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<document id="CAF6771703F9CACF793CF787746E8DBE" ID-DOI="10.5852/ejt.2024.923.2445" ID-GBIF-Dataset="01232941-b1fd-4c18-b269-e97169b18908" ID-ISSN="2118-9773" ID-Zenodo-Dep="10727741" ID-ZooBank="AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1709215085450" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2024" docId="E54587AB38580B14FDD1F9BD5514FE1F" docLanguage="en" docName="EJT.2024.923.1-119.pdf.imf" docOrigin="European Journal of Taxonomy 923" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Emarginula nodulicostata Herbert 2024, sp. nov." docType="treatment" docUuid="DA82AFE9-2C80-4ED8-AEF2-6945D3329814" docUuidSource="ZooBank" docVersion="9" lastPageNumber="8" masterDocId="197CFFD3385E0B1CFF9BFFC25776FFD3" masterDocTitle="The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species" masterLastPageNumber="119" masterPageNumber="1" pageNumber="7" updateTime="1732578736835" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="2C438E20E681C92BEC69F87251D7BBED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="04C269B56F783ABA83BE1223BD02B2E8">
<mods:title id="792814A07B96E9F273E3077B54802B7A">The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species</mods:title>
</mods:titleInfo>
<mods:name id="AAAE57D5BD9782276609D590E31B3ECA" type="personal">
<mods:role id="4AA457105A99EE9C2B56038BEBFEC1E3">
<mods:roleTerm id="F41C17BF7250A99DF50E4689F64151C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8101847958A1DFE727C6A672F3639FFD">Herbert, David G.</mods:namePart>
<mods:nameIdentifier id="43F3BE98B45FBE01983C43E52EA7D4CF" type="LSID">0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
<mods:affiliation id="0CA576B3AD32966C809AFAA54D7789C0">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Institute of Systematics, Evolution, Biodiversity (ISYEB), Muséum national d’histoire naturelle, CNRS, SU, EPHE, UA, CP 51, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
<mods:nameIdentifier id="20B19410D0E4D0A7C5405F3F0B1CF3EC" type="email">phasianella@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="DF09AA4E5E99BA81306001BC84845342">text</mods:typeOfResource>
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<mods:titleInfo id="44354EBEB462B967E248426D2E7FC443">
<mods:title id="1A6C89958C34B185F3EE17E434914B1C">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="CDBCE3A780FCACD6657402F6F69DBF59">
<mods:date id="0D4A34FCFB0936ED8B75678E1A713910">2024</mods:date>
<mods:detail id="383B186F537E71D4BEF934FF6D43B9A0" type="pubDate">
<mods:number id="AE15E20C9C34B5ED0AE1A2F9CE4B318C">2024-02-28</mods:number>
</mods:detail>
<mods:detail id="F464E2F8E28B6BE413B457E8FF63D009" type="volume">
<mods:number id="FDF7F36CBBF975429B64AFE214AC6074">923</mods:number>
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<mods:start id="C97DDB4E7C584EF9D36A89CDE942A4D8">1</mods:start>
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<mods:location id="11C66011D532195419310E52E507B513">
<mods:url id="2AB01C9117A50D134DCC27290589C25F">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835</mods:url>
</mods:location>
<mods:classification id="60B58997210F497B1CACC96AD75FC5FC">journal article</mods:classification>
<mods:identifier id="9A71EE2283F9A633599998BFC5042D1D" type="DOI">10.5852/ejt.2024.923.2445</mods:identifier>
<mods:identifier id="D12ACC48A784ED06C19BA6A9130EA229" type="GBIF-Dataset">01232941-b1fd-4c18-b269-e97169b18908</mods:identifier>
<mods:identifier id="3C378E732AF21BB97A5D912C816B8003" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="D43C46D3D647575C12F5AC76301B5E4E" type="Zenodo-Dep">10727741</mods:identifier>
<mods:identifier id="E9752153D3733355026B52697926E497" type="ZooBank">AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3</mods:identifier>
</mods:mods>
<treatment id="E54587AB38580B14FDD1F9BD5514FE1F" ID-DOI="http://doi.org/10.5281/zenodo.10818314" ID-GBIF-Taxon="223179369" ID-Zenodo-Dep="10818314" LSID="urn:lsid:zoobank.org:act:DA82AFE9-2C80-4ED8-AEF2-6945D3329814" httpUri="http://treatment.plazi.org/id/E54587AB38580B14FDD1F9BD5514FE1F" lastPageId="8" lastPageNumber="8" pageId="6" pageNumber="7" scope_class="Gastropoda" scope_family="Fissurellidae" scope_order="Lepetellida" scope_phylum="Mollusca" scope_subClass="Vetigastropoda" scope_subFamily="Emarginulinae" scope_superFamily="Fissurelloidea">
<subSubSection id="25F6653638580B1AFDD1F9BD549FF949" box="[586,1001,1663,1690]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="6D5336BD38580B1AFDD1F9BD549FF949" blockId="6.[364,1223,1663,1763]" box="[586,1001,1663,1690]" pageId="6" pageNumber="7">
<heading id="361B81D138580B1AFDD1F9BD549FF949" box="[586,1001,1663,1690]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<taxonomicName id="AAEC4D3E38580B1AFDD1F9BD54F1F949" ID-CoL="C6XMY" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[586,903,1663,1690]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="nodulicostata" status="sp. nov.">
<emphasis id="5F98EAAF38580B1AFDD1F9BD54F1F949" bold="true" box="[586,903,1663,1690]" italics="true" pageId="6" pageNumber="7">Emarginula nodulicostata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438580B1AFC14F942549FF949" box="[911,1001,1664,1690]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25F6653638580B1AFEF7F966544AF930" pageId="6" pageNumber="7" type="description">
<paragraph id="6D5336BD38580B1AFEF7F96653B1F96C" blockId="6.[364,1223,1663,1763]" box="[364,1223,1700,1727]" pageId="6" pageNumber="7">
<uri id="197D3ABF38580B1AFEF7F96653B1F96C" box="[364,1223,1700,1727]" pageId="6" pageNumber="7">
urn:lsid:zoobank.org:act:
<uuid id="194A0C6838580B1AFD11F96653B1F96C" box="[650,1223,1700,1727]" pageId="6" pageNumber="7">DA82AFE9-2C80-4ED8-AEF2-6945D3329814</uuid>
</uri>
</paragraph>
<paragraph id="6D5336BD38580B1AFD6CF90B544AF930" blockId="6.[364,1223,1663,1763]" box="[759,828,1737,1763]" pageId="6" pageNumber="7">
<figureCitation id="F5D72A3838580B1AFD6CF90B544AF930" box="[759,828,1737,1763]" captionStart="Fig" captionStartId="8.[189,232,1890,1916]" captionTargetBox="[189,1399,516,1850]" captionTargetId="figure-152@8.[189,1399,516,1850]" captionTargetPageId="8" captionText="Fig. 3. Emarginula nodulicostata sp. nov. A–D. Holotype, base 3.45 ×2.5 mm, height 2.25 mm (MNHN-IM-2000-38252). E. SEM of subadult specimen to show detail of sculpture, projecting lunulae of early selenizone arrowed, stn WS07, basal length 3.15 mm (paratype, MNHN-IM-2000-38256). F. Protoconch, stn WS07, scale bar =100 μm (paratype, MNHN-IM-2000-38256)." figureDoi="http://doi.org/10.5281/zenodo.10727749" httpUri="https://zenodo.org/record/10727749/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="25F6653638580B1AFF26F8CE564AF83B" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="6D5336BD38580B1AFF26F8CE5640F8F5" blockId="6.[189,310,1804,1830]" box="[189,310,1804,1830]" pageId="6" pageNumber="7">
<heading id="361B81D138580B1AFF26F8CE5640F8F5" bold="true" box="[189,310,1804,1830]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="5F98EAAF38580B1AFF26F8CE5640F8F5" bold="true" box="[189,310,1804,1830]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38580B1AFF26F8FE564AF83B" blockId="6.[188,1398,1852,2024]" pageId="6" pageNumber="7">Shell very small, stout, uniformly milky white; apex strongly recurved, extending well beyond posterior margin; dorsal profile strongly convex, anterior slit narrow and deep; sculpture coarsely cancellate, radial and concentric elements of similar strength; radial ribs nodular where crossed by concentric ridges; sculptural interstices simple; basal margin broader at posterior, distinctly crenulated at ends of radial ribs.</paragraph>
</subSubSection>
<subSubSection id="25F6653638590B1BFF26FECB545BFEA3" pageId="7" pageNumber="8" type="etymology">
<paragraph id="6D5336BD38590B1BFF26FECB5632FEF0" blockId="7.[189,324,265,291]" box="[189,324,265,291]" pageId="7" pageNumber="8">
<heading id="361B81D138590B1BFF26FECB5632FEF0" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="5F98EAAF38590B1BFF26FECB5632FEF0" bold="true" box="[189,324,265,291]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38590B1BFF26FEF4545BFEA3" blockId="7.[187,1399,309,369]" pageId="7" pageNumber="8">
From the Latin “
<emphasis id="5F98EAAF38590B1BFEE6FEF756B5FE9C" box="[381,451,309,335]" italics="true" pageId="7" pageNumber="8">nodus</emphasis>
” – “a knot or swelling”, dim. “
<emphasis id="5F98EAAF38590B1BFCB3FEF754F3FE9C" box="[808,901,309,335]" italics="true" pageId="7" pageNumber="8">nodulus</emphasis>
”, and “
<emphasis id="5F98EAAF38590B1BFC47FEF4536CFE9C" box="[988,1050,310,335]" italics="true" pageId="7" pageNumber="8">costa</emphasis>
” – “rib”, “
<emphasis id="5F98EAAF38590B1BFB0DFEF45380FE9C" box="[1174,1270,310,335]" italics="true" pageId="7" pageNumber="8">costatus</emphasis>
” “ribbed”; in reference to the nodose sculpture of the radial ribs.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638590B1BFF26FE54559FFC99" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="6D5336BD38590B1BFF26FE5456DCFE63" blockId="7.[189,426,406,432]" box="[189,426,406,432]" pageId="7" pageNumber="8">
<heading id="361B81D138590B1BFF26FE5456DCFE63" bold="true" box="[189,426,406,432]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="5F98EAAF38590B1BFF26FE5456DCFE63" bold="true" box="[189,426,406,432]" pageId="7" pageNumber="8">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="DD843CE038590B1BFF71FE0155A1FDF3" ID-GBIF-Occurrence="4538174479" collectingDate="2017-04-30" collectionCode="MNHN" elevation="40" latitude="-33.203335" location="WALTERS SHOAL" longLatPrecision="120" longitude="43.84667" pageId="7" pageNumber="8" specimenCode="MNHN-IM-2000-38252" specimenCount="1" typeStatus="holotype">
<paragraph id="6D5336BD38590B1BFF71FE01562AFE0E" blockId="7.[189,1398,451,544]" box="[234,348,451,477]" pageId="7" pageNumber="8">
<emphasis id="5F98EAAF38590B1BFF71FE01562AFE0E" bold="true" box="[234,348,451,477]" pageId="7" pageNumber="8">
<typeStatus id="B257881F38590B1BFF71FE01562AFE0E" box="[234,348,451,477]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD38590B1BFF26FE2655A1FDF3" blockId="7.[189,1398,451,544]" pageId="7" pageNumber="8">
<location id="6833606638590B1BFF26FE2656DBFE2D" LSID="urn:lsid:plazi:treatment:E54587AB38580B14FDD1F9BD5514FE1F:6833606638590B1BFF26FE2656DBFE2D" box="[189,429,484,510]" latitude="-33.203335" longLatPrecision="120" longitude="43.84667" name="WALTERS SHOAL" pageId="7" pageNumber="8">WALTERS SHOAL</location>
• empty shell; summit area south-west, stn WS04;
<geoCoordinate id="08D8507A38590B1BFB83FE2153E0FE2D" box="[1048,1174,483,511]" degrees="33" direction="south" minutes="12.2" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-33.203335">33°12.2′ S</geoCoordinate>
,
<geoCoordinate id="08D8507A38590B1BFB3EFE215251FE2C" box="[1189,1319,483,511]" degrees="43" direction="east" minutes="50.8" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="43.84667">43°50.8′ E</geoCoordinate>
; depth
<quantity id="AA149B5838590B1BFF26FDC4578AFDF3" box="[189,252,518,544]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="m" value="40.0">
<elevation id="E6C1D18E38590B1BFF26FDC4578AFDF3" box="[189,252,518,544]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="m" value="40.0">40 m</elevation>
</quantity>
;
<date id="1952107D38590B1BFE93FDC456D2FDF3" box="[264,420,518,544]" pageId="7" pageNumber="8" value="2017-04-30">
<collectingDate id="0916E99538590B1BFE93FDC456D2FDF3" box="[264,420,518,544]" pageId="7" pageNumber="8" value="2017-04-30">30 Apr. 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638590B1BFE2BFDC755A5FDF3" box="[432,723,517,544]" pageId="7" pageNumber="8">MNHN-IM-2000-38252</specimenCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="DD843CE038590B1BFF71FD8755A8FD70" ID-GBIF-Occurrence="4538174446" collectingDate="2017-04-30" collectionCode="MNHN" elevation="40" latitude="-33.203335" location="WALTERS SHOAL" longLatPrecision="120" longitude="43.84667" pageId="7" pageNumber="8" specimenCode="MNHN-IM-2000-38253" specimenCount="6" typeStatus="paratype">
<paragraph id="6D5336BD38590B1BFF71FD875611FD8C" blockId="7.[189,1399,581,842]" box="[234,359,581,607]" pageId="7" pageNumber="8">
<emphasis id="5F98EAAF38590B1BFF71FD875611FD8C" bold="true" box="[234,359,581,607]" pageId="7" pageNumber="8">
<typeStatus id="B257881F38590B1BFF71FD875611FD8C" box="[234,359,581,607]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD38590B1BFF26FDA5559FFC99" blockId="7.[189,1399,581,842]" pageId="7" pageNumber="8">
<location id="6833606638590B1BFF26FDA556DDFD52" LSID="urn:lsid:plazi:treatment:E54587AB38580B14FDD1F9BD5514FE1F:6833606638590B1BFF26FDA556DDFD52" box="[189,427,615,641]" latitude="-33.203335" longLatPrecision="120" longitude="43.84667" name="WALTERS SHOAL" pageId="7" pageNumber="8">WALTERS SHOAL</location>
•
<specimenCount id="7BEAFD3438590B1BFE58FDA556A4FD52" box="[451,466,615,641]" pageId="7" pageNumber="8" type="generic">6</specimenCount>
empty shells; summit area, south-west, stn WS03;
<geoCoordinate id="08D8507A38590B1BFBB6FDA553EAFD52" box="[1069,1180,615,641]" degrees="33" direction="south" minutes="12.2" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-33.203335">33°12.2S</geoCoordinate>
,
<geoCoordinate id="08D8507A38590B1BFB32FDA7525FFD52" box="[1193,1321,613,641]" degrees="43" direction="east" minutes="50.8" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="43.84667">43°50.8′ E</geoCoordinate>
; depth
<quantity id="AA149B5838590B1BFF26FD4A5789FD70" box="[189,255,648,675]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="m" value="40.0">
<elevation id="E6C1D18E38590B1BFF26FD4A5789FD70" box="[189,255,648,675]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="m" value="40.0">40 m</elevation>
</quantity>
;
<date id="1952107D38590B1BFE95FD4A56D9FD71" box="[270,431,648,674]" pageId="7" pageNumber="8" value="2017-04-30">
<collectingDate id="0916E99538590B1BFE95FD4A56D9FD71" box="[270,431,648,674]" pageId="7" pageNumber="8" value="2017-04-30">30 Apr. 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638590B1BFE25FD4A55A8FD70" box="[446,734,648,675]" pageId="7" pageNumber="8">MNHN-IM-2000-38253</specimenCode>
•
<materialsCitation id="DD843CE038590B1BFD67FD4A5369FD17" ID-GBIF-Occurrence="4538174335" collectingDate="2017-04-30" collectionCode="MNHN" elevation="40" latitude="-33.203335" location="WALTERS SHOAL" longLatPrecision="120" longitude="43.84667" pageId="7" pageNumber="8" specimenCode="MNHN-IM-2000-38253" specimenCount="11" typeStatus="paratype">
11 empty shells; summit area south-west, stn WS04;
<geoCoordinate id="08D8507A38590B1BFF26FD6A564EFD17" box="[189,312,680,708]" degrees="33" direction="south" minutes="12.2" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-33.203335">33°12.2′ S</geoCoordinate>
,
<geoCoordinate id="08D8507A38590B1BFEDFFD6A56B4FD17" box="[324,450,680,708]" degrees="43" direction="east" minutes="50.8" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="43.84667">43°50.8′ E</geoCoordinate>
; depth
<quantity id="AA149B5838590B1BFD8FFD685524FD17" box="[532,594,682,708]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="m" value="40.0">40 m</quantity>
;
<date id="1952107D38590B1BFDC6FD685580FD17" box="[605,758,682,708]" pageId="7" pageNumber="8" value="2017-04-30">30 Apr. 2017</date>
; MNHN-IM-2000-38254
</materialsCitation>
•
<materialsCitation id="DD843CE038590B1BFBADFD68539BFD35" ID-GBIF-Occurrence="4538174540" collectingDate="2017-05-01" collectionCode="MNHN" elevation="26" latitude="-33.251667" location="WALTERS SHOAL" longLatPrecision="120" longitude="43.908333" pageId="7" pageNumber="8" specimenCode="MNHN-IM-2000-38255" specimenCount="1" typeStatus="paratype">
1 empty shell; summit area, south, stn WS06;
<geoCoordinate id="08D8507A38590B1BFE14FD095689FD35" box="[399,511,715,742]" degrees="33" direction="south" minutes="15.1" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-33.251667">33°15.1S</geoCoordinate>
,
<geoCoordinate id="08D8507A38590B1BFD97FD0B55FAFD36" box="[524,652,713,741]" degrees="43" direction="east" minutes="54.5" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="43.908333">43°54.5′ E</geoCoordinate>
; depth
<quantity id="AA149B5838590B1BFD79FD095454FD36" box="[738,802,715,741]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="8" unit="m" value="26.0">
<elevation id="E6C1D18E38590B1BFD79FD095454FD36" box="[738,802,715,741]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="8" unit="m" value="26.0">26 m</elevation>
</quantity>
;
<date id="1952107D38590B1BFCB4FD0954B7FD36" box="[815,961,715,741]" pageId="7" pageNumber="8" value="2017-05-01">
<collectingDate id="0916E99538590B1BFCB4FD0954B7FD36" box="[815,961,715,741]" pageId="7" pageNumber="8" value="2017-05-01">1 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638590B1BFC55FD09539BFD35" box="[974,1261,715,742]" pageId="7" pageNumber="8">MNHN-IM-2000-38255</specimenCode>
</materialsCitation>
•
<materialsCitation id="DD843CE038590B1BFA93FD09560FFCFB" ID-GBIF-Occurrence="4538174387" collectingDate="2017-05-02" collectionCode="MNHN" elevation="32" latitude="-33.256668" location="WALTERS SHOAL" longLatPrecision="120" longitude="43.87" pageId="7" pageNumber="8" specimenCode="MNHN- IM-2000-38256" specimenCount="10" typeStatus="paratype">
<specimenCount id="7BEAFD3438590B1BFA93FD095253FD36" box="[1288,1317,715,741]" pageId="7" pageNumber="8">10</specimenCount>
empty shells; summit area, south, stn WS07;
<geoCoordinate id="08D8507A38590B1BFD09FD2E5477FCD4" box="[658,769,748,775]" degrees="33" direction="south" minutes="15.4" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-33.256668">33°15.4S</geoCoordinate>
,
<geoCoordinate id="08D8507A38590B1BFC8AFD2954E5FCD4" box="[785,915,747,775]" degrees="43" direction="east" minutes="52.2" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="43.87">43°52.2′ E</geoCoordinate>
; depth
<quantity id="AA149B5838590B1BFC75FD2E532AFCD4" box="[1006,1116,748,775]" metricMagnitude="1" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="7" pageNumber="8" unit="m" value="31.5" valueMax="33.0" valueMin="30.0">
<elevation id="E6C1D18E38590B1BFC75FD2E532AFCD4" box="[1006,1116,748,775]" metricMagnitude="1" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="7" pageNumber="8" unit="m" value="31.5" valueMax="33.0" valueMin="30.0">30–33 m</elevation>
</quantity>
;
<date id="1952107D38590B1BFBF7FD2E5272FCD5" box="[1132,1284,748,775]" pageId="7" pageNumber="8" value="2017-05-02">
<collectingDate id="0916E99538590B1BFBF7FD2E5272FCD5" box="[1132,1284,748,775]" pageId="7" pageNumber="8" value="2017-05-02">2 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638590B1BFA88FD2F560FFCFB" pageId="7" pageNumber="8">MNHN- IM-2000-38256</specimenCode>
</materialsCitation>
•
<materialsCitation id="DD843CE038590B1BFE08FCCC559FFC99" ID-GBIF-Occurrence="4538174439" collectingDate="2017-05-03" collectionCode="MNHN" elevation="32" latitude="-33.228333" location="WALTERS SHOAL" longLatPrecision="120" longitude="43.931667" pageId="7" pageNumber="8" specimenCode="MNHN-IM-2000-3824657" specimenCount="3" typeStatus="paratype">
<specimenCount id="7BEAFD3438590B1BFE08FCCC56D4FCFB" box="[403,418,782,808]" pageId="7" pageNumber="8">3</specimenCount>
empty shells; summit area, south-east, stn WS08;
<geoCoordinate id="08D8507A38590B1BFC63FCCC531EFCFB" box="[1016,1128,782,808]" degrees="33" direction="south" minutes="13.7" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="-33.228333">33°13.7S</geoCoordinate>
,
<geoCoordinate id="08D8507A38590B1BFBEFFCCE5382FCFB" box="[1140,1268,780,808]" degrees="43" direction="east" minutes="55.9" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="43.931667">43°55.9′ E</geoCoordinate>
; depth
<quantity id="AA149B5838590B1BFAD1FCCC578AFC9A" metricMagnitude="1" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="7" pageNumber="8" unit="m" value="31.5" valueMax="33.0" valueMin="30.0">
<elevation id="E6C1D18E38590B1BFAD1FCCC578AFC9A" metricMagnitude="1" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="7" pageNumber="8" unit="m" value="31.5" valueMax="33.0" valueMin="30.0">30– 33 m</elevation>
</quantity>
;
<date id="1952107D38590B1BFE93FCED56EFFC9A" box="[264,409,815,841]" pageId="7" pageNumber="8" value="2017-05-03">
<collectingDate id="0916E99538590B1BFE93FCED56EFFC9A" box="[264,409,815,841]" pageId="7" pageNumber="8" value="2017-05-03">3 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638590B1BFE3EFCED5593FC99" box="[421,741,815,842]" pageId="7" pageNumber="8">MNHN-IM-2000-3824657</specimenCode>
.
</materialsCitation>
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="25F6653638590B1BFF26FCAC53F7F959" pageId="7" pageNumber="8" type="description">
<paragraph id="6D5336BD38590B1BFF26FCAC5638FC5B" blockId="7.[189,334,878,904]" box="[189,334,878,904]" pageId="7" pageNumber="8">
<heading id="361B81D138590B1BFF26FCAC5638FC5B" bold="true" box="[189,334,878,904]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="5F98EAAF38590B1BFF26FCAC5638FC5B" bold="true" box="[189,334,878,904]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38590B1BFF26FC59531BFA79" blockId="7.[187,1401,923,1450]" pageId="7" pageNumber="8">
<smallCapsWord id="6BB5A06138590B1BFF26FC595677FC67" baselines="942,943" box="[189,257,923,949]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="7" pageNumber="8">SHELL</smallCapsWord>
. Very small (basal length up to
<quantity id="AA149B5838590B1BFDE3FC595593FC66" box="[632,741,923,950]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" pageId="7" pageNumber="8" unit="mm" value="3.45">3.45 mm</quantity>
), but relatively thick; apex strongly recurved, extending well beyond posterior margin of base; basal outline evenly ovate (L/W=1.37–1.40), slightly concave in side view; height moderate (H/L=0.52–0.65); dorsal profile strongly convex in side view, posterior slope comprising only one-third height of shell in adult; anterior face slightly flattened either side of slit; posterior slope almost vertical in largest specimens, much less so in smaller ones; protoconch tucked under recurved apex, slightly twisted to right. Anterior slit narrow and deep, approx. one-third of anterior slope; selenizone initially forming a broad mid-line rib (margins scarcely raised) with coarse lunulae that project above its margins (
<figureCitation id="F5D72A3838590B1BFDB2FB4755F7FB4C" box="[553,641,1157,1183]" captionStart="Fig" captionStartId="8.[189,232,1890,1916]" captionTargetBox="[189,1399,516,1850]" captionTargetId="figure-152@8.[189,1399,516,1850]" captionTargetPageId="8" captionText="Fig. 3. Emarginula nodulicostata sp. nov. A–D. Holotype, base 3.45 ×2.5 mm, height 2.25 mm (MNHN-IM-2000-38252). E. SEM of subadult specimen to show detail of sculpture, projecting lunulae of early selenizone arrowed, stn WS07, basal length 3.15 mm (paratype, MNHN-IM-2000-38256). F. Protoconch, stn WS07, scale bar =100 μm (paratype, MNHN-IM-2000-38256)." figureDoi="http://doi.org/10.5281/zenodo.10727749" httpUri="https://zenodo.org/record/10727749/files/figure.png" pageId="7" pageNumber="8">Fig. 3E</figureCitation>
arrow), but selenizone becoming somewhat sunken nearer to slit in largest specimens; remaining sculpture relatively coarsely cancellate, comprising ± 16 primary radial ribs with narrower secondary ribs between them; secondaries stronger laterally, those between anterior ribs more slender; radial sculpture crossed by coarse concentric ridges; radial and concentric sculpture of similar strength; radial ribs roundly nodular where crossed by concentric ridges; interstices between ribs and ridges simple, square to concentrically elongate-rectangular, with paired intritacalx pits; basal margin distinctly crenulated at ends of radial ribs and noticeably broader posteriorly (
<figureCitation id="F5D72A3838590B1BFB36FA8F527CFAB4" box="[1197,1290,1357,1383]" captionStart="Fig" captionStartId="8.[189,232,1890,1916]" captionTargetBox="[189,1399,516,1850]" captionTargetId="figure-152@8.[189,1399,516,1850]" captionTargetPageId="8" captionText="Fig. 3. Emarginula nodulicostata sp. nov. A–D. Holotype, base 3.45 ×2.5 mm, height 2.25 mm (MNHN-IM-2000-38252). E. SEM of subadult specimen to show detail of sculpture, projecting lunulae of early selenizone arrowed, stn WS07, basal length 3.15 mm (paratype, MNHN-IM-2000-38256). F. Protoconch, stn WS07, scale bar =100 μm (paratype, MNHN-IM-2000-38256)." figureDoi="http://doi.org/10.5281/zenodo.10727749" httpUri="https://zenodo.org/record/10727749/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
).
<collectingRegion id="AF28F85F38590B1BFA84FA8F5201FAB4" box="[1311,1399,1357,1383]" country="Oman" name="Ad Dakhiliyah" pageId="7" pageNumber="8">Interior</collectingRegion>
with broad, low ridge underlying selenizone posterior to anterior slit, approx. ⅔ length of slit; ridge itself with shallow central furrow; with interior of apex only just visible in basal view.
</paragraph>
<paragraph id="6D5336BD38590B1BFF26FA125334F9D8" blockId="7.[189,1399,1487,1547]" pageId="7" pageNumber="8">
<smallCapsWord id="6BB5A06138590B1BFF26FA12563AFA3B" baselines="1507,1507" box="[189,332,1488,1514]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Protoconch" pageId="7" pageNumber="8">PROTOCONCH</smallCapsWord>
(
<figureCitation id="F5D72A3838590B1BFEFBFA0D56CCFA39" box="[352,442,1487,1514]" captionStart="Fig" captionStartId="8.[189,232,1890,1916]" captionTargetBox="[189,1399,516,1850]" captionTargetId="figure-152@8.[189,1399,516,1850]" captionTargetPageId="8" captionText="Fig. 3. Emarginula nodulicostata sp. nov. A–D. Holotype, base 3.45 ×2.5 mm, height 2.25 mm (MNHN-IM-2000-38252). E. SEM of subadult specimen to show detail of sculpture, projecting lunulae of early selenizone arrowed, stn WS07, basal length 3.15 mm (paratype, MNHN-IM-2000-38256). F. Protoconch, stn WS07, scale bar =100 μm (paratype, MNHN-IM-2000-38256)." figureDoi="http://doi.org/10.5281/zenodo.10727749" httpUri="https://zenodo.org/record/10727749/files/figure.png" pageId="7" pageNumber="8">Fig. 3F</figureCitation>
). Typically emarginuliform; single whorl with sharp, flared terminal lip; surface with finely flocculated superficial granulation, maximum diameter ±140 μm.
</paragraph>
<paragraph id="6D5336BD38590B1BFF26F9F354A2F999" blockId="7.[189,980,1584,1611]" box="[189,980,1584,1611]" pageId="7" pageNumber="8">
<smallCapsWord id="6BB5A06138590B1BFF26F9F3566EF99A" baselines="1604,1604" box="[189,280,1585,1611]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="7" pageNumber="8">COLOUR</smallCapsWord>
. Uniformly milky white, somewhat translucent when fresh.
</paragraph>
<paragraph id="6D5336BD38590B1BFF26F9B353F7F959" blockId="7.[189,1153,1648,1675]" box="[189,1153,1648,1675]" pageId="7" pageNumber="8">
<smallCapsWord id="6BB5A06138590B1BFF26F9B35634F95A" baselines="1668,1668" box="[189,322,1649,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dimensions" pageId="7" pageNumber="8">DIMENSIONS</smallCapsWord>
.
<typeStatus id="B257881F38590B1BFECAF9B256B7F959" box="[337,449,1648,1674]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
, base 3.45 ×
<quantity id="AA149B5838590B1BFDC8F9B255C9F958" box="[595,703,1648,1675]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="2.5">2.50 mm</quantity>
, height
<quantity id="AA149B5838590B1BFC80F9B254F5F958" box="[795,899,1648,1675]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="7" pageNumber="8" unit="mm" value="2.25">2.25 mm</quantity>
(= largest specimen).
</paragraph>
</subSubSection>
<subSubSection id="25F6653638590B1BFF26F96D5689F8CB" pageId="7" pageNumber="8" type="distribution">
<paragraph id="6D5336BD38590B1BFF26F96D5620F91A" blockId="7.[189,342,1711,1737]" box="[189,342,1711,1737]" pageId="7" pageNumber="8">
<heading id="361B81D138590B1BFF26F96D5620F91A" bold="true" box="[189,342,1711,1737]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="5F98EAAF38590B1BFF26F96D5620F91A" bold="true" box="[189,342,1711,1737]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38590B1BFF26F91F5689F8CB" blockId="7.[189,1399,1756,1816]" pageId="7" pageNumber="8">
A shallow-water species known only from the summit area of Walters Shoal, at depths of
<quantity id="AA149B5838590B1BFA9FF91E5207F924" box="[1284,1393,1756,1783]" metricMagnitude="1" metricUnit="m" metricValue="3.3" metricValueMax="4.0" metricValueMin="2.6" pageId="7" pageNumber="8" unit="m" value="33.0" valueMax="40.0" valueMin="26.0">26–40 m</quantity>
; living specimens unknown.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638590B14FF26F8FC5514FE1F" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
<paragraph id="6D5336BD38590B1BFF26F8FC5659F88B" blockId="7.[189,303,1854,1880]" box="[189,303,1854,1880]" pageId="7" pageNumber="8">
<heading id="361B81D138590B1BFF26F8FC5659F88B" bold="true" box="[189,303,1854,1880]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="5F98EAAF38590B1BFF26F8FC5659F88B" bold="true" box="[189,303,1854,1880]" pageId="7" pageNumber="8">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38590B14FF26F8A85514FE1F" blockId="7.[189,1399,1898,2025]" lastBlockId="8.[188,1399,265,460]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Other strongly recurved species in which the apex lies beyond the posterior margin of the base include
<taxonomicName id="AAEC4D3E38590B1BFF26F84955CEF876" authority="Sowerby II, 1863" authorityName="Sowerby II" authorityYear="1863" box="[189,696,1931,1958]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="adamsiana">
<emphasis id="5F98EAAF38590B1BFF26F84956A0F876" box="[189,470,1931,1957]" italics="true" pageId="7" pageNumber="8">Emarginula adamsiana</emphasis>
Sowerby II, 1863
</taxonomicName>
from
<collectingCountry id="15FB762D38590B1BFC91F84E5426F875" box="[778,848,1932,1958]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
,
<taxonomicName id="AAEC4D3E38590B1BFCFBF84E53E5F875" authority="Hedley, 1907" authorityName="Hedley" authorityYear="1907" box="[864,1171,1931,1958]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="convexa">
<emphasis id="5F98EAAF38590B1BFCFBF84E5493F876" box="[864,997,1932,1957]" italics="true" pageId="7" pageNumber="8">E. convexa</emphasis>
Hedley, 1907
</taxonomicName>
from Queensland,
<taxonomicName id="AAEC4D3E38590B1BFF26F86C56B4F814" authority="Thiele, 1913" authorityName="Thiele" authorityYear="1913" box="[189,450,1965,1991]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="dahli">
<emphasis id="5F98EAAF38590B1BFF26F86C5668F814" box="[189,286,1965,1991]" italics="true" pageId="7" pageNumber="8">E. dahli</emphasis>
Thiele, 1913
</taxonomicName>
from the
<collectingCountry id="15FB762D38590B1BFDD9F86F5426F814" box="[578,848,1965,1991]" name="Papua New Guinea" pageId="7" pageNumber="8">Bismarck Archipelago</collectingCountry>
, and
<taxonomicName id="AAEC4D3E38590B1BFC0DF86C5201F814" authority="Poppe &amp; Tagaro, 2020" authorityName="Poppe &amp; Tagaro" authorityYear="2020" box="[918,1399,1965,1991]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="circumalbum">
<emphasis id="5F98EAAF38590B1BFC0DF86C5323F814" box="[918,1109,1965,1991]" italics="true" pageId="7" pageNumber="8">E. circumalbum</emphasis>
<bibRefCitation id="097D4B4C38590B1BFBFAF86F5201F814" author="Poppe G. T. &amp; Tagaro S. P." box="[1121,1399,1965,1991]" pageId="7" pageNumber="8" pagination="1 - 131" refId="ref62653" refString="Poppe G. T. &amp; Tagaro S. P. 2020. The Fissurellidae from the Philippines with description of 26 new species. Visaya Supplement 13: 1 - 131." type="journal article" year="2020">Poppe &amp; Tagaro, 2020</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="AAEC4D3E38590B1BFF69F80D5518F83A" authority="A. Adams 1852" authorityName="A. Adams" authorityYear="1852" box="[242,622,1998,2025]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="galericulata">
<emphasis id="5F98EAAF38590B1BFF69F80D56D3F83B" box="[242,421,1998,2024]" italics="true" pageId="7" pageNumber="8">E. galericulata</emphasis>
A. Adams 1852
</taxonomicName>
from the
<collectingCountry id="15FB762D38590B1BFD70F80C5405F83B" box="[747,883,1998,2024]" name="Philippines" pageId="7" pageNumber="8">Philippines</collectingCountry>
. All except
<taxonomicName id="AAEC4D3E38590B1BFB9CF80D53FDF83B" authorityName="Hedley" authorityYear="1907" box="[1031,1163,1999,2024]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="convexa">
<emphasis id="5F98EAAF38590B1BFB9CF80D53FDF83B" box="[1031,1163,1999,2024]" italics="true" pageId="7" pageNumber="8">E. convexa</emphasis>
</taxonomicName>
attain a larger size than
<taxonomicName id="AAEC4D3E38560B14FF60FEC856B6FEF0" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[251,448,265,291]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="nodulicostata" status="sp. nov.">
<emphasis id="5F98EAAF38560B14FF60FEC856B6FEF0" box="[251,448,265,291]" italics="true" pageId="8" pageNumber="9">E. nodulicostata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438560B14FE50FEC8555CFEF7" box="[459,554,266,292]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
(basal length&gt;
<quantity id="AA149B5838560B14FD74FECB5425FEF0" box="[751,851,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="9" unit="mm" value="4.5">4.5 mm</quantity>
); in addition
<taxonomicName id="AAEC4D3E38560B14FC62FEC853E8FEF0" authorityName="Sowerby II" authorityYear="1863" box="[1017,1182,265,291]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="adamsiana">
<emphasis id="5F98EAAF38560B14FC62FEC853E8FEF0" box="[1017,1182,265,291]" italics="true" pageId="8" pageNumber="9">E. adamsiana</emphasis>
</taxonomicName>
is pale pink, and
<taxonomicName id="AAEC4D3E38560B14FF26FEE95607FE97" authorityName="A. Adams" authorityYear="1852" box="[189,369,298,324]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="galericulata">
<emphasis id="5F98EAAF38560B14FF26FEE95607FE97" box="[189,369,298,324]" italics="true" pageId="8" pageNumber="9">E. galericulata</emphasis>
</taxonomicName>
has finer sculpture.
<taxonomicName id="AAEC4D3E38560B14FDEAFEE8541EFE97" authorityName="Hedley" authorityYear="1907" box="[625,872,298,324]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="convexa">
<emphasis id="5F98EAAF38560B14FDEAFEE8541EFE97" box="[625,872,298,324]" italics="true" pageId="8" pageNumber="9">Emarginula convexa</emphasis>
</taxonomicName>
has an indistinct concentric colour pattern of pale green and grey zones, more numerous radial ribs and finer concentric sculpture.
<taxonomicName id="AAEC4D3E38560B14FB70FE8E5621FE54" authorityName="Poppe &amp; Tagaro" authorityYear="2020" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="circumalbum">
<emphasis id="5F98EAAF38560B14FB70FE8E5621FE54" italics="true" pageId="8" pageNumber="9">Emarginula circumalbum</emphasis>
</taxonomicName>
is a deeper-water species (
<quantity id="AA149B5838560B14FD16FEAF5462FE54" box="[653,788,365,392]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="8" pageNumber="9" unit="m" value="375.0" valueMax="500.0" valueMin="250.0">250–500 m</quantity>
) and has a proportionately broader basal outline and a much shorter slit. The poorly known
<taxonomicName id="AAEC4D3E38560B14FD13FE4D5593FE7B" authorityName="Thiele" authorityYear="1913" box="[648,741,398,424]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="dahli">
<emphasis id="5F98EAAF38560B14FD13FE4D5593FE7B" box="[648,741,398,424]" italics="true" pageId="8" pageNumber="9">E. dahli</emphasis>
</taxonomicName>
is perhaps the most similar species, but this also occurs in much deeper water (
<quantity id="AA149B5838560B14FE51FE735523FE1F" box="[458,597,433,460]" metricMagnitude="2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="8" pageNumber="9" unit="m" value="170.0" valueMax="180.0" valueMin="160.0">160–180 m</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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