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<document id="A12910FCDEB66C247882BF6519F1B305" ID-CLB-Dataset="24859" ID-DOI="10.11646/zootaxa.4700.3.1" ID-GBIF-Dataset="1b0654ea-eecb-4242-ad9a-850602f7d185" ID-ISSN="1175-5326" ID-Zenodo-Dep="3548509" ID-ZooBank="B0712990-7A3B-4042-9A4B-5CE96AFAECF8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1574256400520" checkinUser="plazi" docAuthor="Grogan, William L., Díaz, Florentina, Spinelli, Gustavo R. &amp; Ronderos, Maria M." docDate="2019" docId="DF72B52DDE0FFF83FF541B83FE310A64" docLanguage="en" docName="zootaxa.4700.3.1.pdf" docOrigin="Zootaxa 4700 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyhelea recurva Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos 2019, n. sp." docType="treatment" docVersion="10" lastPageNumber="311" masterDocId="234BCD55DE06FF89FFC31D26FF870D63" masterDocTitle="The Biting Midges of the Caribbean island Curaçao (Diptera: Ceratopogonidae) I. Species in the genus Dasyhelea Kieffer" masterLastPageNumber="325" masterPageNumber="301" pageNumber="310" updateTime="1698701345973" updateUser="plazi">
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<mods:title id="CABBE8A8BC4AC9D517FD9A6F171BCCB9">The Biting Midges of the Caribbean island Curaçao (Diptera: Ceratopogonidae) I. Species in the genus Dasyhelea Kieffer</mods:title>
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<mods:namePart id="4C04798ED1D82D26F038EBEFD42BD297">Grogan, William L.</mods:namePart>
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<mods:namePart id="1F85B794F55FDCAF4227AFA8AD661B81">Díaz, Florentina</mods:namePart>
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<mods:namePart id="45A43DF1ED901ED83C41839C132D3416">Spinelli, Gustavo R.</mods:namePart>
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<mods:namePart id="56F1FFD02FAF75CDDE4FC391CAC5B4CE">Ronderos, Maria M.</mods:namePart>
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<mods:title id="46D6E6B85B4B69B1FCAC6B952BB23D56">Zootaxa</mods:title>
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<subSubSection id="1FC157B0DE0FFF80FF541B83FEF30B80" pageId="9" pageNumber="310" type="nomenclature">
<paragraph id="5764043BDE0FFF80FF541B83FE440BDC" blockId="9.[151,451,1701,1763]" box="[151,451,1701,1728]" pageId="9" pageNumber="310">
<heading id="0C2CB357DE0FFF80FF541B83FE440BDC" bold="true" box="[151,451,1701,1728]" fontSize="11" level="1" pageId="9" pageNumber="310" reason="1">
<emphasis id="65AFD829DE0FFF80FF541B83FE440BDC" bold="true" box="[151,451,1701,1728]" pageId="9" pageNumber="310">
<taxonomicName id="90DB7FB8DE0FFF80FF541B83FEFE0BDC" ID-CoL="8BQWM" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[151,377,1701,1728]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="recurva" status="sp. nov.">
<emphasis id="65AFD829DE0FFF80FF541B83FEFE0BDC" bold="true" box="[151,377,1701,1728]" italics="true" pageId="9" pageNumber="310">Dasyhelea recurva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0FFF80FE431B83FE440BDC" box="[384,451,1701,1727]" pageId="9" pageNumber="310" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<paragraph id="5764043BDE0FFF80FF541BEFFEF30B80" blockId="9.[151,451,1701,1763]" box="[151,372,1736,1763]" pageId="9" pageNumber="310">(Figs. 610, 5557)</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0FFF83FF541A36FE250970" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310" type="diagnosis">
<paragraph id="5764043BDE0FFF80FF541A36FB330AF4" blockId="9.[151,1437,1808,2015]" pageId="9" pageNumber="310">
<emphasis id="65AFD829DE0FFF80FF541A36FE8D0A48" bold="true" box="[151,266,1808,1835]" pageId="9" pageNumber="310">Diagnosis</emphasis>
. The only Neotropical species of the mutabilis group with the following combination of characters. Males aedeagus heavily sclerotized with broad, stout basal arms, and slender posterolateral arms with recurved, bifid apices; gonocoxal apodemes and paramere asymmetrical, the right gonocoxal apodeme slender and narrowly fused to paramere, the left gonocoxal apodeme stout and broadly fused to paramere. Female unknown.
</paragraph>
<paragraph id="5764043BDE0FFF83FF041A87FF530E8C" blockId="9.[151,1437,1808,2015]" lastBlockId="10.[151,1437,153,1799]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
<emphasis id="65AFD829DE0FFF80FF041A87FE830AD8" bold="true" box="[199,260,1953,1979]" pageId="9" pageNumber="310">Male</emphasis>
. Head (Fig. 6). Dark brown. Eyes contiguous, the distance of contiguity not discernible in
<typeStatus id="8860BA99DE0FFF80FACA1A87FAE90AD8" box="[1289,1390,1953,1979]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
, the only known specimen. Antennal flagellum dark brown; flagellomeres 29 rhomboidal, 1013 elongate, only 1013 sculptured; flagellomeres 210 with whorl of long sensillae chaetica comprising the moderately dense plume; antennal ratio 0.95; flagellum length
<quantity id="9023A9DEDE0CFF83FDDD1D9AFD020DB4" box="[542,645,188,215]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="311" unit="mm" value="0.45">0.45 mm</quantity>
. Frontal sclerite elliptical, with slender moderately long, pointed, ventral projection. Clypeus with 5 pairs of setae. Palpus (Fig. 7) brown; segment 3 as long as 4+5 combined with 12 capitate sensillae on distal 1/4 of mesal surface; palpal ratio 3.60. Thorax (Fig. 8). Dark brown; scutellum with 6 large setae. Femora, tibiae dark brown, tarsi light brown, tarsomeres 5 infuscated; hind tibial comb with 6 spines; prothoracic tarsal ratio 1.91, mesothoracic tarsal ratio 2.25, metathoracic tarsal ratio 2.21. Wing (Fig. 9) membrane hyaline with moderately dense macrotrichiae; radial cells without lumen, 2
<superScript id="A0AEA973DE0CFF83FC221C56FC760C1D" attach="left" box="[993,1009,368,382]" fontSize="6" pageId="10" pageNumber="311">nd</superScript>
radial cell very short; apices of costa, radius at perpendicular angle with wing margin; fork of CuA
<subScript id="CB5F067EDE0CFF83FC911C80FCDD0CD7" attach="left" box="[850,858,422,436]" fontSize="6" pageId="10" pageNumber="311">1</subScript>
distad to level of apex of 2
<superScript id="A0AEA973DE0CFF83FB5A1CB2FB2E0CC1" attach="left" box="[1177,1193,404,418]" fontSize="6" pageId="10" pageNumber="311">nd</superScript>
radial cell; anal lobe moderately developed; wing length
<quantity id="9023A9DEDE0CFF83FDEB1C9FFD0A0CB0" box="[552,653,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="311" unit="mm" value="0.62">0.62 mm</quantity>
, width
<quantity id="9023A9DEDE0CFF83FD1C1C9FFCC20CB0" box="[735,837,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="311" unit="mm" value="0.2">0.20 mm</quantity>
; costal ratio 0.44. Halter dark brown. Abdomen. Dark brown. Genitalia (Figs. 10, 5557). Tergite 9 tapering gradually distally to rounded apex that extends to or near apices of gonocoxites; apicolateral processes parallel, slender, elongate with 1 medium-sized apical seta; cercus very small, located at base of apicolateral process with 4 setae, the 2 mesal medium-sized, external ones longer. Sternite 9 (Figs. 10, 55) 0.42 length of greatest width; anterior margin nearly straight; posterior portion tapered with broad, rounded apex that extends nearly to basal arch of aedeagus. Gonocoxite stout, 1.7 x longer than broad with 1012 large setae on apicomesal surface and a slender baso-mesal tubercle; gonostylus as long as gonocoxite, distal portion slightly curved, tapering gradually distally with broader apex and pointed tip. Gonocoxal apodemes and paramere (Figs. 10, 56) asymmetrical; distal-most sections of gonocoxal apodemes broad, foot-shaped, apices rounded; right apodeme nearly straight, proximal portion slender, narrowly fused to paramere; left apodeme much broader than right, proximal portion broadly fused to paramere; paramere moderately stout basally, tapering slightly distally, apex apparently obliquely truncate. Aedeagus (Figs. 10, 57) heavily sclerotized, 0.55 length of greatest width; basal arch concave, extending 0.20 of total length of aedeagus; basal arms short, stout, apices tapered, recurved 60°; medio-posterior margin with 2 short, conical, apicolateral teeth; posterolateral arms convergent proximally, with abruptly recurved apical sections and bifid tips; posteromedian projection short, very lightly sclerotized, with blunt apex.
</paragraph>
<paragraph id="5764043BDE0CFF83FF041EDFFE250970" blockId="10.[151,1437,153,1799]" box="[199,418,1017,1043]" pageId="10" pageNumber="311">
<emphasis id="65AFD829DE0CFF83FF041EDFFE990970" bold="true" box="[199,286,1017,1043]" pageId="10" pageNumber="311">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0CFF83FF04193AFE4B0954" box="[199,460,1052,1079]" pageId="10" pageNumber="311" type="distribution">
<paragraph id="5764043BDE0CFF83FF04193AFE4B0954" blockId="10.[151,1437,153,1799]" box="[199,460,1052,1079]" pageId="10" pageNumber="311">
<emphasis id="65AFD829DE0CFF83FF04193AFEDE0954" bold="true" box="[199,345,1052,1079]" pageId="10" pageNumber="311">Distribution</emphasis>
.
<collectingCountry id="2FCC44ABDE0CFF83FEA4193BFE4F0954" box="[359,456,1053,1079]" name="Curaçao" pageId="10" pageNumber="311">Curaçao</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0CFF83FF041967FB5F09A4" pageId="10" pageNumber="311" type="materials_examined">
<paragraph id="5764043BDE0CFF83FF041967FB27091C" blockId="10.[151,1437,153,1799]" pageId="10" pageNumber="311">
<materialsCitation id="E7B30E66DE0CFF83FF041967FB27091C" ID-GBIF-Occurrence="2556250901" collectingDate="2014-11-09" collectingMethod="BL trap" collectionCode="FSCA" collectorName="M. C. Thomas" country="Curaçao" latitude="12.344619" location="Christoffelberg" pageId="10" pageNumber="311" specimenCount="1" typeStatus="holotype">
<emphasis id="65AFD829DE0CFF83FF041967FEF40938" bold="true" box="[199,371,1088,1115]" pageId="10" pageNumber="311">Type material</emphasis>
.
<typeStatus id="8860BA99DE0CFF83FE451967FE690938" box="[390,494,1089,1115]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
male, labeled “
<collectingCountry id="2FCC44ABDE0CFF83FD691967FC8A0938" box="[682,781,1089,1115]" name="Curaçao" pageId="10" pageNumber="311">Curaçao</collectingCountry>
, Christoffel Nat. Pk.,
<location id="520452E0DE0CFF83FBD61967FB450938" LSID="urn:lsid:plazi:treatment:DF72B52DDE0FFF83FF541B83FE310A64:520452E0DE0CFF83FBD61967FB450938" box="[1045,1218,1089,1115]" country="Curaçao" latitude="12.344619" name="Christoffelberg" pageId="10" pageNumber="311">Christoffelberg</location>
, trailhead parking,
<geoCoordinate id="32EF62FCDE0CFF83FF541942FD8D091C" box="[151,522,1124,1151]" pageId="10" pageNumber="311">12°20ʹ40.63ʹʹ N, 69°67ʹ11.38ʹʹ W</geoCoordinate>
,
<date id="236522FBDE0CFF83FDD61943FD09091C" box="[533,654,1125,1151]" pageId="10" pageNumber="311" value="2014-11-09">
<collectingDate id="3321DB13DE0CFF83FDD61943FD09091C" box="[533,654,1125,1151]" pageId="10" pageNumber="311" value="2014-11-09">9-XI-2014</collectingDate>
</date>
,
<collectorName id="FA2E61EDDE0CFF83FD5A1943FCBA091C" box="[665,829,1125,1151]" pageId="10" pageNumber="311">M. C. Thomas</collectorName>
,
<collectingMethod id="8E9A7C2CDE0CFF83FC8B1943FC27091C" box="[840,928,1125,1151]" pageId="10" pageNumber="311">BL trap</collectingMethod>
” (deposited in
<collectionCode id="31CA9CFEDE0CFF83FB8A1943FB13091C" box="[1097,1172,1125,1151]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="10" pageNumber="311" type="Museum">FSCA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5764043BDE0CFF83FF0419AEFB5F09A4" blockId="10.[151,1437,153,1799]" pageId="10" pageNumber="311">
<emphasis id="65AFD829DE0CFF83FF0419AEFD9D09C0" bold="true" box="[199,538,1160,1187]" pageId="10" pageNumber="311">Derivation of specific epithet</emphasis>
. The specific name is a combination of two Latin terms,
<emphasis id="65AFD829DE0CFF83FB4919ACFB2709C0" box="[1162,1184,1162,1187]" italics="true" pageId="10" pageNumber="311">re</emphasis>
(back, again) and
<emphasis id="65AFD829DE0CFF83FAA919ACFA1B09C0" box="[1386,1436,1162,1187]" italics="true" pageId="10" pageNumber="311">curv</emphasis>
(curved), a reference to the distinctive recurved apices of the posterolateral arms of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0CFF83FF0419F6FE310A64" pageId="10" pageNumber="311" type="discussion">
<paragraph id="5764043BDE0CFF83FF0419F6FD050B14" blockId="10.[151,1437,153,1799]" pageId="10" pageNumber="311">
<emphasis id="65AFD829DE0CFF83FF0419F6FEC20988" bold="true" box="[199,325,1232,1259]" pageId="10" pageNumber="311">Discussion</emphasis>
. At least two other primarily tropical New World species of
<taxonomicName id="90DB7FB8DE0CFF83FC3E19F7FBF40988" baseAuthorityName="Dominiak" baseAuthorityYear="2012" box="[1021,1139,1233,1259]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="genus">
<emphasis id="65AFD829DE0CFF83FC3E19F7FBF40988" box="[1021,1139,1233,1259]" italics="true" pageId="10" pageNumber="311">Dasyhelea</emphasis>
</taxonomicName>
have aedeagal posterolateral arms with recurved apices:
<taxonomicName id="90DB7FB8DE0CFF83FDC019D0FC320873" authority="Wirth &amp; Waugh (1976)" authorityName="Wirth &amp; Waugh" authorityYear="1976" box="[515,949,1269,1296]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="borgmeieri">
<emphasis id="65AFD829DE0CFF83FDC019D0FD90086C" box="[515,535,1270,1295]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FDE419D3FD25086C" box="[551,674,1269,1295]" italics="true" pageId="10" pageNumber="311">borgmeieri</emphasis>
Wirth &amp;
<bibRefCitation id="334A79CADE0CFF83FCD219D3FC320873" author="Waugh, W. T. &amp; Wirth, W. W." box="[785,949,1269,1296]" pageId="10" pageNumber="311" pagination="219 - 247" refId="ref20962" refString="Waugh, W. T. &amp; Wirth, W. W. (1976) A revision of the genus Dasyhelea Kieffer of the eastern United States north of Florida (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 69, 219 - 247. https: // doi. org / 10.1093 / aesa / 69.2.219" type="journal article" year="1976">Waugh (1976)</bibRefCitation>
</taxonomicName>
, that was originally described from
<collectingCountry id="2FCC44ABDE0CFF83FA9219D3FA1F086C" box="[1361,1432,1269,1295]" name="Brazil" pageId="10" pageNumber="311">Brazil</collectingCountry>
,
<collectingCountry id="2FCC44ABDE0CFF83FF54183FFF7E0850" box="[151,249,1305,1331]" name="Grenada" pageId="10" pageNumber="311">Grenada</collectingCountry>
,
<collectingCountry id="2FCC44ABDE0CFF83FEC5183FFED80850" box="[262,351,1305,1331]" name="Panama" pageId="10" pageNumber="311">Panama</collectingCountry>
and
<collectingCountry id="2FCC44ABDE0CFF83FE54183FFE710850" box="[407,502,1305,1331]" name="Trinidad and Tobago" pageId="10" pageNumber="311">Trinidad</collectingCountry>
(
<typeStatus id="8860BA99DE0CFF83FDC4183FFDBE0850" box="[519,569,1305,1331]" pageId="10" pageNumber="311">type</typeStatus>
locality), then subsequently recorded from the
<collectingRegion id="951FCAD9DE0CFF83FB95183FFAA40850" box="[1110,1315,1305,1331]" country="Ecuador" name="Galapagos" pageId="10" pageNumber="311">Galápagos Islands</collectingRegion>
(
<collectingCountry id="2FCC44ABDE0CFF83FAF7183FFA120857" box="[1332,1429,1305,1332]" name="Ecuador" pageId="10" pageNumber="311">Ecuador</collectingCountry>
) by
<bibRefCitation id="334A79CADE0CFF83FF79181BFEEF0834" author="Borkent, A." box="[186,360,1341,1367]" pageId="10" pageNumber="311" pagination="97 - 122" refId="ref18270" refString="Borkent, A. (1991) The Ceratopogonidae (Diptera) of the Galapagos Islands, Ecuador with a discussion of their phylogenetic relationships and zoogeographic origins. Entomologica Scandinavia, 22, 97 - 122. https: // doi. org / 10.1163 / 187631291 X 00336" type="journal article" year="1991">Borkent (1991)</bibRefCitation>
, and it was recently discovered in the
<collectingCountry id="2FCC44ABDE0CFF83FCD1181BFCCF0834" box="[786,840,1341,1367]" name="United States of America" pageId="10" pageNumber="311">USA</collectingCountry>
from
<collectingRegion id="951FCAD9DE0CFF83FC4E181BFC590834" box="[909,990,1341,1367]" country="United States of America" name="Florida" pageId="10" pageNumber="311">Florida</collectingRegion>
(WLG; personal observations) and also from
<collectingCountry id="2FCC44ABDE0CFF83FF171847FE9B0818" box="[212,284,1377,1403]" name="Belize" pageId="10" pageNumber="311">Belize</collectingCountry>
(WLG; in FSCA); and,
<taxonomicName id="90DB7FB8DE0CFF83FDE41844FC1D081F" authority="Wirth &amp; Waugh (1976)" authorityName="Wirth &amp; Waugh" authorityYear="1976" box="[551,922,1377,1404]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="cacaoi">
<emphasis id="65AFD829DE0CFF83FDE41844FDBC0818" box="[551,571,1378,1403]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FD8A1844FD130818" box="[585,660,1378,1403]" italics="true" pageId="10" pageNumber="311">cacaoi</emphasis>
Wirth &amp;
<bibRefCitation id="334A79CADE0CFF83FD381847FC1D081F" author="Waugh, W. T. &amp; Wirth, W. W." box="[763,922,1377,1404]" pageId="10" pageNumber="311" pagination="219 - 247" refId="ref20962" refString="Waugh, W. T. &amp; Wirth, W. W. (1976) A revision of the genus Dasyhelea Kieffer of the eastern United States north of Florida (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 69, 219 - 247. https: // doi. org / 10.1093 / aesa / 69.2.219" type="journal article" year="1976">Waugh (1976)</bibRefCitation>
</taxonomicName>
that is known only from
<collectingCountry id="2FCC44ABDE0CFF83FB731847FA8E0818" box="[1200,1289,1377,1403]" name="Panama" pageId="10" pageNumber="311">Panama</collectingCountry>
and
<collectingCountry id="2FCC44ABDE0CFF83FAFE1847FA1B0818" box="[1341,1436,1377,1403]" name="Trinidad and Tobago" pageId="10" pageNumber="311">Trinidad</collectingCountry>
(
<typeStatus id="8860BA99DE0CFF83FF5C18A3FF5608FC" box="[159,209,1413,1439]" pageId="10" pageNumber="311">type</typeStatus>
locality). The males of these two species differ from
<taxonomicName id="90DB7FB8DE0CFF83FCFA18A0FC3408FC" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[825,947,1413,1439]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="recurva" status="sp. nov.">
<emphasis id="65AFD829DE0CFF83FCFA18A0FCCA08FC" box="[825,845,1414,1439]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FC9E18A0FC3408FC" box="[861,947,1414,1439]" italics="true" pageId="10" pageNumber="311">recurva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0CFF83FC7F18A3FC7D08FC" box="[956,1018,1413,1439]" pageId="10" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
in having less massive aedeagi with apically pointed posterolateral arms, sternite 9 that extends just beyond the basal arch (
<taxonomicName id="90DB7FB8DE0CFF83FBBA188CFA9008A0" authorityName="Wirth &amp; Waugh" authorityYear="1976" box="[1145,1303,1449,1475]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="borgmeieri">
<emphasis id="65AFD829DE0CFF83FBBA188CFB0A08A0" box="[1145,1165,1450,1475]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FB5F188FFA9008A0" box="[1180,1303,1449,1475]" italics="true" pageId="10" pageNumber="311">borgmeieri</emphasis>
</taxonomicName>
) or, is very elongate, tapered and extends more than 1/2 the length of the posterolateral arms (
<taxonomicName id="90DB7FB8DE0CFF83FBEA18E8FB110884" authorityName="Wirth &amp; Waugh" authorityYear="1976" box="[1065,1174,1485,1511]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="cacaoi">
<emphasis id="65AFD829DE0CFF83FBEA18E8FBBA0884" box="[1065,1085,1486,1511]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FB8818E8FB110884" box="[1099,1174,1486,1511]" italics="true" pageId="10" pageNumber="311">cacaoi</emphasis>
</taxonomicName>
). Males of
<taxonomicName id="90DB7FB8DE0CFF83FAD618E8FF310B68" authorityName="Wirth &amp; Waugh" authorityYear="1976" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="borgmeieri">
<emphasis id="65AFD829DE0CFF83FAD618E8FAAE0884" box="[1301,1321,1486,1511]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FAF418EBFF310B68" italics="true" pageId="10" pageNumber="311">borgmeieri</emphasis>
</taxonomicName>
further differ from
<taxonomicName id="90DB7FB8DE0CFF83FE5218D4FD8F0B68" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[401,520,1521,1547]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="recurva" status="sp. nov.">
<emphasis id="65AFD829DE0CFF83FE5218D4FE220B68" box="[401,421,1522,1547]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FE7118D4FD8F0B68" box="[434,520,1522,1547]" italics="true" pageId="10" pageNumber="311">recurva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0CFF83FDCC18D7FDCD0B68" box="[527,586,1521,1547]" pageId="10" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
in having symmetrical gonocoxal apodemes and a short slender paramere, a greatly curved gonostylus and tergite 9 with very short apicolateral processes. Males of
<taxonomicName id="90DB7FB8DE0CFF83FBB41B30FB620B4C" authorityName="Wirth &amp; Waugh" authorityYear="1976" box="[1143,1253,1557,1583]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="cacaoi">
<emphasis id="65AFD829DE0CFF83FBB41B30FB0C0B4C" box="[1143,1163,1558,1583]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FB591B30FB620B4C" box="[1178,1253,1558,1583]" italics="true" pageId="10" pageNumber="311">cacaoi</emphasis>
</taxonomicName>
also differ from
<taxonomicName id="90DB7FB8DE0CFF83FF541B1CFE970B30" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[151,272,1593,1619]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="recurva" status="sp. nov.">
<emphasis id="65AFD829DE0CFF83FF541B1CFF2C0B30" box="[151,171,1594,1619]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FF791B1CFE970B30" box="[186,272,1594,1619]" italics="true" pageId="10" pageNumber="311">recurva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0CFF83FED41B1FFED60B30" box="[279,337,1593,1619]" pageId="10" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
in having more elongate gonocoxites, a longer tergite 9 with a convex apical margin and a longer, slender paramere with pointed curved apex.
</paragraph>
<paragraph id="5764043BDE0CFF83FF041BA7FE310A64" blockId="10.[151,1437,153,1799]" pageId="10" pageNumber="311">
In addition, this species is also similar to
<taxonomicName id="90DB7FB8DE0CFF83FD531BA4FCC20BF8" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[656,837,1665,1691]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="curacaoensis" status="sp. nov.">
<emphasis id="65AFD829DE0CFF83FD531BA4FD230BF8" box="[656,676,1666,1691]" italics="true" pageId="10" pageNumber="311">D</emphasis>
.
<emphasis id="65AFD829DE0CFF83FD721BA4FCC20BF8" box="[689,837,1666,1691]" italics="true" pageId="10" pageNumber="311">curacaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0CFF83FC881BA7FC0F0BF8" box="[843,904,1665,1691]" pageId="10" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
, but the latter species differs from
<taxonomicName id="90DB7FB8DE0CFF83FACA1BA4FA070BF8" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[1289,1408,1666,1691]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="recurva">
<emphasis id="65AFD829DE0CFF83FACA1BA4FA070BF8" box="[1289,1408,1666,1691]" italics="true" pageId="10" pageNumber="311">D. recurva</emphasis>
</taxonomicName>
in having a very short antennal flagellum, longer gonocoxites, shorter nearly straight gonostyli, a much broader tergite 9, and a longer paramere with curved, J-shaped apical section, and, aedeagus with straight, tapered posterolateral arms with pointed apices.
</paragraph>
</subSubSection>
</treatment>
</document>