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<document id="B09DC2AE2F64FA32CED8F077951605F7" ID-ISSN="1175­5334" ID-ZooBank="32126D3A-04BC-4AAC-89C5-F407AE28021C" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629809760349" checkinUser="felipe" docAuthor="Nicholson, Kirsten E., Crother, Brian I., Guyer, Craig &amp; Savage, Jay M." docDate="2012" docId="DA653D5C5251FFE9B1FCC9E0FAF3E2E5" docLanguage="en" docName="zt03477p108.pdf" docOrigin="Zootaxa 3477" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Dactyloa latifrons" docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="265C45245242FFFDB16BC977FF92E212" masterDocTitle="3477" masterLastPageNumber="108" masterPageNumber="1" pageNumber="20" updateTime="1699115282125" updateUser="plazi" zenodo-license-document="CLOSED">
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<treatment id="DA653D5C5251FFE9B1FCC9E0FAF3E2E5" ID-DOI="http://doi.org/10.5281/zenodo.5257463" ID-Zenodo-Dep="5257463" LSID="urn:lsid:plazi:treatment:DA653D5C5251FFE9B1FCC9E0FAF3E2E5" httpUri="http://treatment.plazi.org/id/DA653D5C5251FFE9B1FCC9E0FAF3E2E5" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="1AD6DFC15251FFEEB1FCC9E0FDBEE2A3" box="[151,556,151,177]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="52738C4A5251FFEEB1FCC9E0FDBEE2A3" blockId="19.[151,556,151,177]" box="[151,556,151,177]" pageId="19" pageNumber="20">
<heading id="093B3B265251FFEEB1FCC9E0FDBEE2A3" bold="true" box="[151,556,151,177]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="60B850585251FFEEB1FCC9E0FDBEE2A3" bold="true" box="[151,556,151,177]" pageId="19" pageNumber="20">
<taxonomicName id="95CCF7C95251FFEEB1FCC9E0FEE3E2A3" baseAuthorityName="Berthold" baseAuthorityYear="1846" box="[151,369,151,177]" class="Reptilia" family="Polychrotidae" genus="Dactyloa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="60B850585251FFEEB1FCC9E0FEE3E2A3" bold="true" box="[151,369,151,177]" italics="true" pageId="19" pageNumber="20">Dactyloa latifrons</emphasis>
</taxonomicName>
Species Group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1AD6DFC15251FFEEB1FCC9A9FBCAE04C" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="52738C4A5251FFEEB1FCC9A9FB32E3B8" blockId="19.[151,1436,222,750]" pageId="19" pageNumber="20">
<emphasis id="60B850585251FFEEB1FCC9A9FEBFE2E5" bold="true" box="[151,301,222,247]" pageId="19" pageNumber="20">
<emphasis id="60B850585251FFEEB1FCC9A9FE83E2E5" box="[151,273,222,247]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Diagnosis.</emphasis>
</emphasis>
Support for this group is provided by 54 apomorphies including seven morphological and 47 molecular ones. Six morphological features are unequivocal: maximum male snout-to-vent length increased (1: k to o); ratio of maximum female snout-to-vent length to maximum male snout-to-vent length decreased (2: a to c); length of thigh increased (3: o to w); length of head decreased (4: m to i); scales on dewlap with at least one double row (21: a to z); mean number of scales across snout increased (29: l to m); and scales in supraorbital disc about equal in size (41: 0 to 4). All of the molecular apomorphies are equivocal (see Appendix II).
</paragraph>
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<emphasis id="60B850585251FFEEB1ACC8C1FEF3E3DD" bold="true" box="[199,353,438,463]" pageId="19" pageNumber="20">
<emphasis id="60B850585251FFEEB1ACC8C1FED4E3DD" box="[199,326,438,463]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Definition.</emphasis>
</emphasis>
Species of this group are giant anoles (snout-to-vent length in adult males
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and
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in adult females) and have the following combinations of features: 1) inscriptional rib formula 5:0; 2) lack of caudal autotomy septa (present in one species,
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<emphasis id="60B850585251FFEEB26CC888FC16E004" box="[775,900,511,534]" italics="true" pageId="19" pageNumber="20">D. agassizi</emphasis>
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); 3) large splenial present; 4) rows of multiple scales on the dewlap; 5) double row of middorsal caudal scales.
</paragraph>
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<emphasis id="60B850585251FFEEB1ACCB31FED8E04D" bold="true" box="[199,330,582,607]" pageId="19" pageNumber="20">
<emphasis id="60B850585251FFEEB1ACCB31FEBFE04D" box="[199,301,582,607]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Content.</emphasis>
</emphasis>
Eighteen species are referred to this species group (see Appendix III).
</paragraph>
</subSubSection>
<subSubSection id="1AD6DFC15251FFEEB1ACCB1DFF27E0FC" pageId="19" pageNumber="20" type="distribution">
<paragraph id="52738C4A5251FFEEB1ACCB1DFF27E0FC" blockId="19.[151,1436,222,750]" pageId="19" pageNumber="20">
<emphasis id="60B850585251FFEEB1ACCB1DFEEEE091" bold="true" box="[199,380,618,643]" pageId="19" pageNumber="20">
<emphasis id="60B850585251FFEEB1ACCB1DFEF2E091" box="[199,352,618,643]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Distribution.</emphasis>
</emphasis>
Atlantic and Pacific slopes of
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and
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, south through the
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of
<collectingCountry id="2ADBCCDA5251FFEEB446CB1DFA0EE090" box="[1325,1436,618,642]" name="Colombia" pageId="19" pageNumber="20">Colombia</collectingCountry>
and
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; Malpelo Island; the Caribbean versant of
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, including the major river valleys of the
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Atrato,
<taxonomicName id="95CCF7C95251FFEEB186CBC5FE87E0D8" authority="Janeiro" authorityName="Janeiro" box="[237,277,690,714]" class="Reptilia" family="Dactyloidae" genus="Rio" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="69" phylum="Chordata" rank="genus">Río</taxonomicName>
<collectingRegion id="900842A85251FFEEB076CBC5FEF4E0D8" box="[285,358,690,714]" country="Colombia" name="Cauca" pageId="19" pageNumber="20">Cauca</collectingRegion>
, and
<taxonomicName id="95CCF7C95251FFEEB0CBCBC5FE5AE0D8" authority="Janeiro" authorityName="Janeiro" box="[416,456,690,714]" class="Reptilia" family="Dactyloidae" genus="Rio" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="69" phylum="Chordata" rank="genus">Río</taxonomicName>
<collectingRegion id="900842A85251FFEEB0BBCBC5FDDEE0D8" box="[464,588,690,714]" country="Colombia" name="Magdalena" pageId="19" pageNumber="20">Magdalena</collectingRegion>
and their adjacent slopes, and the Cordillera de la Costa of
<collectingCountry id="2ADBCCDA5251FFEEB587CBC5FACDE0D8" box="[1260,1375,690,714]" name="Venezuela" pageId="19" pageNumber="20">Venezuela</collectingCountry>
(
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).
</paragraph>
</subSubSection>
<caption id="06B3DCC25251FFEEB1FCCEDAFCA6E5D1" box="[151,820,1965,1988]" pageId="19" pageNumber="20" startId="19.[151,250,1965,1988]" targetBox="[176,1409,827,1918]" targetPageId="19">
<paragraph id="52738C4A5251FFEEB1FCCEDAFCA6E5D1" blockId="19.[151,820,1965,1988]" box="[151,820,1965,1988]" pageId="19" pageNumber="20">
<emphasis id="60B850585251FFEEB1FCCEDAFE80E5D6" bold="true" box="[151,274,1965,1988]" pageId="19" pageNumber="20">FIGURE 8.</emphasis>
Distribution of the
<taxonomicName id="95CCF7C95251FFEEB0B1CED9FD01E5D6" baseAuthorityName="Berthold" baseAuthorityYear="1846" box="[474,659,1966,1988]" class="Reptilia" family="Polychrotidae" genus="Dactyloa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="60B850585251FFEEB0B1CED9FD01E5D6" box="[474,659,1966,1988]" italics="true" pageId="19" pageNumber="20">Dactyloa latifrons</emphasis>
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Species Group.
</paragraph>
</caption>
<subSubSection id="1AD6DFC15256FFE9B1ACC9EFFAF3E2E5" pageId="20" pageNumber="21" type="discussion">
<paragraph id="52738C4A5256FFE9B1ACC9EFFAF3E2E5" blockId="20.[151,1436,151,247]" pageId="20" pageNumber="21">
<emphasis id="60B850585256FFE9B1ACC9EFFEC5E2A2" bold="true" box="[199,343,152,176]" pageId="20" pageNumber="21">
<emphasis id="60B850585256FFE9B1ACC9EFFEA8E2A2" box="[199,314,152,176]" inLineHeading="true" pageId="20" pageNumber="21" reason="1">Remarks.</emphasis>
</emphasis>
<bibRefCitation id="365DF1BB5256FFE9B03CC9E0FDD6E2BD" author="Hulebak, E. &amp; Poe, S. &amp; Ibanez, R. &amp; Williams, E. E." box="[343,580,151,176]" pageId="20" pageNumber="21" pagination="5 - 10" refId="ref39107" refString="Hulebak, E., Poe, S., Ibanez, R., &amp; Williams, E. E. (2007) A striking new species of Anolis lizard (Squamata: Iguania) from Panama. Phyllomedusa, 6, 5 - 10." type="journal article" year="2007">
Hulebak
<emphasis id="60B850585256FFE9B0D7C9EEFE7EE2A2" box="[444,492,152,176]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
. (2007)
</bibRefCitation>
in the course of describing
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<emphasis id="60B850585256FFE9B21CC9EFFBCAE2A2" box="[887,1112,152,176]" italics="true" pageId="20" pageNumber="21">Dactyloa kunayalae</emphasis>
</taxonomicName>
suggested that it might be an ally of the Colombian
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<emphasis id="60B850585256FFE9B0F2C9CAFE6FE2C6" box="[409,509,189,212]" italics="true" pageId="20" pageNumber="21">D. mirus</emphasis>
</taxonomicName>
and
<taxonomicName id="95CCF7C95256FFE9B35DC9CAFD3FE2C6" box="[566,685,188,212]" class="Reptilia" family="Polychrotidae" genus="Dactyloa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parrilis">
<emphasis id="60B850585256FFE9B35DC9CAFD3FE2C6" box="[566,685,188,212]" italics="true" pageId="20" pageNumber="21">D. parrilis</emphasis>
</taxonomicName>
. They noted that the three taxa share a unique morphology of the fourth toe, including few lamellae, indistinct toe pad, and especially long claw (see
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Fig.
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in
<bibRefCitation id="365DF1BB5256FFE9B5CEC9A8FAC6E2E5" author="Williams, E. E." box="[1189,1364,223,247]" pageId="20" pageNumber="21" pagination="463 - 480" refId="ref43226" refString="Williams, E. E. (1963) Studies on South American anoles. Description of Anolis mirus, new species from Rio San Juan, Colombia, with comment on digital dilation and dewlap as generic and specific characters in the anoles. Bulletin of the Museum of Comparative Zoology, Harvard, 129, 463 - 480." type="journal article" year="1963">Williams, 1963</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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