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<document id="60588A2A4835DB6B45FFCF5D2FBDA3E2" ID-CLB-Dataset="24859" ID-DOI="10.11646/zootaxa.4700.3.1" ID-GBIF-Dataset="1b0654ea-eecb-4242-ad9a-850602f7d185" ID-ISSN="1175-5326" ID-Zenodo-Dep="3548509" ID-ZooBank="B0712990-7A3B-4042-9A4B-5CE96AFAECF8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1574256400520" checkinUser="plazi" docAuthor="Grogan, William L., Díaz, Florentina, Spinelli, Gustavo R. &amp; Ronderos, Maria M." docDate="2019" docId="DF72B52DDE14FF9CFF541C9FFD910EA8" docLanguage="en" docName="zootaxa.4700.3.1.pdf" docOrigin="Zootaxa 4700 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyhelea rhopaloparamera Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos 2019, n. sp." docType="treatment" docVersion="10" lastPageNumber="322" masterDocId="234BCD55DE06FF89FFC31D26FF870D63" masterDocTitle="The Biting Midges of the Caribbean island Curaçao (Diptera: Ceratopogonidae) I. Species in the genus Dasyhelea Kieffer" masterLastPageNumber="325" masterPageNumber="301" pageNumber="319" updateTime="1698701345973" updateUser="plazi">
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<mods:title id="4C3DABC3620FEDFFBEE4ED13562B5DB7">The Biting Midges of the Caribbean island Curaçao (Diptera: Ceratopogonidae) I. Species in the genus Dasyhelea Kieffer</mods:title>
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<mods:namePart id="075823B5F1B9986715759B22E91572F8">Grogan, William L.</mods:namePart>
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<mods:namePart id="231F807B836FAF65DB1ED1A0C3AFB835">Díaz, Florentina</mods:namePart>
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<mods:namePart id="8C5EA2417A166F6CFDD0C9572DE242D3">Spinelli, Gustavo R.</mods:namePart>
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<mods:namePart id="1A786DEDE83DE4BA9A6D3C788F8A96AF">Ronderos, Maria M.</mods:namePart>
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<paragraph id="5764043BDE14FF9BFF541C9FFDBD0CB0" blockId="18.[151,570,441,504]" box="[151,570,441,468]" pageId="18" pageNumber="319">
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<emphasis id="65AFD829DE14FF9BFF541C9FFDBD0CB0" bold="true" box="[151,570,441,468]" pageId="18" pageNumber="319">
<taxonomicName id="90DB7FB8DE14FF9BFF541C9FFE770CB7" ID-CoL="8BQWP" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[151,496,441,468]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="rhopaloparamera" status="sp. nov.">
<emphasis id="65AFD829DE14FF9BFF541C9FFE770CB7" bold="true" box="[151,496,441,468]" italics="true" pageId="18" pageNumber="319">Dasyhelea rhopaloparamera</emphasis>
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<taxonomicNameLabel id="7E9C6552DE14FF9BFE341C9FFDBD0CB0" box="[503,570,441,467]" pageId="18" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="1FC157B0DE14FF9BFF541CFBFE050C9B" box="[151,386,476,504]" pageId="18" pageNumber="319" type="description">
<paragraph id="5764043BDE14FF9BFF541CFBFE050C9B" blockId="18.[151,570,441,504]" box="[151,386,476,504]" pageId="18" pageNumber="319">
(
<figureCitation id="CFE018BEDE14FF9BFF5C1CFBFEA00C94" box="[159,295,476,503]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Figs. 4251</figureCitation>
,
<figureCitation id="CFE018BEDE14FF9BFEF01CFBFEFD0C9B" box="[307,378,477,504]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="18" pageNumber="319">6769</figureCitation>
)
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<subSubSection id="1FC157B0DE14FF9AFF541F02FC850C7C" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319" type="diagnosis">
<paragraph id="5764043BDE14FF9BFF541F02FAC30EE0" blockId="18.[151,1437,548,2020]" pageId="18" pageNumber="319">
<emphasis id="65AFD829DE14FF9BFF541F02FE8D0F5C" bold="true" box="[151,266,548,575]" pageId="18" pageNumber="319">Diagnosis</emphasis>
. The only Neotropical species with the following combination of characters. Male antennal flagellum short with only flagellomeres 1113 elongate; wing very broad, membrane hyaline without macrotrichiae; gonocoxal apodemes and paramere symmetrical; paramere elongate, club-shaped; aedeagus complex, composed of two sections, a heavily sclerotized H-shaped basal portion and a broad, hyaline membranous quadrate posterior portion attached to the distal section of the basal portion, the apex of which is broad with short, blunt apicolateral extensions and an apicocentral lobe; sternite 9 elongate, distal margin convex with a small, slender, sharply pointed apicocentral projection. Female with flagellomeres 212 moderately short, vasiform, 13 more elongate, antennal ratio 0.70; wing broad, membrane hyaline, macrotrichiae sparse, a few marginal ones in cells r
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and m
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; subgenital plate subcircular posteromedian margin rounded, posterolateral arms stout, nearly straight; spermatheca ovoid, very heavily sclerotized with hyaline punctations, neck short, lightly sclerotized, elongate, very slender, slightly curved.
</paragraph>
<paragraph id="5764043BDE14FF9BFF041EABFC850BF8" blockId="18.[151,1437,548,2020]" pageId="18" pageNumber="319">
<emphasis id="65AFD829DE14FF9BFF041EABFE830EC4" bold="true" box="[199,260,909,935]" pageId="18" pageNumber="319">Male</emphasis>
. Head (
<figureCitation id="CFE018BEDE14FF9BFE991EABFE2A0EC4" box="[346,429,909,935]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Fig. 42</figureCitation>
). Brown. Eyes contiguous for a distance equal to the width of 34 ommatidia.Antennal flagellum short; flagellomeres 1213 extending beyond apex of palpus; 212 with sculpturing, 210 rhomboid, 1213 more elongate, 13 longest, broadest; 210 with single whorl of long sensillae chaetica comprising the dense plume that extends just beyond flagellomere 11; antennal ratio 0.58; flagellum length
<quantity id="9023A9DEDE14FF9BFBD61EDFFBFA0970" box="[1045,1149,1017,1043]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="18" pageNumber="319" unit="mm" value="0.44">0.44 mm</quantity>
. Frontal sclerite distorted and damaged during slide-mounting, but probably elliptical in shape. Clypeus with 4 pairs of setae. Palpus (
<figureCitation id="CFE018BEDE14FF9BFAAC193BFF330938" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Fig. 43</figureCitation>
) brown; segment 3 slender, as long as 4+5 combined with scattered hyaline capitate sensillae on mesal surface; palpal ratio 3.00. Thorax. Brown; scutellum yellowish brown with 6 large, 2 smaller setae. Legs yellowish brown, tarsomeres 5 infuscated. Wing (
<figureCitation id="CFE018BEDE14FF9BFE3B19AFFDCE09C0" box="[504,585,1161,1187]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Fig. 44</figureCitation>
) nearly as broad as female wing; membrane hyaline, without macrotrichiae; veins brown; 2
<superScript id="A0AEA973DE14FF9BFE81198AFED509D9" attach="left" box="[322,338,1196,1210]" fontSize="6" pageId="18" pageNumber="319">nd</superScript>
radial cell short, poorly developed, with gap between apices of costa, radius; fork of CuA
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just beyond level of apex of costa; anal lobe well developed; wing length
<quantity id="9023A9DEDE14FF9BFC5219F6FC720988" box="[913,1013,1232,1259]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="18" pageNumber="319" unit="mm" value="0.85">0.85 mm</quantity>
, width
<quantity id="9023A9DEDE14FF9BFB8619F7FB2C0988" box="[1093,1195,1233,1259]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="18" pageNumber="319" unit="mm" value="0.32">0.32 mm</quantity>
; costal ratio 0.47. Abdomen. Dark brown. Genitalia (
<figureCitation id="CFE018BEDE14FF9BFE3C19D3FDDA086C" box="[511,605,1268,1295]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Figs. 45</figureCitation>
,
<figureCitation id="CFE018BEDE14FF9BFDA919D3FD330873" box="[618,692,1269,1296]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="18" pageNumber="319">6769</figureCitation>
). Tergite 9 short, mid-section thickest, distal 1/2 abruptly tapering to rounded apex; apicolateral process short, conical with stout apical seta; cercus conspicuous with 34 large setae. Sternite 9 (
<figureCitation id="CFE018BEDE14FF9BFED2181BFEE90834" box="[273,366,1340,1367]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Figs. 45</figureCitation>
,
<figureCitation id="CFE018BEDE14FF9BFEB9181BFE10083B" box="[378,407,1341,1368]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="18" pageNumber="319">67</figureCitation>
) 0.44 length of greatest width; base curved antero-laterally; distal margin convex, with slen- der, sharply pointed apicocentral projection (partially folded in
<typeStatus id="8860BA99DE14FF9BFC901847FC3D081F" box="[851,954,1377,1404]" pageId="18" pageNumber="319" type="holotype">holotype</typeStatus>
). Gonocoxite very short, slightly triangular, 1.4 x longer than greatest (distal) width; gonostylus stout proximally, nearly straight, as long as gonocoxite, tapered distally with slightly curved, pointed tip. Paramere and gonocoxal apodemes (
<figureCitation id="CFE018BEDE14FF9BFC38188FFBDF08A0" box="[1019,1112,1448,1475]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Figs. 45</figureCitation>
,
<figureCitation id="CFE018BEDE14FF9BFBA0188FFB0708A7" box="[1123,1152,1449,1476]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="18" pageNumber="319">68</figureCitation>
) symmetrical; gonocoxal apodemes arcuate, heavily sclerotized, basal knob stout; paramere elongate, club-shaped, distal 2/3 lightly sclerotized. Aedeagus (
<figureCitation id="CFE018BEDE14FF9BFE9B18D7FE320B68" box="[344,437,1520,1547]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Figs. 45</figureCitation>
,
<figureCitation id="CFE018BEDE14FF9BFE0218D7FE590B6F" box="[449,478,1521,1548]" captionStart="FIGURES 5269" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 5269. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254 D. aliciae n. sp., 5557 D. recurva n. sp., 5860 D. latiala n. sp., 6163 D. cyrtostyla n. sp., 6466 D. curacaoensis n. sp., 6769 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="18" pageNumber="319">69</figureCitation>
) basal portion heavily sclerotized, 0.50 length of greatest width, basal arch extending 0.3 of total length, basal arms nearly straight, with short, subapical outer tooth, and slender posterolateral arms that are heavily sclerotized basally, hyaline distally with convergent pointed apices; a broad, hyaline membranous quadrate posterior portion attached to the distal section of the basal portion, the apex of which is broad with short, blunt apicolateral extensions and a hyaline apicocentral lobe.
</paragraph>
<paragraph id="5764043BDE14FF9AFF041B83FC850C7C" blockId="18.[151,1437,548,2020]" lastBlockId="19.[151,1437,152,287]" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">
<emphasis id="65AFD829DE14FF9BFF041B83FE990BDC" bold="true" box="[199,286,1701,1727]" pageId="18" pageNumber="319">Female</emphasis>
. Head (
<figureCitation id="CFE018BEDE14FF9BFEB51B83FE4D0BA3" box="[374,458,1701,1728]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Fig. 46</figureCitation>
). Dark brown. Eyes contiguous for a distance equal to the width of 34 ommatidia. Antennal flagellum with flagellomeres 212 vasiform, slightly longer than broad, 13 much longer than broad; antennal ratio 0.70; flagellum length
<quantity id="9023A9DEDE14FF9BFE161BCBFDBA0A64" box="[469,573,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="18" pageNumber="319" unit="mm" value="0.44">0.44 mm</quantity>
. Frontal sclerite distorted and damaged during slide-mounting, but probably elliptical. Clypeus (
<figureCitation id="CFE018BEDE14FF9BFEB31A37FE460A48" box="[368,449,1809,1835]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Fig. 47</figureCitation>
) with 4 pairs of setae. Palpus (
<figureCitation id="CFE018BEDE14FF9BFCD01A37FCE30A48" box="[787,868,1809,1835]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Fig. 48</figureCitation>
) brown; segment 3 with capitate sensillae on entire mesal surface; palpal ratio 3.66. Thorax (
<figureCitation id="CFE018BEDE14FF9BFDAE1A13FD440A2C" box="[621,707,1845,1871]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Fig. 49</figureCitation>
). Brown, humeral angles and prescutellar depression paler; scutellum yellowish brown with 6 large, 2 smaller setae. Legs pale brown, tarsomeres 5 infuscated; hind tibial comb with 5 spines; prothoracic tarsal ratio 2.73, mesothoracic tarsal ratio 3.66, metathoracic tarsal ratio 2.00. Wing (
<figureCitation id="CFE018BEDE14FF9BFAAC1A5BFF330AD8" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="18" pageNumber="319">Fig. 50</figureCitation>
) broad; membrane hyaline, macrotrichiae sparse, a few marginal ones in cells r
<subScript id="CB5F067EDE14FF9BFBFB1A94FBC70AA3" attach="left" box="[1080,1088,1970,1984]" fontSize="6" pageId="18" pageNumber="319">3</subScript>
and m
<subScript id="CB5F067EDE14FF9BFB4D1A94FB110AA3" attach="left" box="[1166,1174,1970,1984]" fontSize="6" pageId="18" pageNumber="319">1</subScript>
; 2
<superScript id="A0AEA973DE14FF9BFB771A86FB430ACD" attach="left" box="[1204,1220,1952,1966]" fontSize="6" pageId="18" pageNumber="319">nd</superScript>
radial cell reduced to a suture; apices of costa, radius at slight acute angle; fork of CuA
<subScript id="CB5F067EDE14FF9BFC571AF0FC1B0A87" attach="left" box="[916,924,2006,2020]" fontSize="6" pageId="18" pageNumber="319">1</subScript>
at level of base of 2
<superScript id="A0AEA973DE14FF9BFB421AE2FB160AB1" attach="left" box="[1153,1169,1988,2002]" fontSize="6" pageId="18" pageNumber="319">nd</superScript>
radial cell; wing length
<quantity id="9023A9DEDE15FF9AFF541DBFFF7A0DD0" box="[151,253,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="19" pageNumber="320" unit="mm" value="0.82">0.82 mm</quantity>
, width
<quantity id="9023A9DEDE15FF9AFE8D1DBEFE320DD0" box="[334,437,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="320" unit="mm" value="0.35">0.35 mm</quantity>
; costal ratio 0.51. Halter brown. Abdomen. Brown. Subgenital plate (
<figureCitation id="CFE018BEDE15FF9AFB7A1DBFFA8D0DD0" box="[1209,1290,152,179]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="19" pageNumber="320">Fig. 51</figureCitation>
) ribbon-like, arcuate, anteromesal margin heavily sclerotized; posterolateral arms stout, nearly straight. Spermatheca (
<figureCitation id="CFE018BEDE15FF9AFA821D9BFA130DB4" box="[1345,1428,188,215]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="19" pageNumber="320">Fig. 51</figureCitation>
) large, ovoid, very heavily sclerotized with hyaline punctations;
<date id="236522FBDE15FF9AFC9E1DC7FC090D98" box="[861,910,225,251]" pageId="19" pageNumber="320" value="1967-10">67 x</date>
<quantity id="9023A9DEDE15FF9AFC561DC7FC660D98" box="[917,993,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="320" unit="mm" value="60.0">60 µm</quantity>
; neck (
<figureCitation id="CFE018BEDE15FF9AFBF31DC7FB070D98" box="[1072,1152,224,251]" captionStart="FIGURES 4251" captionStartId="19.[151,264,1813,1838]" captionTargetBox="[174,1410,326,1775]" captionTargetId="figure@19.[164,1423,309,1788]" captionTargetPageId="19" captionText="FIGURES 4251. Dasyhelea rhopaloparamera n. sp. Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="19" pageNumber="320">Fig. 51</figureCitation>
inset) lightly sclerotized, elongate, very slender, slightly curved, length
<quantity id="9023A9DEDE15FF9AFD5E1C22FD790C7C" box="[669,766,260,287]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="19" pageNumber="320" unit="mm" value="27.5">27.5 µm</quantity>
.
</paragraph>
</subSubSection>
<caption id="03A454B3DE15FF9AFF541A33FAA20A32" ID-DOI="http://doi.org/10.5281/zenodo.3548519" ID-Zenodo-Dep="3548519" httpUri="https://zenodo.org/record/3548519/files/figure.png" pageId="19" pageNumber="320" startId="19.[151,264,1813,1838]" targetBox="[174,1410,326,1775]" targetPageId="19">
<paragraph id="5764043BDE15FF9AFF541A33FAA20A32" blockId="19.[151,1436,1813,1874]" pageId="19" pageNumber="320">
<emphasis id="65AFD829DE15FF9AFF541A33FECA0A4D" bold="true" box="[151,333,1813,1838]" pageId="19" pageNumber="320">FIGURES 4251</emphasis>
.
<taxonomicName id="90DB7FB8DE15FF9AFE981A30FD070A4D" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[347,640,1814,1838]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="rhopaloparamera" status="sp. nov.">
<emphasis id="65AFD829DE15FF9AFE981A30FD070A4D" box="[347,640,1814,1838]" italics="true" pageId="19" pageNumber="320">Dasyhelea rhopaloparamera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE15FF9AFD451A30FD380A4E" box="[646,703,1814,1838]" pageId="19" pageNumber="320" rank="species">
<emphasis id="65AFD829DE15FF9AFD451A30FD130A4D" bold="true" box="[646,660,1814,1838]" pageId="19" pageNumber="320">n</emphasis>
.
<emphasis id="65AFD829DE15FF9AFD621A30FD3E0A4D" bold="true" box="[673,697,1814,1838]" pageId="19" pageNumber="320">sp</emphasis>
.
</taxonomicNameLabel>
Holotype male, 4245, allotype female, 4651. 42, 46 head. 43, 48 palpus. 44, 50 wing. 45 genitalia. 47 clypeus. 49 thorax. 51 apex of abdomen and spermatheca. Scale bars = 0.05 mm.
</paragraph>
</caption>
<subSubSection id="1FC157B0DE15FF9AFF041A59FE4B0AF9" box="[199,460,1919,1947]" pageId="19" pageNumber="320" type="distribution">
<paragraph id="5764043BDE15FF9AFF041A59FE4B0AF9" blockId="19.[151,1436,1919,2019]" box="[199,460,1919,1947]" pageId="19" pageNumber="320">
<emphasis id="65AFD829DE15FF9AFF041A59FEDE0AF9" bold="true" box="[199,345,1919,1946]" pageId="19" pageNumber="320">Distribution</emphasis>
.
<collectingCountry id="2FCC44ABDE15FF9AFEA41AA6FE4F0AF9" box="[359,456,1920,1946]" name="Curaçao" pageId="19" pageNumber="320">Curaçao</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE15FF9CFF041A82FB2D0DB4" lastPageId="21" lastPageNumber="322" pageId="19" pageNumber="320" type="materials_examined">
<paragraph id="5764043BDE15FF9AFF041A82FD670A81" blockId="19.[151,1436,1919,2019]" pageId="19" pageNumber="320">
<emphasis id="65AFD829DE15FF9AFF041A82FEE90ADD" bold="true" box="[199,366,1955,1982]" pageId="19" pageNumber="320">Type material</emphasis>
.
<materialsCitation id="E7B30E66DE15FF9AFEBF1A82FD670A81" ID-GBIF-Occurrence="2454903118" collectingDate="2015-12-05" collectionCode="FSCA" collectorName="M. C. Thomas" country="Curaçao" location="Piscadera Baai" pageId="19" pageNumber="320" specimenCount="2" typeStatus="holotype">
<typeStatus id="8860BA99DE15FF9AFEBF1A82FE630ADD" box="[380,484,1956,1982]" pageId="19" pageNumber="320" type="holotype">Holotype</typeStatus>
male,
<typeStatus id="8860BA99DE15FF9AFDEC1A82FD0E0ADD" box="[559,649,1956,1982]" pageId="19" pageNumber="320" type="allotype">allotype</typeStatus>
female labeled “
<collectingCountry id="2FCC44ABDE15FF9AFC861A82FC2F0ADD" box="[837,936,1956,1982]" name="Curaçao" pageId="19" pageNumber="320">Curaçao</collectingCountry>
,
<location id="520452E0DE15FF9AFC701A82FBDC0ADD" LSID="urn:lsid:plazi:treatment:DF72B52DDE14FF9CFF541C9FFD910EA8:520452E0DE15FF9AFC701A82FBDC0ADD" box="[947,1115,1956,1982]" country="Curaçao" name="Piscadera Baai" pageId="19" pageNumber="320">Piscadera Baai</location>
,
<date id="236522FBDE15FF9AFBA41A85FB6E0ADD" box="[1127,1257,1955,1982]" pageId="19" pageNumber="320" value="2015-12-05">
<collectingDate id="3321DB13DE15FF9AFBA41A85FB6E0ADD" box="[1127,1257,1955,1982]" pageId="19" pageNumber="320" value="2015-12-05">5-XII-2015</collectingDate>
</date>
,
<collectorName id="FA2E61EDDE15FF9AFB371A83FA1F0ADD" box="[1268,1432,1956,1983]" pageId="19" pageNumber="320">M. C. Thomas</collectorName>
,
<collectingMethod id="8E9A7C2CDE15FF9AFF541AEEFEB80A80" box="[151,319,1992,2019]" pageId="19" pageNumber="320">Blacklight trap</collectingMethod>
in mangroves” (deposited in
<collectionCode id="31CA9CFEDE15FF9AFD4A1AEEFD530A81" box="[649,724,1992,2018]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="19" pageNumber="320" type="Museum">FSCA</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="03A454B3DE12FF9DFF541ABEFB0C0A9A" ID-DOI="http://doi.org/10.5281/zenodo.3548521" ID-Zenodo-Dep="3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="20" pageNumber="321" startId="20.[151,264,1944,1969]" targetBox="[187,1405,192,1908]" targetPageId="20">
<paragraph id="5764043BDE12FF9DFF541ABEFB0C0A9A" blockId="20.[151,1436,1944,2041]" pageId="20" pageNumber="321">
<emphasis id="65AFD829DE12FF9DFF541ABEFEC90AD2" bold="true" box="[151,334,1944,1969]" pageId="20" pageNumber="321">FIGURES 5269</emphasis>
. Male
<taxonomicName id="90DB7FB8DE12FF9DFE551ABFFD860AD2" baseAuthorityName="Dominiak" baseAuthorityYear="2012" box="[406,513,1945,1969]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="65AFD829DE12FF9DFE551ABFFD860AD2" box="[406,513,1945,1969]" italics="true" pageId="20" pageNumber="321">Dasyhelea</emphasis>
</taxonomicName>
genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 5254
<taxonomicName id="90DB7FB8DE12FF9DFD9F1A98FD380AB6" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[604,703,1981,2005]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="65AFD829DE12FF9DFD9F1A98FDE90AB6" box="[604,622,1982,2005]" italics="true" pageId="20" pageNumber="321">D</emphasis>
.
<emphasis id="65AFD829DE12FF9DFDB91A9BFD380AB6" box="[634,703,1981,2005]" italics="true" pageId="20" pageNumber="321">aliciae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE12FF9DFD051A9BFD790AB6" box="[710,766,1981,2005]" pageId="20" pageNumber="321" rank="species">
<emphasis id="65AFD829DE12FF9DFD051A9BFD530AB6" bold="true" box="[710,724,1981,2005]" pageId="20" pageNumber="321">n</emphasis>
.
<emphasis id="65AFD829DE12FF9DFD231A9BFD7F0AB6" bold="true" box="[736,760,1981,2005]" pageId="20" pageNumber="321">sp</emphasis>
.
</taxonomicNameLabel>
, 5557
<taxonomicName id="90DB7FB8DE12FF9DFC921A98FC3A0AB6" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[849,957,1982,2005]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="recurva" status="sp. nov.">
<emphasis id="65AFD829DE12FF9DFC921A98FCE40AB6" box="[849,867,1982,2005]" italics="true" pageId="20" pageNumber="321">D</emphasis>
.
<emphasis id="65AFD829DE12FF9DFCAC1A98FC3A0AB6" box="[879,957,1982,2005]" italics="true" pageId="20" pageNumber="321">recurva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE12FF9DFC071A9BFC7B0AB6" box="[964,1020,1981,2005]" pageId="20" pageNumber="321" rank="species">
<emphasis id="65AFD829DE12FF9DFC071A9BFC550AB6" bold="true" box="[964,978,1981,2005]" pageId="20" pageNumber="321">n</emphasis>
.
<emphasis id="65AFD829DE12FF9DFC1D1A9BFC710AB6" bold="true" box="[990,1014,1981,2005]" pageId="20" pageNumber="321">sp</emphasis>
.
</taxonomicNameLabel>
, 5860
<taxonomicName id="90DB7FB8DE12FF9DFB8C1A98FB280AB6" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[1103,1199,1981,2005]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="latiala" status="sp. nov.">
<emphasis id="65AFD829DE12FF9DFB8C1A98FBE60AB6" box="[1103,1121,1982,2005]" italics="true" pageId="20" pageNumber="321">D</emphasis>
.
<emphasis id="65AFD829DE12FF9DFBAE1A9BFB280AB6" box="[1133,1199,1981,2005]" italics="true" pageId="20" pageNumber="321">latiala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE12FF9DFB751A9BFB690AB6" box="[1206,1262,1981,2005]" pageId="20" pageNumber="321" rank="species">
<emphasis id="65AFD829DE12FF9DFB751A9BFB430AB6" bold="true" box="[1206,1220,1981,2005]" pageId="20" pageNumber="321">n</emphasis>
.
<emphasis id="65AFD829DE12FF9DFB131A9BFB6F0AB6" bold="true" box="[1232,1256,1981,2005]" pageId="20" pageNumber="321">sp</emphasis>
.
</taxonomicNameLabel>
, 6163
<taxonomicName id="90DB7FB8DE12FF9DFA821A98FF400A9A" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="cyrtostyla" status="sp. nov.">
<emphasis id="65AFD829DE12FF9DFA821A98FAD40AB6" box="[1345,1363,1982,2005]" italics="true" pageId="20" pageNumber="321">D</emphasis>
.
<emphasis id="65AFD829DE12FF9DFAA31A98FF400A9A" italics="true" pageId="20" pageNumber="321">cyrtostyla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE12FF9DFF0E1AC7FE810A9A" box="[205,262,2017,2041]" pageId="20" pageNumber="321" rank="species">
<emphasis id="65AFD829DE12FF9DFF0E1AC7FF5C0A9A" bold="true" box="[205,219,2017,2041]" pageId="20" pageNumber="321">n</emphasis>
.
<emphasis id="65AFD829DE12FF9DFF2B1AC7FE870A9A" bold="true" box="[232,256,2017,2041]" pageId="20" pageNumber="321">sp</emphasis>
.
</taxonomicNameLabel>
, 6466
<taxonomicName id="90DB7FB8DE12FF9DFE9A1AC4FE790A9A" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[345,510,2018,2041]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="curacaoensis" status="sp. nov.">
<emphasis id="65AFD829DE12FF9DFE9A1AC4FEEC0A9A" box="[345,363,2018,2041]" italics="true" pageId="20" pageNumber="321">D</emphasis>
.
<emphasis id="65AFD829DE12FF9DFEBB1AC4FE790A9A" box="[376,510,2018,2041]" italics="true" pageId="20" pageNumber="321">curacaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE12FF9DFDC71AC7FDBB0A9A" box="[516,572,2017,2041]" pageId="20" pageNumber="321" rank="species">
<emphasis id="65AFD829DE12FF9DFDC71AC7FD950A9A" bold="true" box="[516,530,2017,2041]" pageId="20" pageNumber="321">n</emphasis>
.
<emphasis id="65AFD829DE12FF9DFDDD1AC7FDB10A9A" bold="true" box="[542,566,2017,2041]" pageId="20" pageNumber="321">sp</emphasis>
.
</taxonomicNameLabel>
, 6769
<taxonomicName id="90DB7FB8DE12FF9DFD4C1AC4FCE50A9A" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[655,866,2017,2041]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="rhopaloparamera" status="sp. nov.">
<emphasis id="65AFD829DE12FF9DFD4C1AC4FD260A9A" box="[655,673,2018,2041]" italics="true" pageId="20" pageNumber="321">D</emphasis>
.
<emphasis id="65AFD829DE12FF9DFD6D1AC7FCE50A9A" box="[686,866,2017,2041]" italics="true" pageId="20" pageNumber="321">rhopaloparamera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE12FF9DFCAB1AC7FC260A9A" box="[872,929,2017,2041]" pageId="20" pageNumber="321" rank="species">
<emphasis id="65AFD829DE12FF9DFCAB1AC7FCF10A9A" bold="true" box="[872,886,2017,2041]" pageId="20" pageNumber="321">n</emphasis>
.
<emphasis id="65AFD829DE12FF9DFC401AC7FC1C0A9A" bold="true" box="[899,923,2017,2041]" pageId="20" pageNumber="321">sp</emphasis>
.
</taxonomicNameLabel>
Scale bars = 0.05 mm.
</paragraph>
</caption>
<paragraph id="5764043BDE13FF9CFF041DBEFB2D0DB4" blockId="21.[151,1437,152,971]" pageId="21" pageNumber="322">
<emphasis id="65AFD829DE13FF9CFF041DBEFD9D0DD0" bold="true" box="[199,538,152,179]" pageId="21" pageNumber="322">Derivation of specific epithet</emphasis>
. The specific epithet is a combination of two Greek terms,
<emphasis id="65AFD829DE13FF9CFB661DBFFB7A0DD0" box="[1189,1277,153,179]" italics="true" pageId="21" pageNumber="322">rhopalo</emphasis>
(clubbed) and
<emphasis id="65AFD829DE13FF9CFF541D98FE840DB4" box="[151,259,190,215]" italics="true" pageId="21" pageNumber="322">paramere</emphasis>
, in reference to the club-shaped paramere of the
<typeStatus id="8860BA99DE13FF9CFCEB1D9BFC0D0DB4" box="[808,906,189,215]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
male of this new species.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE13FF9CFF041DC6FD910EA8" pageId="21" pageNumber="322" type="discussion">
<paragraph id="5764043BDE13FF9CFF041DC6FAE20E74" blockId="21.[151,1437,152,971]" pageId="21" pageNumber="322">
<emphasis id="65AFD829DE13FF9CFF041DC6FEC20D98" bold="true" box="[199,325,224,251]" pageId="21" pageNumber="322">Discussion</emphasis>
. The
<typeStatus id="8860BA99DE13FF9CFE441DC7FE6E0D98" box="[391,489,225,251]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
male of this species is similar to males of
<taxonomicName id="90DB7FB8DE13FF9CFC0E1DC4FF640C7C" authority="Diaz &amp; Spinelli, in Diaz et al. (2011)" authorityName="Diaz &amp; Spinelli, in Diaz" authorityYear="2011" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="meloae">
<emphasis id="65AFD829DE13FF9CFC0E1DC4FC660D98" box="[973,993,226,251]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFC331DC7FBC60D98" box="[1008,1089,225,251]" italics="true" pageId="21" pageNumber="322">meloae</emphasis>
Díaz &amp; Spinelli,
<emphasis id="65AFD829DE13FF9CFACA1DC4FA980D98" box="[1289,1311,226,251]" italics="true" pageId="21" pageNumber="322">in</emphasis>
<bibRefCitation id="334A79CADE13FF9CFAE41DC7FF640C7C" author="Diaz, F. &amp; Spinelli, G. R. &amp; Ronderos, M. M." pageId="21" pageNumber="322" pagination="33 - 42" refId="ref18972" refString="Diaz, F., Spinelli, G. R. &amp; Ronderos, M. M. (2011) Six new species of Dasyhelea (Insecta, Diptera, Ceratopogonidae) from Patagonia. Iheringia, Serie Zoologica, 101, 33 - 42. https: // doi. org / 10.1590 / S 0073 - 47212011000100004" type="journal article" year="2011">
Díaz
<emphasis id="65AFD829DE13FF9CFAA01DC4FA120D98" box="[1379,1429,225,251]" italics="true" pageId="21" pageNumber="322">et al</emphasis>
. (2011)
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="90DB7FB8DE13FF9CFF2D1C20FCD70C7C" authority="Diaz &amp; Spinelli, in Diaz et al. (2011)" authorityName="Diaz &amp; Spinelli, in Diaz" authorityYear="2011" box="[238,848,261,287]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="pailemanensis">
<emphasis id="65AFD829DE13FF9CFF2D1C20FE850C7C" box="[238,258,262,287]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFECC1C23FE280C7C" box="[271,431,261,287]" italics="true" pageId="21" pageNumber="322">pailemanensis</emphasis>
Díaz &amp; Spinelli,
<emphasis id="65AFD829DE13FF9CFDB31C20FD010C7C" box="[624,646,262,287]" italics="true" pageId="21" pageNumber="322">in</emphasis>
<bibRefCitation id="334A79CADE13FF9CFD4F1C23FCD70C7C" author="Diaz, F. &amp; Spinelli, G. R. &amp; Ronderos, M. M." box="[652,848,261,287]" pageId="21" pageNumber="322" pagination="33 - 42" refId="ref18972" refString="Diaz, F., Spinelli, G. R. &amp; Ronderos, M. M. (2011) Six new species of Dasyhelea (Insecta, Diptera, Ceratopogonidae) from Patagonia. Iheringia, Serie Zoologica, 101, 33 - 42. https: // doi. org / 10.1590 / S 0073 - 47212011000100004" type="journal article" year="2011">
Díaz
<emphasis id="65AFD829DE13FF9CFD041C20FD700C7C" box="[711,759,261,287]" italics="true" pageId="21" pageNumber="322">et al</emphasis>
. (2011)
</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="90DB7FB8DE13FF9CFC4A1C20FF620C27" authority="Grogan, Diaz, Spinelli &amp; Ronderos (2016)" authorityName="Grogan, Diaz, Spinelli &amp; Ronderos" authorityYear="2016" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="turnbowi">
<emphasis id="65AFD829DE13FF9CFC4A1C20FC1A0C7C" box="[905,925,262,287]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFC691C23FB880C7C" box="[938,1039,261,287]" italics="true" pageId="21" pageNumber="322">turnbowi</emphasis>
<bibRefCitation id="334A79CADE13FF9CFBD61C23FF620C27" author="Grogan, W. L. Jr. &amp; Diaz, F. &amp; Spinelli, G. R. &amp; Ronderos, M. M." pageId="21" pageNumber="322" pagination="201 - 254" refId="ref19484" refString="Grogan, W. L. Jr., Diaz, F., Spinelli, G. R. &amp; Ronderos, M. M. (2016) The biting and predaceous midges of Guadeloupe (Diptera: Ceratopogonidae). II. Species of the subfamily Dasyheleinae. Zootaxa, 4184 (2), 201 - 254." type="journal article" year="2016">Grogan, Díaz, Spinelli &amp; Ronderos (2016)</bibRefCitation>
</taxonomicName>
. Males of
<taxonomicName id="90DB7FB8DE13FF9CFE9D1C0CFE540C20" box="[350,467,297,323]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="meloae">
<emphasis id="65AFD829DE13FF9CFE9D1C0CFEF50C20" box="[350,370,298,323]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFE411C0FFE540C20" box="[386,467,297,323]" italics="true" pageId="21" pageNumber="322">meloae</emphasis>
</taxonomicName>
differ from the
<typeStatus id="8860BA99DE13FF9CFD491C0FFD6B0C20" box="[650,748,297,323]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="90DB7FB8DE13FF9CFC971C0CFBB90C20" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[852,1086,297,323]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="rhopaloparamera" status="sp. nov.">
<emphasis id="65AFD829DE13FF9CFC971C0CFCEF0C20" box="[852,872,298,323]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFCB41C0FFBB90C20" box="[887,1086,297,323]" italics="true" pageId="21" pageNumber="322">rhopaloparamera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE13FF9CFB851C0FFB030C20" box="[1094,1156,297,323]" pageId="21" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
in having slender, much longer flagellomeres 1011; a clypeus with an angular, pointed base; a narrower wing with a poorly developed anal lobe; a shorter sternite 9 with a shallow marginal excavation and a pair of short, triangular submarginal extensions; longer gonocoxites with dense apicomesal patches of long setae and longer, broader gonostyli with broad blunt apices; longer, very slender gonocoxal apodemes and a very short slender paramere. The
<typeStatus id="8860BA99DE13FF9CFB971C9FFB310CB0" box="[1108,1206,441,467]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="90DB7FB8DE13FF9CFAD41C9CFF5A0C94" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="pailemanensis">
<emphasis id="65AFD829DE13FF9CFAD41C9CFAAC0CB0" box="[1303,1323,442,467]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFAFA1C9FFF5A0C94" italics="true" pageId="21" pageNumber="322">pailemanensis</emphasis>
</taxonomicName>
differs from the
<typeStatus id="8860BA99DE13FF9CFE581CFBFE7A0C94" box="[411,509,477,503]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="90DB7FB8DE13FF9CFDA31CF8FCCE0C94" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[608,841,477,503]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="rhopaloparamera" status="sp. nov.">
<emphasis id="65AFD829DE13FF9CFDA31CF8FDF30C94" box="[608,628,478,503]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFD411CFBFCCE0C94" box="[642,841,477,503]" italics="true" pageId="21" pageNumber="322">rhopaloparamera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE13FF9CFC931CFBFC0B0C94" box="[848,908,477,503]" pageId="21" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
in having slender, longer flagellomeres 1011; a shorter wing (wing length
<quantity id="9023A9DEDE13FF9CFE151F27FDBB0F78" box="[470,572,513,539]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="21" pageNumber="322" unit="mm" value="0.68">0.68 mm</quantity>
) with a shorter costa (costal ratio 0.41) and more numerous macrotrichiae in cells r
<subScript id="CB5F067EDE13FF9CFF191F10FF650F27" attach="left" box="[218,226,566,580]" fontSize="6" pageId="21" pageNumber="322">3</subScript>
, m
<subScript id="CB5F067EDE13FF9CFEC51F10FE890F27" attach="left" box="[262,270,566,580]" fontSize="6" pageId="21" pageNumber="322">1</subScript>
and m
<subScript id="CB5F067EDE13FF9CFE991F10FEE50F27" attach="left" box="[346,354,566,580]" fontSize="6" pageId="21" pageNumber="322">2</subScript>
; a similar shaped sternite 9 without a slender, sharply pointed, apico-mesal triangular projection; longer gonocoxites with dense apicomesal patches of long setae and longer gonostyli with broader apices; and narrower, straighter gonocoxal apodemes and a shorter, slender paramere. The
<typeStatus id="8860BA99DE13FF9CFC3B1F4BFBDD0FE4" box="[1016,1114,621,647]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="90DB7FB8DE13FF9CFB011F48FACC0FE4" authorityName="Grogan, Diaz, Spinelli &amp; Ronderos" authorityYear="2016" box="[1218,1355,621,647]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="turnbowi">
<emphasis id="65AFD829DE13FF9CFB011F48FB510FE4" box="[1218,1238,622,647]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFB251F4BFACC0FE4" box="[1254,1355,621,647]" italics="true" pageId="21" pageNumber="322">turnbowi</emphasis>
</taxonomicName>
differs from the
<typeStatus id="8860BA99DE13FF9CFF3C1FB7FEE60FC8" box="[255,353,657,683]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="90DB7FB8DE13FF9CFE061FB4FD280FC8" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[453,687,657,683]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="rhopaloparamera" status="sp. nov.">
<emphasis id="65AFD829DE13FF9CFE061FB4FE5E0FC8" box="[453,473,658,683]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFE2B1FB7FD280FC8" box="[488,687,657,683]" italics="true" pageId="21" pageNumber="322">rhopaloparamera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE13FF9CFD751FB7FD740FC8" box="[694,755,657,683]" pageId="21" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
in having a very long antennal flagellum with slender, elongate flagellomeres 1013; a narrower wing with a longer 2
<superScript id="A0AEA973DE13FF9CFCE41F92FCB00FA1" attach="left" box="[807,823,692,706]" fontSize="6" pageId="21" pageNumber="322">nd</superScript>
radial cell; sternite 9 with a much broader, arrow-head shaped posterior extension; a much longer tergite 9 with longer, cylindrical apicolateral processes; longer gonocoxite and gonostylus; much broader bases of gonocoxal apodemes; and a very short, broader quadrate paramere.
</paragraph>
<paragraph id="5764043BDE13FF9CFF041E07FD910EA8" blockId="21.[151,1437,152,971]" pageId="21" pageNumber="322">
Females of
<taxonomicName id="90DB7FB8DE13FF9CFE8C1E04FE420E58" box="[335,453,801,827]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="meloae">
<emphasis id="65AFD829DE13FF9CFE8C1E04FEE40E58" box="[335,355,802,827]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFEB71E07FE420E58" box="[372,453,801,827]" italics="true" pageId="21" pageNumber="322">meloae</emphasis>
</taxonomicName>
and
<taxonomicName id="90DB7FB8DE13FF9CFDC31E04FD420E58" box="[512,709,801,827]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="pailemanensis">
<emphasis id="65AFD829DE13FF9CFDC31E04FD930E58" box="[512,532,802,827]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFDE61E07FD420E58" box="[549,709,801,827]" italics="true" pageId="21" pageNumber="322">pailemanensis</emphasis>
</taxonomicName>
are unknown. Females of
<taxonomicName id="90DB7FB8DE13FF9CFC381E04FB020E58" authorityName="Grogan, Diaz, Spinelli &amp; Ronderos" authorityYear="2016" box="[1019,1157,801,827]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="turnbowi">
<emphasis id="65AFD829DE13FF9CFC381E04FB880E58" box="[1019,1039,802,827]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFBE31E07FB020E58" box="[1056,1157,801,827]" italics="true" pageId="21" pageNumber="322">turnbowi</emphasis>
</taxonomicName>
differ from the
<typeStatus id="8860BA99DE13FF9CFA811E07FA1B0E58" box="[1346,1436,801,827]" pageId="21" pageNumber="322" type="allotype">allotype</typeStatus>
female of
<taxonomicName id="90DB7FB8DE13FF9CFEC81E60FE710E3C" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[267,502,837,863]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="rhopaloparamera" status="sp. nov.">
<emphasis id="65AFD829DE13FF9CFEC81E60FE980E3C" box="[267,287,838,863]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFEEC1E63FE710E3C" box="[303,502,837,863]" italics="true" pageId="21" pageNumber="322">rhopaloparamera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE13FF9CFE3D1E63FDBB0E3C" box="[510,572,837,863]" pageId="21" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
in having a much longer antennal flagellum with slender, elongate flagellomeres 913; wing with dense macrotrichiae and a longer, broader 2
<superScript id="A0AEA973DE13FF9CFCBD1E4EFC090E15" attach="left" box="[894,910,872,886]" fontSize="6" pageId="21" pageNumber="322">nd</superScript>
radial cell; a broader chalice-shaped subgenital plate with a broad U-shaped posterior margin; and a much smaller (diameter
<quantity id="9023A9DEDE13FF9CFC391EABFBC40EC4" box="[1018,1091,909,935]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="21" pageNumber="322" unit="mm" value="42.0">42 µm</quantity>
vs.
<date id="236522FBDE13FF9CFBB11EABFB240EC4" box="[1138,1187,909,935]" pageId="21" pageNumber="322" value="1967-10">67 x</date>
<quantity id="9023A9DEDE13FF9CFB691EABFB730EC4" box="[1194,1268,909,935]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="322" unit="mm" value="60.0">60 µm</quantity>
in
<taxonomicName id="90DB7FB8DE13FF9CFADB1EA8FE820EA8" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="rhopaloparamera">
<emphasis id="65AFD829DE13FF9CFADB1EA8FAAB0EC4" box="[1304,1324,910,935]" italics="true" pageId="21" pageNumber="322">D</emphasis>
.
<emphasis id="65AFD829DE13FF9CFAF81EABFE820EA8" italics="true" pageId="21" pageNumber="322">rhopaloparamera</emphasis>
</taxonomicName>
) spherical spermatheca.
</paragraph>
</subSubSection>
</treatment>
</document>