145 lines
17 KiB
XML
145 lines
17 KiB
XML
<document id="3C3049B41FBD8EAB6493E1B392F5CCE3" ID-CLB-Dataset="34375" ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1717" ID-GBIF-Dataset="5b86d7da-7fb8-454f-b6e5-267f0fcc216d" ID-PMC="PMC3260776" ID-Pensoft-Pub="1313-2970-130-473" ID-PubMed="22259294" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-473" ModsDocOrigin="ZooKeys 130" ModsDocTitle="Compression fossil Mymaridae (Hymenoptera) from Kishenehn oil shales, with description of two new genera and review of Tertiary amber genera" checkinTime="1451249845677" checkinUser="pensoft" docAuthor="Huber, John T. & Greenwalt, Dale" docDate="2011" docId="DE714A937953DB7FD508DB28A00C2E34" docLanguage="en" docName="ZooKeys 130: 473-494" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1717" docTitle="Eoeustochus Huber, gen. n." docType="treatment" docVersion="6" lastPageNumber="475" masterDocId="FFBAFFB7FFC4FFE6313EB074B028ED08" masterDocTitle="Compression fossil Mymaridae (Hymenoptera) from Kishenehn oil shales, with description of two new genera and review of Tertiary amber genera" masterLastPageNumber="494" masterPageNumber="473" pageNumber="474" updateTime="1732749067596" updateUser="ExternalLinkService">
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<mods:mods id="AF96B5234E7A6C4B450FFFDF16DA7414" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="10B02CF75C1ECD04E906080D379E6C93">
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<mods:title id="C0924B927047FDB060F8723DD48FA225">Compression fossil Mymaridae (Hymenoptera) from Kishenehn oil shales, with description of two new genera and review of Tertiary amber genera</mods:title>
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<mods:name id="91996EB0691EAA512EE0930C8B547A91" type="personal">
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<mods:namePart id="367F26E0D614FC3DBE97622ECF866054">Huber, John T.</mods:namePart>
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</mods:role>
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<mods:namePart id="358FCC07A5C3A95E75F218848B051115">Greenwalt, Dale</mods:namePart>
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<mods:typeOfResource id="2CE979431DB97542D0557079D1593983">text</mods:typeOfResource>
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<mods:title id="B10A26AFE88BA2D5E4F8E6E818212161">ZooKeys</mods:title>
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<mods:part id="83CA212C81670C6F3717B849A2E2E4A4">
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<mods:date id="4AC899D0D5EF17F6D310BCDEB5D3E293">2011</mods:date>
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<mods:detail id="4A18709BE56D58A2316BEFD23FC93AD9" type="volume">
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<mods:number id="B88DD78C2AE0A5CF2E1896B196325311">130</mods:number>
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<mods:start id="9033CB96022EDD6E026A6B21C134A4B3">473</mods:start>
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<mods:end id="019F492451B2E5CF3ED94AF954A7BEEA">494</mods:end>
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<mods:location id="DEBFDA228F7972BB1811614B33F2C453">
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<mods:url id="F86DAA795EE5DFC4FEFF4C65CF98E0C7">http://dx.doi.org/10.3897/zookeys.130.1717</mods:url>
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<mods:classification id="532188854D0AD2FA1C34BD441BD561D1">journal article</mods:classification>
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<mods:identifier id="F4A297E7D62001B52C1C6AF690AE25A7" type="DOI">http://dx.doi.org/10.3897/zookeys.130.1717</mods:identifier>
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<mods:identifier id="E100D7948B4DA6253EC765A5699E3C29" type="Pensoft-Pub">1313-2970-130-473</mods:identifier>
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<treatment id="DE714A937953DB7FD508DB28A00C2E34" ID-GBIF-Taxon="152031022" LSID="urn:lsid:zoobank.org:act:25AE7CE1-EB1D-4B39-8D61-AA86366A4A35" httpUri="http://treatment.plazi.org/id/DE714A937953DB7FD508DB28A00C2E34" lastPageId="2" lastPageNumber="475" pageId="1" pageNumber="474" scope_family="Mymaridae" scope_order="Hymenoptera">
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<subSubSection id="D46D9E4599F320C3EA4CC26BF1C5408E" pageId="1" pageNumber="474" type="nomenclature">
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<paragraph id="78644E2AF980AC138E4045F20162B2E1" pageId="1" pageNumber="474">
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<taxonomicName id="819A7A69A32DFD88BA42497B4927CADF" LSID="urn:lsid:zoobank.org:act:25AE7CE1-EB1D-4B39-8D61-AA86366A4A35" authority="Huber" genus="Eoeustochus" lsidName="Eoeustochus" pageId="1" pageNumber="474" rank="genus">Eoeustochus Huber</taxonomicName>
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<taxonomicNameLabel id="95CDC2A8D8330998E65EC59FC2451044" pageId="1" pageNumber="474">gen. n.</taxonomicNameLabel>
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Figs 18
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</paragraph>
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</subSubSection>
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<subSubSection id="F584D8790FC9E1B0812813676A5A2BBF" pageId="1" pageNumber="474" type="description">
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<paragraph id="25ED27B51D98F577F128D6CB783F5B69" pageId="1" pageNumber="474">Description.</paragraph>
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<paragraph id="E08C02E12E8A6C96027BDF657553D37B" pageId="1" pageNumber="474">
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Female. Body length 718-1133 (seven specimens in total, only four described and named to species). Head. Normal in shape, wider than long and about
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<normalizedToken id="5B472E597767DC16FF1D998186D43F0A" originalValue="¾">3/4</normalizedToken>
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as high as wide. Face slightly convex in lateral view; vertex flat and slightly sloping anteriorly, forming a moderately sharp angle with occiput; back of head slightly concave. Eye higher than wide, about 2/3 head height; malar space about 1/3 eye height. Antenna. Funicle 6-segmented, with each funicle segment longer than wide; clava 3-segmented with the claval sutures perpendicular to claval length. Wings. Fore wing wide, symmetrical, with evenly rounded apex (shape resembling that of the extant genus
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<taxonomicName id="9D16CC3F34D810129D8ED47A011A08F3" class="Insecta" family="Mymaridae" genus="Eustochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustochus" order="Hymenoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">Eustochus</taxonomicName>
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); marginal fringe shorter than fore wing width. Venation about 0.4
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<normalizedToken id="F9618B2729C2051FF863E297625D4890" originalValue="×">x</normalizedToken>
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fore wing length, with long marginal vein and short but distinct stigmal vein. Hind wing narrow; marginal fringe much longer than wing width. Mesosoma. Shorter than gaster. Pronotum length at most about half length of mesoscutum. Mesoscutum length subequal to scutellum. Scutellum with frenum apparently entire, not divided longitudinally. Metanotum much shorter than scutellum. Metasoma. Constricted at base, probably with short petiole. Gastral terga similar in length. Ovipositor moderately short, probably arising near midpoint of gaster and its apex not or barely exserted beyond gastral apex.
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</paragraph>
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</subSubSection>
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<subSubSection id="94479E0B7D27A2D7ADCDD5A46DD8DF2F" pageId="1" pageNumber="474" type="type species">
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<paragraph id="6BB2C6D907D5BB9A5BCDDE13F3D7AD57" pageId="1" pageNumber="474">Type species.</paragraph>
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<paragraph id="AF8046F81B861D115FE6B761906EB89E" pageId="1" pageNumber="474">
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<taxonomicName id="7F27BBF4C5DDC981724288FBE5B2005A" genus="Eoeustochus" lsidName="Eoeustochus kishenehn" pageId="1" pageNumber="474" rank="species" species="kishenehn">Eoeustochus kishenehn</taxonomicName>
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Huber, sp. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="121267233F674E951D60511BAFDD78EA" pageId="1" pageNumber="474" type="derivation of generic name">
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<paragraph id="F93E3C5267BF069A780DD457A56F1273" pageId="1" pageNumber="474">Derivation of generic name.</paragraph>
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<paragraph id="42D02579CEE64BFF8BCED0C8D6F82872" pageId="1" pageNumber="474">
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The name is from eo-, Greek for early, and
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<taxonomicName id="E492E88B00CDDC326CC181C0E3A2371E" class="Insecta" family="Mymaridae" genus="Eustochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustochus" order="Hymenoptera" pageId="1" pageNumber="474" phylum="Arthropoda" rank="genus">Eustochus</taxonomicName>
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, an extant genus. The gender is masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="12735C56080FE6D147EEC1149766F3A5" lastPageId="2" lastPageNumber="475" pageId="1" pageNumber="474" type="discussion">
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<paragraph id="65F3D97B5CEA8628ED0E5300C7F92437" pageId="1" pageNumber="474">Discussion.</paragraph>
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<paragraph id="FB811479B55F9E0E98C92997856ED4E6" pageId="1" pageNumber="474">
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Two species are described, each from two specimens. Three additional specimens (Kishenehn #30,356, 40,410, and 40,023) belong to
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<taxonomicName id="EAD5CDC66B935E080465ACCDF387C65A" genus="Eoeustochus" lsidName="Eoeustochus" pageId="1" pageNumber="474" rank="genus">Eoeustochus</taxonomicName>
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but are not included in the type series of either species because they are not in as good a condition.
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</paragraph>
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<paragraph id="BF3705056F55E3D1FFE7541060C1335D" pageId="2" pageNumber="475">
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<pageBreakToken id="D4BA9C4BF7E8BB251BAC1E7D700E4A35" pageId="2" pageNumber="475" start="start">The</pageBreakToken>
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relationships of
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<taxonomicName id="BF7498776AF637EC2672B69F6A45F6D6" genus="Eoeustochus" lsidName="Eoeustochus" pageId="2" pageNumber="475" rank="genus">Eoeustochus</taxonomicName>
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are with
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<taxonomicName id="219A6BA32DB9E7B484DC4EC90C48DAE6" class="Insecta" family="Mymaridae" genus="Eustochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustochus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Eustochus</taxonomicName>
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, one extant species of which has a 3-segmented clava in females. Although thirteen extant genera have a definite 3-segmented clava (
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<taxonomicName id="95A2F6A1555D977EA761830166DA9098" class="Insecta" family="Mymaridae" genus="Allanagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allanagrus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Allanagrus</taxonomicName>
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,
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<taxonomicName id="914C2E976B8DAAE323B5CD80A52F9557" class="Insecta" family="Mymaridae" genus="Anneckia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anneckia" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Anneckia</taxonomicName>
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,
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<taxonomicName id="66A493A3BC3FC18C36A55AC7CF2A7944" class="Insecta" family="Mymaridae" genus="Idiocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiocentrus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Idiocentrus</taxonomicName>
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,
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<taxonomicName id="C7F5E59DE1ABE7F8B3644A44DC4F3092" class="Insecta" family="Mymaridae" genus="Krokella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krokella" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Krokella</taxonomicName>
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,
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<taxonomicName id="73B7C3E7837AD0A0A551A891AC6FA3A8" class="Insecta" family="Mymaridae" genus="Nesopatasson" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesopatasson (Nesomymar)" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="subGenus" subGenus="Nesomymar">Nesopatasson/Nesomymar</taxonomicName>
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,
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<taxonomicName id="ED6B25527E28D266292DB042BB04DB21" genus="Neostethynium" lsidName="Neostethynium" pageId="2" pageNumber="475" rank="genus">Neostethynium</taxonomicName>
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,
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<taxonomicName id="6867D518E79CA95F42C41E49A240649E" class="Insecta" family="Mymaridae" genus="Notomymar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notomymar" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Notomymar</taxonomicName>
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,
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<taxonomicName id="49A561EFA5F036EAB413A6EFC2D0CD66" class="Insecta" family="Mymaridae" genus="Paracmotemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracmotemnus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Paracmotemnus</taxonomicName>
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,
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<taxonomicName id="E5722C218D6FBE231AD3D85EA5584FE4" genus="Parastethynium" lsidName="Parastethynium" pageId="2" pageNumber="475" rank="genus">Parastethynium</taxonomicName>
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,
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<taxonomicName id="5E227FCA3ACB59A3D987F7C0EA9C10F2" class="Insecta" family="Mymaridae" genus="Polynemoidea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polynemoidea" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Polynemoidea</taxonomicName>
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,
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<taxonomicName id="C8B1E8201E81A79890F3A5F8DF6289F9" class="Insecta" family="Mymaridae" genus="Pseudocleruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocleruchus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Pseudocleruchus</taxonomicName>
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,
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<taxonomicName id="FC5528CD6E70E92F2F0E58E40907FCE5" class="Insecta" family="Mymaridae" genus="Stethynium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stethynium" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Stethynium</taxonomicName>
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), the closest one is
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<taxonomicName id="B2098A66F2AE1253D864F63B3FC30EF1" class="Insecta" family="Mymaridae" genus="Eustochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustochus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Eustochus</taxonomicName>
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, based on the apparent strong constriction in lateral view between meso- and metasoma (none in
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<taxonomicName id="D90C66EEE28686C030ED1CFE2BADE741" class="Insecta" family="Mymaridae" genus="Allanagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allanagrus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Allanagrus</taxonomicName>
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), transverse sutures between claval segments (sutures oblique in
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<taxonomicName id="454E236F46AE816CC959EA7A39636E54" class="Insecta" family="Mymaridae" genus="Stethynium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stethynium" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Stethynium</taxonomicName>
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), shape of fully developed wings (wings almost absent in
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<taxonomicName id="1CDC03AB95B4633DB5D047E05B748B4A" class="Insecta" family="Mymaridae" genus="Nesopatasson" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesopatasson" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Nesopatasson</taxonomicName>
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/
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<taxonomicName id="A93184DB7F99661C93240D722D767BF9" class="Insecta" family="Mymaridae" genus="Nesomymar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesomymar" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Nesomymar</taxonomicName>
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and
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<taxonomicName id="718B2C861DA871557F7988434DD4A4DD" class="Insecta" family="Mymaridae" genus="Notomymar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notomymar" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Notomymar</taxonomicName>
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, and of different shape in
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<taxonomicName id="C328F86AFB760C51CB602F89034B75D9" class="Insecta" family="Mymaridae" genus="Anneckia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anneckia" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Anneckia</taxonomicName>
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,
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<taxonomicName id="0C7BDC7AAE22AC9E68669685832140CA" genus="Neostethynium" lsidName="Neostethynium" pageId="2" pageNumber="475" rank="genus">Neostethynium</taxonomicName>
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,
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<taxonomicName id="B058FBF3EEC0DC4D5A19E8016D0E1EF1" genus="Parastethynium" lsidName="Parastethynium" pageId="2" pageNumber="475" rank="genus">Parastethynium</taxonomicName>
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and
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<taxonomicName id="36108A7B6CFEF5AAAE7C785E51D64230" class="Insecta" family="Mymaridae" genus="Pseudocleruchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocleruchus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Pseudocleruchus</taxonomicName>
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) venation shorter than half fore wing length (venation longer in
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<taxonomicName id="FB818261CEF7713EC2F63B37599B4F18" class="Insecta" family="Mymaridae" genus="Krokella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krokella" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Krokella</taxonomicName>
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and
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<taxonomicName id="CE1E9CE893400329DAB1299D5EDE9CD4" class="Insecta" family="Mymaridae" genus="Paracmotemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracmotemnus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Paracmotemnus</taxonomicName>
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), and ovipositor not distinctly exserted beyond apex of gaster (ovipositor strongly exserted in
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<taxonomicName id="0FE639ABCA2304D64B68E5DB133F77D9" class="Insecta" family="Mymaridae" genus="Polynemoidea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polynemoidea" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Polynemoidea</taxonomicName>
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) or projecting anteriorly under mesosoma (ovipositor strongly projecting anteriorly in
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<taxonomicName id="4D532210DB18E7FAC5B729E002E0C47C" class="Insecta" family="Mymaridae" genus="Idiocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiocentrus" order="Hymenoptera" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">Idiocentrus</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |