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<document ID-CLB-Dataset="54075" ID-DOI="http://dx.doi.org/10.3897/evolsyst.7.85204" ID-GBIF-Dataset="69e3a7a3-d279-4b63-8070-be3b1c813968" ID-Pensoft-Pub="2535-0730-1-133" ID-Pensoft-UUID="CF6673C9B1765D47B848F9C007505762" ID-ZooBank="24DF15ADF8A04086AD8C60AD39C8A4AA" ModsDocID="2535-0730-7-1-133" checkinTime="1683131477234" checkinUser="pensoft" docAuthor="Lehmann, Ingo &amp; Dalsgaard, Thure" docDate="2023" docId="DAAD0A6CA6C553F1A364736AD61D466D" docLanguage="en" docName="EvolutSyst 7(1): 133-182" docOrigin="Evolutionary Systematics 7 (1)" docPubDate="2023-05-03" docSource="http://dx.doi.org/10.3897/evolsyst.7.85204" docTitle="Morondavania Lehmann &amp; Dalsgaard 2023, gen. nov." docType="treatment" docUuid="1AA460B8-26A9-48F1-9311-C1F24A8E0DCB" docUuidSource="ZooBank" docVersion="5" id="CF6673C9B1765D47B848F9C007505762" lastPageNumber="133" masterDocId="CF6673C9B1765D47B848F9C007505762" masterDocTitle="Revision of Saalmulleria Mabille, 1891 (Lepidoptera, Metarbelidae) from Madagascar with the description of three new genera and fifteen new species" masterLastPageNumber="182" masterPageNumber="133" pageNumber="133" updateTime="1732929487722" updateUser="ExternalLinkService">
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<mods:title id="D1D40A72CC444467E2796F0647EB596F">Revision of Saalmulleria Mabille, 1891 (Lepidoptera, Metarbelidae) from Madagascar with the description of three new genera and fifteen new species</mods:title>
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<mods:namePart id="95E8AF02DB2B1A28CC7EED7C0E04A5E1">Lehmann, Ingo</mods:namePart>
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<mods:namePart id="F8C540910BDD06B1286402EE51BDF613">Dalsgaard, Thure</mods:namePart>
<mods:affiliation id="4FFA7F308536EE1C7281F64E825C9AD4">Leibniz-Institute for the Analysis of Biodiversity Change, ztm, Museum der Natur, Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:date id="A5E5F144003ED4C1B6E560C44DFA3D0A">2023</mods:date>
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<treatment id="DAAD0A6CA6C553F1A364736AD61D466D" ID-GBIF-Taxon="209580328" LSID="urn:lsid:zoobank.org:act:1AA460B8-26A9-48F1-9311-C1F24A8E0DCB" httpUri="http://treatment.plazi.org/id/DAAD0A6CA6C553F1A364736AD61D466D" lastPageNumber="133" pageId="0" pageNumber="133" scope_family="Metarbelidae" scope_order="Lepidoptera">
<subSubSection id="6E70FD4A4AD4B043584832B97AC8113E" pageId="0" pageNumber="133" type="nomenclature">
<paragraph id="376458CA87D73DD7FED1BF38DAB4147A" pageId="0" pageNumber="133">
<taxonomicName id="F4BD7CF52B0EAD7C0A34BA5E2813F05F" LSID="https://zoobank.org/1AA460B8-26A9-48F1-9311-C1F24A8E0DCB" authority="Lehmann &amp; Dalsgaard, 2023" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus" status="gen. nov.">Morondavania</taxonomicName>
<taxonomicNameLabel id="384546D16CB6EBCB8DC00C79BDAEF7B3" pageId="0" pageNumber="133">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9D97F54A4A6D920B2759F6596E547976" pageId="0" pageNumber="133" type="type species">
<paragraph id="5E5516433011EDA2BAB1387F011F849B" pageId="0" pageNumber="133">Type species of genus.</paragraph>
<paragraph id="D4452EFAEE2626650BC446EB2D05B064" pageId="0" pageNumber="133">
<taxonomicName id="CD7DDD39604BC2C969CC139A30707B6F" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania mineti" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="mineti">
<emphasis id="513FB74E8E1D5279FD2440BE47AE907C" italics="true" pageId="0" pageNumber="133">Morondavania mineti</emphasis>
</taxonomicName>
sp. nov. is designated as the type species.
</paragraph>
</subSubSection>
<subSubSection id="1E89F0B004B379C0B3A775B9DC033F20" pageId="0" pageNumber="133" type="autapomorphies diagnosis">
<paragraph id="3FCCDF9DF608B9C5AD87B12663DD468A" pageId="0" pageNumber="133">Autapomorphies diagnosis.</paragraph>
<paragraph id="1AFB685D7E069A52708DD570DBC7D848" pageId="0" pageNumber="133">
The genus is defined by the following combination of characters (
<emphasis id="0A5CC6F43F2D3AC54E30CB3D292DBC16" italics="true" pageId="0" pageNumber="133">cf.</emphasis>
<bibRefCitation id="5E116F1AA85C26460E46F6FB503686C0" author="Lehmann, I" journalOrPublisher="Heterocera Sumatrana" pageId="0" pageNumber="133" refId="B60" refString="Lehmann, I, 2019b. First revision of the family Metarbelidae Strand, 1909 (Lepidoptera, Cossoidea Leach, 1815) and a phylogeny based on adult morphology of 60 genera from the Afrotropical and Oriental Region. Doctoral Dissertation, 1-398, Rheinische Friedrich-Wilhelms-Universitaet Bonn. Universitaets- und Landesbibliothek Bonn, published 19th August 2019. [URN: urn:nbn:de:hbz:5n-55423]" title="First revision of the family Metarbelidae Strand, 1909 (Lepidoptera, Cossoidea Leach, 1815) and a phylogeny based on adult morphology of 60 genera from the Afrotropical and Oriental Region. Doctoral Dissertation, 1 - 398, Rheinische Friedrich-Wilhelms-Universitaet Bonn. Universitaets- und Landesbibliothek Bonn, published 19 th August 2019. [URN: urn: nbn: de: hbz: 5 n- 55423]" year="2019 b">Lehmann 2019b</bibRefCitation>
, 102-105): Antennae bipectinate with very long branches, up to 7.0
<normalizedToken id="14DD07DE9474167CA7A35E394D0B67C5" originalValue="×">x</normalizedToken>
longer than width of shaft, suddenly the branches become much shorter at
<emphasis id="762F3D5BE35C4430DC69EA19E068C4F7" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
half of antennae and are up to 3.0
<normalizedToken id="D23E782C3C9AD13FBC1D2CD86B18429E" originalValue="×">x</normalizedToken>
longer than width of shaft. Additionally, triangular-shaped forewings and triangular-shaped hindwings with straight termen, and hence, the termen of hindwing is not bent inwards as in
<taxonomicName id="520810BFEE827FDD5CE72AD090D76A88" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania dubiefi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="dubiefi">
<emphasis id="026F2E92DBBF6AA6787643EAF0BA8E27" italics="true" pageId="0" pageNumber="133">Morondavania dubiefi</emphasis>
</taxonomicName>
(
<bibRefCitation id="7CB3B7D9C1BDF7A89A7686711A6AA554" author="Viette, P" journalOrPublisher="Nouvelle Revue d'Entomologie, Toulouse, IV," pageId="0" pageNumber="133" pagination="211 - 213" refId="B100" refString="Viette, P, 1974. Deux Nouvelles Especes de Cossoidea de Madagascar (Lepidoptera). Nouvelle Revue d'Entomologie, Toulouse, IV, 3: 211 - 213" title="Deux Nouvelles Especes de Cossoidea de Madagascar (Lepidoptera)." volume="3" year="1974">Viette 1974</bibRefCitation>
;
<emphasis id="D1FBEFA0F696574C01998089B5328EA6" italics="true" pageId="0" pageNumber="133">
cf.
<taxonomicName id="1D2D9E926A5B46BCF9421918F1519C74" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania Saalmulleria" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="Saalmulleria">Morondavania Saalmulleria</taxonomicName>
</emphasis>
below). Wings of species of
<taxonomicName id="AC9E844B64305334BBD78FE70B72CDC1" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="0E1C02A338008D004152AD720D74694E" italics="true" pageId="0" pageNumber="133">Morondavania</emphasis>
</taxonomicName>
are largely without a transparent appearance in male (in contrast to
<taxonomicName id="CBFCEA9FA78805C5D0C5D72430CC41FB" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania dubiefi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="dubiefi">
<emphasis id="E6AFE83AB6BCB2AF27D4ACDCA9131646" italics="true" pageId="0" pageNumber="133">Morondavania dubiefi</emphasis>
</taxonomicName>
where forewings and hindwings are largely transparent); a small thorn-like lower gnathal arm has 15-20% the size of valva, and is connected to the base of uncus by a long and broad weakly sclerotized band that has 40-50% the width of the dorsal length of the thorn-like structure; the phallus is at present the largest among
<taxonomicName id="688811045055B7B61E2D4E7A6E383189" authorityName="Strand" authorityYear="1909" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
and with a unique shape, namely banana-like and strongly bent on whole length, with 50-60% as broad as basal width of valva on
<emphasis id="7AE5FF19B5004164B8F7DF9DB12C7CBD" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
80% of its length, very narrow distally, and 30-40% longer than dorsal edge of valva.
</paragraph>
</subSubSection>
<subSubSection id="FC3F4BCF0DE6327C0B07C27B9FBE8BC0" pageId="0" pageNumber="133" type="synapomorphies differential diagnosis shared with shimbania gen. nov. from the african mainland">
<paragraph id="5DBA4132674A538201111BCC82968A6B" pageId="0" pageNumber="133">
Synapomorphies differential diagnosis shared with
<taxonomicName id="B1BDC107996F47C6BB65597C3EB6CF2A" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" genus="Shimbania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shimbania" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="7BDC3E2C3034C5B4A901300C833F41E9" italics="true" pageId="0" pageNumber="133">Shimbania</emphasis>
</taxonomicName>
gen. nov. from the African mainland
</paragraph>
<paragraph id="7E520BE254EA2D051B50030640755F90" pageId="0" pageNumber="133">
(
<emphasis id="D141AA63161F60D7268ECDCF60F7AEF3" italics="true" pageId="0" pageNumber="133">
cf.
<taxonomicName id="22C9D924BA522E2F7279EDC09B466305" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" genus="Shimbania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shimbania" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">Shimbania</taxonomicName>
</emphasis>
gen. nov. above).
</paragraph>
</subSubSection>
<subSubSection id="4C7C7DDE2AB6FE8ACFD2DB5E52D12B11" pageId="0" pageNumber="133" type="description">
<paragraph id="53F76C554018B5924A60C828C5A1496D" pageId="0" pageNumber="133">
Diagnostic characters in males of
<taxonomicName id="98699AC64DDEE32A4EA8A072F1727937" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="F2D4BA7DC380001A9EBE62AE0959F41E" italics="true" pageId="0" pageNumber="133">Morondavania</emphasis>
</taxonomicName>
gen. nov.
</paragraph>
<paragraph id="501E622DFB4C9ED05B94F028E498FB1F" pageId="0" pageNumber="133">
In forewing, the basal point of the fork of R 1+R 2 is much closer to the anterior angle of median cell than the basal point of the fork of R 3+R 4. This is a potential synapomorphy with
<emphasis id="5E9E069F42906E8B90DB32D0735D9FDD" italics="true" pageId="0" pageNumber="133">dubiefi</emphasis>
(
<bibRefCitation id="CC9BB9B0D85DF8675EEE7153E1EFA6E7" author="Viette, P" journalOrPublisher="Nouvelle Revue d'Entomologie, Toulouse, IV," pageId="0" pageNumber="133" pagination="211 - 213" refId="B100" refString="Viette, P, 1974. Deux Nouvelles Especes de Cossoidea de Madagascar (Lepidoptera). Nouvelle Revue d'Entomologie, Toulouse, IV, 3: 211 - 213" title="Deux Nouvelles Especes de Cossoidea de Madagascar (Lepidoptera)." volume="3" year="1974">Viette 1974</bibRefCitation>
,
<emphasis id="87B51C09E39075C1073C2BEF58E8C0CC" italics="true" pageId="0" pageNumber="133">cf.</emphasis>
note to this species in
<taxonomicName id="A5FD37126BCA171764D4141A4DC052E3" authorityName="Mabille" authorityYear="1891" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="01CFEA3D1E3F7C1217F9E3892D1A9FF0" italics="true" pageId="0" pageNumber="133">Saalmulleria</emphasis>
</taxonomicName>
). This forewing venation is in contrast to species of
<taxonomicName id="1EF9A4346A89B63BBA186782D51CCD4B" authorityName="Mabille" authorityYear="1891" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="7B7ADCEA66ABD19F613D9D1318457CF6" italics="true" pageId="0" pageNumber="133">Saalmulleria</emphasis>
</taxonomicName>
and
<taxonomicName id="9281EB6D7DEF59EE2C32C95F8988C8B1" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Eberhardfischeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eberhardfischeria" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="BAEAA084E178E5832D9A709873A05EAD" italics="true" pageId="0" pageNumber="133">Eberhardfischeria</emphasis>
</taxonomicName>
gen. nov. (
<emphasis id="6816BEDC93FE70E97DC4C882BAD46A81" italics="true" pageId="0" pageNumber="133">cf.</emphasis>
below).
</paragraph>
<paragraph id="2754BFD005AD7331CC7E0F58FF0795D0" pageId="0" pageNumber="133">
Cilia of forewing and hindwing extremely short with
<emphasis id="0A1E4655FB9262CB55DA5B221FC23BD1" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
0.2 mm length and among the shortest in Metarbelidae from the Afrotropical and Oriental Region with a similar wingspan.
</paragraph>
<paragraph id="BEFE7DD42DECEB75CC34CFC374FACE5A" pageId="0" pageNumber="133">
The discocellular cell on the hindwing is similar in shape like a fish-tail, with the upper and lower tip in opposite position but with both tips not strongly pointed (in contrast
<emphasis id="D41EFC7B7780B18F06E41668090C258B" italics="true" pageId="0" pageNumber="133">cf.</emphasis>
species of
<taxonomicName id="2293DCEFADABABE95D956E4DF126C99A" authorityName="Mabille" authorityYear="1891" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="EB128AA710B58F04EEEC5D07439E98C1" italics="true" pageId="0" pageNumber="133">Saalmulleria</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E3A2A0EF7F20741F6FB80C3999D7BB15" pageId="0" pageNumber="133" type="description">
<paragraph id="084CC46179A4011D6629C4C5F8824C4C" pageId="0" pageNumber="133">Description.</paragraph>
<paragraph id="FAB29F06C7A2B21A0829660B059860E9" pageId="0" pageNumber="133">
<emphasis id="031A519C2EB58A309FA8301A845984BA" bold="true" italics="true" pageId="0" pageNumber="133">Head</emphasis>
(Figs
<figureCitation id="04DB9919269BB568AE98F1E6BE958303" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. a. Morondavania mineti sp. nov., holotype, male, Madagascar, Western Region, north of Morondava, Marofandilia Forest; b. M. mineti sp. nov., paratype, male, Madagascar, Western Region, north of Morondava, western part of Marofandilia Forest; c. Eberhardfischeria husemanni sp. nov., holotype, female, Madagascar, Western Region, Diego Suarez (today Antsira ṅana) or areas nearby; d. E. husemanni sp. nov., paratype, female, Madagascar, Western Region, Diego Suarez (today Antsira ṅana) or areas nearby; e. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female, Madagascar, Sambirano Region, Nosy Be Island, Lokobe; f. S. analameranaensis sp. nov., holotype, female, Madagascar, Western Region, Analamerana Forest, ca. 10 km to 40 km west of the Indian Ocean coastline; g. S. ampandrandavaensis sp. nov., holotype, female, Madagascar, Central Region, Ampandrandava, ca. 50 km northeast of Bekily." httpUri="https://binary.pensoft.net/fig/845634" pageId="0" pageNumber="133">4a, b</figureCitation>
,
<figureCitation id="E47D11E2ECA48DD81C84975C5640E961" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing venation: a. Eberhardfischeria husemanni sp. nov., paratype, female; b. Morondavania mineti sp. nov., holotype, male; c. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female." httpUri="https://binary.pensoft.net/fig/845635" pageId="0" pageNumber="133">5b</figureCitation>
,
<figureCitation id="8D0B5C52792477BF0BBEE99731B98821" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female postabdominal structures and male genitalia in a not pressed condition: a. Eberhardfischeria husemanni sp. nov., paratype, female; b. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female; c. Shimbania kerstinhempae sp. nov., holotype, female; d. Morondavania mineti sp. nov., paratype, male, with aedeagus below; e. S. wichgrafi (Gruenberg, 1910), &quot; Type &quot;, male, with aedeagus below (all drawings by I. L.)." httpUri="https://binary.pensoft.net/fig/845637" pageId="0" pageNumber="133">7d</figureCitation>
,
<figureCitation id="25135299BFCA9AE62F29B06380A12B0D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Morondavania mineti sp. nov., holotype, male, with unique banana-like large phallus (a), Madagascar, Western Region, north of Morondava, Marofandilia Forest; B. M. mineti sp. nov., paratype, male, Madagascar, Western Region, north of Morondava, western part of Marofandilia Forest, with ventrally connected gnathal arms (b) and below, in horizontal position, with a broad, well sclerotized transtilla (c) that represents a synapomorphy shared with species of Shimbania gen. nov. The transtilla is rarely present among Metarbelidae and if so, it is narrow and often only thinly sclerotized." httpUri="https://binary.pensoft.net/fig/845641" pageId="0" pageNumber="133">11A, B</figureCitation>
,
<figureCitation id="EDAC1A68A40127EC87401E5058D8A41F" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus (phallus) of a. Shimbania tanaensis sp. nov., paratype, male, Kenya, Tana River County, Mchelelo Camp close to the Tana River; b. Morondavania mineti sp. nov., paratype, male, Madagascar, Western Region, north of Morondava, western part of Marofandilia Forest. The size of the aedeagus of the species of these sister-genera is among the largest in Metarbelidae." httpUri="https://binary.pensoft.net/fig/845642" pageId="0" pageNumber="133">12b</figureCitation>
,
<figureCitation id="B2CAEA3E5B13DED2A97E48ABFE1E1D93" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. SEM pictures of diagnostic head structures of Metarbelidae from Madagascar: A. The lower fronto-clypeus of Morondavania mineti sp. nov., holotype, male, is narrow and without a pair of pits as well as without a pair of projections; B, D. The lower fronto-clypeus of Eberhardfischeria husemanni sp. nov., paratype, female, is broad with a strongly sclerotized plate-like structure (a) with a well-defined dorsal ridge (b) and a pair of rudimentary pits (c); C, E, F. The very broad and smooth lower fronto-clypeus of Saalmulleria ampandrandavaensis sp. nov., holotype, female, is without any plate-like structure, but with a pair of well-developed pits (d), a variable character in species of Saalmulleria that is sometimes entirely absent. A pair of pits on the lower fronto-clypeus is a homoplasy among Metarbelidae in both sexes (cf. Lehmann 2019 b) and it is in species of various genera a variable character (SEM pictures taken by Karin Ulmen, ZFMK, Bonn 2013)." httpUri="https://binary.pensoft.net/fig/845646" pageId="0" pageNumber="133">16A</figureCitation>
): Rough-scaled; medium long hair-like scales of dark chestnut and brownish-olive with an olive glint on fronto-clypeus; eyes black; a pair of pits absent on lower fronto-clypeus, a pair of conical projections absent, slits or small oval holes behind labial palpi absent; labial palpi brownish-olive, short, less than half of eye- diameter, narrow, consisting of two segments; 2nd segment longest, elongated oval,
<emphasis id="3C3F2E931B912EB8761B86400B4FF1A3" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
6.0
<normalizedToken id="8907AC9066D9D505707C83F3ED76B2BA" originalValue="×">x</normalizedToken>
longer than 1st (basal) segment that is very short, apical palpomere absent. Antennae bipectinate, basal half of antennae with narrow and very long branches up 7.0
<normalizedToken id="399291824F748EC1F6D4ADB4D0588512" originalValue="×">x</normalizedToken>
longer than width of shaft, suddenly the branches become much shorter at
<emphasis id="2C56770796B6F299552726BCD3043E67" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
half of antennae and are up to 3.0
<normalizedToken id="EA838646C3D254D2718B25E9498E9253" originalValue="×">x</normalizedToken>
longer than width of shaft, branches are widely separated at base with 2.0
<normalizedToken id="656C36ADE855B5FE28E1B8CE06FB0AD6" originalValue="×">x</normalizedToken>
width of branch, dorsal and lateral sides of branches not scaled, but with many setae in pairs ventrally and laterally, dorsal and lateral sides of flagellum scaled brownish-olive.
</paragraph>
<paragraph id="B320A11FDCADBE10724895D175B30F72" pageId="0" pageNumber="133">
<emphasis id="7FE72B46B8FB4472549C866E1B9BC228" bold="true" italics="true" pageId="0" pageNumber="133">Thorax</emphasis>
: Densely covered with hair-like scales of brownish-olive on patagia, often these scales have a light grey or pale olive tip, scales on patagia form no collar ring, scales on tegulae long hair-like, dark chestnut with a light lilac glint; scale crest on metathorax dark chestnut. Fore and mid legs brownish-olive with long dense hair-like scales with a light lilac glint. Epiphyses present, long, up to 2.1 mm, broad and flat. Hind legs brownish-olive, on lower part of tarsus without any darker patch, with two pairs of narrow tibial spurs, lower pair longer, up to 1.3 mm long, upper pair up to 1.0 mm long, all spurs with thorn-like tip. Wingspan is between 47.0 mm up to 49.5 mm. Forewing narrow triangular with a rounded apex, a strongly S-shaped dorsum with extremely short scales, upperside Dresden brown and brownish-olive towards termen with a light golden glint, scale pattern is weak, usually with very narrow lines of sepia from costa towards dorsum, between costa and CuA1 a weak sepia band (sometimes absent), at base of M2 and M3 a small sepia patch, CuA2 and other veins not distinctly marked, termen without lunules, a dark chestnut patch is present below base of 1A+2A, sometimes faded, sometimes up to 40% length of 1A+2A. Hindwing triangular, termen not bent inwards, largely with extremely short scales of Dresden brown with brownish-olive towards termen, some with a light lilac glint, sometimes with a patch at centre that has a slightly vitreous appearance but is still covered with short light brown scales with a light golden glint and is edged towards discal cell by a small sepia spot. Underside with extremely short scales of Dresden brown. Cilia extremely short with
<emphasis id="2F020EAA86694448B71E1E3443E5AC6C" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
0.2 mm length, brownish-olive with a glint. Forewing venation (Fig.
<figureCitation id="5E719403D31D8CEB9E81EE7B8950BE4C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing venation: a. Eberhardfischeria husemanni sp. nov., paratype, female; b. Morondavania mineti sp. nov., holotype, male; c. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female." httpUri="https://binary.pensoft.net/fig/845635" pageId="0" pageNumber="133">5b</figureCitation>
) with 1A+2A only slightly forked at base, or fork absent; CuP absent, represented by a fold on 2/3 of its original length; CuA2 originating from near hind margin of posterior cell; CuA1, M3 and M2 separate and originating from apical angle of posterior cell; M1 originating from distal margin of median cell and near its anterior angle; areole absent; R1+R2 originating from a medium long well visible stalk (the stalk has the length of 25-30% of R3) and initiating from anterior angle of median cell; R3+R4+R5 are very long stalked and originating from anterior angle of median cell; Sc more or less parallel to R1. Hindwing venation with 3A present, 1A+2A present, with or without a small fork at base, CuP represented by a not sclerotized fold on 2/3 of its original length; CuA2 originating from near hind margin of posterior cell; CuA1, M3 and M2 originating from apical angle of posterior cell, separated; M1 and Rs originating from anterior cell, broadly separated, with M1 near ventral end of distal margin of anterior cell; in holotype with a short bar from near base of Rs to Sc+R1 (Fig.
<figureCitation id="6D364A8D0181E36A31CF4CBBE319BA09" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing venation: a. Eberhardfischeria husemanni sp. nov., paratype, female; b. Morondavania mineti sp. nov., holotype, male; c. Saalmulleria stumpffi (Saalmueller, 1884), &quot; Type &quot;, female." httpUri="https://binary.pensoft.net/fig/845635" pageId="0" pageNumber="133">5b</figureCitation>
), a vein in discocellular cell on both fore- and hindwing is weak or absent. The discocellular cell on the hindwing is similar in shape like a fish-tail, but with the upper and lower tip not strongly pointed. Fringe scales extremely short, 0.2 mm, brownish-olive with a glint. Retinaculum and frenulum absent.
</paragraph>
<caption id="9AB56E5BC2F19C6B8EEF2BDDB3E9A2C8" httpUri="https://binary.pensoft.net/fig/845634" pageId="0" pageNumber="133" start="Figure 4" startId="F4">
<paragraph id="A59F67744EDB4F2FB55B91B764256E5A" pageId="0" pageNumber="133">
<emphasis id="A00F1384DBA80C87B9DE75FE38E6A167" bold="true" pageId="0" pageNumber="133">Figure 4.</emphasis>
<emphasis id="DBD5D90A1357783CA43A85E762867790" bold="true" pageId="0" pageNumber="133">a.</emphasis>
<taxonomicName id="CE75B4AD2F7FC571E16DEA33296C2207" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania mineti" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="mineti">
<emphasis id="D887B923C52F46D6D3092A7B1AE9BFBE" italics="true" pageId="0" pageNumber="133">Morondavania mineti</emphasis>
</taxonomicName>
sp. nov., holotype, male, Madagascar, Western Region, north of Morondava, Marofandilia Forest;
<emphasis id="10EF69EFF4D704CE1799DF0B8591C96D" bold="true" pageId="0" pageNumber="133">b.</emphasis>
<taxonomicName id="B16723D7F1C89BBD3FAEE17E7119FAD6" class="Dicotyledoneae" family="Leguminosae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. mineti" order="Fabales" pageId="0" pageNumber="133" phylum="Angiospermae" rank="species" species="mineti">
<emphasis id="F8D6EA70CE5F5DC9DC9C67320D932A43" italics="true" pageId="0" pageNumber="133">M. mineti</emphasis>
</taxonomicName>
sp. nov., paratype, male, Madagascar, Western Region, north of Morondava, western part of Marofandilia Forest;
<emphasis id="AF19E82811E2BAEB7EF8A7775A5A84C8" bold="true" pageId="0" pageNumber="133">c.</emphasis>
<taxonomicName id="97E6CCAE543B2D7969CE2E03E1F8A52E" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Eberhardfischeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eberhardfischeria husemanni" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="husemanni">
<emphasis id="059425715B02CDD8A664B518ED5ACAE8" italics="true" pageId="0" pageNumber="133">Eberhardfischeria husemanni</emphasis>
</taxonomicName>
sp. nov., holotype, female, Madagascar, Western Region,
<normalizedToken id="42E2761A41DB223525D24D67C74DFE26" originalValue="Diégo">Diego</normalizedToken>
Suarez (today Antsiraṅana) or areas nearby;
<emphasis id="84BA2FAD8903D74AF8C7E47ABA7D698E" bold="true" pageId="0" pageNumber="133">d.</emphasis>
<taxonomicName id="2F2221132890C429A0674170452CC997" class="Insecta" kingdom="Animalia" lsidName="E. husemanni" order="Hymenoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="husemanni">
<emphasis id="07941788B6CD33FFF2AB5DA5406BF5E0" italics="true" pageId="0" pageNumber="133">E. husemanni</emphasis>
</taxonomicName>
sp. nov., paratype, female, Madagascar, Western Region,
<normalizedToken id="476CBE93681AEF66A7D72F9E933BBAE6" originalValue="Diégo">Diego</normalizedToken>
Suarez (today Antsiraṅana) or areas nearby;
<emphasis id="6E2CB95F10F6DB08D4BC96587EAB95FF" bold="true" pageId="0" pageNumber="133">e.</emphasis>
<taxonomicName id="93D590B8FFE98B302D20AA4A188BE025" baseAuthorityName="Saalmueller" baseAuthorityYear="1884" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">
<emphasis id="EE3DD4C1091C3B5CB560762488BF0EE0" italics="true" pageId="0" pageNumber="133">Saalmulleria stumpffi</emphasis>
</taxonomicName>
(
<normalizedToken id="1CF0D3CA7A49FBF759571FE195E59764" originalValue="Saalmüller">Saalmueller</normalizedToken>
, 1884),
<normalizedToken id="867B4F556BD017971CB8C705132ACE1E" originalValue="„Type“">&quot;Type&quot;</normalizedToken>
, female, Madagascar, Sambirano Region, Nosy Be Island, Lokobe;
<emphasis id="F38140690E8037069CC4251D6CD369B2" bold="true" pageId="0" pageNumber="133">f.</emphasis>
<taxonomicName id="1F035B1383CA6C65E6FB0BA494D0285C" class="Insecta" kingdom="Animalia" lsidName="S. analameranaensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="analameranaensis">
<emphasis id="7587D51E570B81587C1CEE86212A033C" italics="true" pageId="0" pageNumber="133">S. analameranaensis</emphasis>
</taxonomicName>
sp. nov., holotype, female, Madagascar, Western Region, Analamerana Forest, ca. 10 km to 40 km west of the Indian Ocean coastline;
<emphasis id="114A8540B612B99A75872AD0238FB17B" bold="true" pageId="0" pageNumber="133">g.</emphasis>
<taxonomicName id="74657AD08C1140FC6B13D750B2842729" class="Insecta" kingdom="Animalia" lsidName="S. ampandrandavaensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="ampandrandavaensis">
<emphasis id="2831ADD172D70C90C016F6844F4D48DE" italics="true" pageId="0" pageNumber="133">S. ampandrandavaensis</emphasis>
</taxonomicName>
sp. nov., holotype, female, Madagascar, Central Region, Ampandrandava, ca. 50 km northeast of Bekily.
</paragraph>
</caption>
<caption id="E3E9F299D5CAAAF1E95B7665D2423CBD" httpUri="https://binary.pensoft.net/fig/845635" pageId="0" pageNumber="133" start="Figure 5" startId="F5">
<paragraph id="BCE34E4B3F2C4CE8B4AF88201F670C94" pageId="0" pageNumber="133">
<emphasis id="757EAB338E83F97DA286201FC8D977D5" bold="true" pageId="0" pageNumber="133">Figure 5.</emphasis>
Wing venation:
<emphasis id="A82E51B48808A6D30D74F993BF43B648" bold="true" pageId="0" pageNumber="133">a.</emphasis>
<taxonomicName id="EEFE134C93DEC357292E91D65EEF5EB5" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Eberhardfischeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eberhardfischeria husemanni" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="husemanni">
<emphasis id="A8D5CEA34C4A33FE6034C4A3BF96279E" italics="true" pageId="0" pageNumber="133">Eberhardfischeria husemanni</emphasis>
</taxonomicName>
sp. nov., paratype, female;
<emphasis id="6F30E5A411B2F989FAD1C4B919E50896" bold="true" pageId="0" pageNumber="133">b.</emphasis>
<taxonomicName id="61651610B2E1E6EAD83B8EAF7AD3CDE7" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania mineti" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="mineti">
<emphasis id="FB7F4191836B9A3F74C16A426926BBE1" italics="true" pageId="0" pageNumber="133">Morondavania mineti</emphasis>
</taxonomicName>
sp. nov., holotype, male;
<emphasis id="349A6F2E0071AC02AC9103B6F3EAB676" bold="true" pageId="0" pageNumber="133">c.</emphasis>
<taxonomicName id="B734080B2F18BF80DFC9AE533F4B3E50" baseAuthorityName="Saalmueller" baseAuthorityYear="1884" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">
<emphasis id="16069A0258DD54ED5CB44AD827F50A70" italics="true" pageId="0" pageNumber="133">Saalmulleria stumpffi</emphasis>
</taxonomicName>
(
<normalizedToken id="469088A582E0EA8FCA506E8B42F75D02" originalValue="Saalmüller">Saalmueller</normalizedToken>
, 1884),
<normalizedToken id="124A605BEF334AC53DD506AFE8F6413F" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
, female.
</paragraph>
</caption>
<caption id="B960AB0E06B398AD833CFDB280960A1B" httpUri="https://binary.pensoft.net/fig/845636" pageId="0" pageNumber="133" start="Figure 6" startId="F6">
<paragraph id="C02AA8C769F4E7BF8886522350E75EDE" pageId="0" pageNumber="133">
<emphasis id="78F1C1D8D38D0B3AC59F1B9BF495793B" bold="true" pageId="0" pageNumber="133">Figure 6.</emphasis>
Wing venation:
<emphasis id="5EB651E2330E812B5DCA0DDD877E67A8" bold="true" pageId="0" pageNumber="133">a.</emphasis>
<taxonomicName id="E11A376FCEC2E08E4E0CCD6F8B3CC4BE" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" genus="Shimbania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shimbania kerstinhempae" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="kerstinhempae">
<emphasis id="4BD4802915F955CEA0C4155D6C7815CF" italics="true" pageId="0" pageNumber="133">Shimbania kerstinhempae</emphasis>
</taxonomicName>
sp. nov., holotype, female;
<emphasis id="580F18BFCC7A8DD80365CD45A7DF6157" bold="true" pageId="0" pageNumber="133">b.</emphasis>
<taxonomicName id="229662A06761B669C3D971DB966BA721" class="Insecta" kingdom="Animalia" lsidName="S. wichgrafi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="wichgrafi">
<emphasis id="8297C732571747AA081A955E6BAB29A9" italics="true" pageId="0" pageNumber="133">S. wichgrafi</emphasis>
</taxonomicName>
(
<normalizedToken id="1610F4073368CD089FE494C25818FCD6" originalValue="Grünberg">Gruenberg</normalizedToken>
, 1910),
<normalizedToken id="C2A82482379FE865EEC71DC331E80E63" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
, male (all drawings by I.L.).
</paragraph>
</caption>
<caption id="7472C924B60FB10B4FEF788B81AE8F8F" httpUri="https://binary.pensoft.net/fig/845637" pageId="0" pageNumber="133" start="Figure 7" startId="F7">
<paragraph id="7D989DBA95C9B51F60558C591D2929CA" pageId="0" pageNumber="133">
<emphasis id="945EB99AD852A5DA8D506448BE779402" bold="true" pageId="0" pageNumber="133">Figure 7.</emphasis>
Female postabdominal structures and male genitalia in a not pressed condition:
<emphasis id="E7DDF051F71A58266F3D89ECCAF36B3C" bold="true" pageId="0" pageNumber="133">a.</emphasis>
<taxonomicName id="3D308790B7AA1A63A9A6D47746B8AFF8" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Eberhardfischeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eberhardfischeria husemanni" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="husemanni">
<emphasis id="AB8D984227252CACDB016662D6E1E83F" italics="true" pageId="0" pageNumber="133">Eberhardfischeria husemanni</emphasis>
</taxonomicName>
sp. nov., paratype, female;
<emphasis id="06B1635CAEA481B908F827B349F853F8" bold="true" pageId="0" pageNumber="133">b.</emphasis>
<taxonomicName id="054DF7A7422922C6FBC40B5A96F217C0" baseAuthorityName="Saalmueller" baseAuthorityYear="1884" class="Insecta" family="Leguminosae" genus="Saalmulleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saalmulleria stumpffi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="stumpffi">
<emphasis id="253D3428B6A89DB42D5E76CFA7CF739B" italics="true" pageId="0" pageNumber="133">Saalmulleria stumpffi</emphasis>
</taxonomicName>
(
<normalizedToken id="D3CF2665404F309F67A102449995E6A8" originalValue="Saalmüller">Saalmueller</normalizedToken>
, 1884),
<normalizedToken id="C3C2094FB09103FC978F89EBC32CF2ED" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
, female;
<emphasis id="9DE46368AD3FB4FC4AC40D64ECAE9D67" bold="true" pageId="0" pageNumber="133">c.</emphasis>
<taxonomicName id="7FC1FBC7E0CB5B68DB40EA36524EF28F" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" genus="Shimbania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shimbania kerstinhempae" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="kerstinhempae">
<emphasis id="44A8E675C50E802462BEE04BDB95144D" italics="true" pageId="0" pageNumber="133">Shimbania kerstinhempae</emphasis>
</taxonomicName>
sp. nov., holotype, female;
<emphasis id="B7F7625ADA0F224C6503D67E1F95B9D5" bold="true" pageId="0" pageNumber="133">d.</emphasis>
<taxonomicName id="4B7B7AA267DF3AB2B02F526D8008B48C" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania mineti" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="mineti">
<emphasis id="8E57F3F8D0636E29C4EFB5CAC80B38E8" italics="true" pageId="0" pageNumber="133">Morondavania mineti</emphasis>
</taxonomicName>
sp. nov., paratype, male, with aedeagus below;
<emphasis id="ABDA093B10ADD747BEA40B1D6C451BEC" bold="true" pageId="0" pageNumber="133">e.</emphasis>
<taxonomicName id="0BB58935F704C8EAC0221AEA3F20B4C8" class="Insecta" kingdom="Animalia" lsidName="S. wichgrafi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="wichgrafi">
<emphasis id="8384DFE232E0D371D56B94918005719F" italics="true" pageId="0" pageNumber="133">S. wichgrafi</emphasis>
</taxonomicName>
(
<normalizedToken id="294D3FF7734088108A870E46BF788637" originalValue="Grünberg">Gruenberg</normalizedToken>
, 1910),
<normalizedToken id="D049B84322ABF82DC3DACDAC255EC6E4" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
, male, with aedeagus below (all drawings by I.L.).
</paragraph>
</caption>
<paragraph id="A7AEC81D57DA1E0AC2EB214D37326865" pageId="0" pageNumber="133">
<emphasis id="30C32A6535E48218FF0080F42E15E97F" bold="true" italics="true" pageId="0" pageNumber="133">Abdomen</emphasis>
: With dense hair-like scales of brownish-olive mixed with sepia and short abdominal tuft,
<emphasis id="68F096BE0195C7D353E57149E4D455A2" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
20% of abdomen length. Male genitalia (Figs
<figureCitation id="24D536DCD11653B4D56EC76F020541C0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A. Morondavania mineti sp. nov., holotype, male, with unique banana-like large phallus (a), Madagascar, Western Region, north of Morondava, Marofandilia Forest; B. M. mineti sp. nov., paratype, male, Madagascar, Western Region, north of Morondava, western part of Marofandilia Forest, with ventrally connected gnathal arms (b) and below, in horizontal position, with a broad, well sclerotized transtilla (c) that represents a synapomorphy shared with species of Shimbania gen. nov. The transtilla is rarely present among Metarbelidae and if so, it is narrow and often only thinly sclerotized." httpUri="https://binary.pensoft.net/fig/845641" pageId="0" pageNumber="133">11A, B</figureCitation>
,
<figureCitation id="0B825E229279AB89C15CAB5D07C0C778" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus (phallus) of a. Shimbania tanaensis sp. nov., paratype, male, Kenya, Tana River County, Mchelelo Camp close to the Tana River; b. Morondavania mineti sp. nov., paratype, male, Madagascar, Western Region, north of Morondava, western part of Marofandilia Forest. The size of the aedeagus of the species of these sister-genera is among the largest in Metarbelidae." httpUri="https://binary.pensoft.net/fig/845642" pageId="0" pageNumber="133">12b</figureCitation>
) with tegumen and vinculum fused, forming a firm narrow ring, with tegumen
<emphasis id="631FC1F807EAFD4BE53F8946EC8F75A7" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
2.0-3.0
<normalizedToken id="B6A7FFE78ACD17A06AC4F059C1D94933" originalValue="×">x</normalizedToken>
broader than vinculum, the latter forms a narrow ring ventrally. Saccus present but strongly reduced with a broadly rounded or almost rectangular distal end. Uncus with heavy appearance, narrow elongated, well sclerotized, up to 30-40% of length of whole gnathos, flat dorsally, without a graben-like surface ventrally, not bifurcated at tip, with few tiny setae ventrally and dorsally, tip rounded. Basal edge of uncus well developed, slightly bent towards tip at center. Gnathos has gnathos arms that are medium large, one arm 25-30% the size of valva: upper part of the gnathos arm is a long, broad, weakly sclerotized band, as long as 70-80% of basal width of valva, that is attached to the basal part of uncus, the lower part of the gnathal arm is strongly sclerotized, of triangular shape with a pronounced thorn-like structure and with its ventral base in length
<emphasis id="657E5D38970246B066429CA10841008D" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
50% of the basal width of valva, smaller thorns absent, strong horizontal folds are present; the triangular-shaped gnathos is hollow; the gnathal arms are connected ventrally by a sclerotized long, narrow band that is as broad as 25-30% of the transtilla. The Gnathos is short and ends well above the costa of valva. The valva is strongly bent upwards in lateral view (if not below glass), large, broad at base, elongated and almost triangular-shaped, tip narrowly rounded; sacculus broad, weakly sclerotized, 40-60% of length of ventral edge of valva, costal margin weakly sclerotized towards base of valva; largely the valva is thinly membranous with many soft setae on inner side, but no other structures are present. Juxta well developed, with two broadly almost rectangular lobes and a deep V-shaped emargination in between lobes, the tips of lobes are broadly rounded. Phallus simple, tube-like, at present the largest among
<taxonomicName id="D6950154E2A2CE94F518B7B6C570D20D" authorityName="Strand" authorityYear="1909" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
and with a unique shape, namely banana-like and strongly bent on whole length, with 50-60% as broad as basal width of valva on
<emphasis id="AED5D6D1E51FEA88AC45917AAFB6B9AF" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
80% of its entire length, very narrow distally, and 30-40% longer than costal width of valva.
</paragraph>
<caption id="F6692BDC834A24C914B0FFF0B4EA9ACD" httpUri="https://binary.pensoft.net/fig/845638" pageId="0" pageNumber="133" start="Figure 8" startId="F8">
<paragraph id="7BAE58B00F2835608F4BC56E6303B065" pageId="0" pageNumber="133">
<emphasis id="3A1925687F1A5B181E3C47C46107357E" bold="true" pageId="0" pageNumber="133">Figure 8.</emphasis>
Male and female genitalia in a pressed condition below glass:
<emphasis id="CF5F3626E1AB20AA6FD6E321CF6A0ECD" bold="true" pageId="0" pageNumber="133">a.</emphasis>
<taxonomicName id="43C98C672681907779E69B10898B6953" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" genus="Shimbania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shimbania baginerichardi" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="baginerichardi">
<emphasis id="8DCACF22D6C427FEC8629E22F7207F03" italics="true" pageId="0" pageNumber="133">Shimbania baginerichardi</emphasis>
</taxonomicName>
sp. nov., holotype, male, Kenya, Kwale County, Shimba Hills National Reserve;
<emphasis id="0C9ED04B635C8038FCE86D2F8CAFD515" bold="true" pageId="0" pageNumber="133">b.</emphasis>
<taxonomicName id="B6699597CEE681D112CC073DF29C4ED3" class="Insecta" kingdom="Animalia" lsidName="S. tanaensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="tanaensis">
<emphasis id="E5543204675B24D04E8F964BAA12926A" italics="true" pageId="0" pageNumber="133">S. tanaensis</emphasis>
</taxonomicName>
sp. nov., paratype, male, Kenya, Mombasa County, Shimo la Tewa;
<emphasis id="82EA11511477501CFAEFCB32C4044AFE" bold="true" pageId="0" pageNumber="133">c.</emphasis>
<taxonomicName id="6D63FE23436DA16D3DC8101DE44F1898" class="Insecta" kingdom="Animalia" lsidName="S. budaensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="budaensis">
<emphasis id="77C6013FF359984296918F4605FCF725" italics="true" pageId="0" pageNumber="133">S. budaensis</emphasis>
</taxonomicName>
sp. nov., holotype, male, Kenya, Kwale County, Buda Forest Reserve;
<emphasis id="9C90592163FBC777512D45DCB1CB6A30" bold="true" pageId="0" pageNumber="133">d.</emphasis>
<taxonomicName id="437DB68AAAA22CA44A096A0C5A2EA618" class="Insecta" kingdom="Animalia" lsidName="S. pwaniensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="pwaniensis">
<emphasis id="8D952CEDBF807D8BA05C35B08C6D1889" italics="true" pageId="0" pageNumber="133">S. pwaniensis</emphasis>
</taxonomicName>
sp. nov., holotype, male, Tanzania, Pwani Region, road from Dar es-Salaam to Chalinze, near the railway-crossing, ca. 2 km from the Ruvu River.
</paragraph>
</caption>
<paragraph id="D468E1505D27408DCDE7EC2ED8EC95ED" pageId="0" pageNumber="133">
<emphasis id="482DF0FC746B9E4DB7604212317BC80D" bold="true" pageId="0" pageNumber="133">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C693BD1553EB17D12C9CCD06C11A76FE" pageId="0" pageNumber="133" type="species richness">
<paragraph id="222295A006D0CA94A1D75FF11BBC3247" pageId="0" pageNumber="133">Species richness.</paragraph>
<paragraph id="FDEA54DA28FA364F80D9E82336C9AA1B" pageId="0" pageNumber="133">Currently, this new genus is monotypic including one species new to science.</paragraph>
</subSubSection>
<subSubSection id="EE128B1B42E68240565661ACED645FF1" pageId="0" pageNumber="133" type="distribution">
<paragraph id="072BAD66A5A9AB0979CE68E9B43378D4" pageId="0" pageNumber="133">Distribution.</paragraph>
<paragraph id="474A70200BF0603AEB297074DEC79F19" pageId="0" pageNumber="133">
The only species of
<taxonomicName id="258F8DEDC4EA5D33A579B0416EA2482C" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="C6656CAF31416FD28797ABCA48F42C64" italics="true" pageId="0" pageNumber="133">Morondavania</emphasis>
</taxonomicName>
occurs on Madagascar in the &quot;Western region&quot;
<emphasis id="F263B312BC7B59B7C3D9AD44AFB93305" italics="true" pageId="0" pageNumber="133">sensu</emphasis>
<bibRefCitation id="AE06918517681CC6E03AA4AADEDEDA49" author="Humbert, H" journalOrPublisher="Journal of Applied Ecology" pageId="0" pageNumber="133" refId="B36" refString="Humbert, H, 1955. Les territoires phytogeographiques de Madagascar. In: Colloques internationaux du Centre National de la Recherche Scientifique, LIX: les divisions ecologiques du monde, moyen d'expression, nomenclature, cartographie. Annee Biologique, serie 3(31): 439-448[, Paris]." title="Les territoires phytogeographiques de Madagascar. In: Colloques internationaux du Centre National de la Recherche Scientifique, LIX: les divisions ecologiques du monde, moyen d'expression, nomenclature, cartographie. Annee Biologique, serie 3 (31): 439 - 448 [, Paris]." year="1955">Humbert (1955</bibRefCitation>
,
<bibRefCitation id="B9B48FD1BCF095F63665503076B9C0E4" author="Humbert, H" editor="Humbert, H" journalOrPublisher="Travaux de la Section Scientifique et Technique de l'Institut Franҫais de Pondichery, hors serie, volume 6" pageId="0" pageNumber="133" pagination="46 - 78" refId="B37" refString="Humbert, H, 1965. Description des types de vegetation. In: Humbert, H, Cours-Darne, G, Eds., Notice de la carte de Madagascar. Travaux de la Section Scientifique et Technique de l'Institut Franҫais de Pondichery, hors serie, volume 6: 46 - 78" title="Description des types de vegetation." volumeTitle="Notice de la carte de Madagascar." year="1965">1965</bibRefCitation>
) (
<emphasis id="D202D71968A9A39DD54207A537C5E689" italics="true" pageId="0" pageNumber="133">cf.</emphasis>
distribution map in
<bibRefCitation id="16DE2107C934EC41F59BB65B015DA066" author="Lehmann, I" journalOrPublisher="Heterocera Sumatrana" pageId="0" pageNumber="133" refId="B60" refString="Lehmann, I, 2019b. First revision of the family Metarbelidae Strand, 1909 (Lepidoptera, Cossoidea Leach, 1815) and a phylogeny based on adult morphology of 60 genera from the Afrotropical and Oriental Region. Doctoral Dissertation, 1-398, Rheinische Friedrich-Wilhelms-Universitaet Bonn. Universitaets- und Landesbibliothek Bonn, published 19th August 2019. [URN: urn:nbn:de:hbz:5n-55423]" title="First revision of the family Metarbelidae Strand, 1909 (Lepidoptera, Cossoidea Leach, 1815) and a phylogeny based on adult morphology of 60 genera from the Afrotropical and Oriental Region. Doctoral Dissertation, 1 - 398, Rheinische Friedrich-Wilhelms-Universitaet Bonn. Universitaets- und Landesbibliothek Bonn, published 19 th August 2019. [URN: urn: nbn: de: hbz: 5 n- 55423]" year="2019 b">Lehmann 2019b</bibRefCitation>
, fig. 49). Species of this new genus are most probably restricted to the highly threatened primary dry deciduous forest and woodland patches within the &quot;Madagascar Succulent Woodlands&quot; ecoregion
<emphasis id="35790C9B426E7214DD551E2D9C0BDC9A" italics="true" pageId="0" pageNumber="133">sensu</emphasis>
<bibRefCitation id="BF26F6B64FCFE33AE8C58AAAEFD3B065" author="Crowley, H" editor="Burgess, ND" journalOrPublisher="World Wildlife Fund (United States), Island Press, Washington, 269 - 271, 271 - 273, 276 - 278" pageId="0" pageNumber="133" pagination="417 - 418" refId="B13" refString="Crowley, H, 2004. Madagascar Humid Forests / Madagascar Subhumid Forests / Madagascar Dry Deciduous Forests / Madagascar Succulent Woodlands. In: Burgess, ND, D'Amico Hales, J, Underwood, E, Dinerstein, E, Olson, D, Itoua, I, Schipper, J, Rickketts, T, Newman, K, Eds., Terrestrial ecoregions of Africa and Madagascar: a conservation assessment. World Wildlife Fund (United States), Island Press, Washington, 269-271, 271-273, 276-278: 417 - 418" title="Madagascar Humid Forests / Madagascar Subhumid Forests / Madagascar Dry Deciduous Forests / Madagascar Succulent Woodlands." volumeTitle="Terrestrial ecoregions of Africa and Madagascar: a conservation assessment." year="2004">Crowley (2004)</bibRefCitation>
as well as to the &quot;Madagascar Dry Deciduous Forests&quot; ecoregion
<emphasis id="6901617C6AC685AD12193939D36EF297" italics="true" pageId="0" pageNumber="133">sensu</emphasis>
<bibRefCitation id="CA8A631C5CEDECB98D15BC56A96CA6D4" author="Crowley, H" editor="Burgess, ND" journalOrPublisher="World Wildlife Fund (United States), Island Press, Washington, 269 - 271, 271 - 273, 276 - 278" pageId="0" pageNumber="133" pagination="417 - 418" refId="B13" refString="Crowley, H, 2004. Madagascar Humid Forests / Madagascar Subhumid Forests / Madagascar Dry Deciduous Forests / Madagascar Succulent Woodlands. In: Burgess, ND, D'Amico Hales, J, Underwood, E, Dinerstein, E, Olson, D, Itoua, I, Schipper, J, Rickketts, T, Newman, K, Eds., Terrestrial ecoregions of Africa and Madagascar: a conservation assessment. World Wildlife Fund (United States), Island Press, Washington, 269-271, 271-273, 276-278: 417 - 418" title="Madagascar Humid Forests / Madagascar Subhumid Forests / Madagascar Dry Deciduous Forests / Madagascar Succulent Woodlands." volumeTitle="Terrestrial ecoregions of Africa and Madagascar: a conservation assessment." year="2004">Crowley (2004)</bibRefCitation>
. Due to the definitions of both ecoregions, the distribution range of species of the new genus extends within lowland areas below an altitude of 800 m of the &quot;West Malagasy regional centre of endemism&quot;
<emphasis id="138D69B832589D67D3B3893AD0A929A9" italics="true" pageId="0" pageNumber="133">sensu</emphasis>
<bibRefCitation id="1BFEDF5994646661CE59B08B858A608B" author="White, F" journalOrPublisher="Unesco, Paris" pageId="0" pageNumber="133" refId="B104" refString="White, F, 1983. The Vegetation of Africa: a Descriptive Memoir to Accompany the Unesco/AETFAT/UNSO Vegetation Map of Africa. Natural Resources Research no. XX. Unesco, Paris" title="The Vegetation of Africa: a Descriptive Memoir to Accompany the Unesco / AETFAT / UNSO Vegetation Map of Africa. Natural Resources Research no. XX." year="1983">White (1983)</bibRefCitation>
and might include riverine forests. Both ecoregions are very rich in species of woody
<taxonomicName id="22B1DE3431C7770C98B17BF931EFAC35" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Leguminosae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="family">Leguminosae</taxonomicName>
, comprising often locally endemic species,
<emphasis id="E3D85E1F74A445B9D11B98D1031B446F" italics="true" pageId="0" pageNumber="133">e.g.</emphasis>
species of the genera
<taxonomicName id="07559E747ADF169BEDA9183ABAAFF071" class="Dicotyledoneae" family="Leguminosae" genus="Albizia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Albizia" order="Fabales" pageId="0" pageNumber="133" phylum="Angiospermae" rank="genus">
<emphasis id="A8A9297AE44BF34137F370390EBD392E" italics="true" pageId="0" pageNumber="133">Albizia</emphasis>
</taxonomicName>
Durazz. (
<taxonomicName id="CE58B48DB9BB18D9DC14B3A5AA91D8B7" class="Dicotyledoneae" family="Leguminosae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="133" phylum="Angiospermae" rank="subFamily" subFamily="Mimosoideae">Mimosoideae</taxonomicName>
),
<taxonomicName id="29CFEEE70082A36592123C750FA71801" class="Magnoliopsida" family="Fabaceae" genus="Bauhinia" kingdom="Plantae" lsidName="Bauhinia" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">
<emphasis id="E4CC68C38548FDBA899D6AB25D8B2E1D" italics="true" pageId="0" pageNumber="133">Bauhinia</emphasis>
</taxonomicName>
L.,
<taxonomicName id="DBAF1C77492D914ED66D058F06E3FD81" class="Magnoliopsida" family="Fabaceae" genus="Cynometra" kingdom="Plantae" lsidName="Cynometra" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">
<emphasis id="63337CB0855BDF9F82F68046615AE2AB" italics="true" pageId="0" pageNumber="133">Cynometra</emphasis>
</taxonomicName>
L.,
<taxonomicName id="838D886A3BAD5F3AF47E6BBAD73E7CD3" class="Magnoliopsida" family="Fabaceae" genus="Delonix" kingdom="Plantae" lsidName="Delonix" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">
<emphasis id="0CBFC8A72427C5324E0F24E39FA69D0F" italics="true" pageId="0" pageNumber="133">Delonix</emphasis>
</taxonomicName>
Raf. (
<taxonomicName id="EFF4F30D10F0C92064C756E1A03FAFDD" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="subFamily" subFamily="Caesalpinioideae">Caesalpinioideae</taxonomicName>
),
<taxonomicName id="938D7156EB9ACC41CEE6C2285C8E5AB1" class="Magnoliopsida" family="Fabaceae" genus="Dalbergia" kingdom="Plantae" lsidName="Dalbergia" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">
<emphasis id="297C1662B008F361434C05D4243831E9" italics="true" pageId="0" pageNumber="133">Dalbergia</emphasis>
</taxonomicName>
L. and
<taxonomicName id="C1599BD9A724481459B6D75FBDCC82E0" authorityName="Schubert" authorityYear="1911" class="Dicotyledoneae" family="Leguminosae" genus="Millettia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Millettia" order="Fabales" pageId="0" pageNumber="133" phylum="Angiospermae" rank="genus">
<emphasis id="2FD2EB7AFD16CF36FA70E449F9DE96B2" italics="true" pageId="0" pageNumber="133">Millettia</emphasis>
</taxonomicName>
Wight &amp; Arn. (
<taxonomicName id="C2EF97A8C3331149FA49C7B12952200F" authorityName="Schubert" authorityYear="1911" class="Dicotyledoneae" family="Leguminosae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="133" phylum="Angiospermae" rank="subFamily" subFamily="Papilionoideae">Papilionoideae</taxonomicName>
) with
<taxonomicName id="2453471D1DA64F8E8DE321C862D22D3E" class="Magnoliopsida" family="Fabaceae" genus="Tamarindus" kingdom="Plantae" lsidName="Tamarindus indica" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="FC23A373E1A61B8C0C3462CBE4B32E22" italics="true" pageId="0" pageNumber="133">Tamarindus indica</emphasis>
</taxonomicName>
L. (
<taxonomicName id="1F5ADC6CB3C8C0FB4560ACFFC5A8D4F8" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="subFamily" subFamily="Caesalpinioideae">Caesalpinioideae</taxonomicName>
) in riverine forests (
<bibRefCitation id="3250977BBB74EDF31258F4DF6B14C424" author="Labat, J-N" editor="Goodman, SM" journalOrPublisher="The University of Chicago Press, Chicago and London" pageId="0" pageNumber="133" pagination="346 - 373" refId="B47" refString="Labat, J-N, Moat, J, 2003. Leguminosae (Fabaceae). In: Goodman, SM, Benstead, JP, Eds., The Natural History of Madagascar. The University of Chicago Press, Chicago and London: 346 - 373" title="Leguminosae (Fabaceae)." volumeTitle="The Natural History of Madagascar." year="2003">Labat and Moat 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A42CD681418A8CD2605E948D3C498BBB" pageId="0" pageNumber="133" type="biological traits">
<paragraph id="EDE74ED95E16D05D5AF73B3F6B3A9DF9" pageId="0" pageNumber="133">Biological traits.</paragraph>
<paragraph id="5F93FB5A8EA4263116C25E42B5A723F9" pageId="0" pageNumber="133">
The biology of species of
<taxonomicName id="3F617896865A084163B81E5345A13D4C" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Balsaminaceae" genus="Morondavania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morondavania" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A803E82EE283DF4F23FE7DCC55ACDF65" italics="true" pageId="0" pageNumber="133">Morondavania</emphasis>
</taxonomicName>
is unknown at present. However, lowland tropical
<taxonomicName id="623C42955AE6B228754989F76785EF81" authorityName="Strand" authorityYear="1909" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
species are strongly associated to habitats with woody legumes (
<emphasis id="0D55A6B44646DB86E7BFF928C7571987" italics="true" pageId="0" pageNumber="133">
cf.
<taxonomicName id="AFEAF8E036083E87A723E0ADE58984BF" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" genus="Shimbania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shimbania" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">Shimbania</taxonomicName>
</emphasis>
above;
<bibRefCitation id="098B47D1CE16135542B1EA02772600AA" DOI="https://doi.org/10.2982/0012-8317(2008)97[43:TNSOML]2.0.CO;2" author="Lehmann, I" journalOrPublisher="Nachrichten Entomologischer Verein Apollo" pageId="0" pageNumber="133" refId="B51" refString="Lehmann, I, 2008. Ten new species of Metarbelidae (Lepidoptera: Cossoidea) from the coastal forests and the Eastern Arc Mountains of Kenya and Tanzania, including one species from two upland forests. Journal of East African Natural History 97(1): 43-82. https://doi.org/10.2982/0012-8317(2008)97[43:TNSOML]2.0.CO;2" title="Ten new species of Metarbelidae (Lepidoptera: Cossoidea) from the coastal forests and the Eastern Arc Mountains of Kenya and Tanzania, including one species from two upland forests. Journal of East African Natural History 97 (1): 43 - 82." url="https://doi.org/10.2982/0012-8317(2008)97[43:TNSOML]2.0.CO;2" year="2008">Lehmann 2008</bibRefCitation>
,
<bibRefCitation id="31E0810606590D7E991284CA51E35D74" author="Lehmann, I" journalOrPublisher="Heterocera Sumatrana" pageId="0" pageNumber="133" refId="B60" refString="Lehmann, I, 2019b. First revision of the family Metarbelidae Strand, 1909 (Lepidoptera, Cossoidea Leach, 1815) and a phylogeny based on adult morphology of 60 genera from the Afrotropical and Oriental Region. Doctoral Dissertation, 1-398, Rheinische Friedrich-Wilhelms-Universitaet Bonn. Universitaets- und Landesbibliothek Bonn, published 19th August 2019. [URN: urn:nbn:de:hbz:5n-55423]" title="First revision of the family Metarbelidae Strand, 1909 (Lepidoptera, Cossoidea Leach, 1815) and a phylogeny based on adult morphology of 60 genera from the Afrotropical and Oriental Region. Doctoral Dissertation, 1 - 398, Rheinische Friedrich-Wilhelms-Universitaet Bonn. Universitaets- und Landesbibliothek Bonn, published 19 th August 2019. [URN: urn: nbn: de: hbz: 5 n- 55423]" year="2019 b">2019b</bibRefCitation>
) of the
<taxonomicName id="2A21D8D52E80C945DBFCD6B812F115DE" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="subFamily" subFamily="Papilionoideae">Papilionoideae</taxonomicName>
,
<taxonomicName id="9B652AAF5EFF930D0A6D1E84FB73702B" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="subFamily" subFamily="Mimosoideae">Mimosoideae</taxonomicName>
and Caesalpiniodeae that are most diverse in deciduous vegetation with a marked dry season as well as in lowland humid evergreen forest on western, southwestern and extreme northern Madagascar including many species reflecting a strong affinity with species on the African mainland (
<bibRefCitation id="5DF0AE688D33B075B2CDFD3B50EFDFAE" author="Labat, J-N" editor="Goodman, SM" journalOrPublisher="The University of Chicago Press, Chicago and London" pageId="0" pageNumber="133" pagination="346 - 373" refId="B47" refString="Labat, J-N, Moat, J, 2003. Leguminosae (Fabaceae). In: Goodman, SM, Benstead, JP, Eds., The Natural History of Madagascar. The University of Chicago Press, Chicago and London: 346 - 373" title="Leguminosae (Fabaceae)." volumeTitle="The Natural History of Madagascar." year="2003">Labat and Moat 2003</bibRefCitation>
).
</paragraph>
<caption id="5AF90498691F5EEC764BEAC6D7841C5A" httpUri="https://binary.pensoft.net/fig/845639" pageId="0" pageNumber="133" start="Figure 9" startId="F9">
<paragraph id="A2127255436A2F3763DD06E3E6978898" pageId="0" pageNumber="133">
<emphasis id="1D9372FD10AD1F540A3A1792BD3943CA" bold="true" pageId="0" pageNumber="133">Figure 9.</emphasis>
<emphasis id="117E5C3DC59857CE2F2839EDBD8ABE71" bold="true" pageId="0" pageNumber="133">a.</emphasis>
<taxonomicName id="1FD6803185C6B1622C033AEFFB3790FD" authorityName="Lehmann &amp; Dalsgaard" authorityYear="2023" class="Insecta" family="Bombacaceae" genus="Shimbania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Shimbania puguensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="puguensis">
<emphasis id="816698F8D09DAD94B5EBF0E1CAB5DFD6" italics="true" pageId="0" pageNumber="133">Shimbania puguensis</emphasis>
</taxonomicName>
sp. nov., paratype, male, Tanzania, Pwani Region, Kisarawe Forest;
<emphasis id="4F858ADDE93EE12A87DB5248C91D533D" bold="true" pageId="0" pageNumber="133">b.</emphasis>
<taxonomicName id="C0EE82B05F05EEB39055CEA8A5E442FF" class="Insecta" kingdom="Animalia" lsidName="S. kaguruensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="kaguruensis">
<emphasis id="E213224F43729126D4621963C672BDB6" italics="true" pageId="0" pageNumber="133">S. kaguruensis</emphasis>
</taxonomicName>
sp. nov., holotype, male, Tanzania, Morogoro Region, Kaguru Mountains;
<emphasis id="8EA9C3AC1607028293C5CA228A79B1EF" bold="true" pageId="0" pageNumber="133">c.</emphasis>
<taxonomicName id="8BC1D9ED0CDF955E2708C31D3B85F80D" class="Insecta" kingdom="Animalia" lsidName="S. mbarikaensis" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="mbarikaensis">
<emphasis id="AF086D367FAD39CA14A9C75D65C2CD74" italics="true" pageId="0" pageNumber="133">S. mbarikaensis</emphasis>
</taxonomicName>
sp. nov., holotype, male, Tanzania, Morogoro Region, Mbarika Mountains (also called Mbaraka Mountains), south of Mahenge Forest;
<emphasis id="269EAEA035752ABDB228BBEB8E8B17A7" bold="true" pageId="0" pageNumber="133">d.</emphasis>
<taxonomicName id="5E474F8BE22E052348C88E3E5337EC46" class="Insecta" kingdom="Animalia" lsidName="S. wanjakinuthiaae" order="Lepidoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="wanjakinuthiaae">
<emphasis id="71DA7D713D960C961B2465726C391BDB" italics="true" pageId="0" pageNumber="133">S. wanjakinuthiaae</emphasis>
</taxonomicName>
sp. nov., holotype, male, Republic of South Africa, Province KwaZulu-Natal, ca. 5 km north of Hluhluwe, probably collected on Hluhluwe Farm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="067EF9D6D4506502DAA2014BFF63B8CB" pageId="0" pageNumber="133" type="etymology">
<paragraph id="D6B249431BBD00026B68C773D1BE6E86" pageId="0" pageNumber="133">Etymology.</paragraph>
<paragraph id="D5B00D7ED30EBDAF640EDC74740FC86C" pageId="0" pageNumber="133">
The genus is named after the Morondava River that is located
<emphasis id="09CCEEDE00B562000C53EE5BA0829D79" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
19 km to the South of the type locality and might represent with its riverine forest patches another habitat for species of this new genus extending further inland (0-334 m and
<emphasis id="E83E754E34C843458E5447D6DDDEB4FB" italics="true" pageId="0" pageNumber="133">ca.</emphasis>
20°18'S-20°47'S and 44°14'E-45°15'E).
</paragraph>
</subSubSection>
</treatment>
</document>