319 lines
37 KiB
XML
319 lines
37 KiB
XML
<document id="9E770669C59943F1FFF33CAF6095EE8C" ID-DOI="10.5281/zenodo.177443" ID-GBIF-Dataset="7a8d5548-d535-4437-b87c-3e1ebca275f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="177443" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459861428603" checkinUser="plazi" docAuthor="Ruta, Rafal" docDate="2007" docId="D97187B8A322FF911AAB10476AA3FC22" docLanguage="en" docName="zt01521p062.pdf" docOrigin="Zootaxa 1521" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hydrocyphon kopetdaghensis Ruta, 2007, sp. nov." docType="treatment" docVersion="9" lastPageNumber="62" masterDocId="2548FFC0A322FF921A3C142A6827FFCD" masterDocTitle="A new species of Hydrocyphon Redtenbacher, 1858 (Coleoptera: Scirtidae) from Kopet Dagh Mountains" masterLastPageNumber="62" masterPageNumber="59" pageNumber="59" updateTime="1698226083700" updateUser="plazi">
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<mods:mods id="6B7F01E755CF215560BA7E6CC9588B7B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3979796134726BF8BE05CC68FCB6227C">
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<mods:title id="177C593B709BE416536EA96AD41F715E">A new species of Hydrocyphon Redtenbacher, 1858 (Coleoptera: Scirtidae) from Kopet Dagh Mountains</mods:title>
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</mods:titleInfo>
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<mods:name id="B40EE9057D9251505A0300072275BD2E" type="personal">
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<mods:role id="0FEFCF6EF5870F301710E94C2995E814">
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<mods:roleTerm id="308ECEF77CD1C9437D92EA2CBCD85FDB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="82682D9D8F8DA3B7C3E4EB48005091C8">Ruta, Rafal</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5345DB69560F259EB9302A1DFDB4290A">text</mods:typeOfResource>
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<mods:titleInfo id="25D4E6C2AF6978F9CADC457DA085F46C">
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<mods:title id="61ED8C7AB6F77DFDC192385DAAA02BDE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3032AB4EBC41674F19E9080E240E45AC">
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<mods:date id="13463A336363612ACF3970CC2F3DDFF1">2007</mods:date>
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<mods:detail id="05F87EA646A885F4186C9B7DF5B3D4ED" type="volume">
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<mods:number id="111C3B1E571E10D4E5C5B7CDA321139B">1521</mods:number>
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<mods:start id="C9C39880ECACE3EBB606B4F04E8C9A00">59</mods:start>
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<mods:end id="42720FAD83D3D3D4597DCFFF69FB12A2">62</mods:end>
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</mods:relatedItem>
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<mods:classification id="313B5D0D4E72300E24562EBFD383782B">journal article</mods:classification>
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<mods:identifier id="E2E236C01FA46FB674DFF432B27A92EC" type="DOI">10.5281/zenodo.177443</mods:identifier>
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<mods:identifier id="A9FBAA64461BF74277E6717AC8751D22" type="GBIF-Dataset">7a8d5548-d535-4437-b87c-3e1ebca275f7</mods:identifier>
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<mods:identifier id="5B1BC09EE33350E24B3D3731576A0954" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0CA266CFF8FAB18942240E4B9B261026" type="Zenodo-Dep">177443</mods:identifier>
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</mods:mods>
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<treatment id="D97187B8A322FF911AAB10476AA3FC22" ID-DOI="http://doi.org/10.5281/zenodo.6243983" ID-GBIF-Taxon="119368258" ID-Zenodo-Dep="6243983" LSID="urn:lsid:plazi:treatment:D97187B8A322FF911AAB10476AA3FC22" httpUri="http://treatment.plazi.org/id/D97187B8A322FF911AAB10476AA3FC22" lastPageId="3" lastPageNumber="62" pageId="0" pageNumber="59">
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<subSubSection id="19C26525A322FF921AAB1047692BFB6A" pageId="0" pageNumber="59" type="nomenclature">
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<paragraph id="516736AEA322FF921AAB10476A47FB4A" blockId="0.[151,608,1133,1191]" box="[151,608,1133,1159]" pageId="0" pageNumber="59">
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<heading id="0A2F81C2A322FF921AAB10476A47FB4A" bold="true" box="[151,608,1133,1159]" fontSize="11" level="1" pageId="0" pageNumber="59" reason="1">
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<emphasis id="63ACEABCA322FF921AAB10476A47FB4A" bold="true" box="[151,608,1133,1159]" pageId="0" pageNumber="59">
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<taxonomicName id="96D84D2DA322FF921AAB104769DBFB4A" box="[151,508,1133,1159]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="kopetdaghensis" status="sp. nov.">
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<emphasis id="63ACEABCA322FF921AAB104769DBFB4A" bold="true" box="[151,508,1133,1159]" italics="true" pageId="0" pageNumber="59">Hydrocyphon kopetdaghensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="789F57C7A322FF92183F10476A47FB4A" box="[515,608,1133,1159]" pageId="0" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="516736AEA322FF921AAB10BA692BFB6A" blockId="0.[151,608,1133,1191]" box="[151,268,1168,1191]" pageId="0" pageNumber="59">
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(
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<figureCitation id="C9E32A2BA322FF921AA310BA6924FB6A" box="[159,259,1168,1191]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 7" captionStartId-0="1.[151,255,1559,1583]" captionStartId-1="2.[151,269,1356,1380]" captionTargetBox-1="[209,1415,194,1309]" captionTargetId-1="figure@2.[151,1436,166,1333]" captionTargetPageId-1="2" captionText-0="FIGURE 1. Hydrocyphon kopetdaghensis, holotype." captionText-1="FIGURES 2 – 7. Hydrocyphon kopetdaghensis, male genitalia; scale bar = 0.2 mm. Fig. 2, penis; Fig. 3, tegmen; Fig. 4, tergite VIII; Fig. 5, tergite IX; Fig. 6, sternite VIII; Fig. 7, sternite IX." httpUri-1="https://zenodo.org/record/177444/files/figure.png" pageId="0" pageNumber="59">Figs. 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="19C26525A322FF921AAB10FE6D19FAFC" pageId="0" pageNumber="59" type="materials_examined">
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<paragraph id="516736AEA322FF921AAB10FE6D19FAFC" blockId="0.[151,1437,1236,2023]" pageId="0" pageNumber="59">
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<emphasis id="63ACEABCA322FF921AAB10FE68D8FB21" bold="true" box="[151,255,1236,1260]" pageId="0" pageNumber="59">
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<typeStatus id="8E63880CA322FF921AAB10FE68D8FB21" box="[151,255,1236,1260]" pageId="0" pageNumber="59" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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<collectionCode id="37C9AE6BA322FF921B3710FF6976FB21" box="[267,337,1237,1260]" pageId="0" pageNumber="59">USSR</collectionCode>
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, Turkmenia, Kopet-Dagh / Mts.,
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<quantity id="96209B4BA322FF92188D10FF6B24FB21" box="[689,771,1237,1260]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="59" unit="m" value="1000.0">1000 m</quantity>
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, Kurkulab,
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<quantity id="96209B4BA322FF92194310FF6B92FB21" box="[895,949,1237,1260]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="0" pageNumber="59" unit="km" value="6.0">6 km</quantity>
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W / Germob,
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<geoCoordinate id="34EC5069A322FF921E7010FF6C82FB21" box="[1100,1189,1237,1260]" direction="east" orientation="longitude" pageId="0" pageNumber="59" precision="925" value="57.833332">57º50’E</geoCoordinate>
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,
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<geoCoordinate id="34EC5069A322FF921E8C10FF6D2DFB21" box="[1200,1290,1237,1260]" direction="north" orientation="latitude" pageId="0" pageNumber="59" precision="925" value="38.066666">38º04’N</geoCoordinate>
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/ 0
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<date id="2566106EA322FF921F1710FF6DBFFB21" box="[1323,1432,1237,1260]" pageId="0" pageNumber="59" value="1991-03-10">3.10.1991</date>
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, No. L34 / leg.: A. Podlussány, L. Ronkay & Z. Varga [white label, printed].
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<taxonomicName id="96D84D2DA322FF9219FB10D26D12FAC2" box="[967,1333,1272,1295]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="kopetdaghensis" status="sp. nov.">
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<emphasis id="63ACEABCA322FF9219FB10D26D12FAC2" box="[967,1333,1272,1295]" italics="true" pageId="0" pageNumber="59">HYDROCYPHON kopetdaghensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="789F57C7A322FF921F0710D26DA9FAC2" box="[1339,1422,1272,1295]" pageId="0" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
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/ des. Rafał Ruta,
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<docYear id="49BEDA55A322FF921B741130695AFAFC" box="[328,381,1306,1329]" pageId="0" pageNumber="59">2007</docYear>
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/
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<typeStatus id="8E63880CA322FF921BAE11306A16FAFC" box="[402,561,1306,1329]" pageId="0" pageNumber="59" type="holotype">HOLOTYPUS</typeStatus>
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[red label, printed]. Deposited in Hungarian Museum of Natural History.
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</paragraph>
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</subSubSection>
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<subSubSection id="19C26525A322FF921AFA11166B6AFABA" pageId="0" pageNumber="59" type="diagnosis">
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<paragraph id="516736AEA322FF921AFA11166B6AFABA" blockId="0.[151,1437,1236,2023]" pageId="0" pageNumber="59">
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<emphasis id="63ACEABCA322FF921AFA1116691CFA99" bold="true" box="[198,315,1340,1364]" pageId="0" pageNumber="59">Diagnosis.</emphasis>
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Easily distinguishable on the basis of genital characters: the shape of parameres combined with structural anatomy of the penis are unique for this species of
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<taxonomicName id="96D84D2DA322FF92188A114A6B61FABA" box="[694,838,1376,1399]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="63ACEABCA322FF92188A114A6B61FABA" box="[694,838,1376,1399]" italics="true" pageId="0" pageNumber="59">Hydrocyphon</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="19C26525A322FF931AFA11A86B90F803" lastPageId="1" lastPageNumber="60" pageId="0" pageNumber="59" type="description">
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||
<paragraph id="516736AEA322FF921AFA11A86BA1F9E9" blockId="0.[151,1437,1236,2023]" pageId="0" pageNumber="59">
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||
<emphasis id="63ACEABCA322FF921AFA11A86976FA57" bold="true" box="[198,337,1410,1434]" pageId="0" pageNumber="59">Description.</emphasis>
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<emphasis id="63ACEABCA322FF921B6511A869B0FA54" box="[345,407,1410,1433]" italics="true" pageId="0" pageNumber="59">Male.</emphasis>
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Body oval, slightly depressed, shiny, clothed with bright, yellowish-white hairs. Elytra brownish, pronotum yellow with darkened disc, head dark brown. Venter brown. Legs yellow with slightly darkened tarsi. Maxillary palpi yellowish-brown. Antennomeres 1-4 yellow, 5 brown, 6-11 dark brown. Length (measured from the anterior edge of pronotum to the apex of elytra) 2.0 mm, greatest width of pronotum
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<quantity id="96209B4BA322FF921E2311C06C54F9CC" box="[1055,1139,1514,1537]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="0" pageNumber="59" unit="mm" value="0.9">0.9 mm</quantity>
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||
, greatest width of elytra
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||
<quantity id="96209B4BA322FF921F4611C068E4F9E9" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="0" pageNumber="59" unit="mm" value="1.4">1.4 mm</quantity>
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||
, greatest depth of body
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<quantity id="96209B4BA322FF921BFE12276A31F9E9" box="[450,534,1549,1572]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="59" unit="mm" value="0.7">0.7 mm</quantity>
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, body 1.46 times as long as broad.
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</paragraph>
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<paragraph id="516736AEA322FF921AFA121A69E2F91C" blockId="0.[151,1437,1236,2023]" pageId="0" pageNumber="59">
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Greatest width of the head
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||
<quantity id="96209B4BA322FF921BCC121A6A73F98A" box="[496,596,1584,1607]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="0" pageNumber="59" unit="mm" value="0.65">0.65 mm</quantity>
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. Head 1.86 times as broad as long, 1.6 times wider than width of interocular space, with distinct punctation, and strongly protuberant eyes. Antenna filiform; length ratio of antennomeres: 2.9: 2.3: 1.0: 2.5: 2.1: 2.1: 2.1: 2.1: 2.1: 2.1: 2.1: 3.9; length/width ratio of antennomeres: 1.6, 1.3, 1.1, 1.8, 1.5, 1.5, 1.5, 1.5, 1.5, 1.4, 2.9. Anterior clypeal margin straight, with rounded anterior angles. Labrum with rounded margin. Mandibles reduced, with obtuse apices.
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</paragraph>
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<paragraph id="516736AEA322FF921AFA12F7690DF8F4" blockId="0.[151,1437,1236,2023]" pageId="0" pageNumber="59">Pronotum small, 2.57 times as broad as long, widest at its base. Disc of pronotum with subtle punctation. Posterior pronotal margin almost straight, slightly sinuate. Lateral margins straight, slightly converging anteriorly. Anterior angles not produced.</paragraph>
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<paragraph id="516736AEA322FF921AFA136F6BA9F82A" blockId="0.[151,1437,1236,2023]" pageId="0" pageNumber="59">
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Scutellum triangular, as long as broad, with indistinct punctation. Base of elytra evenly wider than base of pronotum, sides rounded. Elytra without longitudinal ridges, with marked humeri, 1.32 times as long as broad and 5.58 times as long as pronotum. Subtle adsutural strioles are present in proximal half of elytra. Punctation relatively dense, punctures separated by a single diameter. Punctures larger than those on pronotum. Epipleura yellow, gradually narrowing, virtually obsolete in subapical region of elytra.
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<collectingCountry id="29CF763EA322FF92186213FA6AB4F82A" box="[606,659,2000,2023]" name="India" pageId="0" pageNumber="59">Hind</collectingCountry>
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wings fully developed.
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</paragraph>
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<caption id="05A76626A323FF931AAB123D6AEAF9E3" box="[151,717,1559,1583]" pageId="1" pageNumber="60">
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||
<paragraph id="516736AEA323FF931AAB123D6AEAF9E3" blockId="1.[151,717,1559,1583]" box="[151,717,1559,1583]" pageId="1" pageNumber="60">
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<emphasis id="63ACEABCA323FF931AAB123D693EF9E2" bold="true" box="[151,281,1559,1583]" pageId="1" pageNumber="60">FIGURE 1.</emphasis>
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<taxonomicName id="96D84D2DA323FF931B1C12326A7DF9E2" box="[288,602,1560,1583]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="kopetdaghensis">
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<emphasis id="63ACEABCA323FF931B1C12326A7DF9E2" box="[288,602,1560,1583]" italics="true" pageId="1" pageNumber="60">Hydrocyphon kopetdaghensis</emphasis>
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</taxonomicName>
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, holotype.
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</paragraph>
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</caption>
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<paragraph id="516736AEA323FF931AFA12766BDCF916" blockId="1.[151,1437,1628,2034]" pageId="1" pageNumber="60">
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Prosternal process elongated, very narrow, rod-like. Mesocoxae separated by a long subrectangular process (ca. 3 times as long as wide), metaventral discrimen present in posterior 2/3 of metaventrite. Metaventral process reduced, subtriangular, metacoxae touching one another. Five abdominal sternites visible. Ratio of ventrites lengths: 1: 2: 2: 2: 2.5. Right metafemur of the
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<typeStatus id="8E63880CA323FF931BAB12EE69D3F916" box="[407,500,1732,1755]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
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is underdeveloped – shorter than that of the left.
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</paragraph>
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<paragraph id="516736AEA323FF931AFA12CD6CDDF866" blockId="1.[151,1437,1628,2034]" pageId="1" pageNumber="60">
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Genitalia. Penis symmetrical, with long, regularly curved parameroids and wide prosthemes; both parameroids and prosthemes somewhat up-curved; length (L) =
|
||
<quantity id="96209B4BA323FF9318B613206AC9F8EC" box="[650,750,1802,1825]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="60" unit="mm" value="0.64">0.64 mm</quantity>
|
||
., width (W) =
|
||
<quantity id="96209B4BA323FF9319BB13206BCFF8EC" box="[903,1000,1802,1825]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="60" unit="mm" value="0.24">0.24 mm</quantity>
|
||
. Tegmen with parameres wide, produced apically into sharp, excurved hooks; L =
|
||
<quantity id="96209B4BA323FF93187013066A8AF88E" box="[588,685,1836,1859]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="60" unit="mm" value="0.52">0.52 mm</quantity>
|
||
, W =
|
||
<quantity id="96209B4BA323FF9318D213066B77F88E" box="[750,848,1836,1859]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="60" unit="mm" value="0.34">0.34 mm</quantity>
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||
. Tergite VIII with a row of minute setae on apical margin; L =
|
||
<quantity id="96209B4BA323FF931AC913656970F8AB" box="[245,343,1871,1894]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="60" unit="mm" value="0.41">0.41 mm</quantity>
|
||
, W =
|
||
<quantity id="96209B4BA323FF931BA4136569DCF8AB" box="[408,507,1871,1894]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="60" unit="mm" value="0.35">0.35 mm</quantity>
|
||
. Tergite IX membranous in central part of apical margin; L =
|
||
<quantity id="96209B4BA323FF931EAF13656CD3F8AB" box="[1171,1268,1871,1894]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="1" pageNumber="60" unit="mm" value="0.36">0.36 mm</quantity>
|
||
, W =
|
||
<quantity id="96209B4BA323FF931F0A13656DBEF8AB" box="[1334,1433,1871,1894]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="60" unit="mm" value="0.32">0.32 mm</quantity>
|
||
. Sternite VIII v-shaped, L =
|
||
<quantity id="96209B4BA323FF931BFD13586A04F844" box="[449,547,1906,1929]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="60" unit="mm" value="0.25">0.25 mm</quantity>
|
||
, W =
|
||
<quantity id="96209B4BA323FF93185813586AE0F844" box="[612,711,1906,1929]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="60" unit="mm" value="0.28">0.28 mm</quantity>
|
||
. Sternite IX consisting of two hemisternites, each of which consists of a long baculus, with widened apex bearing several setae; L =
|
||
<quantity id="96209B4BA323FF93197813BE6B82F866" box="[836,933,1940,1963]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="60" unit="mm" value="0.46">0.46 mm</quantity>
|
||
, W =
|
||
<quantity id="96209B4BA323FF9319DA13BE6C62F866" box="[998,1093,1940,1963]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="60" unit="mm" value="0.24">0.24 mm</quantity>
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||
(whole sternite).
|
||
</paragraph>
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<paragraph id="516736AEA323FF931AFA139D6B90F803" blockId="1.[151,1437,1628,2034]" box="[198,951,1975,1999]" pageId="1" pageNumber="60">
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||
<emphasis id="63ACEABCA323FF931AFA139D69ABF802" bold="true" box="[198,396,1975,1999]" pageId="1" pageNumber="60">Name derivation.</emphasis>
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Named after
|
||
<emphasis id="63ACEABCA323FF93182113926ABDF802" box="[541,666,1976,1999]" italics="true" pageId="1" pageNumber="60">terra typica</emphasis>
|
||
– Kopet Dagh Mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="19C26525A323FF931AFA13F06C58F83C" box="[198,1151,2010,2034]" pageId="1" pageNumber="60" type="distribution">
|
||
<paragraph id="516736AEA323FF931AFA13F06C58F83C" blockId="1.[151,1437,1628,2034]" box="[198,1151,2010,2034]" pageId="1" pageNumber="60">
|
||
<emphasis id="63ACEABCA323FF931AFA13F0697FF83F" bold="true" box="[198,344,2010,2034]" pageId="1" pageNumber="60">Distribution.</emphasis>
|
||
Known only from its
|
||
<emphasis id="63ACEABCA323FF93187813F06AE8F83C" box="[580,719,2010,2033]" italics="true" pageId="1" pageNumber="60">locus typicus</emphasis>
|
||
in Kopet Dagh Mountains (Turkmenia).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="05A76626A320FF901AAB11666BA5FA4A" httpUri="https://zenodo.org/record/177444/files/figure.png" pageId="2" pageNumber="61" targetBox="[209,1415,194,1309]" targetPageId="2">
|
||
<paragraph id="516736AEA320FF901AAB11666BA5FA4A" blockId="2.[151,1436,1356,1415]" pageId="2" pageNumber="61">
|
||
<emphasis id="63ACEABCA320FF901AAB11666964FAA9" bold="true" box="[151,323,1356,1380]" pageId="2" pageNumber="61">FIGURES 2–7.</emphasis>
|
||
<taxonomicName id="96D84D2DA320FF901B7611676AA1FAA9" box="[330,646,1357,1380]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="kopetdaghensis">
|
||
<emphasis id="63ACEABCA320FF901B7611676AA1FAA9" box="[330,646,1357,1380]" italics="true" pageId="2" pageNumber="61">Hydrocyphon kopetdaghensis</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia; scale bar = 0.2 mm.
|
||
<emphasis id="63ACEABCA320FF901E2911666C7BFAA9" bold="true" box="[1045,1116,1356,1380]" pageId="2" pageNumber="61">Fig. 2,</emphasis>
|
||
penis;
|
||
<emphasis id="63ACEABCA320FF901E9611666CD6FAA9" bold="true" box="[1194,1265,1356,1380]" pageId="2" pageNumber="61">Fig. 3,</emphasis>
|
||
tegmen;
|
||
<emphasis id="63ACEABCA320FF901F6911666DBBFAA9" bold="true" box="[1365,1436,1356,1380]" pageId="2" pageNumber="61">Fig. 4,</emphasis>
|
||
tergite VIII;
|
||
<emphasis id="63ACEABCA320FF901B2111456944FA4A" bold="true" box="[285,355,1391,1415]" pageId="2" pageNumber="61">Fig. 5,</emphasis>
|
||
tergite IX;
|
||
<emphasis id="63ACEABCA320FF901BE211456A02FA4A" bold="true" box="[478,549,1391,1415]" pageId="2" pageNumber="61">Fig. 6,</emphasis>
|
||
sternite VIII;
|
||
<emphasis id="63ACEABCA320FF90188011456B24FA4A" bold="true" box="[700,771,1391,1415]" pageId="2" pageNumber="61">Fig. 7,</emphasis>
|
||
sternite IX.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="05A76626A320FF901AAB138E6C57F876" box="[151,1136,1956,1980]" httpUri="https://zenodo.org/record/177445/files/figure.png" pageId="2" pageNumber="61" targetBox="[384,1203,1469,1932]" targetPageId="2">
|
||
<paragraph id="516736AEA320FF901AAB138E6C57F876" blockId="2.[151,1136,1956,1980]" box="[151,1136,1956,1980]" pageId="2" pageNumber="61">
|
||
<emphasis id="63ACEABCA320FF901AAB138E693DF871" bold="true" box="[151,282,1956,1980]" pageId="2" pageNumber="61">FIGURE 8.</emphasis>
|
||
Zoogeographical distribution of species of the
|
||
<taxonomicName id="96D84D2DA320FF90192D138F6B49F871" box="[785,878,1957,1980]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="63ACEABCA320FF90192D138F6B49F871" box="[785,878,1957,1980]" italics="true" pageId="2" pageNumber="61">australis</emphasis>
|
||
</taxonomicName>
|
||
-group of
|
||
<taxonomicName id="96D84D2DA320FF9019E5138F6C4EF871" box="[985,1129,1957,1980]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="63ACEABCA320FF9019E5138F6C4EF871" box="[985,1129,1957,1980]" italics="true" pageId="2" pageNumber="61">Hydrocyphon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="19C26525A321FF911AFA14BD6AA3FC22" pageId="3" pageNumber="62" type="discussion">
|
||
<paragraph id="516736AEA321FF911AFA14BD699AFCA9" blockId="3.[151,1437,151,1007]" pageId="3" pageNumber="62">
|
||
<emphasis id="63ACEABCA321FF911AFA14BD6962FF62" bold="true" box="[198,325,151,175]" pageId="3" pageNumber="62">Discussion.</emphasis>
|
||
On the basis of structural anatomy of the penis (both parameroides and prosthemes symmetrical) and tegmen (parameres wide), the species is classified within the
|
||
<taxonomicName id="96D84D2DA321FF91196A14906C77FF1C" box="[854,1104,186,209]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="63ACEABCA321FF91196A14906C77FF1C" box="[854,1104,186,209]" italics="true" pageId="3" pageNumber="62">Hydrocyphon australis</emphasis>
|
||
</taxonomicName>
|
||
-group, sensu
|
||
<bibRefCitation id="35494B5FA321FF911ECD14906DBBFF1C" author="Nyholm" box="[1265,1436,186,209]" pageId="3" pageNumber="62" refString="Nyholm, T. (1967) Zur Kenntnis der Gattung Hydrocyphon Redtenbacher (Col. Helodidae). Studien uber die Familie Helodidae. VIII. Opuscula Entomologica, 32 (1 - 2), 9 - 48." type="journal article" year="1967">Nyholm (1967)</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="35494B5FA321FF911AA214F7697BFF39" author="Klausnitzer" box="[158,348,221,244]" pageId="3" pageNumber="62" refString="Klausnitzer, B. (2002) Eine neue Art und Artengruppe der Gattung Hydrocyphon Redtenbacher aus Nepal (Col., Scirtidae). Entomologische Nachrichten und Berichte, 46, 259 - 264." type="journal article" year="2002">Klausnitzer 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="35494B5FA321FF911B5514F76A9BFF39" author="Yoshitomi" box="[361,700,221,244]" pageId="3" pageNumber="62" refString="Yoshitomi, H. & Klausnitzer, B. (2003) SCIRTIDAE: World check list of Hydrocyphon Redtenbacher, and revision of the Chinese species. In: Jaech, M. A. & Ji, L. (eds.), Water Beetles of China, Vol. III. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien. pp. 519 - 537." type="book chapter" year="2003">Yoshitomi & Klausnitzer 2003</bibRefCitation>
|
||
), known from the eastern part of Western Palaearctic Region, with three species occurring in the Mediterranean Subregion, and four in the Middle East. Four of the five species of
|
||
<taxonomicName id="96D84D2DA321FF911F04152A68EBFEF4" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="63ACEABCA321FF911F04152A68EBFEF4" italics="true" pageId="3" pageNumber="62">Hydrocyphon</emphasis>
|
||
</taxonomicName>
|
||
known to occur in the Middle East, belong to the
|
||
<taxonomicName id="96D84D2DA321FF9118CD15086B69FEF4" box="[753,846,290,313]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="63ACEABCA321FF9118CD15086B69FEF4" box="[753,846,290,313]" italics="true" pageId="3" pageNumber="62">australis</emphasis>
|
||
</taxonomicName>
|
||
-group. Their distribution is shown in
|
||
<figureCitation id="C9E32A2BA321FF911ED015086D0CFEF4" box="[1260,1323,290,313]" captionStart="FIGURE 8" captionStartId="2.[151,255,1956,1980]" captionTargetBox="[384,1203,1469,1932]" captionTargetId="figure@2.[384,1203,1469,1932]" captionTargetPageId="2" captionText="FIGURE 8. Zoogeographical distribution of species of the australis - group of Hydrocyphon." httpUri="https://zenodo.org/record/177445/files/figure.png" pageId="3" pageNumber="62">Fig. 8</figureCitation>
|
||
(based on data from:
|
||
<bibRefCitation id="35494B5FA321FF911B2D156F698DFE91" author="Nyholm" box="[273,426,325,348]" pageId="3" pageNumber="62" refString="Nyholm, T. (1967) Zur Kenntnis der Gattung Hydrocyphon Redtenbacher (Col. Helodidae). Studien uber die Familie Helodidae. VIII. Opuscula Entomologica, 32 (1 - 2), 9 - 48." type="journal article" year="1967">Nyholm 1967</bibRefCitation>
|
||
, 1972, 1977;
|
||
<bibRefCitation id="35494B5FA321FF91187E156F6AD9FE91" author="Klausnitzer" box="[578,766,325,348]" pageId="3" pageNumber="62" refString="Klausnitzer, B. (1970) Zur Kenntnis der europaischen Hydrocyphon - Arten (Coleoptera, Helodidae). Reichenbachia, 13 (20), 205 - 209." type="journal article" year="1970">Klausnitzer 1970</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="35494B5FA321FF911937156F6B65FE91" author="Klausnitzer" box="[779,834,325,348]" pageId="3" pageNumber="62" refString="Klausnitzer, B. (1981) Zur Kenntnis der Cyphon - und Hydrocyphon - Fauna des Iran. Faunistische Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 8 (4), 63 - 65." type="journal article" year="1981">1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="35494B5FA321FF91196C156F6BAFFE91" author="Klausnitzer" box="[848,904,325,348]" pageId="3" pageNumber="62" refString="Klausnitzer, B. (1989) Neufunde zur Helodidenfauna der Turkei, mit Beschriebung einer neuen Cyphon-Art (Insecta, Coleoptera: Helodidae). Reichenbachia, 26 (19), 111 - 114." type="journal article" year="1989">1989</bibRefCitation>
|
||
, 1990, 1990b, 1990c). An oriental
|
||
<taxonomicName id="96D84D2DA321FF911F30156F69BFFEB2" authority="Klausnitzer" authorityName="Klausnitzer" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="63ACEABCA321FF911F30156F6931FEB2" italics="true" pageId="3" pageNumber="62">Hydrocyphon bhutanensis</emphasis>
|
||
Klausnitzer
|
||
</taxonomicName>
|
||
was included in the
|
||
<taxonomicName id="96D84D2DA321FF91184B15426AF3FEB2" box="[631,724,360,383]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="63ACEABCA321FF91184B15426AF3FEB2" box="[631,724,360,383]" italics="true" pageId="3" pageNumber="62">australis</emphasis>
|
||
</taxonomicName>
|
||
-group both in the original description (
|
||
<bibRefCitation id="35494B5FA321FF911E4115426D1FFEB2" author="Klausnitzer" box="[1149,1336,360,383]" pageId="3" pageNumber="62" refString="Klausnitzer, B. (1976) Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Helodidae. Deutsche EntomologischeZeitschrift, 23, 213 - 220." type="journal article" year="1976">Klausnitzer 1976</bibRefCitation>
|
||
) and in a recent check-list (
|
||
<bibRefCitation id="35494B5FA321FF911B6B15A06A7BFE6C" author="Yoshitomi" box="[343,604,394,417]" pageId="3" pageNumber="62" refString="Yoshitomi, H. & Sato, M. (2005) Scirtidae of the Oriental Region, Part 8. New Species and Additional Record of the Genus Hydrocyphon (Coleoptera, Scirtidae). Entomological Review of Japan, 60 (2), 153 - 206." type="journal article" year="2005">Yoshitomi & Satô 2005</bibRefCitation>
|
||
), although its placement in
|
||
<emphasis id="63ACEABCA321FF9119B415A06C2EFE6C" box="[904,1033,394,417]" italics="true" pageId="3" pageNumber="62">deflexicollis</emphasis>
|
||
-group does not seem to be well supported. The main difference between these groups is the anatomy of penis—in the
|
||
<emphasis id="63ACEABCA321FF911E2515876CBDFE09" box="[1049,1178,429,452]" italics="true" pageId="3" pageNumber="62">deflexicollis</emphasis>
|
||
-group, the parameroids are symmetrical, in contrast to asymmetrical prosthemes; the structural anatomy of the tegmina is similar in both groups.
|
||
<bibRefCitation id="35494B5FA321FF911AAB15D86946FDC4" author="Klausnitzer" box="[151,353,498,521]" pageId="3" pageNumber="62" refString="Klausnitzer, B. (1976) Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Helodidae. Deutsche EntomologischeZeitschrift, 23, 213 - 220." type="journal article" year="1976">Klausnitzer (1976)</bibRefCitation>
|
||
mentioned that
|
||
<taxonomicName id="96D84D2DA321FF91182C15D86A97FDC4" box="[528,688,498,521]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="63ACEABCA321FF91182C15D86A97FDC4" box="[528,688,498,521]" italics="true" pageId="3" pageNumber="62">H. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
has slightly asymmetrical prosthemes, a feature unusual in
|
||
<taxonomicName id="96D84D2DA321FF911F0A15D86DB4FDC4" box="[1334,1427,498,521]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="63ACEABCA321FF911F0A15D86DB4FDC4" box="[1334,1427,498,521]" italics="true" pageId="3" pageNumber="62">australis</emphasis>
|
||
</taxonomicName>
|
||
- group, nevertheless according to Klausnitzer, it may be regarded as a member of this group, with marked tendency to develop subtle asymmetry. In the author’s opinion, the species in question may also be regarded as a member of
|
||
<emphasis id="63ACEABCA321FF911F70161268F6FDBC" italics="true" pageId="3" pageNumber="62">deflexicollis</emphasis>
|
||
-group with prosthemes not as asymmetrical as usual in this group, and therefore artificially resembling species from the
|
||
<taxonomicName id="96D84D2DA321FF911AC01657697EFD59" box="[252,345,637,660]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="63ACEABCA321FF911AC01657697EFD59" box="[252,345,637,660]" italics="true" pageId="3" pageNumber="62">australis</emphasis>
|
||
</taxonomicName>
|
||
-group. Actually, a few oriental species closely resembling
|
||
<taxonomicName id="96D84D2DA321FF9119E316576CA7FD59" box="[991,1152,637,660]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="63ACEABCA321FF9119E316576CA7FD59" box="[991,1152,637,660]" italics="true" pageId="3" pageNumber="62">H. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
were recently included in the
|
||
<emphasis id="63ACEABCA321FF911AFD168A6965FD7A" box="[193,322,672,695]" italics="true" pageId="3" pageNumber="62">deflexicollis</emphasis>
|
||
-group (e.g.
|
||
<taxonomicName id="96D84D2DA321FF911BF0168A6BC9FD7A" authority="Yoshitomi & Sato, 2005" authorityName="Yoshitomi & Sato" authorityYear="2005" box="[460,1006,672,695]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="lusonensis">
|
||
<emphasis id="63ACEABCA321FF911BF0168A6AF2FD7A" box="[460,725,672,695]" italics="true" pageId="3" pageNumber="62">Hydrocyphon lusonensis</emphasis>
|
||
<bibRefCitation id="35494B5FA321FF9118E3168A6BC9FD7A" author="Yoshitomi" box="[735,1006,672,695]" pageId="3" pageNumber="62" refString="Yoshitomi, H. & Sato, M. (2005) Scirtidae of the Oriental Region, Part 8. New Species and Additional Record of the Genus Hydrocyphon (Coleoptera, Scirtidae). Entomological Review of Japan, 60 (2), 153 - 206." type="journal article" year="2005">Yoshitomi & Satô, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="96D84D2DA321FF9119C7168A6929FD14" authority="Yoshitomi & Sato, 2005" authorityName="Yoshitomi & Sato" authorityYear="2005" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="subcelatus">
|
||
<emphasis id="63ACEABCA321FF9119C7168A6D21FD7A" box="[1019,1286,672,695]" italics="true" pageId="3" pageNumber="62">Hydrocyphon subcelatus</emphasis>
|
||
<bibRefCitation id="35494B5FA321FF911F33168A6929FD14" author="Yoshitomi" pageId="3" pageNumber="62" refString="Yoshitomi, H. & Sato, M. (2005) Scirtidae of the Oriental Region, Part 8. New Species and Additional Record of the Genus Hydrocyphon (Coleoptera, Scirtidae). Entomological Review of Japan, 60 (2), 153 - 206." type="journal article" year="2005">Yoshitomi & Satô, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
). The other hint to exclude
|
||
<taxonomicName id="96D84D2DA321FF91180916E86AF2FD14" box="[565,725,706,729]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="bhutanensis">
|
||
<emphasis id="63ACEABCA321FF91180916E86AF2FD14" box="[565,725,706,729]" italics="true" pageId="3" pageNumber="62">H. bhutanensis</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<taxonomicName id="96D84D2DA321FF91190316E86BBBFD14" box="[831,924,706,729]" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="63ACEABCA321FF91190316E86BBBFD14" box="[831,924,706,729]" italics="true" pageId="3" pageNumber="62">australis</emphasis>
|
||
</taxonomicName>
|
||
-group is its geographical distribution. The
|
||
<taxonomicName id="96D84D2DA321FF911F5216E868E8FD31" class="Insecta" family="Scirtidae" genus="Hydrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="63ACEABCA321FF911F5216E868E8FD31" italics="true" pageId="3" pageNumber="62">australis</emphasis>
|
||
</taxonomicName>
|
||
-group seems to have its distributional centre in the Middle East and Eastern Mediterranean. The existence of a related species in the Oriental Region seem to be doubtful. It is highly plausible, that some of species-groups proposed in the most recent studies (especially the largest groups:
|
||
<emphasis id="63ACEABCA321FF9118D717006B4DFC8C" box="[747,874,810,833]" italics="true" pageId="3" pageNumber="62">pallidicollis</emphasis>
|
||
-group and
|
||
<emphasis id="63ACEABCA321FF9119D717006C4BFC8C" box="[1003,1132,810,833]" italics="true" pageId="3" pageNumber="62">deflexicollis</emphasis>
|
||
-group) are artificial assemblages of unrelated species.
|
||
</paragraph>
|
||
<paragraph id="516736AEA321FF911AFA17456AA3FC22" blockId="3.[151,1437,151,1007]" pageId="3" pageNumber="62">
|
||
<emphasis id="63ACEABCA321FF911AFA1745698DFC4A" bold="true" box="[198,426,879,903]" pageId="3" pageNumber="62">Acknowledgements.</emphasis>
|
||
I am greatly indebted to Dr. Ottó MERKL for allowing me to study the
|
||
<taxonomicName id="96D84D2DA321FF911EA4175A6CD0FC4A" box="[1176,1271,880,903]" class="Insecta" family="Scirtidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Scirtidae</taxonomicName>
|
||
holdings of the Hungarian Museum of Natural History (Budapest,
|
||
<collectingCountry id="29CF763EA321FF9118FA17B86B02FC64" box="[710,805,914,937]" name="Hungary" pageId="3" pageNumber="62">Hungary</collectingCountry>
|
||
) and to Mr. Christopher Majka (Nova
|
||
<collectingCountry id="29CF763EA321FF911EEF17B86D31FC64" box="[1235,1302,914,937]" name="United Kingdom" pageId="3" pageNumber="62">Scotia</collectingCountry>
|
||
Museum of Natural History, Halifax,
|
||
<collectingCountry id="29CF763EA321FF911B94179F69DDFC01" box="[424,506,949,972]" name="Canada" pageId="3" pageNumber="62">Canada</collectingCountry>
|
||
) for linguistic assistance. My studies on
|
||
<taxonomicName id="96D84D2DA321FF91198E179F6C36FC01" box="[946,1041,949,972]" class="Insecta" family="Scirtidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Scirtidae</taxonomicName>
|
||
were supported by Synthesys-grants (FR-TAF-890, BE-TAF-1969, AT-TAF-2276).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |