214 lines
21 KiB
XML
214 lines
21 KiB
XML
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<mods:title id="3FED04AE25D63493BA2ADB5D6D4C0416">Dung beetle fauna from Mount Mabu, Mozambique. Part 3: A new species of Proagoderus van Lansberge, 1883 with an annotated checklist of the Proagoderus dives (Harold, 1877) species-group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="6B5FEDC02982F14B3FABA55D3445C7DD">Daniel, Gimo M.</mods:namePart>
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<mods:affiliation id="DE816E9F26CF994CD5E831026B84C4E5">Department of Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, South Africa & Department of Biological & Environmental Sciences, Walter Sisulu University, Mthatha, South Africa & Field Museum of Natural History, 1400 S DuSable Lake Shore Drive, Chicago, IL 60605, USA</mods:affiliation>
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<mods:affiliation id="3EB36CC681B0D7F4B44147F37109FB22">Faculty of Natural Sciences, Lúrio University, Pemba P. O. Box 958, Mozambique</mods:affiliation>
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<heading id="310F81ABFFA8FF85F5D6E64A3405F93A" bold="true" box="[151,563,1652,1679]" fontSize="11" level="1" pageId="8" pageNumber="183" reason="1">
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<taxonomicName id="ADF84D44FFA8FF85F5D6E64A3405F93A" ID-CoL="4MKX9" authority="(Harold, 1877)" baseAuthorityName="Harold" baseAuthorityYear="1877" box="[151,563,1652,1679]" class="Insecta" family="Scarabaeidae" genus="Proagoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="dives">
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<emphasis id="588CEAD5FFA8FF85F5D6E64A3405F93A" bold="true" box="[151,563,1652,1679]" pageId="8" pageNumber="183">
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<emphasis id="588CEAD5FFA8FF85F5D6E64A3743F93B" bold="true" box="[151,373,1652,1679]" italics="true" pageId="8" pageNumber="183">Proagoderus dives</emphasis>
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(
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<bibRefCitation id="0E694B36FFA8FF85F4C4E64A341CF93A" author="Harold, E." box="[389,554,1652,1678]" pageId="8" pageNumber="183" pagination="97 - 101" refId="ref7281" refString="Harold, E. (1877) Coleopterorum species novae. Mitteilungen der Munchner Entomologischen Gesellschaft, 1, 97 - 101." type="journal article" year="1877">Harold, 1877</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="22E2654CFFA8FF85F5D6E68732BBF944" pageId="8" pageNumber="183" type="description">
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<paragraph id="6A4736C7FFA8FF85F5D6E6873281F965" blockId="8.[151,1436,1721,1868]" box="[151,1207,1721,1745]" pageId="8" pageNumber="183">
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=
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<taxonomicName id="ADF84D44FFA8FF85F5EDE6873413F965" authority="Raffray, 1877" authorityName="Raffray" authorityYear="1877" box="[172,549,1721,1745]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="deyrollei">
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<emphasis id="588CEAD5FFA8FF85F5EDE68737A2F965" box="[172,404,1721,1745]" italics="true" pageId="8" pageNumber="183">Onthophagus deyrollei</emphasis>
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<materialsCitation id="DA903C9AFFA8FF85F4DBE6873413F965" box="[410,549,1721,1745]" country="Tanzania" county="Bagamoyo" location="Zanguebar" municipality="Bagamoyo" pageId="8" pageNumber="183" specimenCount="1">Raffray, 1877</materialsCitation>
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</taxonomicName>
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(
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<typeStatus id="B5438865FFA8FF85F773E6873469F965" box="[562,607,1721,1745]" pageId="8" pageNumber="183">type</typeStatus>
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locality: “
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<location id="6F27601CFFA8FF85F789E687350EF965" LSID="urn:lsid:plazi:treatment:E25187D1FFA8FF85F5D6E64A35FAF868:6F27601CFFA8FF85F789E687350EF965" box="[712,824,1721,1745]" country="Tanzania" county="Bagamoyo" municipality="Bagamoyo" name="Zanguebar" pageId="8" pageNumber="183">Zanguebar</location>
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,
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<collectingMunicipality id="8A23ACBDFFA8FF85F603E6873584F965" box="[834,946,1721,1745]" pageId="8" pageNumber="183">Bagamoyo</collectingMunicipality>
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” [
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<collectingCounty id="83264E4BFFA8FF85F686E687320CF965" box="[967,1082,1721,1745]" pageId="8" pageNumber="183">Bagamoyo</collectingCounty>
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,
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<collectingCountry id="12EF7657FFA8FF85F105E6873290F965" box="[1092,1190,1721,1745]" name="Tanzania" pageId="8" pageNumber="183">Tanzania</collectingCountry>
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]).
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</paragraph>
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<paragraph id="6A4736C7FFA8FF85F5D6E6E632BBF944" blockId="8.[151,1436,1721,1868]" box="[151,1165,1752,1776]" pageId="8" pageNumber="183">
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=
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<taxonomicName id="ADF84D44FFA8FF85F5EDE6E63419F944" authority="Peringuey, 1901" authorityName="Peringuey" authorityYear="1901" box="[172,559,1752,1776]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="optivus">
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<emphasis id="588CEAD5FFA8FF85F5EDE6E637B2F95B" box="[172,388,1752,1776]" italics="true" pageId="8" pageNumber="183">Onthophagus optivus</emphasis>
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<materialsCitation id="DA903C9AFFA8FF85F4CBE6E63419F944" box="[394,559,1752,1776]" country="Zimbabwe" location="Mtoko" pageId="8" pageNumber="183" specimenCount="1">Péringuey, 1901</materialsCitation>
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</taxonomicName>
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||
(
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<typeStatus id="B5438865FFA8FF85F77DE6E6345FF944" box="[572,617,1752,1776]" pageId="8" pageNumber="183">type</typeStatus>
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locality: “Southern Rhodesia (
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<location id="6F27601CFFA8FF85F6E0E6E635DFF944" LSID="urn:lsid:plazi:treatment:E25187D1FFA8FF85F5D6E64A35FAF868:6F27601CFFA8FF85F6E0E6E635DFF944" box="[929,1001,1752,1776]" country="Zimbabwe" name="Mtoko" pageId="8" pageNumber="183">Mtoko</location>
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)” [
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<collectingCountry id="12EF7657FFA8FF85F147E6E6324AF944" box="[1030,1148,1752,1776]" name="Zimbabwe" pageId="8" pageNumber="183">Zimbabwe</collectingCountry>
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]).
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</paragraph>
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</subSubSection>
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<subSubSection id="22E2654CFFA8FF85F5D6E6C932BAF8F8" pageId="8" pageNumber="183" type="reference_group">
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<paragraph id="6A4736C7FFA8FF85F5D6E6C9370CF899" blockId="8.[151,1436,1721,1868]" pageId="8" pageNumber="183">
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=
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<taxonomicName id="ADF84D44FFA8FF85F5F1E6C83568F8BA" ID-CoL="7HM97" authority="d'Orbigny, 1902" authorityName="d'Orbigny" authorityYear="1902" box="[176,862,1782,1806]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="variety" species="dives" subGenus="Proagoderus" variety="funereus">
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<emphasis id="588CEAD5FFA8FF85F5F1E6C8370FF8BA" box="[176,313,1782,1806]" italics="true" pageId="8" pageNumber="183">Onthophagus</emphasis>
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(
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<emphasis id="588CEAD5FFA8FF85F40DE6C837F9F8BA" box="[332,463,1782,1806]" italics="true" pageId="8" pageNumber="183">Proagoderus</emphasis>
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)
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<emphasis id="588CEAD5FFA8FF85F4A3E6C83420F8BA" box="[482,534,1782,1806]" italics="true" pageId="8" pageNumber="183">dives</emphasis>
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<materialsCitation id="DA903C9AFFA8FF85F760E6C93568F8BA" box="[545,862,1782,1806]" country="Tanzania" location="l'Ouzigoua" municipality="Mhonda" pageId="8" pageNumber="183" specimenCount="1">
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var.
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<emphasis id="588CEAD5FFA8FF85F710E6C83491F8BA" box="[593,679,1782,1806]" italics="true" pageId="8" pageNumber="183">funereus</emphasis>
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<bibRefCitation id="0E694B36FFA8FF85F7F3E6C83568F8BA" author="d'Orbigny, H." box="[690,862,1782,1806]" pageId="8" pageNumber="183" pagination="1 - 324" refId="ref7060" refString="d'Orbigny, H. (1902) Memoire sur les Onthophagides d'Afrique. Annales de la Societe Entomologique de France, 71, 1 - 324." type="journal article" year="1902">d’Orbigny, 1902</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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(
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||
<typeStatus id="B5438865FFA8FF85F62EE6C835AAF8BA" box="[879,924,1782,1806]" pageId="8" pageNumber="183">type</typeStatus>
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locality: “M’honda dans
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<location id="6F27601CFFA8FF85F1F0E6C83313F8BA" LSID="urn:lsid:plazi:treatment:E25187D1FFA8FF85F5D6E64A35FAF868:6F27601CFFA8FF85F1F0E6C83313F8BA" box="[1201,1317,1782,1806]" country="Tanzania" municipality="Mhonda" name="l'Ouzigoua" pageId="8" pageNumber="183">l’Ouzigoua</location>
|
||
” [
|
||
<collectingMunicipality id="8A23ACBDFFA8FF85F07EE6C833AFF8BA" box="[1343,1433,1782,1806]" pageId="8" pageNumber="183">Mhonda</collectingMunicipality>
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,
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<collectingCountry id="12EF7657FFA8FF85F586E72B371FF899" box="[199,297,1813,1837]" name="Tanzania" pageId="8" pageNumber="183">Tanzania</collectingCountry>
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]).
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</paragraph>
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<paragraph id="6A4736C7FFA8FF85F5D6E70A32BAF8F8" blockId="8.[151,1436,1721,1868]" box="[151,1164,1844,1868]" pageId="8" pageNumber="183">
|
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=
|
||
<taxonomicName id="ADF84D44FFA8FF85F5EDE70A3546F8F8" ID-CoL="7HLWW" authority="d'Orbigny, 1913" authorityName="d'Orbigny" authorityYear="1913" box="[172,880,1844,1868]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="variety" species="dives" subGenus="Proagoderus" variety="semicuprinus">
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<emphasis id="588CEAD5FFA8FF85F5EDE70A3703F8F8" box="[172,309,1844,1868]" italics="true" pageId="8" pageNumber="183">Onthophagus</emphasis>
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(
|
||
<emphasis id="588CEAD5FFA8FF85F402E70A37F0F8F8" box="[323,454,1844,1868]" italics="true" pageId="8" pageNumber="183">Proagoderus</emphasis>
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||
)
|
||
<emphasis id="588CEAD5FFA8FF85F494E70A343FF8F8" box="[469,521,1844,1868]" italics="true" pageId="8" pageNumber="183">dives</emphasis>
|
||
var.
|
||
<emphasis id="588CEAD5FFA8FF85F77AE70A34F4F8FF" box="[571,706,1844,1867]" italics="true" pageId="8" pageNumber="183">semicuprinus</emphasis>
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||
<bibRefCitation id="0E694B36FFA8FF85F788E70A3546F8F8" author="d'Orbigny, H." box="[713,880,1844,1868]" pageId="8" pageNumber="183" pagination="1 - 742" refId="ref7205" refString="d'Orbigny, H. (1913) Synopsis des Onthophagides d'Afrique. Annales de la Societe Entomologique de France, 82, 1 - 742." type="journal article" year="1913">d’Orbigny, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="B5438865FFA8FF85F63CE70A359CF8F8" box="[893,938,1844,1868]" pageId="8" pageNumber="183">type</typeStatus>
|
||
locality: not defined).
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="22E2654CFFA8FF85F5D6E744355BF820" box="[151,877,1913,1940]" pageId="8" pageNumber="183" type="materials_examined">
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<paragraph id="6A4736C7FFA8FF85F5D6E744355BF820" blockId="8.[151,1437,1913,2012]" box="[151,877,1913,1940]" pageId="8" pageNumber="183">
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<materialsCitation id="DA903C9AFFA8FF85F5D6E744355CF820" box="[151,874,1913,1940]" collectorName="Nyassa Afric. Or. Trop." country="Malawi" location="Lake" pageId="8" pageNumber="183" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="588CEAD5FFA8FF85F5D6E7443706F827" bold="true" box="[151,304,1913,1940]" pageId="8" pageNumber="183">
|
||
<typeStatus id="B5438865FFA8FF85F5D6E74436E4F820" box="[151,210,1914,1940]" pageId="8" pageNumber="183">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
: “
|
||
<collectorName id="C70D5311FFA8FF85F408E7443465F820" box="[329,595,1914,1940]" pageId="8" pageNumber="183">Nyassa Afric. Or. Trop.</collectorName>
|
||
” [
|
||
<location id="6F27601CFFA8FF85F72BE7443493F820" LSID="urn:lsid:plazi:treatment:E25187D1FFA8FF85F5D6E64A35FAF868:6F27601CFFA8FF85F72BE7443493F820" box="[618,677,1914,1940]" country="Malawi" name="Lake" pageId="8" pageNumber="183">Lake</location>
|
||
<collectingCountry id="12EF7657FFA8FF85F7EDE7443534F820" box="[684,770,1914,1940]" name="Malawi" pageId="8" pageNumber="183">Malawi</collectingCountry>
|
||
region]”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22E2654CFFA8FF85F586E7A335FAF868" pageId="8" pageNumber="183" type="distribution">
|
||
<paragraph id="6A4736C7FFA8FF85F586E7A335FAF868" blockId="8.[151,1437,1913,2012]" pageId="8" pageNumber="183">
|
||
<emphasis id="588CEAD5FFA8FF85F586E7A3376FF803" bold="true" box="[199,345,1949,1975]" pageId="8" pageNumber="183">Distribution</emphasis>
|
||
:
|
||
<collectingCountry id="12EF7657FFA8FF85F428E7A037ECF80C" box="[361,474,1950,1976]" name="Botswana" pageId="8" pageNumber="183">Botswana</collectingCountry>
|
||
,
|
||
<collectingCountry id="12EF7657FFA8FF85F4A4E7A03407F80C" box="[485,561,1950,1976]" name="Kenya" pageId="8" pageNumber="183">Kenya</collectingCountry>
|
||
,
|
||
<collectingCountry id="12EF7657FFA8FF85F77CE7A03496F80C" box="[573,672,1950,1976]" name="Namibia" pageId="8" pageNumber="183">Namibia</collectingCountry>
|
||
,
|
||
<collectingCountry id="12EF7657FFA8FF85F7EAE7A03525F80C" box="[683,787,1950,1976]" name="Tanzania" pageId="8" pageNumber="183">Tanzania</collectingCountry>
|
||
,
|
||
<collectingCountry id="12EF7657FFA8FF85F65FE7A03540F80C" box="[798,886,1950,1976]" name="Zambia" pageId="8" pageNumber="183">Zambia</collectingCountry>
|
||
,
|
||
<collectingCountry id="12EF7657FFA8FF85F6C3E7A035ECF80C" box="[898,986,1950,1976]" name="Malawi" pageId="8" pageNumber="183">Malawi</collectingCountry>
|
||
,
|
||
<collectingCountry id="12EF7657FFA8FF85F6A4E7A0324AF80C" box="[997,1148,1950,1976]" name="Mozambique" pageId="8" pageNumber="183">Mozambique</collectingCountry>
|
||
,
|
||
<collectingCountry id="12EF7657FFA8FF85F1C9E7A03335F80C" box="[1160,1283,1950,1976]" name="Zimbabwe" pageId="8" pageNumber="183">Zimbabwe</collectingCountry>
|
||
,
|
||
<collectingCountry id="12EF7657FFA8FF85F04FE7A033ABF80C" box="[1294,1437,1950,1976]" name="South Africa" pageId="8" pageNumber="183">South Africa</collectingCountry>
|
||
(
|
||
<bibRefCitation id="0E694B36FFA8FF85F5DEE7FC3706F868" author="Harold, E." box="[159,304,1986,2012]" pageId="8" pageNumber="183" pagination="97 - 101" refId="ref7281" refString="Harold, E. (1877) Coleopterorum species novae. Mitteilungen der Munchner Entomologischen Gesellschaft, 1, 97 - 101." type="journal article" year="1877">Harold 1877</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="0E694B36FFA8FF85F47DE7FC37C5F868" author="d'Orbigny, H." box="[316,499,1986,2012]" pageId="8" pageNumber="183" pagination="1 - 324" refId="ref7060" refString="d'Orbigny, H. (1902) Memoire sur les Onthophagides d'Afrique. Annales de la Societe Entomologique de France, 71, 1 - 324." type="journal article" year="1902">d’Orbigny 1902</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="0E694B36FFA8FF85F4BFE7FC340FF868" author="d'Orbigny, H." box="[510,569,1986,2012]" pageId="8" pageNumber="183" pagination="1 - 742" refId="ref7205" refString="d'Orbigny, H. (1913) Synopsis des Onthophagides d'Afrique. Annales de la Societe Entomologique de France, 82, 1 - 742." type="journal article" year="1913">1913</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="0E694B36FFA8FF85F705E7FC34C6F868" author="Palestrini, C." box="[580,752,1986,2012]" pageId="8" pageNumber="183" pagination="1 - 358" refId="ref7748" refString="Palestrini, C. (1992) Sistematica e zoogeografia del genere Onthophagus sottogenere Proagoderus Lansberge. Memorie della Societa Entomologica Italiana, 71 (1), 1 - 358." type="journal article" year="1992">Palestrini 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="0E694B36FFA8FF85F7BDE7FC358DF868" author="Davis, A. L. V. & Deschodt, C. M. & Scholtz, C. H." box="[764,955,1986,2012]" pageId="8" pageNumber="183" refId="ref6997" refString="Davis, A. L. V., Deschodt, C. M. & Scholtz, C. H. (2020) Conservation assessment of Scarabaeine dung beetles in South Africa, Botswana and Namibia: IUCN Red List categories, atlas and ecological notes. Suricata 6. South African National Biodiversity Institute, Pretoria, 799 pp." type="book" year="2020">
|
||
Davis
|
||
<emphasis id="588CEAD5FFA8FF85F605E7FD354BF868" box="[836,893,1986,2012]" italics="true" pageId="8" pageNumber="183">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |