165 lines
18 KiB
XML
165 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.97.115682" ID-Pensoft-Pub="2747-8106-97-53" ID-Pensoft-UUID="4A3318772B185D35B1E7EC9578EF4A47" ID-ZooBank="933EC356F21C45AF9CFA563E64D27953" ModsDocID="2747-8106-97-53" checkinTime="1702462880200" checkinUser="pensoft" docAuthor="Neubauer, Thomas A. & Wesselingh, Frank P." docDate="2023" docId="DE219B80EC72550F9BE03190D68B3CA7" docLanguage="en" docName="Zitteliana 97: 53-88" docOrigin="Zitteliana 97" docPubDate="2023-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.97.115682" docTitle="Ecrobia Stimpson 1865" docType="treatment" docVersion="1" id="4A3318772B185D35B1E7EC9578EF4A47" lastPageNumber="53" masterDocId="4A3318772B185D35B1E7EC9578EF4A47" masterDocTitle="The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms" masterLastPageNumber="88" masterPageNumber="53" pageNumber="53" updateTime="1702462880200" updateUser="pensoft">
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<mods:title id="419567657763ECE5CB930D5A583FB00D">The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms</mods:title>
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<mods:name id="3E080F7F240846AB8C5649764841B788" type="personal">
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<mods:roleTerm id="287B74F21032D9D61D5E664FDB9CC2A9">Author</mods:roleTerm>
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<mods:namePart id="DF4B2CF7A66CFC2ED3F6E8AD2E90A90B">Neubauer, Thomas A.</mods:namePart>
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<mods:nameIdentifier id="3D0DD8BD981CDAFB01B56C35C712CD55" type="ORCID">https://orcid.org/0000-0002-1398-9941</mods:nameIdentifier>
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<mods:affiliation id="81DDAB5E92F301CC367DB5D8879A56A8">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich, Germany & Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:nameIdentifier id="E7463A77B42D18F3D5138518AD6CF2B2" type="email">neubauer@snsb.de</mods:nameIdentifier>
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<mods:namePart id="68A461A94FFE0302D18FE45CE388D060">Wesselingh, Frank P.</mods:namePart>
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<mods:nameIdentifier id="7C43C9A72FA22ADA0AB8AE01D7809264" type="ORCID">https://orcid.org/0000-0003-3655-0701</mods:nameIdentifier>
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<mods:affiliation id="A9461EB8D953F29ECFA128B099B238E1">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands & Department of Earth Sciences, Utrecht University, Princetonlaan 8 a, 3584 CB Utrecht, Netherlands</mods:affiliation>
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<mods:title id="C00CA26F099906CA655AD3BF5F0FBEAE">Zitteliana</mods:title>
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<mods:part id="6DA43C9C53448E9A3A137FE49A6C195F">
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<mods:date id="249D18A9E32A0B1621CA5AAB73C84955">2023</mods:date>
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<mods:number id="14FD9FB2A2AF6A2E04806A05285DB9C9">2023-12-12</mods:number>
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<mods:number id="B4E640A0850D95906EED2B411885C4C6">97</mods:number>
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<mods:start id="38550031826F828291F61859C3C6A52C">53</mods:start>
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<mods:url id="1821A0B473E38798FC580D86F20D2DDB">http://dx.doi.org/10.3897/zitteliana.97.115682</mods:url>
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<mods:classification id="8A6811C03415C4935845B7A3A77FBAE7">journal article</mods:classification>
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<mods:identifier id="F73BCC98C8C99E285C846DE6EAB8D577" type="DOI">http://dx.doi.org/10.3897/zitteliana.97.115682</mods:identifier>
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<treatment id="DE219B80EC72550F9BE03190D68B3CA7" LSID="urn:lsid:plazi:treatment:DE219B80EC72550F9BE03190D68B3CA7" httpUri="http://treatment.plazi.org/id/DE219B80EC72550F9BE03190D68B3CA7" lastPageNumber="53" pageId="0" pageNumber="53">
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<subSubSection id="23BBE23F4F0F15254ECDBD3005BCAB66" pageId="0" pageNumber="53" type="nomenclature">
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<paragraph id="3D64ABB9F3AB648919A82E08AD59092C" pageId="0" pageNumber="53">
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<taxonomicName id="C9E2763999C52426BE46E57D6396BC86" ID-CoL="7NY3T" LSID="DE219B80-EC72-550F-9BE0-3190D68B3CA7" authority="sp." authorityName="Stimpson" authorityYear="1865" class="Gastropoda" family="Hydrobiidae" genus="Ecrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecrobia" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">Ecrobia sp.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="4EEFD5B105B94190D6A0B80B2C2854C8" pageId="0" pageNumber="53" type="description">
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<paragraph id="87EEBB3DED96303B604FAB1C3DDD6785" pageId="0" pageNumber="53">
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<figureCitation id="4E1B81121602E723EA590029B5A40F82" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Hydrobiinae (Hydrobiidae) of the Kolankaya I fauna. A-C, Q. Ecrobia sp. RGM 962612, sample 2. D-F. Ecrobia sp. SNSB-BSPG 2023 XII 6, sample 2. G-I. P. Harzhauseria schizopleura gen. et sp. nov., holotype, RGM 962613, sample 2. J-L. Harzhauseria schizopleura gen. et sp. nov., paratype, RGM 962614, sample 2. M-O, R-T. Hydrobiinae sp. indet., RGM 1365350, sample 1. Scale bars: 1 mm (A-O); 100 μm (P-T)." figureDoi="10.3897/zitteliana.97.115682.figure8" httpUri="https://binary.pensoft.net/fig/950745" pageId="0" pageNumber="53">Fig. 8A-F, Q</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="23A0EA9D5E504B5F3DD9FBA7EEB47A00" pageId="0" pageNumber="53" type="materials_examined">
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<paragraph id="A3BF7A06841B213B5B550A4018DF6B91" pageId="0" pageNumber="53">Material.</paragraph>
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<paragraph id="0B86B2BF370C26559D19DDC6E218FE63" pageId="0" pageNumber="53">
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<specimenCount id="C94F58BFFE19E8E527A754865A8952F1" type="generic">1 specimen</specimenCount>
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(RGM 962612),
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<specimenCount id="B6336E9E8CFB49869BA9B9F8C916B044" type="generic">4 specimens</specimenCount>
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(RGM 1310850),
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<specimenCount id="810EC2AE96C14E23AEEDE2B9F96AA889" type="generic">1 specimen</specimenCount>
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SNSB-BSPG
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<date id="05B61388831801380CD3864D0E373594" value="2023-12-06">2023 XII 6</date>
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); all from sample 2.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6523D1A4B0C73DA3D6D485401218098" pageId="0" pageNumber="53" type="description">
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<paragraph id="DC2AB17C8874E353AAE06FD770B6F8D3" pageId="0" pageNumber="53">Dimensions.</paragraph>
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<paragraph id="C574BF57C19338187D4BC7BD6168DB48" pageId="0" pageNumber="53">
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5.54
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<normalizedToken id="7B5326A21DA04D29D233645F89C6C54B" originalValue="×">x</normalizedToken>
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3.55 mm (RGM 962612; Fig.
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<figureCitation id="1A0129A2B7F138A9F4FBD5A8690E6A87" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Hydrobiinae (Hydrobiidae) of the Kolankaya I fauna. A-C, Q. Ecrobia sp. RGM 962612, sample 2. D-F. Ecrobia sp. SNSB-BSPG 2023 XII 6, sample 2. G-I. P. Harzhauseria schizopleura gen. et sp. nov., holotype, RGM 962613, sample 2. J-L. Harzhauseria schizopleura gen. et sp. nov., paratype, RGM 962614, sample 2. M-O, R-T. Hydrobiinae sp. indet., RGM 1365350, sample 1. Scale bars: 1 mm (A-O); 100 μm (P-T)." figureDoi="10.3897/zitteliana.97.115682.figure8" httpUri="https://binary.pensoft.net/fig/950745" pageId="0" pageNumber="53">8A-C, Q</figureCitation>
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), 4.61
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<normalizedToken id="148F721A5C8527D24C3A4B7B7797DE01" originalValue="×">x</normalizedToken>
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3.01 mm (SNSB-BSPG 2023 XII 6, Fig.
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<figureCitation id="1261BAF27EF26533E2BDB25A8E3BA592" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Hydrobiinae (Hydrobiidae) of the Kolankaya I fauna. A-C, Q. Ecrobia sp. RGM 962612, sample 2. D-F. Ecrobia sp. SNSB-BSPG 2023 XII 6, sample 2. G-I. P. Harzhauseria schizopleura gen. et sp. nov., holotype, RGM 962613, sample 2. J-L. Harzhauseria schizopleura gen. et sp. nov., paratype, RGM 962614, sample 2. M-O, R-T. Hydrobiinae sp. indet., RGM 1365350, sample 1. Scale bars: 1 mm (A-O); 100 μm (P-T)." figureDoi="10.3897/zitteliana.97.115682.figure8" httpUri="https://binary.pensoft.net/fig/950745" pageId="0" pageNumber="53">8D-F</figureCitation>
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).
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</paragraph>
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<caption id="7193EBA15E06C67E2997EDE41AC62EBC" doi="10.3897/zitteliana.97.115682.figure8" httpUri="https://binary.pensoft.net/fig/950745" pageId="0" pageNumber="53" start="Figure 8" startId="F8">
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<paragraph id="5C03C37D54D082FA34BF6C588D2192EC" pageId="0" pageNumber="53">
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<emphasis id="029525B34AAB007E57C9CC4CFA83402C" bold="true" pageId="0" pageNumber="53">Figure 8.</emphasis>
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<taxonomicName id="C636523AB04DD43C4DA572C27123E09B" authorityName="W.Stimpson" authorityYear="1865" lsidName="" pageId="0" pageNumber="53" rank="subFamily" subFamily="Hydrobiinae">Hydrobiinae</taxonomicName>
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(
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<taxonomicName id="8471D89585E8B058F6AC43665AE232EE" class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
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) of the Kolankaya I fauna.
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<emphasis id="9783EEAB2F870805318C2D27EEBF6DDA" bold="true" pageId="0" pageNumber="53">A-C, Q.</emphasis>
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<taxonomicName id="48B5FA055F54DB43E365BBA398864C96" authorityName="Stimpson" authorityYear="1865" class="Gastropoda" family="Hydrobiidae" genus="Ecrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecrobia" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="D57AC36ECEB975CCF9B3B204F14D30B9" italics="true" pageId="0" pageNumber="53">Ecrobia</emphasis>
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</taxonomicName>
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sp. RGM 962612, sample 2.
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<emphasis id="041CB0658DC76B41A3DF07A8303AD12D" bold="true" pageId="0" pageNumber="53">D-F.</emphasis>
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<taxonomicName id="1834B6B78776D0B0EB7E4A995561DA6B" authorityName="Stimpson" authorityYear="1865" class="Gastropoda" family="Hydrobiidae" genus="Ecrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecrobia" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="2E183D2520C800B917017A57CE5F6D3D" italics="true" pageId="0" pageNumber="53">Ecrobia</emphasis>
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</taxonomicName>
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sp. SNSB-BSPG 2023 XII 6, sample 2.
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<emphasis id="CD4D1D4748D7FAE9C3095F5E838D4CEE" bold="true" pageId="0" pageNumber="53">G-I.</emphasis>
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P.
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<taxonomicName id="71CA92368CF51908610E99D79515E05C" authorityName="gen. et" authorityYear="2023" family="Hydrobiidae" genus="Harzhauseria" kingdom="Animalia" lsidName="Harzhauseria schizopleura" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="schizopleura">
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<emphasis id="F039FBB2FF642BA3891029B35310A7AC" italics="true" pageId="0" pageNumber="53">Harzhauseria schizopleura</emphasis>
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</taxonomicName>
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gen. et sp. nov., holotype, RGM 962613, sample 2.
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<emphasis id="0EA6C61071CAA3737A658F9C3DD62F62" bold="true" pageId="0" pageNumber="53">J-L.</emphasis>
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<taxonomicName id="9459EDA3F7A1B4D658412583C79DA0A4" authorityName="gen. et" authorityYear="2023" family="Hydrobiidae" genus="Harzhauseria" kingdom="Animalia" lsidName="Harzhauseria schizopleura" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="schizopleura">
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<emphasis id="0432A7A3E75BCFCC8B041E71A1407BDA" italics="true" pageId="0" pageNumber="53">Harzhauseria schizopleura</emphasis>
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</taxonomicName>
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gen. et sp. nov., paratype, RGM 962614, sample 2.
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<emphasis id="2DE3F589EB82F0B7089D0FEF9A87A117" bold="true" pageId="0" pageNumber="53">M-O, R-T.</emphasis>
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<taxonomicName id="84F9817AE025B579219FF237F3C76B81" authorityName="W.Stimpson" authorityYear="1865" lsidName="" pageId="0" pageNumber="53" rank="subFamily" subFamily="Hydrobiinae">Hydrobiinae</taxonomicName>
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sp. indet., RGM 1365350, sample 1. Scale bars: 1 mm (
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<emphasis id="A0208C06606DA855421B9403450A5C11" bold="true" pageId="0" pageNumber="53">A-O</emphasis>
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); 100
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<normalizedToken id="2BE6CAB3F84790D0E6C71DDB04CCD04F" originalValue="μm">μm</normalizedToken>
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(
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<emphasis id="71E7FAAD3CFEDDE379E2C040569CA145" bold="true" pageId="0" pageNumber="53">P-T</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="6DD8DA9D96EBB4C2D46649603B3E8540" pageId="0" pageNumber="53" type="remarks">
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<paragraph id="C3533C52A30BA023C5F4B92104D258B9" pageId="0" pageNumber="53">Remarks.</paragraph>
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<paragraph id="5523CE293C35A6C401506478BD4898C3" pageId="0" pageNumber="53">
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The shape of the shell and the aperture and the type of growth lines are typical of the genus
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<taxonomicName id="A9BBFBB8E1C8156311129C129E85468D" authorityName="Stimpson" authorityYear="1865" class="Gastropoda" family="Hydrobiidae" genus="Ecrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecrobia" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="61B823E23C839EE5ED1CD4AD0B4DF9DF" italics="true" pageId="0" pageNumber="53">Ecrobia</emphasis>
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</taxonomicName>
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. The protoconch includes ~1.1 whorls and appears to be smooth (the preservation of the apex is, however, rather poor). The P/T boundary is marked by the onset of growth lines. Shape-wise our specimens fall well into the morphological range of extant
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<taxonomicName id="7C188ADAF35358F59156DFCB468EECE4" lsidName="E. grimmi" pageId="0" pageNumber="53" rank="species" species="grimmi">
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<emphasis id="A095F5562E2A779976EAB9CC77D3DD36" italics="true" pageId="0" pageNumber="53">E. grimmi</emphasis>
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</taxonomicName>
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(Clessin in Dybowski, 1887) from the Caspian Sea, but it also resembles stout forms of the widespread European
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<taxonomicName id="80EC85F7FBA8800F2DD1DC6AB37C621D" lsidName="E. ventrosa" pageId="0" pageNumber="53" rank="species" species="ventrosa">
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<emphasis id="F43D658AD89AC7979192359C7FC112D4" italics="true" pageId="0" pageNumber="53">E. ventrosa</emphasis>
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</taxonomicName>
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(Montagu, 1803) and the Black Sea/eastern Mediterranean species
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<taxonomicName id="BA15891EE61A410FDE80098B2C71915D" lsidName="E. maritima" pageId="0" pageNumber="53" rank="species" species="maritima">
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<emphasis id="AFB45979F41DF57FF40D26A150A4AEA8" italics="true" pageId="0" pageNumber="53">E. maritima</emphasis>
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</taxonomicName>
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(Milaschewitsch, 1916). Only, our material is exceptionally large for
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<taxonomicName id="B94AD78C9C0DA4DD9580A74F9A05F5C4" authorityName="Stimpson" authorityYear="1865" class="Gastropoda" family="Hydrobiidae" genus="Ecrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecrobia" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="A4FAB41DB6C5BAFFE976A1604656C665" italics="true" pageId="0" pageNumber="53">Ecrobia</emphasis>
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</taxonomicName>
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and has a slightly thickened aperture, which is untypical as well. It might well represent an undescribed species, but since
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<taxonomicName id="F211034D1845EE88FE76D9DBB9C3CE61" authorityName="Stimpson" authorityYear="1865" class="Gastropoda" family="Hydrobiidae" genus="Ecrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecrobia" order="Littorinimorpha" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
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<emphasis id="FDC563E8AA69CB3356B6065AF214246D" italics="true" pageId="0" pageNumber="53">Ecrobia</emphasis>
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</taxonomicName>
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species are known to be extremely variable (
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<bibRefCitation id="EC7DA6A41221CE1C76AF92AA6909C3E6" author="Kantor, YuI" journalOrPublisher="KMK Scientific Press, Moscow" pageId="0" pageNumber="53" refId="B73" refString="Kantor, YuI, Sysoev, AV, 2006. Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow" title="Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog." year="2006">Kantor and Sysoev 2006</bibRefCitation>
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;
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<bibRefCitation id="173BE2961B68E4D470770C3DC39B1A47" DOI="https://doi.org/10.3897/zookeys.770.25365" author="Neubauer, TA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="53" pagination="43 - 103" refId="B91" refString="Neubauer, TA, van de Velde, S, Yanina, T, Wesselingh, FP, 2018. A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys 770: 43 - 103, DOI: https://doi.org/10.3897/zookeys.770.25365" title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy." url="https://doi.org/10.3897/zookeys.770.25365" volume="770" year="2018">Neubauer et al. 2018</bibRefCitation>
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;
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<bibRefCitation id="AF95334C731DDE37BF803E484AB3809D" DOI="https://doi.org/10.1002/ece3.5602" author="Vandendorpe, J" journalOrPublisher="Ecology and Evolution" pageId="0" pageNumber="53" pagination="10816 - 10827" refId="B159" refString="Vandendorpe, J, van Baak, CGC, Stelbrink, B, Delicado, D, Albrecht, C, Wilke, T, 2019. Historical faunal exchange between the Pontocaspian Basin and North America. Ecology and Evolution 9: 10816 - 10827, DOI: https://doi.org/10.1002/ece3.5602" title="Historical faunal exchange between the Pontocaspian Basin and North America." url="https://doi.org/10.1002/ece3.5602" volume="9" year="2019">Vandendorpe et al. 2019</bibRefCitation>
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) and we have only limited material available for comparison, we refrain from introducing a new species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |