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<document ID-CLB-Dataset="272969" ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.73.e110233" ID-GBIF-Dataset="05e2f800-086d-4e82-93c3-a816091e6340" ID-Pensoft-Pub="2511-6428-2-137" ID-Pensoft-UUID="125D003AED0C534FB5909DF2AA72900D" ID-ZooBank="F14284E8457D49978287F6F91AC96285" ModsDocID="2511-6428-73-2-137" checkinTime="1697446522815" checkinUser="pensoft" docAuthor="Mey, Wolfram &amp; Wichard, Wilfried" docDate="2023" docId="D77A4F5B6192544591C3FC0133F85E8D" docLanguage="en" docName="ContribEntomol 73(2): 137-146" docOrigin="Contributions to Entomology 73 (2)" docPubDate="2023-10-16" docSource="http://dx.doi.org/10.3897/contrib.entomol.73.e110233" docTitle="Kinitocelis patrickmuelleri Mey &amp; Wichard 2023, sp. nov." docType="treatment" docVersion="2" id="125D003AED0C534FB5909DF2AA72900D" lastPageNumber="137" masterDocId="125D003AED0C534FB5909DF2AA72900D" masterDocTitle="Tarachoptera: The extinct and enigmatic cousins of Trichoptera and Lepidoptera, with descriptions of two new species" masterLastPageNumber="146" masterPageNumber="137" pageNumber="137" updateTime="1697447294263" updateUser="ExternalLinkService">
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<mods:title id="3EF8C8B39E670A6A0A602EB9CF4C64F7">Tarachoptera: The extinct and enigmatic cousins of Trichoptera and Lepidoptera, with descriptions of two new species</mods:title>
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<mods:namePart id="CB29B9BAD57CDE153E01418020408FB1">Mey, Wolfram</mods:namePart>
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<mods:affiliation id="BEB5D5C1F4269D3D57E93680E6C4CAA9">Museum fuer Naturkunde, Leibniz Institute of Evolution and Biodiversity Research at the Humboldt University, Invalidenstr. 43, D - 10115 Berlin, Germany</mods:affiliation>
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<mods:namePart id="2F187779BDA33E06856975DBABE2A04C">Wichard, Wilfried</mods:namePart>
<mods:nameIdentifier id="14C8BD46F9C7D9F11CCAA78EC78BDDA3" type="ORCID">https://orcid.org/0000-0002-5933-6292</mods:nameIdentifier>
<mods:affiliation id="C4A42BC64BBBFEE2C44DB021AADB3AD9">University of Koeln, Institute of Biology, Koeln, Germany</mods:affiliation>
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<mods:title id="C3B1553BE3DE8A9485EC90D85B902418">Contributions to Entomology</mods:title>
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<mods:date id="D00FBA1B470D0A4D9CAC211C9F20CD28">2023</mods:date>
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<mods:number id="73D6D0F562A64F1B089BC9BF233A8C4A">73</mods:number>
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<treatment id="D77A4F5B6192544591C3FC0133F85E8D" LSID="urn:lsid:zoobank.org:act:/DFE7DCA8-D788-4792-B7FE-F1CBF5034281" httpUri="http://treatment.plazi.org/id/D77A4F5B6192544591C3FC0133F85E8D" lastPageNumber="137" pageId="0" pageNumber="137">
<subSubSection id="D872CAB269ECFBAF34C0A7EC4AD0D0F0" pageId="0" pageNumber="137" type="nomenclature">
<paragraph id="92D7716D2D289FF98AE7450F6F54E7EA" pageId="0" pageNumber="137">
<taxonomicName id="B28A43B6A5A4DCDA3F2C8C3A1020F1A4" LSID="https://zoobank.org//DFE7DCA8-D788-4792-B7FE-F1CBF5034281" authority="Mey &amp; Wichard, 2023" authorityName="Mey &amp; Wichard" authorityYear="2023" class="Insecta" family="Tarachocelidae" genus="Kinitocelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kinitocelis patrickmuelleri" order="Tarachoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="sp. nov.">Kinitocelis patrickmuelleri</taxonomicName>
<taxonomicNameLabel id="517D0933A6D0C7589A591E97D61D47E4" pageId="0" pageNumber="137">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="41C95929CA9DF167F8ABD77BCF4E7205" pageId="0" pageNumber="137" type="description">
<paragraph id="B7C28D72AE2453CAF06EF8432B5346AA" pageId="0" pageNumber="137">
<figureCitation id="A80E232708177BD107BD81200E50E485" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Tarachoptera new species, 1, 2. Kinitocelis patrickmuelleri sp. nov., male holotype: 1. Head and antennae, dorsal view; 2. Tip of abdomen, left ventrolateral view. 3 - 5. Tarachocelis emmarossae sp. nov., male holotype: 3. Head, frontal view; 4. Right antenna; 5. Terminal segment with ventral comb, ventral view. Scale bar: 0.5 mm (1 - 4)." figureDoi="10.3897/contrib.entomol.73.e110233.figures1-5" httpUri="https://binary.pensoft.net/fig/921058" pageId="0" pageNumber="137">Figs 1</figureCitation>
<figureCitation id="047C243161A5D5DC7999F44D08D45603" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Tarachoptera new species, 1, 2. Kinitocelis patrickmuelleri sp. nov., male holotype: 1. Head and antennae, dorsal view; 2. Tip of abdomen, left ventrolateral view. 3 - 5. Tarachocelis emmarossae sp. nov., male holotype: 3. Head, frontal view; 4. Right antenna; 5. Terminal segment with ventral comb, ventral view. Scale bar: 0.5 mm (1 - 4)." figureDoi="10.3897/contrib.entomol.73.e110233.figures1-5" httpUri="https://binary.pensoft.net/fig/921058" pageId="0" pageNumber="137">, 2</figureCitation>
<figureCitation id="327942C03AE8303B59EDF2A9C298C9F3" captionStart="Figure 6" captionStartId="F2" captionText="Figure 6. Microphotographs of Kinitocelis patrickmuelleri sp. nov., male holotype. a. Male habitus, dorsal view; b. Abdominal tip, left ventrolateral view; c. Apical part of antenna; d. Basal part of antenna." figureDoi="10.3897/contrib.entomol.73.e110233.figure6" httpUri="https://binary.pensoft.net/fig/921059" pageId="0" pageNumber="137">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="529872E78915CEE067A302C60FD410D3" pageId="0" pageNumber="137" type="materials_examined">
<paragraph id="23FE81B0C8391CC54A21E41110491FD8" pageId="0" pageNumber="137">Material.</paragraph>
<paragraph id="13E60104FBB4EB495A70F4E003E17FB6" pageId="0" pageNumber="137">
<materialsCitation id="9EBC49A0B74FC1108E0EA65BFBC43A21" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Patrick Mueller" country="Germany" location="Germany" specimenCount="1" typeStatus="Holotype">
<emphasis id="A743C178BF6A34BDAA031D4D0D0CA46B" bold="true" italics="true" pageId="0" pageNumber="137">
<typeStatus id="265315B6411267DA4F75897C25017586">Holotype</typeStatus>
</emphasis>
, male, Burmese Amber, deposited in the Zoological Research Museum Alexander Koenig, Bonn,
<collectingCountry id="ACBDBF24B5227C663045BFB0AAF768AB" name="Germany">Germany</collectingCountry>
, inventory no.: ZFMK-TRI000838 (ex coll.
<collectorName id="77532CBD48426361EEA1B968093A65FB">
Patrick
<normalizedToken id="414497D1B9E6E5CEB5B9D06A90162754" originalValue="Müller">Mueller</normalizedToken>
</collectorName>
, BUB 4498).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AA624FA3B39B835596668744A32A158D" pageId="0" pageNumber="137" type="preservation">
<paragraph id="1C22EADA2E25E2EFF4AD87E9E9EEA094" pageId="0" pageNumber="137">Preservation.</paragraph>
<paragraph id="FC7D3DF4F6B8861300C965EB52CC4DC0" pageId="0" pageNumber="137">
The fossil is embedded in a flattened, oval piece of amber. The male is completely preserved, but in an unfavorable position with wings overlapping and covering the body (Fig.
<figureCitation id="D530924764E65FB3A405763C5D741806" captionStart="Figure 6" captionStartId="F2" captionText="Figure 6. Microphotographs of Kinitocelis patrickmuelleri sp. nov., male holotype. a. Male habitus, dorsal view; b. Abdominal tip, left ventrolateral view; c. Apical part of antenna; d. Basal part of antenna." figureDoi="10.3897/contrib.entomol.73.e110233.figure6" httpUri="https://binary.pensoft.net/fig/921059" pageId="0" pageNumber="137">6</figureCitation>
). Head, thorax, and abdomen partly macerated. Several air bubbles are present in the inclusion.
</paragraph>
</subSubSection>
<subSubSection id="49F677A9E72F35D7ECBC9BAEFD25D607" pageId="0" pageNumber="137" type="etymology">
<paragraph id="D14F4D3CB5705147C9E55FFCB0B5727A" pageId="0" pageNumber="137">Etymology.</paragraph>
<paragraph id="C42ADCB66568AF94FE0ACF191EA0EDAA" pageId="0" pageNumber="137">
The species is named in honor of Patrick
<normalizedToken id="CBBDFD9B56AF4872E6FF36BA1D741CD2" originalValue="Müller">Mueller</normalizedToken>
, collector and promoter of research on Burma amber inclusions.
</paragraph>
</subSubSection>
<subSubSection id="F7B6F3D9A81357703679C9AEDE8F861F" pageId="0" pageNumber="137" type="description">
<paragraph id="81D2EC0BC5E1ADA334C48BBF1CF548DB" pageId="0" pageNumber="137">Description.</paragraph>
<paragraph id="1DE25C4A67206F14E1F6F4DFF0C233C9" pageId="0" pageNumber="137">
Length of body 2.2 mm, forewings 2.9 mm; head with a triangular, frontal process; eyes hemispherical; scape and pedicellus slender, not as broad as following flagellomeres (Fig.
<figureCitation id="E4E8C5DBD50E42BBF412B7728E350744" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Tarachoptera new species, 1, 2. Kinitocelis patrickmuelleri sp. nov., male holotype: 1. Head and antennae, dorsal view; 2. Tip of abdomen, left ventrolateral view. 3 - 5. Tarachocelis emmarossae sp. nov., male holotype: 3. Head, frontal view; 4. Right antenna; 5. Terminal segment with ventral comb, ventral view. Scale bar: 0.5 mm (1 - 4)." figureDoi="10.3897/contrib.entomol.73.e110233.figures1-5" httpUri="https://binary.pensoft.net/fig/921058" pageId="0" pageNumber="137">1</figureCitation>
); 24 flagellomeres, broad, flat, quadrangular, with hairs or scales shorter than flagellomere diameter (Figs
<figureCitation id="B137E878277AAC6287B4F4DDFD5DDDCF" captionStart="Figure 6" captionStartId="F2" captionText="Figure 6. Microphotographs of Kinitocelis patrickmuelleri sp. nov., male holotype. a. Male habitus, dorsal view; b. Abdominal tip, left ventrolateral view; c. Apical part of antenna; d. Basal part of antenna." figureDoi="10.3897/contrib.entomol.73.e110233.figure6" httpUri="https://binary.pensoft.net/fig/921059" pageId="0" pageNumber="137">6c</figureCitation>
,
<figureCitation id="93298EE7C1E08794B0E8F4B29DCEA373" captionStart="Figure 6" captionStartId="F2" captionText="Figure 6. Microphotographs of Kinitocelis patrickmuelleri sp. nov., male holotype. a. Male habitus, dorsal view; b. Abdominal tip, left ventrolateral view; c. Apical part of antenna; d. Basal part of antenna." figureDoi="10.3897/contrib.entomol.73.e110233.figure6" httpUri="https://binary.pensoft.net/fig/921059" pageId="0" pageNumber="137">6d</figureCitation>
).
</paragraph>
<paragraph id="319D112D1A5E18258FAA95258C96904A" pageId="0" pageNumber="137">
<emphasis id="CE2A7DCE04062C25E582CB66F47FDC31" bold="true" italics="true" pageId="0" pageNumber="137">Male genitalia</emphasis>
(Figs
<figureCitation id="37FD1B5F92E61D8D26B7B0DEC841D8B3" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Tarachoptera new species, 1, 2. Kinitocelis patrickmuelleri sp. nov., male holotype: 1. Head and antennae, dorsal view; 2. Tip of abdomen, left ventrolateral view. 3 - 5. Tarachocelis emmarossae sp. nov., male holotype: 3. Head, frontal view; 4. Right antenna; 5. Terminal segment with ventral comb, ventral view. Scale bar: 0.5 mm (1 - 4)." figureDoi="10.3897/contrib.entomol.73.e110233.figures1-5" httpUri="https://binary.pensoft.net/fig/921058" pageId="0" pageNumber="137">2</figureCitation>
,
<figureCitation id="958B47869D7953178A4CA8B834D7A30B" captionStart="Figure 6" captionStartId="F2" captionText="Figure 6. Microphotographs of Kinitocelis patrickmuelleri sp. nov., male holotype. a. Male habitus, dorsal view; b. Abdominal tip, left ventrolateral view; c. Apical part of antenna; d. Basal part of antenna." figureDoi="10.3897/contrib.entomol.73.e110233.figure6" httpUri="https://binary.pensoft.net/fig/921059" pageId="0" pageNumber="137">6</figureCitation>
): dorsal plate long, tapering to round apex; ventral comb on sternum IX with 10 stiff spines.
</paragraph>
</subSubSection>
<subSubSection id="51727EF6023DB1987829480CF92708BC" pageId="0" pageNumber="137" type="diagnosis">
<paragraph id="D8AE6F1062B3C2A9ADC312E60821DE11" pageId="0" pageNumber="137">Diagnosis.</paragraph>
<paragraph id="3EE08FCF598308F43C3C730E602B8196" pageId="0" pageNumber="137">
By using the identification key of
<bibRefCitation id="C4FB7BE43872E215DC42A47049749147" DOI="https://doi.org/10.1016/j.cretres.2020.104681" author="Wichard, W" journalOrPublisher="Current Biology" pageId="0" pageNumber="137" refId="B14" refString="Wichard, W, Mey, W, 2021. Order Tarachoptera in mid-Cretaceous Burmese amber (Insecta, Amphiesmenoptera). Cretaceous Research 119: 104681. https://doi.org/10.1016/j.cretres.2020.104681" title="Order Tarachoptera in mid-Cretaceous Burmese amber (Insecta, Amphiesmenoptera). Cretaceous Research 119: 104681." url="https://doi.org/10.1016/j.cretres.2020.104681" year="2021">Wichard and Mey (2021)</bibRefCitation>
, the new species comes out as
<taxonomicName id="CAC0E001A866ECC8EE4C06945985BC0F" class="Insecta" family="Tarachocelidae" genus="Kinitocelis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Kinitocelis" order="Tarachoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="A08D946DBAF998FF5CE60EF4B4F7D8FA" italics="true" pageId="0" pageNumber="137">Kinitocelis</emphasis>
</taxonomicName>
. The new species can be distinguished from all congeners by the triangular, pointed anterior margin of the head and by the very broad flagellomeres of the antennae.
</paragraph>
</subSubSection>
</treatment>
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