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<document id="151203C5B1F15D88F715CD563456D5EF" ID-DOI="10.5852/ejt.2024.923.2445" ID-GBIF-Dataset="01232941-b1fd-4c18-b269-e97169b18908" ID-ISSN="2118-9773" ID-Zenodo-Dep="10727741" ID-ZooBank="AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1709215085450" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2024" docId="E54587AB38010B78FDC3F9AB5638F93F" docLanguage="en" docName="EJT.2024.923.1-119.pdf.imf" docOrigin="European Journal of Taxonomy 923" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Kaiparathina monticola Herbert 2024, sp. nov." docType="treatment" docUuid="C4A503AD-4E0D-4DF7-8A82-ECF326086C36" docUuidSource="ZooBank" docVersion="6" lastPageNumber="101" masterDocId="197CFFD3385E0B1CFF9BFFC25776FFD3" masterDocTitle="The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species" masterLastPageNumber="119" masterPageNumber="1" pageNumber="96" updateTime="1732578736835" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C6ABBFB763139254EBD574B30B1F2967">The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species</mods:title>
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<mods:name id="403801CF1EB6132E1A257BFE79B0627E" type="personal">
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<mods:namePart id="EED05A97D66C2C648C1EB187B338598C">Herbert, David G.</mods:namePart>
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<mods:affiliation id="D766474F73DAF2D181F29BEDC14A303F">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Institute of Systematics, Evolution, Biodiversity (ISYEB), Muséum national dhistoire naturelle, CNRS, SU, EPHE, UA, CP 51, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
<mods:nameIdentifier id="8382AEEA9B26142B5978C202D5C7134B" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="945F2E8126472D1CB41537E1E26E096B">European Journal of Taxonomy</mods:title>
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<mods:date id="569BBFCE4779C19498AC02D6A47112D0">2024</mods:date>
<mods:detail id="D86E45E341E20006AC51F3CE7CA6BAFB" type="pubDate">
<mods:number id="ABEAA2E794937071E5304757AFC83CD5">2024-02-28</mods:number>
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<mods:number id="3CDD4DBCC8509848A25AE8C0DDB0374D">923</mods:number>
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<mods:identifier id="04949F2CB2958845E53A772C8E777385" type="DOI">10.5852/ejt.2024.923.2445</mods:identifier>
<mods:identifier id="29C1D59D4AD98A795E3EC01B6C3D61AF" type="GBIF-Dataset">01232941-b1fd-4c18-b269-e97169b18908</mods:identifier>
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<treatment id="E54587AB38010B78FDC3F9AB5638F93F" ID-DOI="http://doi.org/10.5281/zenodo.10818420" ID-GBIF-Taxon="223179259" ID-Zenodo-Dep="10818420" LSID="urn:lsid:zoobank.org:act:C4A503AD-4E0D-4DF7-8A82-ECF326086C36" httpUri="http://treatment.plazi.org/id/E54587AB38010B78FDC3F9AB5638F93F" lastPageId="100" lastPageNumber="101" pageId="95" pageNumber="96" scope_class="Gastropoda" scope_family="Trochidae" scope_order="Trochida" scope_phylum="Mollusca" scope_subClass="Vetigastropoda" scope_subFamily="Kaiparathininae" scope_superFamily="Trochoidea">
<subSubSection id="25F6653638010B43FDC3F9AB54AAF957" box="[600,988,1641,1668]" pageId="95" pageNumber="96" type="nomenclature">
<paragraph id="6D5336BD38010B43FDC3F9AB54AAF957" blockId="95.[361,1226,1641,1738]" box="[600,988,1641,1668]" pageId="95" pageNumber="96">
<heading id="361B81D138010B43FDC3F9AB54AAF957" box="[600,988,1641,1668]" centered="true" fontSize="11" level="2" pageId="95" pageNumber="96" reason="2">
<taxonomicName id="AAEC4D3E38010B43FDC3F9AB540CF957" ID-CoL="C7JC3" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[600,890,1641,1668]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="95" pageNumber="96" phylum="Mollusca" rank="species" species="monticola" status="sp. nov.">
<emphasis id="5F98EAAF38010B43FDC3F9AB540CF957" bold="true" box="[600,890,1641,1668]" italics="true" pageId="95" pageNumber="96">Kaiparathina monticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438010B43FC1AF9A854AAF957" box="[897,988,1642,1668]" pageId="95" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25F6653638010B43FEF2F94F54FAF919" pageId="95" pageNumber="96" type="description">
<paragraph id="6D5336BD38010B43FEF2F94F53BCF97B" blockId="95.[361,1226,1641,1738]" box="[361,1226,1677,1704]" pageId="95" pageNumber="96">
<uri id="197D3ABF38010B43FEF2F94F53BCF97B" box="[361,1226,1677,1704]" pageId="95" pageNumber="96">
urn:lsid:zoobank.org:act:
<uuid id="194A0C6838010B43FD1DF94F53BCF97B" box="[646,1226,1677,1704]" pageId="95" pageNumber="96">C4A503AD-4E0D-4DF7-8A82-ECF326086C36</uuid>
</uri>
</paragraph>
<paragraph id="6D5336BD38010B43FD33F97254FAF919" blockId="95.[361,1226,1641,1738]" box="[680,908,1712,1738]" pageId="95" pageNumber="96">
<figureCitation id="F5D72A3838010B43FD33F9725440F919" box="[680,822,1712,1738]" captionStart="Fig" captionStartId="97.[188,231,1860,1886]" captionTargetBox="[189,1399,461,1819]" captionTargetId="figure-127@97.[189,1399,461,1819]" captionTargetPageId="97" captionText="Fig. 38. AB. Calliostoma pantopunctatum sp. nov., holotype, living animal, shell diameter 5.8 mm (MNHN-IM-2013-67275). CE. Kaiparathina monticola sp. nov., holotype, living animal, anterolateral structure arrowed, shell diameter 6.8 mm (MNHN-IM-2013-67168). F. Stomatella multilirata sp. nov., Walters Shoal, stn WS03, greatest shell dimension 6.0 mm (MNHN). Images courtesy of Alain Barrère/ MNHN." figureDoi="http://doi.org/10.5281/zenodo.10727854" httpUri="https://zenodo.org/record/10727854/files/figure.png" pageId="95" pageNumber="96">Figs 38CE</figureCitation>
,
<figureCitation id="F5D72A3838010B43FCD9F97254FAF919" box="[834,908,1712,1738]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="98.[189,232,1892,1918]" captionStartId-1="99.[189,232,1855,1881]" captionTargetBox-0="[189,1399,1030,1849]" captionTargetBox-1="[188,1398,745,1817]" captionTargetId-0="figure-286@98.[189,1399,1030,1849]" captionTargetId-1="figure-248@99.[188,1398,745,1817]" captionTargetPageId-0="98" captionTargetPageId-1="99" captionText-0="Fig. 39. Kaiparathina monticola sp. nov. A. Holotype, diameter 6.8 mm (MNHN-IM-2013-67168). B. Specimen with zig-zag colour pattern, diameter 6.0 mm (paratype, MNHN-IM-2000-35726). CF. Live-taken specimen with blotched colour pattern, diameter 6.4 mm (paratype, MNHN- IM-2000-35727)." captionText-1="Fig. 40. Kaiparathina monticola sp. nov. AC. Paratypes (MNHN-IM-2000-35726). A. Oblique view of apex. B. Sculptural detail of protoconch. C. Basal view of juvenile showing lack of spiral lirae. DG. Jaw and radula (paratype, MNHN-IM-2013-67260). D. Ventral (inner) surface of jaw, anterior edge below. E. Detail of jaw structure. F. Radula, entire width. G. Radula, detail of rachidian, lateral and inner marginal teeth. Scale bars: A, D =200 μm; B, F = 100 μm; C=1.0 mm; E, G =50 μm." figureDoi-0="http://doi.org/10.5281/zenodo.10727857" figureDoi-1="http://doi.org/10.5281/zenodo.10727861" httpUri-0="https://zenodo.org/record/10727857/files/figure.png" httpUri-1="https://zenodo.org/record/10727861/files/figure.png" pageId="95" pageNumber="96">3940</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="25F6653638010B43FF26F933546EF83A" pageId="95" pageNumber="96" type="diagnosis">
<paragraph id="6D5336BD38010B43FF26F9335640F8D8" blockId="95.[189,310,1777,1803]" box="[189,310,1777,1803]" pageId="95" pageNumber="96">
<heading id="361B81D138010B43FF26F9335640F8D8" bold="true" box="[189,310,1777,1803]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
<emphasis id="5F98EAAF38010B43FF26F9335640F8D8" bold="true" box="[189,310,1777,1803]" pageId="95" pageNumber="96">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38010B43FF26F8DD546EF83A" blockId="95.[189,1399,1823,2025]" pageId="95" pageNumber="96">Shell elevated-turbiniform, height up to 8.0 mm; whorls strongly convex, periphery often weakly angular, well below mid-whorl; early teleoconch whorls with fine, close-set, spiral threads, but shell smooth and glossy thereafter; anomphalous; 13 indistinct spiral cords adjacent to umbilical region; columella noticeably thickened, reflected over umbilical region; shell pale corneous brown, variously marked with white blotches often in two spiral bands, occasionally merging into zig-zag axial flames; protoconch white, its tip sometimes tinged maroon.</paragraph>
</subSubSection>
<subSubSection id="25F66536383E0B7CFF26FECB566FFEA7" pageId="96" pageNumber="97" type="etymology">
<paragraph id="6D5336BD383E0B7CFF26FECB5632FEF0" blockId="96.[189,324,265,291]" box="[189,324,265,291]" pageId="96" pageNumber="97">
<heading id="361B81D1383E0B7CFF26FECB5632FEF0" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis id="5F98EAAF383E0B7CFF26FECB5632FEF0" bold="true" box="[189,324,265,291]" pageId="96" pageNumber="97">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD383E0B7CFF26FEFA566FFEA7" blockId="96.[189,1399,311,372]" pageId="96" pageNumber="97">
From the Latin “
<emphasis id="5F98EAAF383E0B7CFE1BFEFA56C8FE82" box="[384,446,312,337]" italics="true" pageId="96" pageNumber="97">mons</emphasis>
“a mountain” and “
<emphasis id="5F98EAAF383E0B7CFD51FEF55473FE82" box="[714,773,311,337]" italics="true" pageId="96" pageNumber="97">-cola</emphasis>
“a dweller”; reflecting the seamount habitat of the species.
</paragraph>
</subSubSection>
<subSubSection id="25F66536383E0B7CFF26FE5955EFFB6D" pageId="96" pageNumber="97" type="materials_examined">
<paragraph id="6D5336BD383E0B7CFF26FE5956DCFE66" blockId="96.[189,426,411,437]" box="[189,426,411,437]" pageId="96" pageNumber="97">
<heading id="361B81D1383E0B7CFF26FE5956DCFE66" bold="true" box="[189,426,411,437]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis id="5F98EAAF383E0B7CFF26FE5956DCFE66" bold="true" box="[189,426,411,437]" pageId="96" pageNumber="97">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="DD843CE0383E0B7CFF71FE0853D1FDF9" ID-GBIF-Occurrence="4538174533" collectingDate="2017-05-02" collectionCode="MNHN" elevation="380" latitude="-33.266666" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.833332" pageId="96" pageNumber="97" specimenCode="MNHN-IM-2013-67168" specimenCount="1" typeStatus="holotype">
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<typeStatus id="B257881F383E0B7CFF71FE08562AFE37" box="[234,348,458,484]" pageId="96" pageNumber="97" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD383E0B7CFF26FE2C53D1FDF9" blockId="96.[189,1399,458,555]" pageId="96" pageNumber="97">
<location id="68336066383E0B7CFF26FE2C56D8FDDB" LSID="urn:lsid:plazi:treatment:E54587AB38010B78FDC3F9AB5638F93F:68336066383E0B7CFF26FE2C56D8FDDB" box="[189,430,494,520]" latitude="-33.266666" longLatPrecision="1205" longitude="43.833332" name="WALTERS SHOAL" pageId="96" pageNumber="97">WALTERS SHOAL</location>
• living
<specimenCount id="7BEAFD34383E0B7CFD87FE2F55F9FDD4" box="[540,655,493,519]" pageId="96" pageNumber="97" type="generic">specimen</specimenCount>
; slopes, stn DW4881;
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,
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; depth
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</quantity>
;
<date id="1952107D383E0B7CFF26FDD35638FDF8" box="[189,334,529,555]" pageId="96" pageNumber="97" value="2017-05-02">
<collectingDate id="0916E995383E0B7CFF26FDD35638FDF8" box="[189,334,529,555]" pageId="96" pageNumber="97" value="2017-05-02">2 May 2017</collectingDate>
</date>
; DNA tissue sample, photo alive (
<figureCitation id="F5D72A38383E0B7CFD44FDD3541DFDF8" box="[735,875,529,555]" captionStart="Fig" captionStartId="97.[188,231,1860,1886]" captionTargetBox="[189,1399,461,1819]" captionTargetId="figure-127@97.[189,1399,461,1819]" captionTargetPageId="97" captionText="Fig. 38. AB. Calliostoma pantopunctatum sp. nov., holotype, living animal, shell diameter 5.8 mm (MNHN-IM-2013-67275). CE. Kaiparathina monticola sp. nov., holotype, living animal, anterolateral structure arrowed, shell diameter 6.8 mm (MNHN-IM-2013-67168). F. Stomatella multilirata sp. nov., Walters Shoal, stn WS03, greatest shell dimension 6.0 mm (MNHN). Images courtesy of Alain Barrère/ MNHN." figureDoi="http://doi.org/10.5281/zenodo.10727854" httpUri="https://zenodo.org/record/10727854/files/figure.png" pageId="96" pageNumber="97">Fig. 38CE</figureCitation>
);
<specimenCode id="3D4A9EC6383E0B7CFC1BFDD253D4FDF9" box="[896,1186,528,554]" pageId="96" pageNumber="97">MNHN-IM-2013-67168</specimenCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFF71FD905536FD61" ID-GBIF-Occurrence="4538174494" collectingDate="2017-05-02" collectionCode="MNHN" elevation="380" latitude="-33.266666" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.833332" pageId="96" pageNumber="97" specimenCode="MNHN-IM-2013-67167" specimenCount="1" typeStatus="paratype">
<paragraph id="6D5336BD383E0B7CFF71FD905611FDBF" blockId="96.[189,1399,594,938]" box="[234,359,594,620]" pageId="96" pageNumber="97">
<emphasis id="5F98EAAF383E0B7CFF71FD905611FDBF" bold="true" box="[234,359,594,620]" pageId="96" pageNumber="97">
<typeStatus id="B257881F383E0B7CFF71FD905611FDBF" box="[234,359,594,620]" pageId="96" pageNumber="97" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD383E0B7CFF26FDB75428FC79" blockId="96.[189,1399,594,938]" pageId="96" pageNumber="97">
<location id="68336066383E0B7CFF26FDB756DBFD5C" LSID="urn:lsid:plazi:treatment:E54587AB38010B78FDC3F9AB5638F93F:68336066383E0B7CFF26FDB756DBFD5C" box="[189,429,629,655]" name="WALTERS SHOAL" pageId="96" pageNumber="97">WALTERS SHOAL</location>
<emphasis id="5F98EAAF383E0B7CFE4BFDB7556DFD5C" bold="true" box="[464,539,629,655]" pageId="96" pageNumber="97">slopes</emphasis>
• 1 living
<specimenCount id="7BEAFD34383E0B7CFD39FDB75463FD5C" box="[674,789,629,655]" pageId="96" pageNumber="97" type="generic">specimen</specimenCount>
; same collection data as for holotype; DNA tissue sample;
<specimenCode id="3D4A9EC6383E0B7CFEBAFD5A5536FD61" box="[289,576,664,690]" pageId="96" pageNumber="97">MNHN-IM-2013-67167</specimenCode>
<materialsCitation id="DD843CE0383E0B7CFDC7FD5B560FFD05" ID-GBIF-Occurrence="4538174373" collectingDate="2017-05-02" collectionCode="MNHN-" elevation="380" latitude="-33.266666" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.833332" pageId="96" pageNumber="97" specimenCode="MNHN- IM-2000-35726" specimenCount="4" typeStatus="paratype">
<specimenCount id="7BEAFD34383E0B7CFDC7FD5B5587FD61" box="[604,753,664,691]" count="4" pageId="96" pageNumber="97" type="generic">4 specimens</specimenCount>
, living; same collection data as for holotype;
<specimenCode id="3D4A9EC6383E0B7CFA88FD5B560FFD05" pageId="96" pageNumber="97">MNHN- IM-2000-35726</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFE09FD7E55A1FD29" ID-GBIF-Occurrence="4538174431" collectingDate="2017-05-04" collectionCode="MNHN" elevation="540" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="43.983334" pageId="96" pageNumber="97" specimenCode="MNHN-IM-2013-67258" specimenCount="1" typeStatus="paratype">
1 living
<specimenCount id="7BEAFD34383E0B7CFE6FFD7E5511FD05" box="[500,615,700,726]" pageId="96" pageNumber="97" type="generic">specimen</specimenCount>
; stn DW4890;
<geoCoordinate id="08D8507A383E0B7CFC83FD785409FD05" box="[792,895,698,726]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFC17FD785483FD05" box="[908,1013,698,726]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.983334">43°59 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFBD0FD7953A5FD05" box="[1099,1235,699,726]" metricMagnitude="2" metricUnit="m" metricValue="5.4" metricValueMax="5.88" metricValueMin="4.92" pageId="96" pageNumber="97" unit="m" value="540.0" valueMax="588.0" valueMin="492.0">
<elevation id="E6C1D18E383E0B7CFBD0FD7953A5FD05" box="[1099,1235,699,726]" metricMagnitude="2" metricUnit="m" metricValue="5.4" metricValueMax="5.88" metricValueMin="4.92" pageId="96" pageNumber="97" unit="m" value="540.0" valueMax="588.0" valueMin="492.0">492588 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFB7BFD7E5204FD05" box="[1248,1394,700,726]" pageId="96" pageNumber="97" value="2017-05-04">
<collectingDate id="0916E995383E0B7CFB7BFD7E5204FD05" box="[1248,1394,700,726]" pageId="96" pageNumber="97" value="2017-05-04">4 May 2017</collectingDate>
</date>
; DNA tissue sample;
<specimenCode id="3D4A9EC6383E0B7CFE22FD1D55A1FD29" box="[441,727,735,762]" pageId="96" pageNumber="97">MNHN-IM-2013-67258</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFD6CFD1D5304FCCE" ID-GBIF-Occurrence="4538174429" collectingDate="2017-05-04" collectionCode="MNHN" elevation="540" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="43.983334" pageId="96" pageNumber="97" specimenCode="MNHN-IM-2013-67259" specimenCount="1" typeStatus="paratype">
1 living
<specimenCount id="7BEAFD34383E0B7CFCC4FD1D54A4FD2A" box="[863,978,735,761]" pageId="96" pageNumber="97" type="generic">specimen</specimenCount>
; stn DW4890;
<geoCoordinate id="08D8507A383E0B7CFB10FD1F5383FD2A" box="[1163,1269,733,761]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFA9EFD1F5207FD2A" box="[1285,1393,733,761]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.983334">43°59 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFE91FCC056E1FCCE" box="[266,407,770,797]" metricMagnitude="2" metricUnit="m" metricValue="5.4" metricValueMax="5.88" metricValueMin="4.92" pageId="96" pageNumber="97" unit="m" value="540.0" valueMax="588.0" valueMin="492.0">
<elevation id="E6C1D18E383E0B7CFE91FCC056E1FCCE" box="[266,407,770,797]" metricMagnitude="2" metricUnit="m" metricValue="5.4" metricValueMax="5.88" metricValueMin="4.92" pageId="96" pageNumber="97" unit="m" value="540.0" valueMax="588.0" valueMin="492.0">492588 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFE33FCC05534FCCF" box="[424,578,770,797]" pageId="96" pageNumber="97" value="2017-05-04">
<collectingDate id="0916E995383E0B7CFE33FCC05534FCCF" box="[424,578,770,797]" pageId="96" pageNumber="97" value="2017-05-04">4 May 2017</collectingDate>
</date>
; DNA tissue sample;
<specimenCode id="3D4A9EC6383E0B7CFCC8FCC05304FCCE" box="[851,1138,770,797]" pageId="96" pageNumber="97">MNHN-IM-2013-67259</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFB0FFCC0560EFCB1" ID-GBIF-Occurrence="4538174503" collectingDate="2017-05-05" collectionCode="MNHN" elevation="537" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="43.983334" pageId="96" pageNumber="97" specimenCode="MNHN- IM-2013-67260" specimenCount="1" typeStatus="paratype">
1 living
<specimenCount id="7BEAFD34383E0B7CFA9BFCC05205FCCF" box="[1280,1395,770,796]" pageId="96" pageNumber="97" type="generic">specimen</specimenCount>
; stn DW4897;
<geoCoordinate id="08D8507A383E0B7CFEF5FCE656ACFCEC" box="[366,474,804,832]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFE76FCE6552BFC93" box="[493,605,804,832]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.983334">43°59 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFD25FCE7543AFC93" box="[702,844,805,832]" metricMagnitude="2" metricUnit="m" metricValue="5.37" metricValueMax="5.84" metricValueMin="4.9" pageId="96" pageNumber="97" unit="m" value="537.0" valueMax="584.0" valueMin="490.0">
<elevation id="E6C1D18E383E0B7CFD25FCE7543AFC93" box="[702,844,805,832]" metricMagnitude="2" metricUnit="m" metricValue="5.37" metricValueMax="5.84" metricValueMin="4.9" pageId="96" pageNumber="97" unit="m" value="537.0" valueMax="584.0" valueMin="490.0">490584 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFCC4FCE7548AFCEC" box="[863,1020,805,832]" pageId="96" pageNumber="97" value="2017-05-05">
<collectingDate id="0916E995383E0B7CFCC4FCE7548AFCEC" box="[863,1020,805,832]" pageId="96" pageNumber="97" value="2017-05-05">5 May 2017</collectingDate>
</date>
; DNA tissue sample;
<specimenCode id="3D4A9EC6383E0B7CFA88FCE4560EFCB1" pageId="96" pageNumber="97">MNHN- IM-2013-67260</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFE0BFC8A55B3FC55" ID-GBIF-Occurrence="4538174309" collectingDate="2017-05-05" collectionCode="MNHN" elevation="537" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="43.983334" pageId="96" pageNumber="97" specimenCode="MNHN-IM-2000-35727" specimenCount="5" typeStatus="paratype">
<specimenCount id="7BEAFD34383E0B7CFE0BFC8A5554FCB1" box="[400,546,840,867]" count="5" pageId="96" pageNumber="97" type="generic">5 specimens</specimenCount>
, living; stn DW4897;
<geoCoordinate id="08D8507A383E0B7CFCB8FC8554FEFCB0" box="[803,904,839,867]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFC0FFC85548AFCB0" box="[916,1020,839,867]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.983334">43°59 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFBCBFC8A53A1FCB0" box="[1104,1239,840,867]" metricMagnitude="2" metricUnit="m" metricValue="5.37" metricValueMax="5.84" metricValueMin="4.9" pageId="96" pageNumber="97" unit="m" value="537.0" valueMax="584.0" valueMin="490.0">
<elevation id="E6C1D18E383E0B7CFBCBFC8A53A1FCB0" box="[1104,1239,840,867]" metricMagnitude="2" metricUnit="m" metricValue="5.37" metricValueMax="5.84" metricValueMin="4.9" pageId="96" pageNumber="97" unit="m" value="537.0" valueMax="584.0" valueMin="490.0">490584 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFB79FC8A5204FCB0" box="[1250,1394,840,867]" pageId="96" pageNumber="97" value="2017-05-05">
<collectingDate id="0916E995383E0B7CFB79FC8A5204FCB0" box="[1250,1394,840,867]" pageId="96" pageNumber="97" value="2017-05-05">5 May 2017</collectingDate>
</date>
; radula SEM prep.;
<specimenCode id="3D4A9EC6383E0B7CFE3DFCA955B3FC55" box="[422,709,875,902]" pageId="96" pageNumber="97">MNHN-IM-2000-35727</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFD7FFCAE5428FC79" ID-GBIF-Occurrence="4538174577" collectingDate="2017-05-06" collectionCode="MNHN" elevation="660" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="44.016666" pageId="96" pageNumber="97" specimenCode="MNHN-IM-2000-35728" specimenCount="2" typeStatus="paratype">
<specimenCount id="7BEAFD34383E0B7CFD7FFCAE540CFC55" box="[740,890,876,902]" count="2" pageId="96" pageNumber="97" type="generic">2 specimens</specimenCount>
, living; stn DW4898;
<geoCoordinate id="08D8507A383E0B7CFB10FCA85383FC55" box="[1163,1269,874,902]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFA9EFCA85207FC55" box="[1285,1393,874,902]" degrees="44" direction="east" minutes="01" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="44.016666">44°01 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFE9EFC4D56FBFC7A" box="[261,397,911,938]" metricMagnitude="2" metricUnit="m" metricValue="6.6" metricValueMax="6.68" metricValueMin="6.52" pageId="96" pageNumber="97" unit="m" value="660.0" valueMax="668.0" valueMin="652.0">
<elevation id="E6C1D18E383E0B7CFE9EFC4D56FBFC7A" box="[261,397,911,938]" metricMagnitude="2" metricUnit="m" metricValue="6.6" metricValueMax="6.68" metricValueMin="6.52" pageId="96" pageNumber="97" unit="m" value="660.0" valueMax="668.0" valueMin="652.0">652668 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFE01FC4D555DFC7A" box="[410,555,911,937]" pageId="96" pageNumber="97" value="2017-05-06">
<collectingDate id="0916E995383E0B7CFE01FC4D555DFC7A" box="[410,555,911,937]" pageId="96" pageNumber="97" value="2017-05-06">6 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC6383E0B7CFDACFC4D542CFC79" box="[567,858,911,938]" pageId="96" pageNumber="97">MNHN-IM-2000-35728</specimenCode>
.
</materialsCitation>
</paragraph>
</materialsCitation>
<paragraph id="6D5336BD383E0B7CFF69FC1256DBFC39" blockId="96.[188,1399,976,1214]" box="[242,429,976,1002]" pageId="96" pageNumber="97">
<emphasis id="5F98EAAF383E0B7CFF69FC1256DBFC39" bold="true" box="[242,429,976,1002]" pageId="96" pageNumber="97">Other material</emphasis>
</paragraph>
<paragraph id="6D5336BD383E0B7CFF26FC3655EFFB6D" blockId="96.[188,1399,976,1214]" pageId="96" pageNumber="97">
<materialsCitation id="DD843CE0383E0B7CFF26FC365274FBDD" ID-GBIF-Occurrence="4538174334" box="[189,1282,1011,1038]" collectingDate="2017-05-02" collectionCode="MNHN" elevation="380" latitude="-33.266666" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.833332" pageId="96" pageNumber="97" specimenCode="MNHN-IM-2013-67168" specimenCount="40">
<location id="68336066383E0B7CFF26FC3656DFFBDD" LSID="urn:lsid:plazi:treatment:E54587AB38010B78FDC3F9AB5638F93F:68336066383E0B7CFF26FC3656DFFBDD" box="[189,425,1012,1038]" name="WALTERS SHOAL" pageId="96" pageNumber="97">WALTERS SHOAL</location>
<emphasis id="5F98EAAF383E0B7CFE59FC36557BFBDD" bold="true" box="[450,525,1012,1038]" pageId="96" pageNumber="97">slopes</emphasis>
<specimenCount id="7BEAFD34383E0B7CFDBEFC365535FBDD" box="[549,579,1012,1038]" pageId="96" pageNumber="97">40</specimenCount>
+ empty shells; same collection data as for holotype;
<collectionCode id="0BFDAE78383E0B7CFB33FC365274FBDD" box="[1192,1282,1012,1038]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="96" pageNumber="97">MNHN</collectionCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFA83FC3653EAFBE2" ID-GBIF-Occurrence="4538174534" collectingDate="2017-05-03" collectionCode="MNHN" elevation="326" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.916668" pageId="96" pageNumber="97" specimenCount="1">
1 empty shell; stn DW4885;
<geoCoordinate id="08D8507A383E0B7CFE2EFBD75569FBE2" box="[437,543,1045,1073]" degrees="33" direction="south" minutes="17" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.283333">33°17 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFDABFBD755E8FBE2" box="[560,670,1045,1073]" degrees="43" direction="east" minutes="55" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.916668">43°55 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFD60FBD554F1FBE2" box="[763,903,1047,1073]" metricMagnitude="2" metricUnit="m" metricValue="3.26" metricValueMax="3.8" metricValueMin="2.7199999999999998" pageId="96" pageNumber="97" unit="m" value="326.0" valueMax="380.0" valueMin="272.0">
<elevation id="E6C1D18E383E0B7CFD60FBD554F1FBE2" box="[763,903,1047,1073]" metricMagnitude="2" metricUnit="m" metricValue="3.26" metricValueMax="3.8" metricValueMin="2.7199999999999998" pageId="96" pageNumber="97" unit="m" value="326.0" valueMax="380.0" valueMin="272.0">272380 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFC03FBD55347FBE2" box="[920,1073,1047,1073]" pageId="96" pageNumber="97" value="2017-05-03">
<collectingDate id="0916E995383E0B7CFC03FBD55347FBE2" box="[920,1073,1047,1073]" pageId="96" pageNumber="97" value="2017-05-03">3 May 2017</collectingDate>
</date>
;
<collectionCode id="0BFDAE78383E0B7CFBD9FBD553EAFBE2" box="[1090,1180,1047,1073]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="96" pageNumber="97">MNHN</collectionCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFB27FBD55379FB87" ID-GBIF-Occurrence="4538174648" collectingDate="2017-05-03" collectionCode="MNHN" elevation="578" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.933334" pageId="96" pageNumber="97" specimenCount="6">
<specimenCount id="7BEAFD34383E0B7CFB27FBD553BDFBE2" box="[1212,1227,1047,1073]" pageId="96" pageNumber="97">6</specimenCount>
empty shells; stn DW4886;
<geoCoordinate id="08D8507A383E0B7CFEC4FBFA56B2FB87" box="[351,452,1080,1108]" degrees="33" direction="south" minutes="17" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.283333">33°17 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFE4BFBFA554EFB87" box="[464,568,1080,1108]" degrees="43" direction="east" minutes="56" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.933334">43°56 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFD11FBF85467FB87" box="[650,785,1082,1109]" metricMagnitude="2" metricUnit="m" metricValue="5.775" metricValueMax="5.82" metricValueMin="5.7299999999999995" pageId="96" pageNumber="97" unit="m" value="577.5" valueMax="582.0" valueMin="573.0">
<elevation id="E6C1D18E383E0B7CFD11FBF85467FB87" box="[650,785,1082,1109]" metricMagnitude="2" metricUnit="m" metricValue="5.775" metricValueMax="5.82" metricValueMin="5.7299999999999995" pageId="96" pageNumber="97" unit="m" value="577.5" valueMax="582.0" valueMin="573.0">573582 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFC87FBF854DCFB87" box="[796,938,1082,1108]" pageId="96" pageNumber="97" value="2017-05-03">
<collectingDate id="0916E995383E0B7CFC87FBF854DCFB87" box="[796,938,1082,1108]" pageId="96" pageNumber="97" value="2017-05-03">3 May 2017</collectingDate>
</date>
;
<collectionCode id="0BFDAE78383E0B7CFC2EFBF85379FB87" box="[949,1039,1082,1108]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="96" pageNumber="97">MNHN</collectionCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFBBEFBF854F7FBAB" ID-GBIF-Occurrence="4538174671" collectingDate="2017-05-03" collectionCode="MNHN" elevation="620" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.95" pageId="96" pageNumber="97" specimenCount="2">
<specimenCount id="7BEAFD34383E0B7CFBBEFBF85342FB87" box="[1061,1076,1082,1108]" pageId="96" pageNumber="97">2</specimenCount>
empty shells; stn DW4887;
<geoCoordinate id="08D8507A383E0B7CFF26FB995652FBA4" box="[189,292,1115,1143]" degrees="33" direction="south" minutes="17" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.283333">33°17 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFEA9FB9956EDFBAB" box="[306,411,1115,1144]" degrees="43" direction="east" minutes="57" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.95">43°57 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFE69FB9F550DFBAB" box="[498,635,1117,1144]" metricMagnitude="2" metricUnit="m" metricValue="6.195" metricValueMax="6.4" metricValueMin="5.99" pageId="96" pageNumber="97" unit="m" value="619.5" valueMax="640.0" valueMin="599.0">
<elevation id="E6C1D18E383E0B7CFE69FB9F550DFBAB" box="[498,635,1117,1144]" metricMagnitude="2" metricUnit="m" metricValue="6.195" metricValueMax="6.4" metricValueMin="5.99" pageId="96" pageNumber="97" unit="m" value="619.5" valueMax="640.0" valueMin="599.0">599640 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFD13FB9F546CFBA4" box="[648,794,1117,1144]" pageId="96" pageNumber="97" value="2017-05-03">
<collectingDate id="0916E995383E0B7CFD13FB9F546CFBA4" box="[648,794,1117,1144]" pageId="96" pageNumber="97" value="2017-05-03">3 May 2017</collectingDate>
</date>
;
<collectionCode id="0BFDAE78383E0B7CFCBCFB9C54F7FBAB" box="[807,897,1118,1144]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="96" pageNumber="97">MNHN</collectionCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFC00FB9F547DFB48" ID-GBIF-Occurrence="4538174434" collectingDate="2017-05-04" collectionCode="MNHN" elevation="540" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="43.983334" pageId="96" pageNumber="97" specimenCount="11">
<specimenCount id="7BEAFD34383E0B7CFC00FB9F54C1FBA4" box="[923,951,1117,1143]" pageId="96" pageNumber="97">11</specimenCount>
empty shells; stn DW4890;
<geoCoordinate id="08D8507A383E0B7CFA91FB995207FBA4" box="[1290,1393,1115,1143]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFF26FBBD5651FB48" box="[189,295,1151,1179]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.983334">43°59 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFEE6FB425573FB48" box="[381,517,1152,1179]" metricMagnitude="2" metricUnit="m" metricValue="5.4" metricValueMax="5.88" metricValueMin="4.92" pageId="96" pageNumber="97" unit="m" value="540.0" valueMax="588.0" valueMin="492.0">
<elevation id="E6C1D18E383E0B7CFEE6FB425573FB48" box="[381,517,1152,1179]" metricMagnitude="2" metricUnit="m" metricValue="5.4" metricValueMax="5.88" metricValueMin="4.92" pageId="96" pageNumber="97" unit="m" value="540.0" valueMax="588.0" valueMin="492.0">492588 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFD89FB4255D2FB49" box="[530,676,1152,1179]" pageId="96" pageNumber="97" value="2017-05-04">
<collectingDate id="0916E995383E0B7CFD89FB4255D2FB49" box="[530,676,1152,1179]" pageId="96" pageNumber="97" value="2017-05-04">4 May 2017</collectingDate>
</date>
;
<collectionCode id="0BFDAE78383E0B7CFD2AFB43547DFB48" box="[689,779,1153,1179]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="96" pageNumber="97">MNHN</collectionCode>
</materialsCitation>
<materialsCitation id="DD843CE0383E0B7CFCBEFB4255E3FB6D" ID-GBIF-Occurrence="4538174364" collectingDate="2017-05-05" collectionCode="MNHN" elevation="537" latitude="-33.15" location="WALTERS SHOAL" longLatPrecision="1206" longitude="43.983334" pageId="96" pageNumber="97" specimenCount="15">
<specimenCount id="7BEAFD34383E0B7CFCBEFB425434FB48" box="[805,834,1152,1179]" pageId="96" pageNumber="97">15</specimenCount>
empty shells; stn DW4897;
<geoCoordinate id="08D8507A383E0B7CFB08FBBD538CFB49" box="[1171,1274,1151,1179]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="96" pageNumber="97" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A383E0B7CFA93FBBD5207FB48" box="[1288,1393,1151,1179]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="96" pageNumber="97" precision="925" value="43.983334">43°59 E</geoCoordinate>
; depth
<quantity id="AA149B58383E0B7CFE9EFB6156FBFB6D" box="[261,397,1187,1214]" metricMagnitude="2" metricUnit="m" metricValue="5.37" metricValueMax="5.84" metricValueMin="4.9" pageId="96" pageNumber="97" unit="m" value="537.0" valueMax="584.0" valueMin="490.0">
<elevation id="E6C1D18E383E0B7CFE9EFB6156FBFB6D" box="[261,397,1187,1214]" metricMagnitude="2" metricUnit="m" metricValue="5.37" metricValueMax="5.84" metricValueMin="4.9" pageId="96" pageNumber="97" unit="m" value="537.0" valueMax="584.0" valueMin="490.0">490584 m</elevation>
</quantity>
;
<date id="1952107D383E0B7CFE01FB61555DFB6D" box="[410,555,1187,1214]" pageId="96" pageNumber="97" value="2017-05-05">
<collectingDate id="0916E995383E0B7CFE01FB61555DFB6D" box="[410,555,1187,1214]" pageId="96" pageNumber="97" value="2017-05-05">5 May 2017</collectingDate>
</date>
;
<collectionCode id="0BFDAE78383E0B7CFDACFB6655E3FB6D" box="[567,661,1188,1214]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="96" pageNumber="97">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="25F66536383E0B78FF26FB2756D8FE95" lastPageId="100" lastPageNumber="101" pageId="96" pageNumber="97" type="description">
<paragraph id="6D5336BD383E0B7CFF26FB275638FB2C" blockId="96.[189,334,1253,1279]" box="[189,334,1253,1279]" pageId="96" pageNumber="97">
<heading id="361B81D1383E0B7CFF26FB275638FB2C" bold="true" box="[189,334,1253,1279]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis id="5F98EAAF383E0B7CFF26FB275638FB2C" bold="true" box="[189,334,1253,1279]" pageId="96" pageNumber="97">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD383E0B7CFF26FAD653C4F8C9" blockId="96.[188,1400,1299,1819]" pageId="96" pageNumber="97">
<smallCapsWord id="6BB5A061383E0B7CFF26FAD65677FAFF" baselines="1319,1319" box="[189,257,1300,1326]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="96" pageNumber="97">SHELL</smallCapsWord>
. Relatively large for the genus (height up to 8.0 mm), elevated-turbiniform, slightly higher than wide; spire prominent with strongly convex whorls; periphery often weakly angular, well below mid-whorl; suture distinctly indented; sculpture weak, shell essentially smooth and glossy; base less strongly convex, umbilicus lacking; teleoconch of up to 5.0 whorls. First two teleoconch whorls with pair of fine, close-set, spiral threads midway between sutures (
<figureCitation id="F5D72A38383E0B7CFCD9FA6254D9FA69" box="[834,943,1440,1466]" captionStart="Fig" captionStartId="99.[189,232,1855,1881]" captionTargetBox="[188,1398,745,1817]" captionTargetId="figure-248@99.[188,1398,745,1817]" captionTargetPageId="99" captionText="Fig. 40. Kaiparathina monticola sp. nov. AC. Paratypes (MNHN-IM-2000-35726). A. Oblique view of apex. B. Sculptural detail of protoconch. C. Basal view of juvenile showing lack of spiral lirae. DG. Jaw and radula (paratype, MNHN-IM-2013-67260). D. Ventral (inner) surface of jaw, anterior edge below. E. Detail of jaw structure. F. Radula, entire width. G. Radula, detail of rachidian, lateral and inner marginal teeth. Scale bars: A, D =200 μm; B, F = 100 μm; C=1.0 mm; E, G =50 μm." figureDoi="http://doi.org/10.5281/zenodo.10727861" httpUri="https://zenodo.org/record/10727861/files/figure.png" pageId="96" pageNumber="97">Fig. 40A</figureCitation>
), second weaker pair below this, lower one of which close to or level with abapical suture; upper pair each developing shallow adapical groove during second whorl; spiral sculpture evanescing near start of third whorl and later whorls with at most traces of spiral sculpture, though in live-taken specimens spiral patterning in underlying nacre visible by transparency; axial sculpture of weak growth-lines and close-set, microscopic, collabral threads; juvenile and subadult shells with more angular periphery, delineated by low spiral cord; insertion of suture level with cord; cord weak or absent on much of last whorl in adult specimens; base sculptured as above, often with 13 indistinct spiral cords adjacent to umbilical region; growth-lines sigmoidal below periphery and somewhat stronger medially; umbilicus lacking. Aperture subcircular; columella noticeably thickened, reflected over umbilical region; outer lip thin, concave below periphery, convex near columella, basal lip thus sinuous; interior nacreous, lacking labral ornamentation.
</paragraph>
<paragraph id="6D5336BD383E0B7CFF26F8805517F815" blockId="96.[189,1399,1857,1990]" pageId="96" pageNumber="97">
<smallCapsWord id="6BB5A061383E0B7CFF26F880563AF889" baselines="1877,1877" box="[189,332,1858,1884]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Protoconch" pageId="96" pageNumber="97">PROTOCONCH</smallCapsWord>
(
<figureCitation id="F5D72A38383E0B7CFEF9F8805680F88F" box="[354,502,1858,1884]" captionStart="Fig" captionStartId="99.[189,232,1855,1881]" captionTargetBox="[188,1398,745,1817]" captionTargetId="figure-248@99.[188,1398,745,1817]" captionTargetPageId="99" captionText="Fig. 40. Kaiparathina monticola sp. nov. AC. Paratypes (MNHN-IM-2000-35726). A. Oblique view of apex. B. Sculptural detail of protoconch. C. Basal view of juvenile showing lack of spiral lirae. DG. Jaw and radula (paratype, MNHN-IM-2013-67260). D. Ventral (inner) surface of jaw, anterior edge below. E. Detail of jaw structure. F. Radula, entire width. G. Radula, detail of rachidian, lateral and inner marginal teeth. Scale bars: A, D =200 μm; B, F = 100 μm; C=1.0 mm; E, G =50 μm." figureDoi="http://doi.org/10.5281/zenodo.10727861" httpUri="https://zenodo.org/record/10727861/files/figure.png" pageId="96" pageNumber="97">Fig. 40AB</figureCitation>
). Globose, somewhat elevated, comprising approx. 1.25 whorls, diameter 350375 μm; tip of apex roundly angled, terminal lip convex, lacking terminal or subterminal varix; surface smooth save for three fine, widely spaced, subspiral threads, lowest one close to abapical suture, a fourth one delineating tip of apex.
</paragraph>
<paragraph id="6D5336BD383F0B7EFF26FECB5598FE00" blockId="97.[188,1399,265,397]" lastBlockId="98.[188,1398,265,467]" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98">
<smallCapsWord id="6BB5A061383F0B7DFF26FECB566EFEF2" baselines="284,284" box="[189,280,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="97" pageNumber="98">COLOUR</smallCapsWord>
. Protoconch white, its tip sometimes tinged maroon; teleoconch with pale corneous brown ground colour, translucent in live-taken specimens, with pink/green iridescence of underlying nacre showing through. Colour pattern variable; commonly with two spiral rows of white blotches (
<figureCitation id="F5D72A38383F0B7DFB65FE8D521FFEBA" box="[1278,1385,335,361]" captionStart="Fig" captionStartId="98.[189,232,1892,1918]" captionTargetBox="[189,1399,1030,1849]" captionTargetId="figure-286@98.[189,1399,1030,1849]" captionTargetPageId="98" captionText="Fig. 39. Kaiparathina monticola sp. nov. A. Holotype, diameter 6.8 mm (MNHN-IM-2013-67168). B. Specimen with zig-zag colour pattern, diameter 6.0 mm (paratype, MNHN-IM-2000-35726). CF. Live-taken specimen with blotched colour pattern, diameter 6.4 mm (paratype, MNHN- IM-2000-35727)." figureDoi="http://doi.org/10.5281/zenodo.10727857" httpUri="https://zenodo.org/record/10727857/files/figure.png" pageId="97" pageNumber="98">Fig. 39C</figureCitation>
), one at periphery, the other between this and adapical suture, sometimes a third row near middle of base; intervals between white blotches somewhat darker brown, particularly at periphery, on spire whorls this creates a thin supra-sutural line of alternating white/brown dashes; often also with thin subsutural white line; columella and medial area of base white; variation in white and brown markings considerable, the latter sometimes merging into zig-zag axial flames (
<figureCitation id="F5D72A38383C0B7EFCD5FEB054CCFE5E" box="[846,954,370,397]" captionStart="Fig" captionStartId="98.[189,232,1892,1918]" captionTargetBox="[189,1399,1030,1849]" captionTargetId="figure-286@98.[189,1399,1030,1849]" captionTargetPageId="98" captionText="Fig. 39. Kaiparathina monticola sp. nov. A. Holotype, diameter 6.8 mm (MNHN-IM-2013-67168). B. Specimen with zig-zag colour pattern, diameter 6.0 mm (paratype, MNHN-IM-2000-35726). CF. Live-taken specimen with blotched colour pattern, diameter 6.4 mm (paratype, MNHN- IM-2000-35727)." figureDoi="http://doi.org/10.5281/zenodo.10727857" httpUri="https://zenodo.org/record/10727857/files/figure.png" pageId="98" pageNumber="99">Fig. 39B</figureCitation>
); some specimens with broad whitish subsutural band, others with only alternating light and dark peripheral blotches or with blotches absent on spire whorls or present only on spire whorls.
</paragraph>
<caption id="39936635383F0B7DFF27F8865668F83F" ID-DOI="http://doi.org/10.5281/zenodo.10727854" ID-Zenodo-Dep="10727854" httpUri="https://zenodo.org/record/10727854/files/figure.png" pageId="97" pageNumber="98" startId="97.[188,231,1860,1886]" targetBox="[189,1399,461,1819]" targetPageId="97">
<paragraph id="6D5336BD383F0B7DFF27F8865668F83F" blockId="97.[188,1399,1860,2028]" pageId="97" pageNumber="98">
<emphasis id="5F98EAAF383F0B7DFF27F8865628F88D" bold="true" box="[188,350,1860,1886]" pageId="97" pageNumber="98">Fig. 38. AB</emphasis>
.
<taxonomicName id="AAEC4D3E383F0B7DFEF4F88655B1F88D" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[367,711,1860,1886]" class="Gastropoda" family="Calliostomatidae" genus="Calliostoma" kingdom="Animalia" order="Trochida" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="pantopunctatum" status="sp. nov.">
<emphasis id="5F98EAAF383F0B7DFEF4F88655B1F88D" box="[367,711,1860,1886]" italics="true" pageId="97" pageNumber="98">Calliostoma pantopunctatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4383F0B7DFD4AF8875447F88C" box="[721,817,1861,1887]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, living animal, shell diameter 5.8 mm (MNHN-IM-2013-67275).
<emphasis id="5F98EAAF383F0B7DFE66F8A55542F852" bold="true" box="[509,564,1895,1921]" pageId="97" pageNumber="98">CE</emphasis>
.
<taxonomicName id="AAEC4D3E383F0B7DFDD9F8AA542DF851" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[578,859,1896,1922]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="monticola" status="sp. nov.">
<emphasis id="5F98EAAF383F0B7DFDD9F8AA542DF851" box="[578,859,1896,1922]" italics="true" pageId="97" pageNumber="98">Kaiparathina monticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4383F0B7DFCFAF8AA54CBF851" box="[865,957,1896,1922]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, living animal, anterolateral structure arrowed, shell diameter 6.8 mm (MNHN-IM-2013-67168).
<emphasis id="5F98EAAF383F0B7DFC71F849548AF876" bold="true" box="[1002,1020,1931,1957]" pageId="97" pageNumber="98">F</emphasis>
.
<taxonomicName id="AAEC4D3E383F0B7DFB90F8495278F876" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[1035,1294,1931,1957]" class="Gastropoda" family="Trochidae" genus="Stomatella" kingdom="Animalia" order="Trochida" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="multilirata" status="sp. nov.">
<emphasis id="5F98EAAF383F0B7DFB90F8495278F876" box="[1035,1294,1931,1957]" italics="true" pageId="97" pageNumber="98">Stomatella multilirata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4383F0B7DFA8EF8495204F876" box="[1301,1394,1931,1957]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
, Walters Shoal, stn WS03, greatest shell dimension 6.0 mm (MNHN). Images courtesy of Alain Barrère/ MNHN.
</paragraph>
</caption>
<paragraph id="6D5336BD383C0B7EFF26FE395668FDEB" blockId="98.[189,1399,506,568]" pageId="98" pageNumber="99">
<smallCapsWord id="6BB5A061383C0B7EFF26FE395634FDC0" baselines="526,526" box="[189,322,507,533]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dimensions" pageId="98" pageNumber="99">DIMENSIONS</smallCapsWord>
.
<typeStatus id="B257881F383C0B7EFECFFE3856B2FDC7" box="[340,452,506,532]" pageId="98" pageNumber="99" type="holotype">Holotype</typeStatus>
, height
<quantity id="AA149B58383C0B7EFDBDFE3855F0FDC6" box="[550,646,506,533]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="98" pageNumber="99" unit="mm" value="6.9">6.9 mm</quantity>
, diameter
<quantity id="AA149B58383C0B7EFC9DFE385410FDC6" box="[774,870,506,533]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="98" pageNumber="99" unit="mm" value="6.8">6.8 mm</quantity>
; largest specimen, height 8.0 mm, diameter
<quantity id="AA149B58383C0B7EFF26FDDF566CFDEB" box="[189,282,541,568]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="98" pageNumber="99" unit="mm" value="7.6">7.6 mm</quantity>
.
</paragraph>
<paragraph id="6D5336BD383C0B7EFF26FD9D5238FDAA" blockId="98.[189,1358,607,634]" box="[189,1358,607,634]" pageId="98" pageNumber="99">
<smallCapsWord id="6BB5A061383C0B7EFF26FD9D5634FDAB" baselines="626,627" box="[189,322,607,633]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Operculum" pageId="98" pageNumber="99">OPERCULUM</smallCapsWord>
. Corneous, thin, light yellowish-brown, tightly multispiral with a short growing margin.
</paragraph>
<paragraph id="6D5336BD383C0B7EFF26FD635545FCD2" blockId="98.[189,1401,672,769]" pageId="98" pageNumber="99">
<smallCapsWord id="6BB5A061383C0B7EFF26FD63579FFD6A" baselines="692,692" box="[189,233,673,699]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Jaw" pageId="98" pageNumber="99">JAW</smallCapsWord>
(
<figureCitation id="F5D72A38383C0B7EFF67FD6256FAFD68" box="[252,396,672,699]" captionStart="Fig" captionStartId="99.[189,232,1855,1881]" captionTargetBox="[188,1398,745,1817]" captionTargetId="figure-248@99.[188,1398,745,1817]" captionTargetPageId="99" captionText="Fig. 40. Kaiparathina monticola sp. nov. AC. Paratypes (MNHN-IM-2000-35726). A. Oblique view of apex. B. Sculptural detail of protoconch. C. Basal view of juvenile showing lack of spiral lirae. DG. Jaw and radula (paratype, MNHN-IM-2013-67260). D. Ventral (inner) surface of jaw, anterior edge below. E. Detail of jaw structure. F. Radula, entire width. G. Radula, detail of rachidian, lateral and inner marginal teeth. Scale bars: A, D =200 μm; B, F = 100 μm; C=1.0 mm; E, G =50 μm." figureDoi="http://doi.org/10.5281/zenodo.10727861" httpUri="https://zenodo.org/record/10727861/files/figure.png" pageId="98" pageNumber="99">Fig. 40DE</figureCitation>
). Oval-quadrate, lateral portions rounded and dipping anteriorly, posterior edge thus weakly convex and anterior edge weakly concave; surface composed of minute, elongate hexagonal units (occasionally pentagonal).
</paragraph>
<paragraph id="6D5336BD383C0B7FFF26FCEB5446FEBA" blockId="98.[189,1398,808,976]" lastBlockId="99.[189,1401,265,362]" lastPageId="99" lastPageNumber="100" pageId="98" pageNumber="99">
<smallCapsWord id="6BB5A061383C0B7EFF26FCEB566FFC92" baselines="828,828" box="[189,281,809,835]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Radula" pageId="98" pageNumber="99">RADULA</smallCapsWord>
(
<figureCitation id="F5D72A38383C0B7EFEB0FCEA56CFFC91" box="[299,441,808,835]" captionStart="Fig" captionStartId="99.[189,232,1855,1881]" captionTargetBox="[188,1398,745,1817]" captionTargetId="figure-248@99.[188,1398,745,1817]" captionTargetPageId="99" captionText="Fig. 40. Kaiparathina monticola sp. nov. AC. Paratypes (MNHN-IM-2000-35726). A. Oblique view of apex. B. Sculptural detail of protoconch. C. Basal view of juvenile showing lack of spiral lirae. DG. Jaw and radula (paratype, MNHN-IM-2013-67260). D. Ventral (inner) surface of jaw, anterior edge below. E. Detail of jaw structure. F. Radula, entire width. G. Radula, detail of rachidian, lateral and inner marginal teeth. Scale bars: A, D =200 μm; B, F = 100 μm; C=1.0 mm; E, G =50 μm." figureDoi="http://doi.org/10.5281/zenodo.10727861" httpUri="https://zenodo.org/record/10727861/files/figure.png" pageId="98" pageNumber="99">Fig. 40FG</figureCitation>
). Formula ∞+7+1+7+∞, with ± 35 transverse rows of teeth; rachidian tooth very large and robust, its cusp basally trigonal but with convex sides; cutting edge dominated by large, narrowly pointed central denticle; basal part of cusp with finer lateral denticles; anterior margin of cusp concave, abutting convex posterior edge of base-plate of next anterior tooth. Lateral teeth slender, scythe-like, cusp elongate and slightly curved, its outer base flanged and bearing small denticles; shaft face with thickened pilaster; laterals 35 largest. Marginal teeth numerous, progressively reducing in size toward edge of radula, shafts of all but innermost one seemingly fused; cusps of inner marginals narrowly spathulate, outer edge minutely denticulate.
</paragraph>
<caption id="39936635383C0B7EFF26F8A656FCF83B" ID-DOI="http://doi.org/10.5281/zenodo.10727857" ID-Zenodo-Dep="10727857" httpUri="https://zenodo.org/record/10727857/files/figure.png" pageId="98" pageNumber="99" startId="98.[189,232,1892,1918]" targetBox="[189,1399,1030,1849]" targetPageId="98">
<paragraph id="6D5336BD383C0B7EFF26F8A656FCF83B" blockId="98.[189,1399,1892,2024]" pageId="98" pageNumber="99">
<emphasis id="5F98EAAF383C0B7EFF26F8A6566BF8AD" bold="true" box="[189,285,1892,1918]" pageId="98" pageNumber="99">Fig. 39.</emphasis>
<taxonomicName id="AAEC4D3E383C0B7EFEB3F8A65530F8AD" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[296,582,1892,1918]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="98" pageNumber="99" phylum="Mollusca" rank="species" species="monticola" status="sp. nov.">
<emphasis id="5F98EAAF383C0B7EFEB3F8A65530F8AD" box="[296,582,1892,1918]" italics="true" pageId="98" pageNumber="99">Kaiparathina monticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4383C0B7EFDC9F8A755C6F8AC" box="[594,688,1893,1919]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5F98EAAF383C0B7EFD20F8A655A6F8AD" bold="true" box="[699,720,1892,1918]" pageId="98" pageNumber="99">A</emphasis>
. Holotype, diameter 6.8 mm (MNHN-IM-2013-67168).
<emphasis id="5F98EAAF383C0B7EFF26F84557A7F872" bold="true" box="[189,209,1927,1953]" pageId="98" pageNumber="99">B</emphasis>
. Specimen with zig-zag colour pattern, diameter 6.0 mm (paratype, MNHN-IM-2000-35726).
<emphasis id="5F98EAAF383C0B7EFF26F8685785F817" bold="true" box="[189,243,1962,1988]" pageId="98" pageNumber="99">CF</emphasis>
. Live-taken specimen with blotched colour pattern, diameter 6.4 mm (paratype, MNHN- IM-2000-35727).
</paragraph>
</caption>
<paragraph id="6D5336BD383D0B78FF26FE5356D8FE95" blockId="99.[189,1400,401,674]" lastBlockId="100.[189,1398,265,326]" lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100">
<smallCapsWord id="6BB5A061383D0B7FFF26FE535659FE7A" baselines="420,420" box="[189,303,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="External" pageId="99" pageNumber="100">EXTERNAL</smallCapsWord>
<smallCapsWord id="6BB5A061383D0B7FFEA1FE5456D6FE7A" baselines="420" box="[314,416,406,425]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="anatomy" pageId="99" pageNumber="100">ANATOMY</smallCapsWord>
(
<figureCitation id="F5D72A38383D0B7FFE2DFE535533FE78" box="[438,581,401,427]" captionStart="Fig" captionStartId="97.[188,231,1860,1886]" captionTargetBox="[189,1399,461,1819]" captionTargetId="figure-127@97.[189,1399,461,1819]" captionTargetPageId="97" captionText="Fig. 38. AB. Calliostoma pantopunctatum sp. nov., holotype, living animal, shell diameter 5.8 mm (MNHN-IM-2013-67275). CE. Kaiparathina monticola sp. nov., holotype, living animal, anterolateral structure arrowed, shell diameter 6.8 mm (MNHN-IM-2013-67168). F. Stomatella multilirata sp. nov., Walters Shoal, stn WS03, greatest shell dimension 6.0 mm (MNHN). Images courtesy of Alain Barrère/ MNHN." figureDoi="http://doi.org/10.5281/zenodo.10727854" httpUri="https://zenodo.org/record/10727854/files/figure.png" pageId="99" pageNumber="100">Fig. 38CE</figureCitation>
). As described for
<taxonomicName id="AAEC4D3E383D0B7FFCAAFE535267FE78" authority="(Marshall 1993)" baseAuthorityName="Marshall" baseAuthorityYear="1993" box="[817,1297,401,427]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="99" pageNumber="100" phylum="Mollusca" rank="species" species="boucheti">
<emphasis id="5F98EAAF383D0B7FFCAAFE535336FE78" box="[817,1088,401,427]" italics="true" pageId="99" pageNumber="100">Kaiparathina boucheti</emphasis>
(
<bibRefCitation id="097D4B4C383D0B7FFBC8FE53527EFE78" author="Marshall B. A." box="[1107,1288,401,427]" pageId="99" pageNumber="100" pagination="185 - 198" refId="ref61761" refString="Marshall B. A. 1993. A review of the genus Kaiparathina Laws, 1941 (Mollusca: Gastropoda: Trochidae). The Veliger 36 (2): 185 - 198. Available from https: // www. biodiversitylibrary. org / page / 42465795 [accessed 10 Feb. 2024]." type="journal article" year="1993">Marshall 1993</bibRefCitation>
)
</taxonomicName>
. Animal for the most part milky-white; head and neck tinged with pale apricot-orange; cephalic tentacles well developed, micro-papillate and with a faint orange-brown mid-dorsal line; eyes prominent, pigmented and with open aperture; anterolateral structures pale, coarsely pustulose (arrowed in
<figureCitation id="F5D72A38383D0B7FFB22FE385252FDC6" box="[1209,1316,506,533]" captionStart="Fig" captionStartId="97.[188,231,1860,1886]" captionTargetBox="[189,1399,461,1819]" captionTargetId="figure-127@97.[189,1399,461,1819]" captionTargetPageId="97" captionText="Fig. 38. AB. Calliostoma pantopunctatum sp. nov., holotype, living animal, shell diameter 5.8 mm (MNHN-IM-2013-67275). CE. Kaiparathina monticola sp. nov., holotype, living animal, anterolateral structure arrowed, shell diameter 6.8 mm (MNHN-IM-2013-67168). F. Stomatella multilirata sp. nov., Walters Shoal, stn WS03, greatest shell dimension 6.0 mm (MNHN). Images courtesy of Alain Barrère/ MNHN." figureDoi="http://doi.org/10.5281/zenodo.10727854" httpUri="https://zenodo.org/record/10727854/files/figure.png" pageId="99" pageNumber="100">Fig. 38E</figureCitation>
), lying beneath non-digitate neck lobes; three unpigmented, micro-papillate epipodial tentacles on each side, each with basal epipodial sense organ; sides of foot beneath epipodium orange-brown, colour progressively deepening ventrally, but then ceasing abruptly, lower portion comprising uniformly pale band, encircled by thinner orange-brown band (evanescing prior to propodial horns) and finally pale peripodial fringe; colour pattern continues onto metapodium, which has white mid-line band at extreme posterior (
<figureCitation id="F5D72A38383A0B78FEAEFEEE56D6FE95" box="[309,416,300,326]" captionStart="Fig" captionStartId="97.[188,231,1860,1886]" captionTargetBox="[189,1399,461,1819]" captionTargetId="figure-127@97.[189,1399,461,1819]" captionTargetPageId="97" captionText="Fig. 38. AB. Calliostoma pantopunctatum sp. nov., holotype, living animal, shell diameter 5.8 mm (MNHN-IM-2013-67275). CE. Kaiparathina monticola sp. nov., holotype, living animal, anterolateral structure arrowed, shell diameter 6.8 mm (MNHN-IM-2013-67168). F. Stomatella multilirata sp. nov., Walters Shoal, stn WS03, greatest shell dimension 6.0 mm (MNHN). Images courtesy of Alain Barrère/ MNHN." figureDoi="http://doi.org/10.5281/zenodo.10727854" httpUri="https://zenodo.org/record/10727854/files/figure.png" pageId="100" pageNumber="101">Fig. 38C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="39936635383D0B7FFF26F8FD5396F834" ID-DOI="http://doi.org/10.5281/zenodo.10727861" ID-Zenodo-Dep="10727861" httpUri="https://zenodo.org/record/10727861/files/figure.png" pageId="99" pageNumber="100" startId="99.[189,232,1855,1881]" targetBox="[188,1398,745,1817]" targetPageId="99">
<paragraph id="6D5336BD383D0B7FFF26F8FD5396F834" blockId="99.[188,1399,1855,2023]" pageId="99" pageNumber="100">
<emphasis id="5F98EAAF383D0B7FFF26F8FD566DF88A" bold="true" box="[189,283,1855,1881]" pageId="99" pageNumber="100">Fig. 40.</emphasis>
<taxonomicName id="AAEC4D3E383D0B7FFEB8F8FD5549F88A" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[291,575,1855,1881]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="99" pageNumber="100" phylum="Mollusca" rank="species" species="monticola" status="sp. nov.">
<emphasis id="5F98EAAF383D0B7FFEB8F8FD5549F88A" box="[291,575,1855,1881]" italics="true" pageId="99" pageNumber="100">Kaiparathina monticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4383D0B7FFDD2F88255D5F889" box="[585,675,1856,1882]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5F98EAAF383D0B7FFD36F8FD5593F88A" bold="true" box="[685,741,1855,1881]" pageId="99" pageNumber="100">AC</emphasis>
. Paratypes (MNHN-IM-2000-35726).
<emphasis id="5F98EAAF383D0B7FFB29F8FD53B0F88A" bold="true" box="[1202,1222,1855,1881]" pageId="99" pageNumber="100">A</emphasis>
. Oblique view of apex.
<emphasis id="5F98EAAF383D0B7FFEB1F8A15648F8AE" bold="true" box="[298,318,1891,1917]" pageId="99" pageNumber="100">B</emphasis>
. Sculptural detail of protoconch.
<emphasis id="5F98EAAF383D0B7FFD40F8A05586F8AF" bold="true" box="[731,752,1890,1916]" pageId="99" pageNumber="100">C</emphasis>
. Basal view of juvenile showing lack of spiral lirae.
<emphasis id="5F98EAAF383D0B7FFF26F8445781F873" bold="true" box="[189,247,1926,1952]" pageId="99" pageNumber="100">DG</emphasis>
. Jaw and radula (paratype, MNHN-IM-2013-67260).
<emphasis id="5F98EAAF383D0B7FFC19F84454E1F873" bold="true" box="[898,919,1926,1952]" pageId="99" pageNumber="100">D</emphasis>
. Ventral (inner) surface of jaw, anterior edge below.
<emphasis id="5F98EAAF383D0B7FFED7F86B5616F810" bold="true" box="[332,352,1961,1987]" pageId="99" pageNumber="100">E</emphasis>
. Detail of jaw structure.
<emphasis id="5F98EAAF383D0B7FFDE5F86B55E6F810" bold="true" box="[638,656,1961,1987]" pageId="99" pageNumber="100">F</emphasis>
. Radula, entire width.
<emphasis id="5F98EAAF383D0B7FFC0CF86B54D8F810" bold="true" box="[919,942,1961,1987]" pageId="99" pageNumber="100">G</emphasis>
. Radula, detail of rachidian, lateral and inner marginal teeth. Scale bars: A, D =200 μm; B, F = 100 μm; C=1.0 mm; E, G =50 μm.
</paragraph>
</caption>
<subSubSection id="25F66536383A0B78FF26FEAF53B7FE0A" pageId="100" pageNumber="101" type="distribution">
<paragraph id="6D5336BD383A0B78FF26FEAF5620FE54" blockId="100.[189,342,365,391]" box="[189,342,365,391]" pageId="100" pageNumber="101">
<heading id="361B81D1383A0B78FF26FEAF5620FE54" bold="true" box="[189,342,365,391]" fontSize="11" level="3" pageId="100" pageNumber="101" reason="3">
<emphasis id="5F98EAAF383A0B78FF26FEAF5620FE54" bold="true" box="[189,342,365,391]" pageId="100" pageNumber="101">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD383A0B78FF26FE5F53B7FE0A" blockId="100.[188,1398,412,474]" pageId="100" pageNumber="101">
Known only from the slopes of Walters Shoal, at depths of
<quantity id="AA149B58383A0B78FCF2FE5E549BFE64" box="[873,1005,412,439]" metricMagnitude="2" metricUnit="m" metricValue="5.16" metricValueMax="6.52" metricValueMin="3.8" pageId="100" pageNumber="101" unit="m" value="516.0" valueMax="652.0" valueMin="380.0">380652 m</quantity>
(live-taken material
<quantity id="AA149B58383A0B78FB7BFE5E521EFE65" box="[1248,1384,412,439]" metricMagnitude="2" metricUnit="m" metricValue="5.17" metricValueMax="6.52" metricValueMin="3.82" pageId="100" pageNumber="101" unit="m" value="517.0" valueMax="652.0" valueMin="382.0">382652 m</quantity>
); dredged on substrata of coarse sand and fine gravel, with ophiuroids and solitary corals.
</paragraph>
</subSubSection>
<subSubSection id="25F66536383A0B78FF26FDC35638F93F" pageId="100" pageNumber="101" type="discussion">
<paragraph id="6D5336BD383A0B78FF26FDC35659FDC8" blockId="100.[189,303,513,539]" box="[189,303,513,539]" pageId="100" pageNumber="101">
<heading id="361B81D1383A0B78FF26FDC35659FDC8" bold="true" box="[189,303,513,539]" fontSize="11" level="3" pageId="100" pageNumber="101" reason="3">
<emphasis id="5F98EAAF383A0B78FF26FDC35659FDC8" bold="true" box="[189,303,513,539]" pageId="100" pageNumber="101">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD383A0B78FF26FDF256FDFAA0" blockId="100.[187,1400,559,1395]" pageId="100" pageNumber="101">
In terms of its relatively large size,
<taxonomicName id="AAEC4D3E383A0B78FDEBFDED54FBFD9A" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[624,909,559,585]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola" status="sp. nov.">
<emphasis id="5F98EAAF383A0B78FDEBFDED54FBFD9A" box="[624,909,559,585]" italics="true" pageId="100" pageNumber="101">Kaiparathina monticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4383A0B78FC0CFDF25483FD99" box="[919,1013,560,586]" pageId="100" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="AAEC4D3E383A0B78FBE5FDF2578EFDBE" authority="Marshall, 1993" authorityName="Marshall" authorityYear="1993" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="coriolis">
<emphasis id="5F98EAAF383A0B78FBE5FDF2538BFD9A" box="[1150,1277,559,585]" italics="true" pageId="100" pageNumber="101">K. coriolis</emphasis>
<bibRefCitation id="097D4B4C383A0B78FA9CFDED578EFDBE" author="Marshall B. A." pageId="100" pageNumber="101" pagination="185 - 198" refId="ref61761" refString="Marshall B. A. 1993. A review of the genus Kaiparathina Laws, 1941 (Mollusca: Gastropoda: Trochidae). The Veliger 36 (2): 185 - 198. Available from https: // www. biodiversitylibrary. org / page / 42465795 [accessed 10 Feb. 2024]." type="journal article" year="1993">Marshall, 1993</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="AAEC4D3E383A0B78FEACFD9155F2FDBE" authority="Marshall, 1993" authorityName="Marshall" authorityYear="1993" box="[311,644,594,621]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="boucheti">
<emphasis id="5F98EAAF383A0B78FEACFD9156B4FDBF" box="[311,450,594,620]" italics="true" pageId="100" pageNumber="101">K. boucheti</emphasis>
<bibRefCitation id="097D4B4C383A0B78FE56FD9055F2FDBE" author="Marshall B. A." box="[461,644,594,621]" pageId="100" pageNumber="101" pagination="185 - 198" refId="ref61761" refString="Marshall B. A. 1993. A review of the genus Kaiparathina Laws, 1941 (Mollusca: Gastropoda: Trochidae). The Veliger 36 (2): 185 - 198. Available from https: // www. biodiversitylibrary. org / page / 42465795 [accessed 10 Feb. 2024]." type="journal article" year="1993">Marshall, 1993</bibRefCitation>
</taxonomicName>
, both from the New Caledonian region.
<taxonomicName id="AAEC4D3E383A0B78FBEFFD905201FDBF" authorityName="Marshall" authorityYear="1993" box="[1140,1399,594,620]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="coriolis">
<emphasis id="5F98EAAF383A0B78FBEFFD905201FDBF" box="[1140,1399,594,620]" italics="true" pageId="100" pageNumber="101">Kaiparathina coriolis</emphasis>
</taxonomicName>
has more strongly convex spire whorls, retains an umbilical chink at maturity and has a different coloration. In addition, the terminal cusp of the rachidian tooth is proportionately much smaller than it is in
<taxonomicName id="AAEC4D3E383A0B78FE8AFD7F56DFFD05" authorityName="Herbert" authorityYear="2024" box="[273,425,700,726]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="5F98EAAF383A0B78FE8AFD7F56DFFD05" box="[273,425,700,726]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
.
<taxonomicName id="AAEC4D3E383A0B78FE23FD7E55B5FD05" authorityName="Marshall" authorityYear="1993" box="[440,707,700,726]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="boucheti">
<emphasis id="5F98EAAF383A0B78FE23FD7E55B5FD05" box="[440,707,700,726]" italics="true" pageId="100" pageNumber="101">Kaiparathina boucheti</emphasis>
</taxonomicName>
has stronger spiral sculpture on the apical whorls, is more uniform in colour and has a reddish-brown protoconch; the rachidian tooth also has a more distinctly triangular cusp. Perhaps the most similar species is
<taxonomicName id="AAEC4D3E383A0B78FC95FCC15330FCCE" authority="Marshall, 1993" authorityName="Marshall" authorityYear="1993" box="[782,1094,770,797]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="daedala">
<emphasis id="5F98EAAF383A0B78FC95FCC154F8FCCF" box="[782,910,770,796]" italics="true" pageId="100" pageNumber="101">K. daedala</emphasis>
<bibRefCitation id="097D4B4C383A0B78FC0EFCC05330FCCE" author="Marshall B. A." box="[917,1094,770,797]" pageId="100" pageNumber="101" pagination="185 - 198" refId="ref61761" refString="Marshall B. A. 1993. A review of the genus Kaiparathina Laws, 1941 (Mollusca: Gastropoda: Trochidae). The Veliger 36 (2): 185 - 198. Available from https: // www. biodiversitylibrary. org / page / 42465795 [accessed 10 Feb. 2024]." type="journal article" year="1993">Marshall, 1993</bibRefCitation>
</taxonomicName>
from off
<collectingCountry id="15FB762D383A0B78FB29FCC05263FCCF" box="[1202,1301,770,796]" name="Réunion" pageId="100" pageNumber="101">Réunion</collectingCountry>
and this is also the species most geographically proximate to Walters Shoal. Like the present species,
<taxonomicName id="AAEC4D3E383A0B78FB6DFCE45200FC93" authorityName="Marshall" authorityYear="1993" box="[1270,1398,806,832]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="daedala">
<emphasis id="5F98EAAF383A0B78FB6DFCE45200FC93" box="[1270,1398,806,832]" italics="true" pageId="100" pageNumber="101">K. daedala</emphasis>
</taxonomicName>
has a narrow supra-sutural band of reddish-brown marks (also present in
<taxonomicName id="AAEC4D3E383A0B78FB97FC8853F1FCB0" authorityName="Marshall" authorityYear="1993" box="[1036,1159,841,867]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="coriolis">
<emphasis id="5F98EAAF383A0B78FB97FC8853F1FCB0" box="[1036,1159,841,867]" italics="true" pageId="100" pageNumber="101">K. coriolis</emphasis>
</taxonomicName>
) and a narrow white subsutural line. Elsewhere, the colour pattern in the only two available specimens is variable (
<bibRefCitation id="097D4B4C383A0B78FA90FCAE578DFC7A" author="Marshall B. A." pageId="100" pageNumber="101" pagination="185 - 198" refId="ref61761" refString="Marshall B. A. 1993. A review of the genus Kaiparathina Laws, 1941 (Mollusca: Gastropoda: Trochidae). The Veliger 36 (2): 185 - 198. Available from https: // www. biodiversitylibrary. org / page / 42465795 [accessed 10 Feb. 2024]." type="journal article" year="1993">Marshall 1993</bibRefCitation>
), but neither exhibit the spiral lines of white spots or zig-zag brown axial flames commonly seen in
<taxonomicName id="AAEC4D3E383A0B78FF26FC715622FC1F" authorityName="Herbert" authorityYear="2024" box="[189,340,946,972]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="5F98EAAF383A0B78FF26FC715622FC1F" box="[189,340,946,972]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
. In addition, a distinctive feature of
<taxonomicName id="AAEC4D3E383A0B78FC9AFC7154F5FC1F" authorityName="Marshall" authorityYear="1993" box="[769,899,946,972]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="daedala">
<emphasis id="5F98EAAF383A0B78FC9AFC7154F5FC1F" box="[769,899,946,972]" italics="true" pageId="100" pageNumber="101">K. daedala</emphasis>
</taxonomicName>
is the presence of numerous spiral threads on the inner two-thirds of the base. Specimens of
<taxonomicName id="AAEC4D3E383A0B78FC83FC1454C4FC23" authorityName="Herbert" authorityYear="2024" box="[792,946,982,1008]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="5F98EAAF383A0B78FC83FC1454C4FC23" box="[792,946,982,1008]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
of a similar size (
<quantity id="AA149B58383A0B78FB17FC175384FC23" box="[1164,1266,981,1008]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="100" pageNumber="101" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
) may have 12 such spiral threads, but in most individuals they are absent (
<figureCitation id="F5D72A38383A0B78FC57FC3B534AFBC0" box="[972,1084,1017,1043]" captionStart="Fig" captionStartId="99.[189,232,1855,1881]" captionTargetBox="[188,1398,745,1817]" captionTargetId="figure-248@99.[188,1398,745,1817]" captionTargetPageId="99" captionText="Fig. 40. Kaiparathina monticola sp. nov. AC. Paratypes (MNHN-IM-2000-35726). A. Oblique view of apex. B. Sculptural detail of protoconch. C. Basal view of juvenile showing lack of spiral lirae. DG. Jaw and radula (paratype, MNHN-IM-2013-67260). D. Ventral (inner) surface of jaw, anterior edge below. E. Detail of jaw structure. F. Radula, entire width. G. Radula, detail of rachidian, lateral and inner marginal teeth. Scale bars: A, D =200 μm; B, F = 100 μm; C=1.0 mm; E, G =50 μm." figureDoi="http://doi.org/10.5281/zenodo.10727861" httpUri="https://zenodo.org/record/10727861/files/figure.png" pageId="100" pageNumber="101">Fig. 40C</figureCitation>
). The radula described for
<taxonomicName id="AAEC4D3E383A0B78FF26FBDF564BFBE5" authorityName="Marshall" authorityYear="1993" box="[189,317,1052,1078]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="daedala">
<emphasis id="5F98EAAF383A0B78FF26FBDF564BFBE5" box="[189,317,1052,1078]" italics="true" pageId="100" pageNumber="101">K. daedala</emphasis>
</taxonomicName>
has only five pairs of lateral teeth in each row (vs seven in
<taxonomicName id="AAEC4D3E383A0B78FC44FBDF5302FBE5" authorityName="Herbert" authorityYear="2024" box="[991,1140,1052,1078]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="5F98EAAF383A0B78FC44FBDF5302FBE5" box="[991,1140,1052,1078]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
), but this difference is undoubtedly size-related and reflects the fact that the
<typeStatus id="B257881F383A0B78FCBEFB82542EFB89" box="[805,856,1088,1114]" pageId="100" pageNumber="101">type</typeStatus>
material of
<taxonomicName id="AAEC4D3E383A0B78FC7FFB825312FB8A" authorityName="Marshall" authorityYear="1993" box="[996,1124,1087,1113]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="daedala">
<emphasis id="5F98EAAF383A0B78FC7FFB825312FB8A" box="[996,1124,1087,1113]" italics="true" pageId="100" pageNumber="101">K. daedala</emphasis>
</taxonomicName>
was probably subadult, the number of lateral teeth increasing progressively through in-column transformation of marginal teeth during ontogeny (
<bibRefCitation id="097D4B4C383A0B78FE11FB44554FFB73" author="Marshall B. A." box="[394,569,1158,1184]" pageId="100" pageNumber="101" pagination="185 - 198" refId="ref61761" refString="Marshall B. A. 1993. A review of the genus Kaiparathina Laws, 1941 (Mollusca: Gastropoda: Trochidae). The Veliger 36 (2): 185 - 198. Available from https: // www. biodiversitylibrary. org / page / 42465795 [accessed 10 Feb. 2024]." type="journal article" year="1993">Marshall 1993</bibRefCitation>
). There are, nonetheless, differences in the shape of the radula teeth, the terminal denticle of the rachidian and lateral teeth being proportionately longer in
<taxonomicName id="AAEC4D3E383A0B78FBEAFB685271FB10" authorityName="Herbert" authorityYear="2024" box="[1137,1287,1193,1219]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="5F98EAAF383A0B78FBEAFB685271FB10" box="[1137,1287,1193,1219]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
, whereas the subterminal denticles on these teeth are coarser and more distinct in
<taxonomicName id="AAEC4D3E383A0B78FBB1FB0F53DDFB35" authorityName="Marshall" authorityYear="1993" box="[1066,1195,1228,1254]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="daedala">
<emphasis id="5F98EAAF383A0B78FBB1FB0F53DDFB35" box="[1066,1195,1228,1254]" italics="true" pageId="100" pageNumber="101">K. daedala</emphasis>
</taxonomicName>
. Furthermore, in
<taxonomicName id="AAEC4D3E383A0B78FF24FB325622FADA" authorityName="Herbert" authorityYear="2024" box="[191,340,1263,1289]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="5F98EAAF383A0B78FF24FB325622FADA" box="[191,340,1263,1289]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
the cusp of the rachidian is laterally more convex at its base than the more evenly trigonal rachidian cusp of
<taxonomicName id="AAEC4D3E383A0B78FE0FFAD15563FAFF" authorityName="Marshall" authorityYear="1993" box="[404,533,1298,1324]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="daedala">
<emphasis id="5F98EAAF383A0B78FE0FFAD15563FAFF" box="[404,533,1298,1324]" italics="true" pageId="100" pageNumber="101">K. daedala</emphasis>
</taxonomicName>
. Although likely to be closely related to
<taxonomicName id="AAEC4D3E383A0B78FC64FAD153F6FAFF" authorityName="Marshall" authorityYear="1993" box="[1023,1152,1298,1324]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="daedala">
<emphasis id="5F98EAAF383A0B78FC64FAD153F6FAFF" box="[1023,1152,1298,1324]" italics="true" pageId="100" pageNumber="101">K. daedala</emphasis>
</taxonomicName>
, these differences in coloration, basal sculpture and radula tooth morphology justify the separation of the present material as a distinct species.
</paragraph>
<paragraph id="6D5336BD383A0B78FF26FA585491F9CD" blockId="100.[188,1399,1434,1566]" pageId="100" pageNumber="101">
The shell of
<taxonomicName id="AAEC4D3E383A0B78FEF9FA595689FA67" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[354,511,1434,1460]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola" status="sp. nov.">
<emphasis id="5F98EAAF383A0B78FEF9FA595689FA67" box="[354,511,1434,1460]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4383A0B78FD96FA595518FA66" box="[525,622,1435,1461]" pageId="100" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
superficially resembles that of some brown-spotted species of
<taxonomicName id="AAEC4D3E383A0B78FF26FA7F5614FA04" authorityName="Dall" authorityYear="1881" box="[189,354,1469,1495]" class="Gastropoda" family="Solariellidae" genus="Bathymophila" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="genus">
<emphasis id="5F98EAAF383A0B78FF26FA7F5614FA04" box="[189,354,1469,1495]" italics="true" pageId="100" pageNumber="101">Bathymophila</emphasis>
</taxonomicName>
(
<bibRefCitation id="097D4B4C383A0B78FEE8FA7F5519FA0B" author="Williams S. T. &amp; Kano Y. &amp; Waren A. &amp; Herbert D. G." box="[371,623,1469,1496]" pageId="100" pageNumber="101" pagination="1 - 26" refId="ref64534" refString="Williams S. T., Kano Y., Waren A. &amp; Herbert D. G. 2020. Marrying molecules and morphology: first steps towards a reevaluation of solariellid genera (Gastropoda: Trochoidea) in the light of molecular phylogenetic studies. Journal of Molluscan Studies 86 (1): 1 - 26. https: // doi. org / 10.1093 / mollus / eyz 038" type="journal article" year="2020">
Williams
<emphasis id="5F98EAAF383A0B78FE72FA7C5569FA04" box="[489,543,1469,1495]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. 2020
</bibRefCitation>
: fig.
<quantity id="AA149B58383A0B78FD36FA7C55BBFA0B" box="[685,717,1470,1496]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="100" pageNumber="101" unit="g" value="7.0">7g</quantity>
), which
<bibRefCitation id="097D4B4C383A0B78FCADFA7F5349FA0B" author="Williams S. T. &amp; Noone E. S. &amp; Smith L. M. &amp; Sumner-Rooney L." box="[822,1087,1469,1496]" pageId="100" pageNumber="101" pagination="3026 - 3040" refId="ref64607" refString="Williams S. T., Noone E. S., Smith L. M. &amp; Sumner-Rooney L. 2022. Evolutionary loss of shell pigmentation, pattern, and eye structure in deep-sea snails in the dysphotic zone. Evolution 76 (12): 3026 - 3040. https: // doi. org / 10.1111 / evo. 14647" type="journal article" year="2022">
Williams
<emphasis id="5F98EAAF383A0B78FC31FA7C5496FA04" box="[938,992,1469,1495]" italics="true" pageId="100" pageNumber="101">et al</emphasis>
. (2022)
</bibRefCitation>
have shown belong to an undescribed
<taxonomicName id="AAEC4D3E383A0B78FECAFA235680FA28" authorityName="Dall" authorityYear="1881" box="[337,502,1505,1531]" class="Gastropoda" family="Solariellidae" genus="Bathymophila" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="genus">
<emphasis id="5F98EAAF383A0B78FECAFA235680FA28" box="[337,502,1505,1531]" italics="true" pageId="100" pageNumber="101">Bathymophila</emphasis>
</taxonomicName>
-like genus. These differ from
<taxonomicName id="AAEC4D3E383A0B78FCCEFA20549AFA28" authorityName="Herbert" authorityYear="2024" box="[853,1004,1505,1531]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="5F98EAAF383A0B78FCCEFA20549AFA28" box="[853,1004,1505,1531]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
in having a patent umbilicus with a heavy flange around the umbilicus, and a less acutely pointed apex.
</paragraph>
<paragraph id="6D5336BD383A0B78FF26F9875638F93F" blockId="100.[187,1399,1605,1772]" pageId="100" pageNumber="101">
Mention should also be made of
<taxonomicName id="AAEC4D3E383A0B78FDA1F987530CF98C" authority="Poppe, Tagaro &amp; Dekker, 2006" authorityName="Poppe, Tagaro &amp; H. Dekker" authorityYear="2006" box="[570,1146,1605,1632]" class="Gastropoda" family="Trochidae" genus="Cantharidus" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="nolfi">
<emphasis id="5F98EAAF383A0B78FDA1F987547EF98C" box="[570,776,1605,1631]" italics="true" pageId="100" pageNumber="101">Cantharidus nolfi</emphasis>
<bibRefCitation id="097D4B4C383A0B78FC94F984530CF98C" author="Poppe G. T. &amp; Tagaro S. P. &amp; Dekker H." box="[783,1146,1605,1632]" pageId="100" pageNumber="101" pagination="1 - 228" refId="ref62686" refString="Poppe G. T., Tagaro S. P. &amp; Dekker H. 2006. The Seguenziidae, Chilodontidae, Trochidae, Calliostomatidae and Solariellidae of the Philippine Islands. Visaya, Supplement 2: 1 - 228." type="journal article" year="2006">Poppe, Tagaro &amp; Dekker, 2006</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="15FB762D383A0B78FB70F9875205F98C" box="[1259,1395,1605,1631]" name="Philippines" pageId="100" pageNumber="101">Philippines</collectingCountry>
. Although not known anatomically, I believe the shell of this species exhibits all the features characteristic of
<taxonomicName id="AAEC4D3E383A0B78FF47F94E560CF975" authorityName="Laws" authorityYear="1841" box="[220,378,1676,1702]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="genus">
<emphasis id="5F98EAAF383A0B78FF47F94E560CF975" box="[220,378,1676,1702]" italics="true" pageId="100" pageNumber="101">Kaiparathina</emphasis>
</taxonomicName>
and I propose that it be transferred thereto. It is of similar size as
<taxonomicName id="AAEC4D3E383A0B78FBECF94F5278F975" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[1143,1294,1676,1702]" class="Gastropoda" family="Trochidae" genus="Kaiparathina" kingdom="Animalia" order="Trochida" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="monticola" status="sp. nov.">
<emphasis id="5F98EAAF383A0B78FBECF94F5278F975" box="[1143,1294,1676,1702]" italics="true" pageId="100" pageNumber="101">K. monticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4383A0B78FA8EF94E5204F975" box="[1301,1394,1676,1702]" pageId="100" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
, but differs in having a colour pattern that typically includes fine, close-set and strongly oblique gold or brown lines.
</paragraph>
</subSubSection>
</treatment>
</document>