5FB83" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315BEAF21DFC44" bold="true" box="[199,323,943,968]" pageId="29" pageNumber="430">Abdomen.</emphasis>
|
||
Middorsal light blue line on S9 present or absent, light blue spots on S9–10 present or absent, or dorsum entirely black with only distal margin of segments pale (probably gynochrome females) (
|
||
<figureCitation id="F7FE68C6D000FFF3FB275B91F673FC60" box="[1233,1325,980,1005]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="29" pageNumber="430">
|
||
Figs.
|
||
<quantity id="A83DD9A6D000FFF3FAE65B91F673FC60" box="[1296,1325,980,1004]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="29" pageNumber="430" unit="g" value="10.0">10</quantity>
|
||
</figureCitation>
|
||
g, 13a–b). Total length
|
||
<quantity id="A83DD9A6D000FFF3FED25BBDF2FEFB83" box="[292,416,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="29" pageNumber="430" unit="mm" value="30.0" valueMax="31.0" valueMin="29.0">29–31 mm</quantity>
|
||
. Abdomen length
|
||
<quantity id="A83DD9A6D000FFF3FD9B5BBDF1B7FB83" box="[621,745,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="29" pageNumber="430" unit="mm" value="23.0" valueMax="24.0" valueMin="22.0">22–24 mm</quantity>
|
||
. FW length
|
||
<quantity id="A83DD9A6D000FFF3FC875BBDF0B3FB83" box="[881,1005,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="29" pageNumber="430" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 mm</quantity>
|
||
. HW length
|
||
<quantity id="A83DD9A6D000FFF3FB8D5BBDF7A9FB83" box="[1147,1271,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="29" pageNumber="430" unit="mm" value="17.5" valueMax="18.0" valueMin="17.0">17–18 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D000FFF3FF315C59F157FBF0" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315C59F1F5FBB9" bold="true" box="[199,683,1052,1077]" pageId="29" pageNumber="430">
|
||
Variation in immature female
|
||
<typeStatus id="B07ECAE1D000FFF3FDC35C59F1F5FBB9" box="[565,683,1052,1077]" pageId="29" pageNumber="430" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
. As allotype but light brown instead of dark brown or black, light green mesepisternal stripe, dorsum of S1–3 red except blue distal margin of S1, black distal margin of S2, and black distal half of S3 (
|
||
<figureCitation id="F7FE68C6D000FFF3FE6A5C21F2B0FBF0" box="[412,494,1124,1149]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="29" pageNumber="430">Fig. 10</figureCitation>
|
||
c).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="27DF27C8D000FFF3FF315CC2F2A6F931" pageId="29" pageNumber="430" type="diagnosis">
|
||
<paragraph id="6F7A7443D000FFF3FF315CC2F2A6F931" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315CC2F21FFB2C" bold="true" box="[199,321,1159,1184]" pageId="29" pageNumber="430">Diagnosis.</emphasis>
|
||
The dorsal process of male cercus longer than the ventrobasal external process (
|
||
<figureCitation id="F7FE68C6D000FFF3FB245CC2F64BFB2C" box="[1234,1301,1159,1184]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="29" pageNumber="430">Fig. 9</figureCitation>
|
||
c–d) groups
|
||
<taxonomicName id="A8C50FC0D000FFF3FF615CE8F247FB49" box="[151,281,1196,1221]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="nataliae">
|
||
<emphasis id="5DB1A851D000FFF3FF615CE8F247FB49" box="[151,281,1196,1221]" italics="true" pageId="29" pageNumber="430">M. nataliae</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<emphasis id="5DB1A851D000FFF3FEAD5CE8F7EEFB49" box="[347,1200,1196,1221]" italics="true" pageId="29" pageNumber="430">
|
||
<taxonomicName id="A8C50FC0D000FFF3FEAD5CE8F2B0FB48" box="[347,494,1197,1220]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="tamaense">M. tamaense</taxonomicName>
|
||
,
|
||
<taxonomicName id="A8C50FC0D000FFF3FE0D5CE8F1F8FB49" box="[507,678,1196,1221]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="demarmelsi">M. demarmelsi</taxonomicName>
|
||
,
|
||
<taxonomicName id="A8C50FC0D000FFF3FD455CE8F0C4FB49" box="[691,922,1196,1221]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="gaudiimontanum">M. gaudiimontanum</taxonomicName>
|
||
,
|
||
<taxonomicName id="A8C50FC0D000FFF3FC505CE8F741FB48" box="[934,1055,1197,1220]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="gaianii">M. gaianii</taxonomicName>
|
||
,
|
||
<taxonomicName id="A8C50FC0D000FFF3FBDD5CE8F7F2FB49" box="[1067,1196,1196,1221]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="laterale">M. laterale</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="A8C50FC0D000FFF3FB1E5CE8F63EFB49" box="[1256,1376,1196,1221]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="dunklei">
|
||
<emphasis id="5DB1A851D000FFF3FB1E5CE8F63EFB49" box="[1256,1376,1196,1221]" italics="true" pageId="29" pageNumber="430">M. dunklei</emphasis>
|
||
</taxonomicName>
|
||
. The well developed inner ventroapical process of male cercus, curving distally and representing the most distal point of the body (
|
||
<figureCitation id="F7FE68C6D000FFF3FEF85CB1F20DFA81" box="[270,339,1268,1293]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="29" pageNumber="430">Fig. 9</figureCitation>
|
||
c–d), clearly separates this species from
|
||
<emphasis id="5DB1A851D000FFF3FCDA5CB0F7FCFA80" box="[812,1186,1268,1293]" italics="true" pageId="29" pageNumber="430">
|
||
<taxonomicName id="A8C50FC0D000FFF3FCDA5CB0F74BFA81" box="[812,1045,1268,1293]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="gaudiimontanum">M. gaudiimontanum</taxonomicName>
|
||
,
|
||
<taxonomicName id="A8C50FC0D000FFF3FBD25CB0F7C0FA80" box="[1060,1182,1269,1292]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="gaianii">M. gaianii</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="A8C50FC0D000FFF3FB295CB0F601FA81" box="[1247,1375,1268,1293]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="laterale">
|
||
<emphasis id="5DB1A851D000FFF3FB295CB0F601FA81" box="[1247,1375,1268,1293]" italics="true" pageId="29" pageNumber="430">M. laterale</emphasis>
|
||
</taxonomicName>
|
||
. The shape of genital ligula (
|
||
<figureCitation id="F7FE68C6D000FFF3FE685D52F2BEFABC" box="[414,480,1303,1328]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="29" pageNumber="430">Fig. 9</figureCitation>
|
||
a–b) clearly separates it from all known species in the genus, resembling the ligula of
|
||
<taxonomicName id="A8C50FC0D000FFF3FF615D78F21AFAD9" box="[151,324,1340,1365]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="demarmelsi">
|
||
<emphasis id="5DB1A851D000FFF3FF615D78F21AFAD9" box="[151,324,1340,1365]" italics="true" pageId="29" pageNumber="430">M. demarmelsi</emphasis>
|
||
</taxonomicName>
|
||
but with lateroapical lobes longer and projected parallel to the body axis, versus shorter and projected oblique to the body axis in
|
||
<taxonomicName id="A8C50FC0D000FFF3FDC35D24F183FAF4" box="[565,733,1375,1400]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="demarmelsi">
|
||
<emphasis id="5DB1A851D000FFF3FDC35D24F183FAF4" box="[565,733,1375,1400]" italics="true" pageId="29" pageNumber="430">M. demarmelsi</emphasis>
|
||
</taxonomicName>
|
||
. The medial lobe of posterior prothoracic lobe relatively broad (
|
||
<figureCitation id="F7FE68C6D000FFF3FF695DC1F3BFFA10" box="[159,225,1412,1437]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="29" pageNumber="430">Fig. 9</figureCitation>
|
||
e) separates it from
|
||
<emphasis id="5DB1A851D000FFF3FE4B5DC0F133FA11" box="[445,621,1412,1437]" italics="true" pageId="29" pageNumber="430">
|
||
<taxonomicName id="A8C50FC0D000FFF3FE4B5DC0F137FA11" box="[445,617,1412,1437]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="demarmelsi">M. demarmelsi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The color pattern of mature males (
|
||
<figureCitation id="F7FE68C6D000FFF3FC0D5DC1F714FA10" box="[1019,1098,1412,1437]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="29" pageNumber="430">Fig. 10</figureCitation>
|
||
a) is almost identical to that of mature males of
|
||
<taxonomicName id="A8C50FC0D000FFF3FEA05DECF165FA4C" box="[342,571,1447,1472]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="gaudiimontanum">
|
||
<emphasis id="5DB1A851D000FFF3FEA05DECF165FA4C" box="[342,571,1447,1472]" italics="true" pageId="29" pageNumber="430">M. gaudiimontanum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F7FE68C6D000FFF3FDBD5DE2F1C7FA4C" box="[587,665,1447,1472]" captionStart="FIGURE 3" captionStartId="17.[151,250,1955,1977]" captionTargetBox="[204,1382,193,1920]" captionTargetId="figure@17.[204,1382,193,1921]" captionTargetPageId="17" captionText="FIGURE 3. Mesamphiagrion santainense (Antioquia, Belmira): (a) Male holotype; (b) Female paratype. M. gaudiimontanum (Antioquia, Belmira) Mature males: (c) Dorsal view; (d) Lateral view." httpUri="https://zenodo.org/record/246831/files/figure.png" pageId="29" pageNumber="430">Figs. 3</figureCitation>
|
||
c–d, 5b), but juvenile males of
|
||
<taxonomicName id="A8C50FC0D000FFF3FC085DECF7DEFA4C" box="[1022,1152,1447,1472]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="nataliae">
|
||
<emphasis id="5DB1A851D000FFF3FC085DECF7DEFA4C" box="[1022,1152,1447,1472]" italics="true" pageId="29" pageNumber="430">M. nataliae</emphasis>
|
||
</taxonomicName>
|
||
may have red coloration on S2–3 (
|
||
<figureCitation id="F7FE68C6D000FFF3FEF55D89F20AFA69" box="[259,340,1484,1509]" captionStart="FIGURE 10" captionStartId="25.[151,250,1798,1820]" captionTargetBox="[161,1423,197,1769]" captionTargetId="figure@25.[151,1436,193,1776]" captionTargetPageId="25" captionText="FIGURE 10. Mesamphiagrion nataliae (Cundinamarca, Guasca): (a) Male holotype; (f) Juvenile male paratype; (b) Androchrome female allotype; (g) Gynochrome female paratype; (c) Juvenile female paratype. (h) M. rosseri female paratype (Antioquia, Medellín). (d) M. ovigerum female (Cundinamarca, Bogotá) and (j) dorsal view of posterior lobe of pronotum and mesostigmal plates. (i) M. occultum female (Cundinamarca, PNN Chingaza) and (e) dorsal view of posterior lobe of pronotum and mesostigmal plates. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe." httpUri="https://zenodo.org/record/246838/files/figure.png" pageId="29" pageNumber="430">Fig. 10</figureCitation>
|
||
f, but not seen in life), whereas in
|
||
<taxonomicName id="A8C50FC0D000FFF3FD235D88F0E7FA69" box="[725,953,1484,1509]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="gaudiimontanum">
|
||
<emphasis id="5DB1A851D000FFF3FD235D88F0E7FA69" box="[725,953,1484,1509]" italics="true" pageId="29" pageNumber="430">M. gaudiimontanum</emphasis>
|
||
</taxonomicName>
|
||
there is red on S1–4. The posterior lobe of female pronotum, dorsally concave with distal edge about as long as 1.5x the length of a lateral edge, is unique among species of the genus (
|
||
<figureCitation id="F7FE68C6D000FFF3FE145E51F179F9A0" box="[482,551,1556,1581]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="29" pageNumber="430">Fig. 9</figureCitation>
|
||
e). Also unique is the carina extending from distal tip of mesostigmal plate to middorsal suture on each side (
|
||
<figureCitation id="F7FE68C6D000FFF3FDE15E72F13FF9DC" box="[535,609,1591,1616]" captionStart="FIGURE 9" captionStartId="24.[151,250,1856,1878]" captionTargetBox="[271,1310,193,1838]" captionTargetId="figure@24.[271,1315,193,1838]" captionTargetPageId="24" captionText="FIGURE 9. Mesamphiagrion nataliae (Cuandinamarca, Guasca): Male holotype: (a) Lateral view of genital ligula; (b) Ectal view of genital ligula; (c) Lateral view of caudal appendages; (d) Medio-dorsal view of right cercus. Female Paratype: (e) Dorsal view of posterior lobe of pronotum and mesostigmal plates. c. m.: Carina mesostigmal. p. m.: Mesostigmal plates. l. p.: Ponotum posterior lobe. l. l-ap.: Lateroapical lobe. l. l-ap. d.: Latero apical lobe distal process. l. l-m.: Lateromedial lobe. p. d.: Dorsal process. p. v-a. e.: Ventroapical external process. p. v-a. i.: Ventroapical inner process. p. v-b.: Ventrobasal process. p. i.: Innner process." httpUri="https://zenodo.org/record/246837/files/figure.png" pageId="29" pageNumber="430">Fig. 9</figureCitation>
|
||
e). Females present two color patterns: androchrome (
|
||
<figureCitation id="F7FE68C6D000FFF3FB195E72F618F9DC" box="[1263,1350,1591,1616]" captionStart="FIGURE 13. S 7 – 10" captionStartId="32.[151,250,1386,1408]" captionTargetBox="[378,1208,193,1364]" captionTargetId="figure@32.[378,1209,193,1364]" captionTargetPageId="32" captionText="FIGURE 13. S 7 – 10 color patterns of males: (a) and (b) Mesamphiagrion gaudiimontanum; (c) and (d) M. risi; (e) M. rosseri; (f) and (g) M. santainense; (h) M. tamaense; (i) and (j) M. occultum; (k) M. demarmelsi; (l) M. laterale; (m) M. ovigerum; (n) M. nataliae." httpUri="https://zenodo.org/record/246841/files/figure.png" pageId="29" pageNumber="430">Fig. 13</figureCitation>
|
||
c) with abdominal color pattern similar to females of
|
||
<taxonomicName id="A8C50FC0D000FFF3FD5E5E18F078F9F9" box="[680,806,1628,1653]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="laterale">
|
||
<emphasis id="5DB1A851D000FFF3FD5E5E18F078F9F9" box="[680,806,1628,1653]" italics="true" pageId="29" pageNumber="430">M. laterale</emphasis>
|
||
</taxonomicName>
|
||
, and gynochrome (
|
||
<figureCitation id="F7FE68C6D000FFF3FBF05E19F706F9F9" box="[1030,1112,1628,1653]" captionStart="FIGURE 13. S 7 – 10" captionStartId="32.[151,250,1386,1408]" captionTargetBox="[378,1208,193,1364]" captionTargetId="figure@32.[378,1209,193,1364]" captionTargetPageId="32" captionText="FIGURE 13. S 7 – 10 color patterns of males: (a) and (b) Mesamphiagrion gaudiimontanum; (c) and (d) M. risi; (e) M. rosseri; (f) and (g) M. santainense; (h) M. tamaense; (i) and (j) M. occultum; (k) M. demarmelsi; (l) M. laterale; (m) M. ovigerum; (n) M. nataliae." httpUri="https://zenodo.org/record/246841/files/figure.png" pageId="29" pageNumber="430">Fig. 13</figureCitation>
|
||
a–b), with abdomen entirely dark, which is shared with other gynochrome females in the genus, i.e.
|
||
<taxonomicName id="A8C50FC0D000FFF3FC365EC4F713F914" box="[960,1101,1663,1688]" class="Insecta" family="Coenagrionidae" genus="Enallagma" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="occultum">
|
||
<emphasis id="5DB1A851D000FFF3FC365EC4F713F914" box="[960,1101,1663,1688]" italics="true" pageId="29" pageNumber="430">M. occultum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F7FE68C6D000FFF3FBAB5E3AF7F3F914" box="[1117,1197,1663,1688]" captionStart="FIGURE 13. S 7 – 10" captionStartId="32.[151,250,1386,1408]" captionTargetBox="[378,1208,193,1364]" captionTargetId="figure@32.[378,1209,193,1364]" captionTargetPageId="32" captionText="FIGURE 13. S 7 – 10 color patterns of males: (a) and (b) Mesamphiagrion gaudiimontanum; (c) and (d) M. risi; (e) M. rosseri; (f) and (g) M. santainense; (h) M. tamaense; (i) and (j) M. occultum; (k) M. demarmelsi; (l) M. laterale; (m) M. ovigerum; (n) M. nataliae." httpUri="https://zenodo.org/record/246841/files/figure.png" pageId="29" pageNumber="430">Fig. 13</figureCitation>
|
||
n),
|
||
<emphasis id="5DB1A851D000FFF3FB225EC4F635F914" box="[1236,1387,1665,1688]" italics="true" pageId="29" pageNumber="430">
|
||
<taxonomicName id="A8C50FC0D000FFF3FB225EC4F639F914" box="[1236,1383,1665,1688]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="tamaense">M. tamaense</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="A8C50FC0D000FFF3FF615EE0F224F931" box="[151,378,1700,1725]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="gaudiimontanum">
|
||
<emphasis id="5DB1A851D000FFF3FF615EE0F224F931" box="[151,378,1700,1725]" italics="true" pageId="29" pageNumber="430">M. gaudiimontanum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F7FE68C6D000FFF3FE7F5EE1F287F930" box="[393,473,1700,1725]" captionStart="FIGURE 13. S 7 – 10" captionStartId="32.[151,250,1386,1408]" captionTargetBox="[378,1208,193,1364]" captionTargetId="figure@32.[378,1209,193,1364]" captionTargetPageId="32" captionText="FIGURE 13. S 7 – 10 color patterns of males: (a) and (b) Mesamphiagrion gaudiimontanum; (c) and (d) M. risi; (e) M. rosseri; (f) and (g) M. santainense; (h) M. tamaense; (i) and (j) M. occultum; (k) M. demarmelsi; (l) M. laterale; (m) M. ovigerum; (n) M. nataliae." httpUri="https://zenodo.org/record/246841/files/figure.png" pageId="29" pageNumber="430">Fig. 13</figureCitation>
|
||
l).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="27DF27C8D000FFF3FF315E82F713F8A4" pageId="29" pageNumber="430" type="discussion">
|
||
<paragraph id="6F7A7443D000FFF3FF315E82F713F8A4" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315E82F265F96C" bold="true" box="[199,315,1735,1760]" pageId="29" pageNumber="430">Remarks.</emphasis>
|
||
There are only two known populations of this species, both located in a geographic area relatively well sampled, suggesting that this species may be threatened. The high variability observed for female coloration probably indicates polymorphism. Some specimens were found parasitized by mites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="27DF27C8D000FFF3FF315F71F0BEF850" pageId="29" pageNumber="430" type="distribution">
|
||
<paragraph id="6F7A7443D000FFF3FF315F71F7CEF8C1" blockId="29.[151,1437,151,2013]" box="[199,1168,1844,1869]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315F71F23EF8C1" bold="true" box="[199,352,1844,1869]" pageId="29" pageNumber="430">Distribution.</emphasis>
|
||
Cerros Orientales of the Sabana de Bogotá, Cordillera Oriental (
|
||
<figureCitation id="F7FE68C6D000FFF3FBC05F71F7DCF8C1" box="[1078,1154,1844,1869]" captionStart="FIGURE 15" captionStartId="35.[151,250,1933,1955]" captionTargetBox="[151,1436,949,1912]" captionTargetId="figure@35.[151,1436,949,1912]" captionTargetPageId="35" captionText="FIGURE 15. New distributional records of Mesamphiagrion laterale, M. nataliae, M. occultum, M. rosseri, M. santainense, and M. tamaense in Colombia." httpUri="https://zenodo.org/record/246843/files/figure.png" pageId="29" pageNumber="430">Fig.15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6F7A7443D000FFF3FF315F12F0BEF850" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="430">
|
||
<emphasis id="5DB1A851D000FFF3FF315F12F2E1F8FC" bold="true" box="[199,447,1879,1904]" pageId="29" pageNumber="430">Habitat and biology.</emphasis>
|
||
Adults were found at an artificial reservoir in pasture land; according to the collector "The population was quite large and spiders eat enough of these damselflies, I found a lot trapped in the cobwebs" (Sánchez
|
||
<emphasis id="5DB1A851D000FFF3FEF35FE4F2D0F834" box="[261,398,1953,1976]" italics="true" pageId="29" pageNumber="430">pers. comm.</emphasis>
|
||
). Other odonates found at the same locality included (Sánchez
|
||
<emphasis id="5DB1A851D000FFF3FBAF5FE4F7BCF834" box="[1113,1250,1953,1976]" italics="true" pageId="29" pageNumber="430">pers. comm.</emphasis>
|
||
)
|
||
<emphasis id="5DB1A851D000FFF3FB045FE4F27EF851" italics="true" pageId="29" pageNumber="430">
|
||
<taxonomicName id="A8C50FC0D000FFF3FB045FE4F62CF835" box="[1266,1394,1952,1977]" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="laterale">M. laterale</taxonomicName>
|
||
,
|
||
<taxonomicName id="A8C50FC0D000FFF3FA8B5FE4F242F851" class="Insecta" family="Coenagrionidae" genus="Mesamphiagrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="demarmelsi">M. demarmelsi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="A8C50FC0D000FFF3FED15F81F1CBF850" authority="Realpe, 2010" authorityName="Realpe" authorityYear="2010" box="[295,661,1988,2013]" class="Insecta" family="Coenagrionidae" genus="Ischnura" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="chingaza">
|
||
<emphasis id="5DB1A851D000FFF3FED15F81F2A9F851" box="[295,503,1988,2013]" italics="true" pageId="29" pageNumber="430">Ischnura chingaza</emphasis>
|
||
Realpe, 2010
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A8C50FC0D000FFF3FD395F81F087F851" box="[719,985,1988,2013]" class="Insecta" family="Aeshnidae" genus="Rhionaeschna" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="marchali">
|
||
<emphasis id="5DB1A851D000FFF3FD395F81F087F851" box="[719,985,1988,2013]" italics="true" pageId="29" pageNumber="430">Rhionaeschna marchali</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |