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<document id="799A8DBC746823311310F8849E468C45" ID-CLB-Dataset="116165" ID-DOI="10.11646/zootaxa.5171.1.1" ID-GBIF-Dataset="9923ae54-fea7-4394-9c56-f5f4db376e0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6966207" ID-ZooBank="5B00C374-33B0-4433-95A0-DC9B5FFC5B0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1659689746521" checkinUser="plazi" docAuthor="Borowiec, Lech &amp; Świętojańska, Jolanta" docDate="2022" docId="E417E526B17CA908FF6CFAE40F939236" docLanguage="en" docName="zootaxa.5171.1.1.pdf" docOrigin="Zootaxa 5171 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5171.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cassida benguelica Spaeth 1933" docType="treatment" docVersion="7" lastPageNumber="36" masterDocId="182E9D5EB15FA92BFFFBFA730C63955A" masterDocTitle="A monograph of the Afrotropical Cassidinae (Coleoptera: Chrysomelidae). Part 6. Revision of the tribe Cassidini 3, the genus Cassida L." masterLastPageNumber="250" masterPageNumber="1" pageNumber="36" updateTime="1699364714896" updateUser="ExternalLinkService">
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<mods:title id="D7488C1116D5CA3D9396EFC58AAA98FE">A monograph of the Afrotropical Cassidinae (Coleoptera: Chrysomelidae). Part 6. Revision of the tribe Cassidini 3, the genus Cassida L.</mods:title>
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<subSubSection id="24A407BBB17CA908FF6CFAE40E4B95EB" box="[151,552,151,178]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="6C015430B17CA908FF6CFAE40E4B95EB" blockId="35.[151,552,151,213]" box="[151,552,151,178]" pageId="35" pageNumber="36">
<heading id="3749E35CB17CA908FF6CFAE40E4B95EB" bold="true" box="[151,552,151,178]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<taxonomicName id="ABBE2FB3B17CA908FF6CFAE40E4B95EB" ID-CoL="7RYNQ" authority="Spaeth, 1933" authorityName="Spaeth" authorityYear="1933" box="[151,552,151,178]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FF6CFAE40E4B95EB" bold="true" box="[151,552,151,178]" pageId="35" pageNumber="36">
<emphasis id="5ECA8822B17CA908FF6CFAE40DE295E8" bold="true" box="[151,385,151,178]" italics="true" pageId="35" pageNumber="36">Cassida benguelica</emphasis>
Spaeth, 1933
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="24A407BBB17CA908FF6CFAC80D43958F" box="[151,288,187,213]" pageId="35" pageNumber="36" type="description">
<paragraph id="6C015430B17CA908FF6CFAC80D43958F" blockId="35.[151,552,151,213]" box="[151,288,187,213]" pageId="35" pageNumber="36">
(
<figureCitation id="F48548B5B17CA908FF5BFAC80C82958F" box="[160,225,187,213]" captionStart="FIGURE 1" captionStartId="22.[151,250,1967,1991]" captionTargetBox="[159,1426,753,1933]" captionTargetId="figure-256@22.[151,1436,741,1944]" captionTargetPageId="22" captionText="FIGURE 1. Distribution of Cassida aethiopica (black circles) and Cassida benguelica (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6966248" httpUri="https://zenodo.org/record/6966248/files/figure.png" pageId="35" pageNumber="36">figs.1</figureCitation>
,
<figureCitation id="F48548B5B17CA908FF15FAC80D7B958F" box="[238,280,187,213]" captionStart="FIGURE 119" captionStartId="198.[151,250,1616,1640]" captionTargetBox="[168,1411,205,1575]" captionTargetId="figure-36@198.[151,1435,181,1592]" captionTargetPageId="198" captionText="FIGURE 119. Cassida benguelica. FIGURES 120121. Cassida leleupi. FIGURES 122123. Cassida flavosignata. 119120, 122. Dorsal; 121, 123. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966444" httpUri="https://zenodo.org/record/6966444/files/figure.png" pageId="35" pageNumber="36">119</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="24A407BBB17CA908FF6CFB730F7B9442" box="[151,792,255,280]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="6C015430B17CA908FF6CFB730F7B9442" blockId="35.[151,792,255,280]" box="[151,792,255,280]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="4CAE731EB17CA908FF6CFB730F7B9442" box="[151,792,255,280]" pageId="35" pageNumber="36">
<treatmentCitation id="ED1F7221B17CA908FF6CFB730E52944D" author="Spaeth, F." box="[151,561,255,280]" page="345" pageId="35" pageNumber="36" year="1933">
<taxonomicName id="ABBE2FB3B17CA908FF6CFB730E52944D" ID-CoL="7RYNQ" authority="Spaeth, 1933 a: 345" authorityName="Spaeth" authorityPageNumber="345" authorityYear="1933" box="[151,561,255,280]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FF6CFB730D3D9442" box="[151,350,256,280]" italics="true" pageId="35" pageNumber="36">Cassida benguelica</emphasis>
<bibRefCitation id="082F29C1B17CA908FE9EFB730E52944D" author="Spaeth, F." box="[357,561,255,280]" pageId="35" pageNumber="36" pagination="345 - 359" refId="ref188213" refString="Spaeth, F. (1933 a) Neue Beitrage zur Kenntnis der Afrikanischen Cassidinen (Col. Chrys.). Revue de Zoologie et de Botanique Africaines, 22, 345 - 359." type="journal article" year="1933">Spaeth, 1933 a: 345</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="ED1F7221B17CA908FDC0FB730F709442" author="Borowiec, L." box="[571,787,256,280]" page="239" pageId="35" pageNumber="36" year="1999">
<bibRefCitation id="082F29C1B17CA908FDC0FB730F709442" author="Borowiec, L." box="[571,787,256,280]" pageId="35" pageNumber="36" refId="ref184613" refString="Borowiec, L. (1999) A world catalogue of the Cassidinae (Coleoptera: Chrysomelidae). Biologica Silesiae, Wroclaw, 476 pp." type="book" year="1999">Borowiec, 1999: 239</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="24A407BBB17CA908FF6CFB360D0590C2" pageId="35" pageNumber="36" type="description">
<paragraph id="6C015430B17CA908FF6CFB360CB294DE" blockId="35.[151,1437,325,1900]" pageId="35" pageNumber="36">
<emphasis id="5ECA8822B17CA908FF6CFB360D4A9405" bold="true" box="[151,297,325,351]" pageId="35" pageNumber="36">Description.</emphasis>
L: 4.70 mm, W: 3.65 mm, Lp: 1.60 mm, Wp: 2.
<quantity id="AB46F9D5B17CA908FC9CFB350FD9943A" box="[871,954,326,352]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="35" pageNumber="36" unit="mm" value="70.0">70 mm</quantity>
, L/W: 1.29, Wp/Lp: 1.69. Body oval (
<figureCitation id="F48548B5B17CA908FA8EFB350CA794DE" captionStart="FIGURE 119" captionStartId="198.[151,250,1616,1640]" captionTargetBox="[168,1411,205,1575]" captionTargetId="figure-36@198.[151,1435,181,1592]" captionTargetPageId="198" captionText="FIGURE 119. Cassida benguelica. FIGURES 120121. Cassida leleupi. FIGURES 122123. Cassida flavosignata. 119120, 122. Dorsal; 121, 123. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966444" httpUri="https://zenodo.org/record/6966444/files/figure.png" pageId="35" pageNumber="36">fig. 119</figureCitation>
).
</paragraph>
<paragraph id="6C015430B17CA908FF3CFBFD084197FE" blockId="35.[151,1437,325,1900]" pageId="35" pageNumber="36">
Pronotum yellow, disc with large, black, triangular spot occupying almost entire surface of disc except sides, insides the black spot, close to scutellum two large, yellow spots. Apex of the triangular black spot rounded. Scutellum black. Elytral disc with black ground colour and pattern of yellow spots. In humeral and posterolateral area the black ground colour extending to submarginal row but in the middle and on apex margin of the black spot emarginate. Yellow pattern forms two small, round yellow spots close to base of scutellum, two large, irregular, elevated spots at sides of postscutellar point, two smaller, elevated spots in the mid length of disc close to suture, two irregular spots in ¾ length of disc close to suture, and two small spots in posterolateral part of disc (
<figureCitation id="F48548B5B17CA908FACFF81509F397DA" box="[1332,1424,614,640]" captionStart="FIGURE 119" captionStartId="198.[151,250,1616,1640]" captionTargetBox="[168,1411,205,1575]" captionTargetId="figure-36@198.[151,1435,181,1592]" captionTargetPageId="198" captionText="FIGURE 119. Cassida benguelica. FIGURES 120121. Cassida leleupi. FIGURES 122123. Cassida flavosignata. 119120, 122. Dorsal; 121, 123. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966444" httpUri="https://zenodo.org/record/6966444/files/figure.png" pageId="35" pageNumber="36">fig. 119</figureCitation>
). Ventrites, legs and antennae yellow only apex of last antennal segment infuscate.
</paragraph>
<paragraph id="6C015430B17CA908FF3CF8DD0E49964A" blockId="35.[151,1437,325,1900]" pageId="35" pageNumber="36">Pronotum elliptical, with maximum width in the middle, sides rounded. Disc indistinctly margined from explanate margin, without distinct lateral lobes, surface smooth and shiny. Explanate margin broad, with honeycomb structure, surface smooth and shiny.</paragraph>
<paragraph id="6C015430B17CA908FF3CF969089F9122" blockId="35.[151,1437,325,1900]" pageId="35" pageNumber="36">Base of elytra much wider than base of pronotum, humeral angles moderately protruding anterad, angulate, elytral margins behind humeral angle very shallowly emarginate. Disc moderately, regularly convex in profile, with shallow postscutellar and principal impressions emphasized by elevated yellow spots. Punctation moderately coarse and sparse, distance between punctures mostly wider than puncture diameter. Rows regular but interrupted by yellow spots. Three pairs of central yellow spots elevated, first and third pair of the spots marked with 12 coarse punctures. Marginal row distinct, its punctures only slightly coarser than in submarginal row, dense with distance between punctures mostly narrower than puncture diameter. Intervals 1.52.5 times as wide as rows, flat (except elevated yellow spots), their surface slightly alutaceous but shiny. Marginal interval without humeral fold, lateral fold very narrow, hardly visible. Explanate margin moderately declivous, broad, in the widest part approximately three times narrower than disc, its surface shallowly and densely punctate, appears slightly irregular.</paragraph>
<paragraph id="6C015430B17CA908FF3CFEF10F459076" blockId="35.[151,1437,325,1900]" pageId="35" pageNumber="36">Eyes very large, gena obsolete. Clypeus moderately broad, approximately as long as wide, flat. Clypeal grooves fine, in basal part run very close to margin of eye, apically converging in arch. Surface of clypeal plate impunctate, shiny. Labrum emarginate to ¼ length. Antennae slim, segment 9 approximately 1.8 times as long as wide, segment 10 twice longer than wide, segment 11 very long, 1.8 times as long as wide. Segment 3 approximately 1.2 times longer than segment 2 and slightly shorter than segment 4.</paragraph>
<paragraph id="6C015430B17CA908FF3CFF450D96902E" blockId="35.[151,1437,325,1900]" pageId="35" pageNumber="36">Prosternal process narrow, flat and impunctate between coxae, very broad apically, central part of rhomboidal apex shiny with few punctures.</paragraph>
<paragraph id="6C015430B17CA908FF3CFF0D0D0590C2" blockId="35.[151,1437,325,1900]" box="[199,358,1406,1432]" pageId="35" pageNumber="36">Claws simple.</paragraph>
</subSubSection>
<subSubSection id="24A407BBB17CA908FF3CFFD20E7790E6" box="[199,532,1441,1468]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="6C015430B17CA908FF3CFFD20E7790E6" blockId="35.[151,1437,325,1900]" box="[199,532,1441,1468]" pageId="35" pageNumber="36">
<emphasis id="5ECA8822B17CA908FF3CFFD20D0290E1" bold="true" box="[199,353,1441,1467]" pageId="35" pageNumber="36">Distribution.</emphasis>
<collectingCountry id="14A914A0B17CA908FE9DFFD10DDB90E6" box="[358,440,1442,1468]" name="Angola" pageId="35" pageNumber="36">Angola</collectingCountry>
(
<figureCitation id="F48548B5B17CA908FE33FFD10E6090E6" box="[456,515,1442,1468]" captionStart="FIGURE 1" captionStartId="22.[151,250,1967,1991]" captionTargetBox="[159,1426,753,1933]" captionTargetId="figure-256@22.[151,1436,741,1944]" captionTargetPageId="22" captionText="FIGURE 1. Distribution of Cassida aethiopica (black circles) and Cassida benguelica (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6966248" httpUri="https://zenodo.org/record/6966248/files/figure.png" pageId="35" pageNumber="36">fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="24A407BBB17CA908FF3CFFB50FE39212" pageId="35" pageNumber="36" type="discussion">
<paragraph id="6C015430B17CA908FF3CFFB50FE39212" blockId="35.[151,1437,325,1900]" pageId="35" pageNumber="36">
<emphasis id="5ECA8822B17CA908FF3CFFB50D5890BA" bold="true" box="[199,315,1478,1504]" pageId="35" pageNumber="36">Remarks.</emphasis>
A member of the
<taxonomicName id="ABBE2FB3B17CA908FDDEFFB50F7490BA" authorityName="Spaeth" authorityYear="1932" box="[549,791,1478,1504]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="flavosignata">
<emphasis id="5ECA8822B17CA908FDDEFFB50F7490BA" box="[549,791,1478,1504]" italics="true" pageId="35" pageNumber="36">Cassida flavosignata</emphasis>
</taxonomicName>
speciesgroup. General shape, dorsal colouration and sculpture place this species near to only
<taxonomicName id="ABBE2FB3B17CA908FD9AFF990EAC935E" authorityName="Borowiec &amp; Świętojańska" authorityYear="2022" box="[609,719,1514,1540]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="leleupi">
<emphasis id="5ECA8822B17CA908FD9AFF990EAC935E" box="[609,719,1514,1540]" italics="true" pageId="35" pageNumber="36">C. leleupi</emphasis>
</taxonomicName>
but it differs in body more circular, humeral angles more acute with the elytral margin distinctly emarginate behind the humeral angle (figs. 120, 121) while in
<taxonomicName id="ABBE2FB3B17CA908FB34FC7D090F9372" authorityName="Spaeth" authorityYear="1933" box="[1231,1388,1550,1576]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FB34FC7D090F9372" box="[1231,1388,1550,1576]" italics="true" pageId="35" pageNumber="36">C. benguelica</emphasis>
</taxonomicName>
, the elytral margin behind the humeral angle is very shallowly emarginate thus the angles appear rather angulate than acute (
<figureCitation id="F48548B5B17CA908FF19FC250D5D932A" box="[226,318,1622,1648]" captionStart="FIGURE 119" captionStartId="198.[151,250,1616,1640]" captionTargetBox="[168,1411,205,1575]" captionTargetId="figure-36@198.[151,1435,181,1592]" captionTargetPageId="198" captionText="FIGURE 119. Cassida benguelica. FIGURES 120121. Cassida leleupi. FIGURES 122123. Cassida flavosignata. 119120, 122. Dorsal; 121, 123. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966444" httpUri="https://zenodo.org/record/6966444/files/figure.png" pageId="35" pageNumber="36">fig. 119</figureCitation>
). In
<taxonomicName id="ABBE2FB3B17CA908FE8AFC250DBD932A" authorityName="Borowiec &amp; Świętojańska" authorityYear="2022" box="[369,478,1622,1648]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="leleupi">
<emphasis id="5ECA8822B17CA908FE8AFC250DBD932A" box="[369,478,1622,1648]" italics="true" pageId="35" pageNumber="36">C. leleupi</emphasis>
</taxonomicName>
two small, yellow spots occur at apex of scutellum (in
<taxonomicName id="ABBE2FB3B17CA908FBAAFC25088D932A" authorityName="Spaeth" authorityYear="1933" box="[1105,1262,1622,1648]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FBAAFC25088D932A" box="[1105,1262,1622,1648]" italics="true" pageId="35" pageNumber="36">C. benguelica</emphasis>
</taxonomicName>
without spots), elytral spots not elevated and impunctate (in
<taxonomicName id="ABBE2FB3B17CA908FD77FC090F4B93CE" authorityName="Spaeth" authorityYear="1933" box="[652,808,1658,1684]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FD77FC090F4B93CE" box="[652,808,1658,1684]" italics="true" pageId="35" pageNumber="36">C. benguelica</emphasis>
</taxonomicName>
the large, central spots are elevated and with 12 coarse punctures), the black pronotal spot with acute apex (in
<taxonomicName id="ABBE2FB3B17CA908FCF9FCED0FFD93E2" authorityName="Spaeth" authorityYear="1933" box="[770,926,1694,1720]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FCF9FCED0FFD93E2" box="[770,926,1694,1720]" italics="true" pageId="35" pageNumber="36">C. benguelica</emphasis>
</taxonomicName>
the apex of the pronotal spot is rounded) and the black ground colour behind the humeral callus extending to the marginal row (in
<taxonomicName id="ABBE2FB3B17CA908FBBCFCB108819386" authorityName="Spaeth" authorityYear="1933" box="[1095,1250,1730,1756]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FBBCFCB108819386" box="[1095,1250,1730,1756]" italics="true" pageId="35" pageNumber="36">C. benguelica</emphasis>
</taxonomicName>
extending to the submarginal row). Pronotal sides in
<taxonomicName id="ABBE2FB3B17CA908FDCFFC950EC1925A" authorityName="Borowiec &amp; Świętojańska" authorityYear="2022" box="[564,674,1766,1792]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="leleupi">
<emphasis id="5ECA8822B17CA908FDCFFC950EC1925A" box="[564,674,1766,1792]" italics="true" pageId="35" pageNumber="36">C. leleupi</emphasis>
</taxonomicName>
are more broadly rounded than in
<taxonomicName id="ABBE2FB3B17CA908FBD4FC9508AF925A" authorityName="Spaeth" authorityYear="1933" box="[1071,1228,1766,1792]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FBD4FC9508AF925A" box="[1071,1228,1766,1792]" italics="true" pageId="35" pageNumber="36">C. benguelica</emphasis>
</taxonomicName>
, the punctation of disc is approximately two times smaller than in
<taxonomicName id="ABBE2FB3B17CA908FD56FD790F2B927E" authorityName="Spaeth" authorityYear="1933" box="[685,840,1802,1828]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FD56FD790F2B927E" box="[685,840,1802,1828]" italics="true" pageId="35" pageNumber="36">C. benguelica</emphasis>
</taxonomicName>
, and the surface of the disc is more shiny with mirror brilliance while in
<taxonomicName id="ABBE2FB3B17CA908FE91FD5D0E659212" authorityName="Spaeth" authorityYear="1933" box="[362,518,1838,1864]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="benguelica">
<emphasis id="5ECA8822B17CA908FE91FD5D0E659212" box="[362,518,1838,1864]" italics="true" pageId="35" pageNumber="36">C. benguelica</emphasis>
</taxonomicName>
the surface is slightly alutaceous.
</paragraph>
</subSubSection>
<subSubSection id="24A407BBB17CA908FF3CFD210F939236" box="[199,1008,1873,1900]" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="6C015430B17CA908FF3CFD210F939236" blockId="35.[151,1437,325,1900]" box="[199,1008,1873,1900]" pageId="35" pageNumber="36">
<emphasis id="5ECA8822B17CA908FF3CFD210DE09231" bold="true" box="[199,387,1873,1900]" pageId="35" pageNumber="36">Type examined.</emphasis>
<materialsCitation id="DCD65E6DB17CA908FE71FD210F939236" ID-GBIF-Occurrence="3881325122" box="[394,1008,1874,1900]" collectionCode="MM" country="Angola" location="Benguela" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Benguela" typeStatus="holotype">
<typeStatus id="B305EA92B17CA908FE71FD210D949236" box="[394,503,1874,1900]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
: [
<collectingCountry id="14A914A0B17CA908FDFCFD210EE49236" box="[519,647,1874,1900]" name="Angola" pageId="35" pageNumber="36">ANGOLA</collectingCountry>
]:
<collectingRegion id="AE7A9AD2B17CA908FD62FD210F659236" box="[665,774,1874,1900]" country="Angola" name="Benguela" pageId="35" pageNumber="36">Benguela</collectingRegion>
, 1914, Staud. (
<collectionCode id="0AAFCCF5B17CA908FC57FD210F809236" box="[940,995,1874,1900]" country="France" name="University of Montpellier" pageId="35" pageNumber="36">MM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>