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<taxonomicName id="9EC3D2A3FFAFFFAE8386FD4AFD83C3EF" ID-CoL="84W7R" authority="Gibbes, 1850" authorityName="Gibbes" authorityYear="1850" box="[151,538,711,737]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosus">
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<bibRefCitation id="3D52D4D1FFAFFFAE8268FD4AFD83C3EF" author="Gibbes" box="[377,538,711,737]" pageId="4" pageNumber="19" refString="Gibbes, L. R. (1850) On the carcinological collections of the cabinets of natural history in the United States: with an enumeration of the species contained therein and descriptions of new species. Proceedings of third meeting of American Association for Advancement of Science, 3, 165 - 201." type="journal article" year="1850">Gibbes, 1850</bibRefCitation>
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<figureCitation id="C1F8B5A5FFAFFFAE8386FD67FE9FC20A" box="[151,262,746,772]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="5.[151,255,1497,1521]" captionStartId-1="7.[151,255,1467,1491]" captionStartId-2="9.[151,255,1899,1923]" captionStartId-3="10.[151,255,1608,1632]" captionTargetBox-0="[172,1407,211,1464]" captionTargetBox-1="[155,1431,196,1402]" captionTargetBox-2="[198,1376,158,1870]" captionTargetBox-3="[177,1430,752,1577]" captionTargetId-0="figure@5.[172,1416,194,1473]" captionTargetId-1="figure@7.[151,1436,194,1443]" captionTargetId-2="figure@9.[191,1397,145,1883]" captionTargetId-3="figure@10.[151,1436,742,1584]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." captionText-1="FIGURE 2. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." captionText-2="FIGURE 3. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." captionText-3="FIGURE 4. Alpheus formosus Gibbes, 1850, neotype female from Big Pine Key, Florida Keys (USNM 1100697): a, frontal region, dorsal; b, major cheliped, mesial; c, same, detail of distomesial tooth; d, minor chela and carpus, mesial; e, detail of distolateral spine of uropod, dorsal. Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/180923/files/figure.png" httpUri-1="https://zenodo.org/record/180924/files/figure.png" httpUri-2="https://zenodo.org/record/180925/files/figure.png" httpUri-3="https://zenodo.org/record/180926/files/figure.png" pageId="4" pageNumber="5">Figs. 14</figureCitation>
,
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ce
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<treatmentCitation id="D8628F31FFAFFFAE800DFCB7FC72C25F" author="Heilprin" box="[796,1003,826,849]" page="322" pageId="4" pageNumber="5" year="1888">
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<bibRefCitation id="3D52D4D1FFAFFFAE80E9FCB7FB08C25F" author="Heilprin" box="[1016,1169,826,849]" pageId="4" pageNumber="20" refString="Heilprin, A. (1889) The Bermuda Islands: A contribution to the physical history and zoology of the Somers Archipelago. With an examination of the structure of coral reefs. Researches undertaken under the auspices of the Academy of Natural Sciences of Philadelphia, 1 - 231, pls. 1 - 17. Heilprin, Philadelphia." type="book chapter" year="1889">Heilprin 1889</bibRefCitation>
: 151
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(for page numbers see
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<bibRefCitation id="3D52D4D1FFAFFFAE8195FCD7FCB9C27F" author="Coutiere" box="[644,800,858,881]" pageId="4" pageNumber="19" refString="Coutiere, H. (1910) The snapping shrimps (Alpheidae) of the Dry Tortugas, Florida. Proceedings of the United States National Museum, 37, 485 - 487." type="journal article" year="1910">Coutière 1910</bibRefCitation>
: 485
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;
<treatmentCitation id="D8628F31FFAFFFAE8071FCD7FB80C27F" author="Verrill" box="[864,1049,858,881]" page="579" pageId="4" pageNumber="5" year="1900">
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: 579
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;
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: 152
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: 106
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;
<treatmentCitation id="D8628F31FFAFFFAE83EFFCF7FE32C29F" author="Verrill" box="[254,427,890,913]" page="84" pageId="4" pageNumber="5" year="1922">
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: 84
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<treatmentCitation id="D8628F31FFAFFFAE82A9FCF7FDF0C29F" author="Balss" box="[440,617,890,913]" page="177" pageId="4" pageNumber="5" year="1924">
<bibRefCitation id="3D52D4D1FFAFFFAE82A9FCF7FDABC29F" author="Balss" box="[440,562,890,913]" pageId="4" pageNumber="18" refString="Balss, H. (1924) Westindische Decapoden. Zoologischer Anzeiger, 61, 177 - 182." type="journal article" year="1924">Balss 1924</bibRefCitation>
: 177
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<treatmentCitation id="D8628F31FFAFFFAE8164FCF7FCABC29F" author="Boone" box="[629,818,890,913]" page="166" pageId="4" pageNumber="5" year="1930">
<bibRefCitation id="3D52D4D1FFAFFFAE8164FCF7FD62C29F" author="Boone" box="[629,763,890,913]" pageId="4" pageNumber="18" refString="Boone, L. (1930) Crustacea: Anomura, Macrura, Schizopoda, Isopoda, Amphipoda, Mysidacea, Cirripedia, and Copepoda. In: Scientific Results of Cruises of the yachts &quot; Eagle &quot; and &quot; Ara &quot;, 1921 - 1928, William K. Vanderbilt Commanding. Bulletin Vanderbilt Marine Museum, 3, 1 - 221, pls. 1 - 83." type="journal article" year="1930">Boone 1930</bibRefCitation>
: 166
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<treatmentCitation id="D8628F31FFAFFFAE802FFCF7FC6EC29F" author="Chace" box="[830,1015,890,913]" page="146" pageId="4" pageNumber="5" year="1956">
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: 82
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<treatmentCitation id="D8628F31FFAFFFAE87D7FCF7FE82C2BF" author="Bullis" page="8" pageId="4" pageNumber="5" year="1965">
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<bibRefCitation id="3D52D4D1FFAFFFAE8019FBF7FC54C59F" author="Posey" box="[776,973,1146,1169]" pageId="4" pageNumber="21" refString="Posey, M. H., Troy, D. A., Banner, S., Vose, F. &amp; Lindberg, W. (1998) Temporal variability, diversity and guild structure of a benthic community in the notheastern Gulf of Mexico. Bulletin of Marine Science, 63, 143 - 155." type="journal article" year="1998">
Posey
<emphasis id="6BB77532FFAFFFAE8043FBF7FC1AC59F" box="[850,899,1146,1169]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 1998
</bibRefCitation>
: 154
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE8705FBF7FABEC59F" author="Christoffersen" box="[1044,1319,1146,1169]" page="358" pageId="4" pageNumber="5" year="1998">
<bibRefCitation id="3D52D4D1FFAFFFAE8705FBF7FB77C59F" author="Christoffersen" box="[1044,1262,1146,1169]" pageId="4" pageNumber="19" refString="Christoffersen, M. L. (1998) Malacostraca. Eucarida. Caridea, Crangonoidea and Alpheoidea (except Glyphocrangonidae and Crangonidae). In Young, P. S. (ed.), Catalogue of Crustacea of Brazil, pp. 351 - 372. Museu Nacional, Rio de Janeiro." type="book chapter" year="1998">Christoffersen 1998</bibRefCitation>
: 358
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE8624FBF7FE18C5BF" author="Vargas" page="901" pageId="4" pageNumber="5" year="1999">
<bibRefCitation id="3D52D4D1FFAFFFAE8624FBF7FED5C5BF" author="Vargas" pageId="4" pageNumber="21" refString="Vargas, R. &amp; Cortes, J. (1999) Biodiversidad marina de Costa Rica: Crustacea: Decapoda (Penaeoidea, Sergestoidea, Stenopodidea, Caridea, Thalassinidea, Palinura) del Caribe. Revista de Biologia Tropical, 47, 887 - 911." type="journal article" year="1999">Vargas &amp; Cortés 1999</bibRefCitation>
: 901
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE829DFB17FD48C5BF" author="Morgado" box="[396,721,1178,1201]" page="177" pageId="4" pageNumber="5" year="2001">
<bibRefCitation id="3D52D4D1FFAFFFAE829DFB17FD05C5BF" author="Morgado" box="[396,668,1178,1201]" pageId="4" pageNumber="21" refString="Morgado, E. H. &amp; Tanaka, N. O. (2001) The macrofauna associated with the bryozoan Schizoporella errata (Walters) in southeastern Brazil. Scientia Marina, 65, 173 - 181." type="journal article" year="2001">Morgado &amp; Tanaka 2001</bibRefCitation>
: 177
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE81CDFB17FC55C5BF" author="Coelho" box="[732,972,1178,1201]" page="51" pageId="4" pageNumber="5" year="2006">
<bibRefCitation id="3D52D4D1FFAFFFAE81CDFB17FC3CC5BF" author="Coelho" box="[732,933,1178,1201]" pageId="4" pageNumber="19" refString="Coelho, P. A., de Almeida, A. O., de Souza-Filho, J. F., Bezerra, L. E. A. &amp; Giraldes, B. W. (2006) Diversity and distribution of the marine and estuarine shrimps (Dendrobranchiata, Stenopodidea and Caridea) from North and Northeast Brazil. Zootaxa, 1221, 41 - 62." type="journal article" year="2006">
Coelho
<emphasis id="6BB77532FFAFFFAE8021FB17FCC6C5BF" box="[816,863,1178,1201]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2006
</bibRefCitation>
: 51
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE80C9FB17FA91C5BF" author="Manning" box="[984,1288,1178,1201]" page="289" pageId="4" pageNumber="5" year="2006">
<bibRefCitation id="3D52D4D1FFAFFFAE80C9FB17FB48C5BF" author="Manning" box="[984,1233,1178,1201]" pageId="4" pageNumber="20" refString="Manning, R. B. &amp; Reed, K. (2006) Decapod crustaceans deposited in the zoological museum of Copenhagen by William Stimpson in 1859. Raffles Bulletin of Zoology, 54, 283 - 293." type="journal article" year="2006">Manning &amp; Reed 2006</bibRefCitation>
: 289
</treatmentCitation>
.*
</treatmentCitationGroup>
</paragraph>
<paragraph id="597CA920FFAFFFAE8386FB31FD33C5DD" blockId="4.[151,1437,826,1639]" box="[151,682,1210,1236]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="79D38E0EFFAFFFAE8386FB31FD33C5DD" box="[151,682,1210,1236]" pageId="4" pageNumber="5">
<taxonomicName id="9EC3D2A3FFAFFFAE8386FB31FECCC5DD" ID-CoL="84W7R" box="[151,341,1212,1235]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFAFFFAE8386FB31FECCC5DD" box="[151,341,1212,1235]" italics="true" pageId="4" pageNumber="5">Alpheus formosus</emphasis>
</taxonomicName>
-a—Knowlton &amp;
<treatmentCitation id="D8628F31FFAFFFAE8105FB31FD3FC5DD" author="Mills" box="[532,678,1212,1235]" page="2" pageId="4" pageNumber="5" year="1992">
<bibRefCitation id="3D52D4D1FFAFFFAE8105FB31FD15C5DD" author="Mills" box="[532,652,1212,1235]" pageId="4" pageNumber="20" refString="Knowlton, N. &amp; Mills D. K. (1992) The systematic importance of color and color pattern: evidence for complexes of sibling species of snapping shrimp (Caridea: Alpheidae: Alpheus) from the Caribbean and Pacific coasts of Panama. Proceedings of the San Diego Society of Natural History, 18, 1 - 5." type="journal article" year="1992">Mills 1992</bibRefCitation>
: 2
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="11D9FAABFFAFFFAE8386FB52FC01C435" pageId="4" pageNumber="5" type="description">
<paragraph id="597CA920FFAFFFAE8386FB52FD4EC5F8" blockId="4.[151,1437,826,1639]" box="[151,727,1245,1271]" pageId="4" pageNumber="5">
<taxonomicName id="9EC3D2A3FFAFFFAE8386FB52FECCC5F8" box="[151,341,1247,1270]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFAFFFAE8386FB52FECCC5F8" box="[151,341,1247,1270]" italics="true" pageId="4" pageNumber="5">Alpheus formosus</emphasis>
</taxonomicName>
sp. a—Knowlton
<emphasis id="6BB77532FFAFFFAE810AFB52FDD0C5F8" box="[539,585,1247,1270]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 1993: 1630.
</paragraph>
<paragraph id="597CA920FFAFFFAE8386FA8FFD3BC416" blockId="4.[151,1437,826,1639]" box="[151,674,1280,1306]" pageId="4" pageNumber="5">
<taxonomicName id="9EC3D2A3FFAFFFAE8386FA8FFECCC417" box="[151,341,1282,1305]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFAFFFAE8386FA8FFECCC417" box="[151,341,1282,1305]" italics="true" pageId="4" pageNumber="5">Alpheus formosus</emphasis>
</taxonomicName>
A—Williams
<emphasis id="6BB77532FFAFFFAE82E5FA8FFDBAC417" box="[500,547,1282,1305]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2001: 377.
</paragraph>
<paragraph id="597CA920FFAFFFAE8386FAA9FC01C435" blockId="4.[151,1437,826,1639]" box="[151,920,1314,1340]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="79D38E0EFFAFFFAE8386FAA9FC01C435" box="[151,920,1314,1340]" pageId="4" pageNumber="5">
<taxonomicName id="9EC3D2A3FFAFFFAE8386FAA9FED6C435" box="[151,335,1316,1339]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosis">
<emphasis id="6BB77532FFAFFFAE8386FAA9FED6C435" box="[151,335,1316,1339]" italics="true" pageId="4" pageNumber="5">Alpheus formosis</emphasis>
</taxonomicName>
(lap. cal.)—
<treatmentCitation id="D8628F31FFAFFFAE82C9FAA9FD32C435" author="McClure" box="[472,683,1316,1339]" page="142" pageId="4" pageNumber="5" year="2005">
<bibRefCitation id="3D52D4D1FFAFFFAE82C9FAA9FDEEC435" author="McClure" box="[472,631,1316,1339]" pageId="4" pageNumber="20" refString="McClure, M. (2005) Alpheidae. In: Hernandez Aguilera, J. L., Ruiz Nuno, J. A., Toral Almazan, R. E. &amp; Arenas Fuentes, V. (eds.), Camarones, langostas y cangrejos de la costa este de Mexico. Volume 1, pp. 119 - 202. Estudio y Conservacion de la Naturaleza, A. C. y CONABIO, Mexico. 350 pp, pls. 1 - 8." type="book chapter" year="2005">McClure 2005</bibRefCitation>
: 142
</treatmentCitation>
, fig. 14, pl. 4, fig. A.*
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="11D9FAABFFAFFFAE8386FACAFC7FC769" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="597CA920FFAFFFAE8386FACAFB33C471" blockId="4.[151,1437,826,1639]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="79D38E0EFFAFFFAE8386FACAFB33C471" pageId="4" pageNumber="5">
<taxonomicName id="9EC3D2A3FFAFFFAE8386FACAFEC6C450" box="[151,351,1351,1374]" class="Malacostraca" family="Crangonidae" genus="Crangon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFAFFFAE8386FACAFEC6C450" box="[151,351,1351,1374]" italics="true" pageId="4" pageNumber="5">Crangon formosus</emphasis>
</taxonomicName>
<treatmentCitation id="D8628F31FFAFFFAE826DFACAFD1AC450" author="Hay" box="[380,643,1351,1374]" page="384" pageId="4" pageNumber="5" year="1918">
<bibRefCitation id="3D52D4D1FFAFFFAE826DFACAFDD5C450" author="Hay" box="[380,588,1351,1374]" pageId="4" pageNumber="19" refString="Hay, W. P. &amp; Shore, C. A. (1918) The decapod crustaceans of Beaufort, North Carolina, and surrounding regions. Bulletin of Bureau of Fisheries, Washington, 35, 371 - 475, pls. 25 - 39." type="journal article" year="1918">Hay &amp; Shore 1918</bibRefCitation>
: 384
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE819EFACAFCCEC450" author="Schmitt" box="[655,855,1351,1374]" page="65" pageId="4" pageNumber="5" year="1924">
<bibRefCitation id="3D52D4D1FFAFFFAE819EFACAFCB6C450" author="Schmitt" box="[655,815,1351,1374]" pageId="4" pageNumber="21" refString="Schmitt, W. L. (1924 b) Report on Macrura, Anomura and Stomatopoda collected by the Barbados-Antigua Expedition from the University of Iowa in 1918. Studies of Natural History, Iowa University, 10, 65 - 99, pls. 1 - 5." type="journal article" year="1924" yearSuffix="a">Schmitt 1924a</bibRefCitation>
: 65
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE8075FACAFBB7C450" author="Schmitt" box="[868,1070,1351,1374]" page="73" pageId="4" pageNumber="5" year="1924">
<bibRefCitation id="3D52D4D1FFAFFFAE8075FACAFB9FC450" author="Schmitt" box="[868,1030,1351,1374]" pageId="4" pageNumber="21" refString="Schmitt, W. L. (1924 b) Report on Macrura, Anomura and Stomatopoda collected by the Barbados-Antigua Expedition from the University of Iowa in 1918. Studies of Natural History, Iowa University, 10, 65 - 99, pls. 1 - 5." type="journal article" year="1924" yearSuffix="b">Schmitt 1924b</bibRefCitation>
: 73
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE872AFACAFB61C450" author="Pearse" box="[1083,1272,1351,1374]" page="107" pageId="4" pageNumber="5" year="1932">
<bibRefCitation id="3D52D4D1FFAFFFAE872AFACAFB5BC450" author="Pearse" box="[1083,1218,1351,1374]" pageId="4" pageNumber="21" refString="Pearse, A. S. (1932) Observations on the parasites and commensals found associated with crustaceans and fishes at Dry Tortugas, Florida. Papers from the Tortugas Laboratory, Carnegie Institution of Washington, 28, 103 - 115." type="journal article" year="1932">Pearse 1932</bibRefCitation>
: 107
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE8615FACAFF6BC471" author="Schmitt" page="144" pageId="4" pageNumber="5" year="1935">
<bibRefCitation id="3D52D4D1FFAFFFAE8615FACAFA01C450" author="Schmitt" box="[1284,1432,1351,1374]" pageId="4" pageNumber="21" refString="Schmitt, W. L. (1935) Crustacea Macrura and Anomura of Porto Rico and the Virgin Islands. Science Survey of Porto Rico (New York Academy of Sciences), 15, 125 - 227." type="journal article" year="1935">Schmitt 1935</bibRefCitation>
: 144
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE83ECFAE5FE5CC471" author="Schmitt" box="[253,453,1384,1407]" page="368" pageId="4" pageNumber="5" year="1936">
<bibRefCitation id="3D52D4D1FFAFFFAE83ECFAE5FE09C471" author="Schmitt" box="[253,400,1384,1407]" pageId="4" pageNumber="21" refString="Schmitt, W. L. (1936) Macruran and anomuran Crustacea from Bonaire, Curacao and Aruba. Zoologische Ergebnisse einer Reise nach Bonaire, Curacao und Aruba im Jahre 1930. Nr 16. Zoologische Jahrbucher, Abteilung fur Systematik, Oekologie und Geographie der Tiere, 67, 363 - 378, pls. 11 - 13." type="journal article" year="1936">Schmitt 1936</bibRefCitation>
: 368
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE82C1FAE5FD12C471" author="Schmitt" box="[464,651,1384,1407]" page="28" pageId="4" pageNumber="5" year="1939">
<bibRefCitation id="3D52D4D1FFAFFFAE82C1FAE5FDFAC471" author="Schmitt" box="[464,611,1384,1407]" pageId="4" pageNumber="21" refString="Schmitt, W. L. (1939) Decapod and other Crustacea collected on the Presidential Cruise of 1938 (with introduction and station data). Smithsonian Miscellaneous Collections, 98, 1 - 29, 3 pls." type="journal article" year="1939">Schmitt 1939</bibRefCitation>
: 28
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE8187FAE5FCC9C471" author="Pearse" box="[662,848,1384,1407]" page="150" pageId="4" pageNumber="5" year="1950">
<bibRefCitation id="3D52D4D1FFAFFFAE8187FAE5FC82C471" author="Pearse" box="[662,795,1384,1407]" pageId="4" pageNumber="21" refString="Pearse, A. S. (1950) Notes on the inhabitants of certain sponges at Bimini. Ecology, 31, 149 - 151." type="journal article" year="1950">Pearse 1950</bibRefCitation>
: 150
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE804AFAE5FB03C471" author="Pearse" box="[859,1178,1384,1407]" page="143" pageId="4" pageNumber="5" year="1951">
<bibRefCitation id="3D52D4D1FFAFFFAE804AFAE5FBFDC471" author="Pearse" box="[859,1124,1384,1407]" pageId="4" pageNumber="21" refString="Pearse, A. S. &amp; Williams, L. G. (1951) The biota of the reefs off the Carolinas. Journal of Elisha Mitchell Scientific Society, 67, 133 - 161." type="journal article" year="1951">Pearse &amp; Williams 1951</bibRefCitation>
: 143
</treatmentCitation>
.*
</treatmentCitationGroup>
</paragraph>
<paragraph id="597CA920FFAFFFAE8386FA07FDCEC4AE" blockId="4.[151,1437,826,1639]" box="[151,599,1416,1442]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="79D38E0EFFAFFFAE8386FA07FDCEC4AE" box="[151,599,1416,1442]" pageId="4" pageNumber="5">
<taxonomicName id="9EC3D2A3FFAFFFAE8386FA07FEFEC4AF" box="[151,359,1418,1441]" class="Malacostraca" family="Crangonidae" genus="Crangon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosum">
<emphasis id="6BB77532FFAFFFAE8386FA07FEFEC4AF" box="[151,359,1418,1441]" italics="true" pageId="4" pageNumber="5">Crangon formosum</emphasis>
</taxonomicName>
<treatmentCitation id="D8628F31FFAFFFAE8295FA04FDDEC4AE" author="Manter" box="[388,583,1417,1440]" page="177" pageId="4" pageNumber="5" year="1933">
<bibRefCitation id="3D52D4D1FFAFFFAE8295FA04FD88C4AE" author="Manter" box="[388,529,1417,1440]" pageId="4" pageNumber="20" refString="Manter, W. H. (1933) The genus Helicometra and related trematodes from Tortugas, Florida. Papers from the Tortugas Laboratory, Carnegie Institution of Washington, 28, 169 - 180, 3 pls." type="journal article" year="1933">Manter 1933</bibRefCitation>
: 177
</treatmentCitation>
.*
</treatmentCitationGroup>
</paragraph>
<paragraph id="597CA920FFAFFFAE8386FA27FDF8C4CF" blockId="4.[151,1437,826,1639]" box="[151,609,1450,1473]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="79D38E0EFFAFFFAE8386FA27FDF8C4CF" box="[151,609,1450,1473]" pageId="4" pageNumber="5">
<taxonomicName id="9EC3D2A3FFAFFFAE8386FA27FDC4C4CF" ID-CoL="84LB5" authority="Poeyi Guerin-Meneville 1857: 19" authorityName="Poeyi Guerin-Meneville" authorityPageNumber="19" authorityYear="1857" box="[151,605,1450,1473]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="6BB77532FFAFFFAE8386FA27FEA9C4CF" box="[151,304,1450,1473]" italics="true" pageId="4" pageNumber="5">Alpheus Poeyi</emphasis>
<treatmentCitation id="D8628F31FFAFFFAE8226FA27FDC4C4CF" author="Guerin-Meneville" box="[311,605,1450,1473]" page="19" pageId="4" pageNumber="5" year="1857">
<bibRefCitation id="3D52D4D1FFAFFFAE8226FA27FDACC4CF" author="Guerin-Meneville" box="[311,565,1450,1473]" pageId="4" pageNumber="19" refString="Guerin-Meneville, F. E. (1857) Crustaces. Animaux articules a pieds articules. In: de la Sagra, R. (ed.), Histoire physique, politique et naturelle de l'ile de Cuba, pp. 1 - 87, pl. 2." type="journal article" year="1857">Guérin-Méneville 1857</bibRefCitation>
: 19
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="597CA920FFAFFFAE8386FA41FADFC4ED" blockId="4.[151,1437,826,1639]" box="[151,1350,1482,1508]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="79D38E0EFFAFFFAE8386FA41FADFC4ED" box="[151,1350,1482,1508]" pageId="4" pageNumber="5">
<taxonomicName id="9EC3D2A3FFAFFFAE8386FA41FEE8C4ED" ID-CoL="85LMF" box="[151,369,1484,1507]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="6BB77532FFAFFFAE8386FA41FEE8C4ED" box="[151,369,1484,1507]" italics="true" pageId="4" pageNumber="5">Alpheus panamensis</emphasis>
</taxonomicName>
(not
<emphasis id="6BB77532FFAFFFAE82B8FA41FE7DC4ED" box="[425,484,1484,1507]" italics="true" pageId="4" pageNumber="5">sensu</emphasis>
Kingsley 1878)—
<treatmentCitation id="D8628F31FFAFFFAE81BFFA41FCE8C4ED" author="Pocock" box="[686,881,1484,1507]" page="519" pageId="4" pageNumber="5" year="1890">
<bibRefCitation id="3D52D4D1FFAFFFAE81BFFA41FCA5C4ED" author="Pocock" box="[686,828,1484,1507]" pageId="4" pageNumber="21" refString="Pocock, R. I. (1890) Crustacea. In: Ridley, H. N. (ed.), Notes on the zoology of Fernando Noronha. Journal of the Linnean Society, Zoology, 20, 506 - 526." type="journal article" year="1890">Pocock 1890</bibRefCitation>
: 519
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE806DFA41FBDEC4ED" author="Zimmer" box="[892,1095,1484,1507]" page="391" pageId="4" pageNumber="5" year="1913">
<bibRefCitation id="3D52D4D1FFAFFFAE806DFA41FB8BC4ED" author="Zimmer" box="[892,1042,1484,1507]" pageId="4" pageNumber="22" refString="Zimmer, C. (1913) Westindische Decapoden, 1: Die Familie Alpheidae. Zoologische Jahrbucher, Supplement 11, 381 - 412." type="journal article" year="1913">Zimmer 1913</bibRefCitation>
: 391
</treatmentCitation>
;
<treatmentCitation id="D8628F31FFAFFFAE8743FA41FAADC4ED" author="Fausto-Filho" box="[1106,1332,1484,1507]" page="5" pageId="4" pageNumber="5" year="1974">
<bibRefCitation id="3D52D4D1FFAFFFAE8743FA41FA83C4ED" author="Fausto-Filho" box="[1106,1306,1484,1507]" pageId="4" pageNumber="19" refString="Fausto-Filho, J. (1974) Stomatopod and decapod crustaceans of the Archipelago of Fernando do Noronha, northeast Brazil. Arquivos de Ciencias do Mar, 14, 1 - 35." type="journal article" year="1974">Fausto-Filho 1974</bibRefCitation>
: 5
</treatmentCitation>
.*
</treatmentCitationGroup>
</paragraph>
<paragraph id="597CA920FFAFFFAE8386FA62FD77C729" blockId="4.[151,1437,826,1639]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="79D38E0EFFAFFFAE8386FA62FD77C729" pageId="4" pageNumber="5">
Not
<taxonomicName id="9EC3D2A3FFAFFFAE83D6FA62FE1FC708" ID-CoL="84W7R" box="[199,390,1519,1542]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFAFFFAE83D6FA62FE1FC708" box="[199,390,1519,1542]" italics="true" pageId="4" pageNumber="5">Alpheus formosus</emphasis>
</taxonomicName>
<treatmentCitation id="D8628F31FFAFFFAE82B5FA62FD29C708" author="Christoffersen" box="[420,688,1519,1542]" page="315" pageId="4" pageNumber="5" year="1979">
<bibRefCitation id="3D52D4D1FFAFFFAE82B5FA62FDE5C708" author="Christoffersen" box="[420,636,1519,1542]" pageId="4" pageNumber="19" refString="Christoffersen, M. L. (1979) Decapod Crustacea: Alpheoidea. Campagnes de la Calypso au large des cotes atlantiques de l'Amerique du Sud (1961 - 1962). I. Resultats Scientifiques de la Campagne de la Calypso, 11, 297 - 377." type="journal article" year="1979">Christoffersen 1979</bibRefCitation>
: 315
</treatmentCitation>
(part., specimens from E Pacific) (=
<taxonomicName id="9EC3D2A3FFAFFFAE8755FA62FA16C708" ID-CoL="85LMF" authority="Kingsley, 1878" authorityName="Kingsley" authorityYear="1878" box="[1092,1423,1519,1542]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="6BB77532FFAFFFAE8755FA62FB78C708" box="[1092,1249,1519,1542]" italics="true" pageId="4" pageNumber="5">A. panamensis</emphasis>
Kingsley, 1878
</taxonomicName>
);
<treatmentCitation id="D8628F31FFAFFFAE83D6F99DFEE3C729" author="Frische" box="[199,378,1552,1575]" page="38" pageId="4" pageNumber="5" year="2003">
<bibRefCitation id="3D52D4D1FFAFFFAE83D6F99DFECCC729" author="Frische" box="[199,341,1552,1575]" pageId="4" pageNumber="19" refString="Frische, J. (2003) Garnelen und Krebse im Meerwasseraquarium. Bede-Verlag, Ruhmannsfelden, 79 pp." type="book" year="2003">Frische 2003</bibRefCitation>
: 38
</treatmentCitation>
(color photograph, =
<taxonomicName id="9EC3D2A3FFAFFFAE8172F99DFD20C729" ID-CoL="84LB5" box="[611,697,1552,1575]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="6BB77532FFAFFFAE8172F99DFD20C729" box="[611,697,1552,1575]" italics="true" pageId="4" pageNumber="5">Alpheus</emphasis>
</taxonomicName>
sp.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="597CA920FFAFFFAE8386F9BDFC7FC769" blockId="4.[151,1437,826,1639]" pageId="4" pageNumber="5">
*Remark: Two species were previously confused under the name “
<taxonomicName id="9EC3D2A3FFAFFFAE8068F9BDFBA2C749" ID-CoL="84W7R" box="[889,1083,1584,1607]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFAFFFAE8068F9BDFBA2C749" box="[889,1083,1584,1607]" italics="true" pageId="4" pageNumber="5">Alpheus formosus</emphasis>
</taxonomicName>
”; therefore, some of the abovelisted records may actually refer to the new species and not to
<taxonomicName id="9EC3D2A3FFAFFFAE804EF9DDFC46C769" ID-CoL="84W7R" box="[863,991,1616,1639]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFAFFFAE804EF9DDFC46C769" box="[863,991,1616,1639]" italics="true" pageId="4" pageNumber="5">A. formosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="11D9FAABFFAFFFA28386F91AFF5BC2DC" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="597CA920FFAFFFAE8386F91AFEABC60C" blockId="4.[151,1436,1687,1873]" pageId="4" pageNumber="5">
<emphasis id="6BB77532FFAFFFAE8386F91AFEDFC7BF" bold="true" box="[151,326,1687,1713]" pageId="4" pageNumber="5">
<typeStatus id="86781782FFAFFFAE8386F91AFF4BC7BF" box="[151,210,1687,1713]" pageId="4" pageNumber="5">Type</typeStatus>
material
</emphasis>
.—
<collectingCountry id="21D4E9B0FFAFFFAE827DF915FE33C7BC" box="[364,426,1688,1714]" name="United States of America" pageId="4" pageNumber="5">USA</collectingCountry>
, Florida.
<typeStatus id="86781782FFAFFFAE810EF915FD1CC7BC" box="[543,645,1688,1714]" pageId="4" pageNumber="5" type="neotype">Neotype</typeStatus>
, ovig. female (CL 9.7),
<collectionCode id="3FD231E5FFAFFFAE80BDF915FB98C7BC" LSID="urn:lsid:biocol.org:col:34871" box="[940,1025,1688,1714]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5">USNM</collectionCode>
1100697,
<collectingCountry id="21D4E9B0FFAFFFAE8790F915FB26C7BC" box="[1153,1215,1688,1714]" name="United States of America" pageId="4" pageNumber="5">USA</collectingCountry>
, Florida, Big Pine Key, Bahia Honda State Park, shallow subtidal (less than
<quantity id="9E3B04C5FFAFFFAE805AF94DFC0DC7D4" box="[843,916,1728,1754]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="m" value="1.5">1.5 m</quantity>
), under rocks, coll. A. Anker,
<date id="2D7D8FE0FFAFFFAE8612F94DFA05C7D4" box="[1283,1436,1728,1754]" pageId="4" pageNumber="5" value="2004-04-18">18 Apr 2004</date>
[fcn 04-010].
</paragraph>
<paragraph id="597CA920FFAFFFAC83D7F89DFCA0C3D8" blockId="4.[151,1436,1687,1873]" lastBlockId="6.[151,1437,152,1988]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
Additional material examined.—
<collectingCountry id="21D4E9B0FFAFFFAE815BF89DFD30C624" box="[586,681,1808,1834]" name="Panama" pageId="4" pageNumber="5">Panama</collectingCountry>
(Caribbean coast).
<specimenCount id="4FC562A9FFAFFFAE8083F89DFC7BC624" box="[914,994,1808,1834]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
(CL 5.6),
<specimenCount id="4FC562A9FFAFFFAE8770F89DFB50C624" box="[1121,1225,1808,1834]" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
(CL 6.0),
<collectionCode id="3FD231E5FFAFFFAE8656F89DFA02C624" LSID="urn:lsid:biocol.org:col:34871" box="[1351,1435,1808,1834]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="5">USNM</collectionCode>
1100699, Bocas del Toro,
<collectingCountry id="21D4E9B0FFAFFFAE82DBF8BAFE6DC65F" box="[458,500,1847,1873]" name="Iceland" pageId="4" pageNumber="5">Isla</collectingCountry>
Bastimentos, shallow subtidal (
<quantity id="9E3B04C5FFAFFFAE807AF8BAFC56C65F" box="[875,975,1847,1873]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="4" pageNumber="5" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.51 m</quantity>
), from coral rocks, coll. A. Anker,
<date id="2D7D8FE0FFAFFFAF867BF8BAFE9DC7AB" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" value="2005-10-21">21 Oct 2005</date>
[fcn 05-103];
<specimenCount id="4FC562A9FFAEFFAF82BEF906FE67C7AB" box="[431,510,1675,1701]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(CL 7.8)
<collectionCode id="3FD231E5FFAEFFAF8165F906FD50C7AB" LSID="urn:lsid:biocol.org:col:34871" box="[628,713,1675,1701]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
1100700,
<collectingCountry id="21D4E9B0FFAEFFAF8054F906FCF6C7AB" box="[837,879,1675,1701]" name="Iceland" pageId="5" pageNumber="6">Isla</collectingCountry>
Grande, southern shore, shallow subtidal (
<quantity id="9E3B04C5FFAEFFAF8676F906FF46C7C2" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="5" pageNumber="6" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.5 1.5 m</quantity>
), from coral rubble, coll. A. Anker,
<date id="2D7D8FE0FFAEFFAF8181F93FFCB1C7C2" box="[656,808,1714,1740]" pageId="5" pageNumber="6" value="2006-04-21">21 Apr 2006</date>
[fcn 06-425];
<specimenCount id="4FC562A9FFAEFFAF80C6F93FFBD4C7C2" box="[983,1101,1714,1740]" pageId="5" pageNumber="6" type="juvenile">1 juvenile</specimenCount>
(CL not measured),
<collectionCode id="3FD231E5FFAEFFAF8656F93FFA05C7C2" LSID="urn:lsid:biocol.org:col:34871" box="[1351,1436,1714,1740]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
1100701,
<collectingCountry id="21D4E9B0FFAEFFAF821AF957FEACC7FA" box="[267,309,1754,1780]" name="Iceland" pageId="5" pageNumber="6">Isla</collectingCountry>
Grande, western point, from coral rocks, shallow subtidal (
<quantity id="9E3B04C5FFAEFFAF80FAF957FBD0C7FA" box="[1003,1097,1754,1780]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="6" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.51m</quantity>
), coll. A. Anker and C. Hurt,
<date id="2D7D8FE0FFAEFFAF8386F88CFE85C615" box="[151,284,1793,1819]" pageId="5" pageNumber="6" value="2005-10-06">6 Oct 2005</date>
[fcn 05-101];
<specimenCount id="4FC562A9FFAEFFAF82DBF88CFD83C615" box="[458,538,1793,1819]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(CL 6.9),
<collectionCode id="3FD231E5FFAEFFAF8188F88CFD77C615" LSID="urn:lsid:biocol.org:col:34871" box="[665,750,1793,1819]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
1100702, same collection data as previous specimen [fcn 06-392].
<collectingCountry id="21D4E9B0FFAEFFAF821BF8A5FE13C64C" box="[266,394,1832,1858]" name="Costa Rica" pageId="5" pageNumber="6">Costa Rica</collectingCountry>
(Caribbean coast).
<specimenCount id="4FC562A9FFAEFFAF8161F8A5FD53C64C" box="[624,714,1832,1858]" pageId="5" pageNumber="6" type="male">3 males</specimenCount>
(CL 5.8, 6.6, 6.5), 1 ovig. female (CL 6.7),
<collectionCode id="3FD231E5FFAEFFAF87C2F8A5FAB1C64C" LSID="urn:lsid:biocol.org:col:34871" box="[1235,1320,1832,1858]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
1100698, Cahuita, shallow subtidal (
<quantity id="9E3B04C5FFAEFFAF82C2F8C2FDD6C667" box="[467,591,1871,1897]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="5" pageNumber="6" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.51.5 m</quantity>
), from rock crevices and
<taxonomicName id="9EC3D2A3FFAEFFAF8091F8DDFC6AC667" box="[896,1011,1872,1897]" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="5" pageNumber="6" phylum="Chlorophyta" rank="genus">
<emphasis id="6BB77532FFAEFFAF8091F8DDFC6AC667" box="[896,1011,1872,1897]" italics="true" pageId="5" pageNumber="6">Halimeda</emphasis>
</taxonomicName>
clumps, coll. A. Anker, I.S. Wehrtmann and L. Harris,
<date id="2D7D8FE0FFAEFFAF8296F8FBFDD5C69E" box="[391,588,1910,1936]" pageId="5" pageNumber="6" value="2005-11-26" valueMax="2005-11-27" valueMin="2005-11-26">2627 Nov 2005</date>
[fcn 05-100];
<specimenCount id="4FC562A9FFAEFFAF81E2F8FBFCDBC69E" box="[755,834,1910,1936]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(CL 10.3), MNHN-Na 16385, same collection data as previous specimens [fcn 05-102].
<collectingCountry id="21D4E9B0FFAEFFAF8158F813FD0BC6B6" box="[585,658,1950,1976]" name="Aruba" pageId="5" pageNumber="6">Aruba</collectingCountry>
.
<specimenCount id="4FC562A9FFAEFFAF81B1F813FD63C6B6" box="[672,762,1950,1976]" pageId="5" pageNumber="6" type="male">2 males</specimenCount>
(CL 5.3, 6.3),
<specimenCount id="4FC562A9FFAEFFAF80B4F813FB93C6B6" box="[933,1034,1950,1976]" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
(CL 6.2), 1 ovig. female (CL 7.1), MNHN-Na 16360, Baby Beach, shallow subtidal (
<quantity id="9E3B04C5FFAEFFAF81E1F848FCCFC6D1" box="[752,854,1989,2015]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="6" unit="m" value="1.25" valueMax="1.5" valueMin="1.0">11.5 m</quantity>
), from crevices in coral rubble, coll. A. Anker,
<date id="2D7D8FE0FFAEFFAC869CF848FE90C1BC" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" value="2003-12-06">6 Dec 2003</date>
[fcn 03-011].
<collectingCountry id="21D4E9B0FFADFFAC82AAFF15FDD1C1BC" box="[443,584,152,178]" name="Guadeloupe" pageId="6" pageNumber="7">Guadeloupe</collectingCountry>
.
<specimenCount id="4FC562A9FFADFFAC8149FF15FD26C1BC" box="[600,703,152,178]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL not measured), MNHN-Na 13715, Le Moule, depth
<quantity id="9E3B04C5FFADFFAC8676FF15FA01C1BC" box="[1383,1432,152,178]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="2.0">2 m</quantity>
, coll. F. Fasquel,
<date id="2D7D8FE0FFADFFAC8247FF32FE70C1D7" box="[342,489,191,217]" pageId="6" pageNumber="7" value="1999-10-29">29 Oct 1999</date>
;
<specimenCount id="4FC562A9FFADFFAC82E4FF32FDDAC1D7" box="[501,579,191,217]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL not measured), MNHN-Na 16386, same collection data as for previous specimen.
<collectingCountry id="21D4E9B0FFADFFAC825DFF6BFE0DC00E" box="[332,404,230,256]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
.
<specimenCount id="4FC562A9FFADFFAC82B2FF6BFE64C00E" box="[419,509,230,256]" pageId="6" pageNumber="7" type="male">2 males</specimenCount>
(CL 6.7, 7.8), 1 ovig. female (CL 7.9),
<collectionCode id="3FD231E5FFADFFAC80C0FF6BFB86C00E" box="[977,1055,230,256]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
20755, LT 702, Atol das Rocas, between Piscina das Tartarugas and Piscina das Rocas, in pools, among calcareous algae, coll. P.S. Young, P.C. Paiva and A.A. Aguiar,
<date id="2D7D8FE0FFADFFAC82F8FEB9FDE2C040" box="[489,635,308,334]" pageId="6" pageNumber="7" value="2000-10-28">28 Oct 2000</date>
[
<specimenCount id="4FC562A9FFADFFAC819CFEB9FD45C040" box="[653,732,308,334]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
dissected];
<specimenCount id="4FC562A9FFADFFAC8079FEB9FC21C040" box="[872,952,308,334]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 5.7),
<collectionCode id="3FD231E5FFADFFAC8725FEB9FB1BC040" box="[1076,1154,308,334]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
20756, Atol das Rocas, Piscina do Cemitério, in calcareous algae, coll. P.S. Young
<emphasis id="6BB77532FFADFFAC805FFED1FC1BC07B" box="[846,898,348,373]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
.,
<date id="2D7D8FE0FFADFFAC8089FED1FB82C078" box="[920,1051,348,374]" pageId="6" pageNumber="7" value="2001-01-01">1 Jan 2001</date>
;
<specimenCount id="4FC562A9FFADFFAC8736FED1FBEEC078" box="[1063,1143,348,374]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 9.5), 1 ovig. female (CL 9.4),
<collectionCode id="3FD231E5FFADFFAC821BFE0EFEC1C093" box="[266,344,387,413]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
20757, Atol das Rocas, near Pedra do Garapira, in tide pool, coll. P.S. Young, P.C. Paiva and A.A. Aguiar,
<date id="2D7D8FE0FFADFFAC8226FE27FE55C0CA" box="[311,460,426,452]" pageId="6" pageNumber="7" value="2000-10-14">14 Oct 2000</date>
;
<specimenCount id="4FC562A9FFADFFAC82C9FE27FDB1C0CA" box="[472,552,426,452]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 11.0),
<collectionCode id="3FD231E5FFADFFAC81A0FE27FD66C0CA" box="[689,767,426,452]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
20758, Atol das Rocas, Piscina das Rocas, in calcareous alga, coll. P.S. Young
<emphasis id="6BB77532FFADFFAC82B3FE5CFE4EC0E4" box="[418,471,465,490]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
.,
<date id="2D7D8FE0FFADFFAC82FEFE5CFD11C0E5" box="[495,648,465,491]" pageId="6" pageNumber="7" value="2000-10-26">26 Oct 2000</date>
;
<specimenCount id="4FC562A9FFADFFAC8187FE5CFD71C0E5" box="[662,744,465,491]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL not measured), MNHN-Na 16359, Pernambuco, Rio Formozo, coll. A.C. Branner Hartt,
<date id="2D7D8FE0FFADFFAC8152FE74FD2FC31D" box="[579,694,505,531]" pageId="6" pageNumber="7" value="1875-12">Dec 1875</date>
[specimen lacking all pereiopods].
<collectingCountry id="21D4E9B0FFADFFAC8773FE74FB39C31D" box="[1122,1184,505,531]" name="United States of America" pageId="6" pageNumber="7">USA</collectingCountry>
, Florida.
<specimenCount id="4FC562A9FFADFFAC8602FE74FAFDC31D" box="[1299,1380,505,531]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 11.5), OUMNH-ZC
<date id="2D7D8FE0FFADFFAC8297FDADFE71C334" box="[390,488,544,570]" pageId="6" pageNumber="7" value="2007" valueMax="2013">2007-13</date>
-034,
<collectingCountry id="21D4E9B0FFADFFAC813DFDADFDF3C334" box="[556,618,544,570]" name="United States of America" pageId="6" pageNumber="7">USA</collectingCountry>
, Florida, near Marineland,
<quantity id="9E3B04C5FFADFFAC80B7FDADFC78C334" box="[934,993,544,570]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="km" value="5.0">5 km</quantity>
south of Sebastian Inlet, shallow subtidal, coll. J.A. Baeza,
<date id="2D7D8FE0FFADFFAC828EFDCAFDBEC36F" box="[415,551,583,609]" pageId="6" pageNumber="7" value="2007-07-15">15 Jul 2007</date>
[fcn 07-240; specimen somewhat dessicated during transport, rehydrated in EtOH].
<collectingCountry id="21D4E9B0FFADFFAC83EDFDE3FE63C386" box="[252,506,622,648]" name="British Virgin Islands" pageId="6" pageNumber="7">British Virgin Islands</collectingCountry>
.
<specimenCount id="4FC562A9FFADFFAC811BFDE3FDE7C386" box="[522,638,622,648]" pageId="6" pageNumber="7" type="juvenile">1 juvenile</specimenCount>
(CL not measured), Guana
<collectingCountry id="21D4E9B0FFADFFAC80DBFDE3FB8CC386" box="[970,1045,622,648]" name="Iceland" pageId="6" pageNumber="7">Island</collectingCountry>
, coll. Guana Marine Invertebrate Project, 2000 [fcn Vc 0600] [specimen missing minor cheliped, identification tentative based on length of adrostral furrows; specimen will be deposited in
<collectionCode id="3FD231E5FFADFFAC81C2FD31FCB7C3D8" LSID="urn:lsid:biocol.org:col:34802" box="[723,814,700,726]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="6" pageNumber="7">LACM</collectionCode>
].
</paragraph>
<caption id="0DBCF9A8FFAEFFAF8386FA54FCF1C756" httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="5" pageNumber="6" targetBox="[172,1407,211,1464]" targetPageId="5">
<paragraph id="597CA920FFAEFFAF8386FA54FCF1C756" blockId="5.[151,1436,1497,1624]" pageId="5" pageNumber="6">
<emphasis id="6BB77532FFAEFFAF8386FA54FE80C4FF" bold="true" box="[151,281,1497,1521]" pageId="5" pageNumber="6">FIGURE 1.</emphasis>
<taxonomicName id="9EC3D2A3FFAEFFAF8231FA57FDECC4FE" authority="Gibbes, 1850" authorityName="Gibbes" authorityYear="1850" box="[288,629,1497,1521]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFAEFFAF8231FA57FE47C4FF" box="[288,478,1498,1521]" italics="true" pageId="5" pageNumber="6">Alpheus formosus</emphasis>
Gibbes, 1850
</taxonomicName>
, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="597CA920FFADFFAC83D7FD6EFA05C51E" blockId="6.[151,1437,152,1988]" pageId="6" pageNumber="7">
<emphasis id="6BB77532FFADFFAC83D7FD6EFEEDC3F3" bold="true" box="[198,372,739,765]" pageId="6" pageNumber="7">Redescription</emphasis>
.—Carapace glabrous, non-setose, somewhat compressed laterally. Rostrum well developed, dorsally flattened; margins fringed with setae directed up- or sidewards (
<figureCitation id="C1F8B5A5FFADFFAC8729FC86FBE5C22B" box="[1080,1148,779,805]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a, b); tip acute, reaching to or slightly beyond distal margin of first segment of antennular peduncle (
<figureCitation id="C1F8B5A5FFADFFAC8720FCBFFBEFC242" box="[1073,1142,818,844]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a); rostral carina absent; orbito-rostral process broad, round W-shaped. Orbital hoods with acute teeth inserted at some distance from anterior margin (
<figureCitation id="C1F8B5A5FFADFFAC8276FC0CFE36C295" box="[359,431,897,923]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a). Adrostral furrows deep, narrow, abruptly delimited from rostrum, posteriorly extending almost to level of cornea base (
<figureCitation id="C1F8B5A5FFADFFAC816EFC25FD5BC2CC" box="[639,706,936,962]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a). Pterygostomial angle rounded, somewhat protruding anteriorly (
<figureCitation id="C1F8B5A5FFADFFAC83FBFC42FEB7C2E7" box="[234,302,975,1001]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
b, c); cardiac notch well developed. Eyes completely concealed in dorsal, lateral and partly in frontal view. Ocellar beak vertically protruding towards rostrum, more or less visible in lateral view (
<figureCitation id="C1F8B5A5FFADFFAC8628FC7BFAE2C51E" box="[1337,1403,1014,1040]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
c).
</paragraph>
<paragraph id="597CA920FFADFFAC83D7FB90FE31C47F" blockId="6.[151,1437,152,1988]" pageId="6" pageNumber="7">
Antennular peduncles relatively stout, second segment longer than dorsally visible portion of first, about 1.5 times as long as wide (
<figureCitation id="C1F8B5A5FFADFFAC82C3FBC9FD8CC550" box="[466,533,1092,1118]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a); stylocerite with acute tip, slightly overreaching distal margin of first segment (
<figureCitation id="C1F8B5A5FFADFFAC83F5FBE1FEB0C588" box="[228,297,1132,1158]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a, b); mesioventral carina of first segment with large, bluntly protruding tooth (
<figureCitation id="C1F8B5A5FFADFFAC87F2FBE1FAB1C588" box="[1251,1320,1132,1158]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
c); lateral flagellum with numerous tufts of aesthetascs, accessory ramus rudimentary (
<figureCitation id="C1F8B5A5FFADFFAC8726FB1EFBE7C5A3" box="[1079,1150,1171,1197]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
b). Antenna with stout basicerite bearing strong acute ventrolateral tooth, its tip not reaching level of stylocerite tip (
<figureCitation id="C1F8B5A5FFADFFAC87F2FB37FABFC5DA" box="[1251,1318,1210,1236]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
b); carpocerite stout, reaching slightly beyond distolateral spine of scaphocerite; scaphocerite with very strong distolateral tooth separated from narrow blade by deep cleft (
<figureCitation id="C1F8B5A5FFADFFAC8030FA84FCFEC42D" box="[801,871,1289,1315]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a), tip of distolateral tooth reaching far beyond blade and exceeding end of antennular peduncle (
<figureCitation id="C1F8B5A5FFADFFAC81C6FABDFC80C444" box="[727,793,1328,1354]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a); lateral margin of scaphocerite very slightly concave at mid-length (
<figureCitation id="C1F8B5A5FFADFFAC8255FADAFE1EC47F" box="[324,391,1367,1393]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a).
</paragraph>
<paragraph id="597CA920FFADFFAC83D7FAF3FB10C73B" blockId="6.[151,1437,152,1988]" pageId="6" pageNumber="7">
Mouthparts (mandible, maxillule, maxilla, first and second maxillipeds) typical for
<taxonomicName id="9EC3D2A3FFADFFAC8785FAF3FB6BC499" box="[1172,1266,1406,1431]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="6BB77532FFADFFAC8785FAF3FB6BC499" box="[1172,1266,1406,1431]" italics="true" pageId="6" pageNumber="7">Alpheus</emphasis>
</taxonomicName>
. Third maxilliped moderately stout; coxa with distally subacute lateral plate; exopod reaching penultimate segment; antepenultimate segment slightly flattened, ventral margin straight; penultimate segment about 2.5 times as long as wide; arthrobranch well developed (third maxilliped generally very similar to that of
<taxonomicName id="9EC3D2A3FFADFFAC87AEFA79FAF5C703" box="[1215,1388,1524,1549]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="6BB77532FFADFFAC87AEFA79FAF5C703" box="[1215,1388,1524,1549]" italics="true" pageId="6" pageNumber="7">A. panamensis</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="3D52D4D1FFADFFAC8386F996FE1DC73B" author="Kim" box="[151,388,1563,1589]" pageId="6" pageNumber="20" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
: fig. 15d, and to that of below-described new species, cf.
<figureCitation id="C1F8B5A5FFADFFAC8734F996FBF3C73B" box="[1061,1130,1563,1589]" captionStart="FIGURE 5" captionStartId="12.[151,255,1764,1788]" captionTargetBox="[152,1416,189,1743]" captionTargetId="figure@12.[152,1437,184,1757]" captionTargetPageId="12" captionText="FIGURE 5. Alpheus paraformosus n. sp., holotype male from Bocas del Toro, Panama (USNM 1100703) [a j]; paratype female from Bocas del Toro, Panama (USNM 1100704) [k]: a, frontal region, dorsal; b, same, lateral; c, scaphocerite, dorsal; d, third maxilliped, lateral; e, second pereiopod, lateral; f, third pereiopod, lateral; g, second pleopod, detail of appendix interna and appendix masculina; h, uropod, dorsal (setae omitted); i, distal margin of endopod (setae omitted); j, telson, dorsal; k, rostrum and orbital hoods, dorsal. Scale bars = 1 mm." httpUri="https://zenodo.org/record/180927/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
d).
</paragraph>
<paragraph id="597CA920FFADFFAC83D7F9CFFE18C6CA" blockId="6.[151,1437,152,1988]" pageId="6" pageNumber="7">
<collectingCountry id="21D4E9B0FFADFFAC83D7F9CFFE9BC752" box="[198,258,1602,1628]" name="Malta" pageId="6" pageNumber="7">Male</collectingCountry>
major cheliped (
<figureCitation id="C1F8B5A5FFADFFAC82D9F9CFFD97C752" box="[456,526,1602,1628]" captionStart="FIGURE 2" captionStartId="7.[151,255,1467,1491]" captionTargetBox="[155,1431,196,1402]" captionTargetId="figure@7.[151,1436,194,1443]" captionTargetPageId="7" captionText="FIGURE 2. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180924/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
) with short, stout ischium; merus stout, about 2.5 times as long as wide proximally, dorsal margin distally protruding (
<figureCitation id="C1F8B5A5FFADFFAC8197F9E7FD57C78A" box="[646,718,1642,1668]" captionStart="FIGURE 2" captionStartId="7.[151,255,1467,1491]" captionTargetBox="[155,1431,196,1402]" captionTargetId="figure@7.[151,1436,194,1443]" captionTargetPageId="7" captionText="FIGURE 2. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180924/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
a, c); ventrolateral margin straight, ventromesial margin rugose, with spinules, distally with acute tooth (
<figureCitation id="C1F8B5A5FFADFFAC81FEF91FFCD8C7A2" box="[751,833,1682,1708]" captionStart="FIGURE 2" captionStartId="7.[151,255,1467,1491]" captionTargetBox="[155,1431,196,1402]" captionTargetId="figure@7.[151,1436,194,1443]" captionTargetPageId="7" captionText="FIGURE 2. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180924/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
c); carpus very short, cup-shaped; chela large, subcylindrical, broadly oval in cross-section; palm smooth, moderately setose distally; distomesial margin with strong acute tooth (
<figureCitation id="C1F8B5A5FFADFFAC82ACF96FFD9CC7F2" box="[445,517,1762,1788]" captionStart="FIGURE 2" captionStartId="7.[151,255,1467,1491]" captionTargetBox="[155,1431,196,1402]" captionTargetId="figure@7.[151,1436,194,1443]" captionTargetPageId="7" captionText="FIGURE 2. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180924/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
b); linea impressa conspicuous (
<figureCitation id="C1F8B5A5FFADFFAC8094F96FFC57C7F2" box="[901,974,1762,1788]" captionStart="FIGURE 2" captionStartId="7.[151,255,1467,1491]" captionTargetBox="[155,1431,196,1402]" captionTargetId="figure@7.[151,1436,194,1443]" captionTargetPageId="7" captionText="FIGURE 2. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180924/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
a); adhesive discs relatively large (
<figureCitation id="C1F8B5A5FFADFFAC867CF96FFF3EC62A" captionStart="FIGURE 2" captionStartId="7.[151,255,1467,1491]" captionTargetBox="[155,1431,196,1402]" captionTargetId="figure@7.[151,1436,194,1443]" captionTargetPageId="7" captionText="FIGURE 2. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180924/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
a); fingers more than half-length of palm, dactylus usually reaching beyond pollex, with large plunger, latter distally with stamen-shaped sensillae (
<figureCitation id="C1F8B5A5FFADFFAC8176F8BFFD28C642" box="[615,689,1842,1868]" captionStart="FIGURE 2" captionStartId="7.[151,255,1467,1491]" captionTargetBox="[155,1431,196,1402]" captionTargetId="figure@7.[151,1436,194,1443]" captionTargetPageId="7" captionText="FIGURE 2. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180924/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
a). Female major cheliped similar to that of male, except for somewhat smaller size and different proportions of fingers to palm (
<figureCitation id="C1F8B5A5FFADFFAC80C4F8D7FBB9C67A" box="[981,1056,1882,1908]" captionStart="FIGURE 4" captionStartId="10.[151,255,1608,1632]" captionTargetBox="[177,1430,752,1577]" captionTargetId="figure@10.[151,1436,742,1584]" captionTargetPageId="10" captionText="FIGURE 4. Alpheus formosus Gibbes, 1850, neotype female from Big Pine Key, Florida Keys (USNM 1100697): a, frontal region, dorsal; b, major cheliped, mesial; c, same, detail of distomesial tooth; d, minor chela and carpus, mesial; e, detail of distolateral spine of uropod, dorsal. Scale bars = 1 mm." httpUri="https://zenodo.org/record/180926/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
b). Mesiodistal tooth on merus sometimes (
<figureCitation id="C1F8B5A5FFADFFAC8236F80FFEE8C692" box="[295,369,1922,1948]" captionStart="FIGURE 4" captionStartId="10.[151,255,1608,1632]" captionTargetBox="[177,1430,752,1577]" captionTargetId="figure@10.[151,1436,742,1584]" captionTargetPageId="10" captionText="FIGURE 4. Alpheus formosus Gibbes, 1850, neotype female from Big Pine Key, Florida Keys (USNM 1100697): a, frontal region, dorsal; b, major cheliped, mesial; c, same, detail of distomesial tooth; d, minor chela and carpus, mesial; e, detail of distolateral spine of uropod, dorsal. Scale bars = 1 mm." httpUri="https://zenodo.org/record/180926/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
b, c), independently of sex, with two acute teeth: one larger and stronger tooth, and one much smaller tooth.
</paragraph>
<caption id="0DBCF9A8FFACFFAD8386FA36FAE0C4F7" httpUri="https://zenodo.org/record/180924/files/figure.png" pageId="7" pageNumber="8" targetBox="[155,1431,196,1402]" targetPageId="7">
<paragraph id="597CA920FFACFFAD8386FA36FAE0C4F7" blockId="7.[151,1436,1467,1529]" pageId="7" pageNumber="8">
<emphasis id="6BB77532FFACFFAD8386FA36FE83C4DD" bold="true" box="[151,282,1467,1491]" pageId="7" pageNumber="8">FIGURE 2.</emphasis>
<taxonomicName id="9EC3D2A3FFACFFAD8233FA31FDE5C4DD" authority="Gibbes, 1850" authorityName="Gibbes" authorityYear="1850" box="[290,636,1468,1491]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFACFFAD8233FA31FE78C4DD" box="[290,481,1468,1491]" italics="true" pageId="7" pageNumber="8">Alpheus formosus</emphasis>
Gibbes, 1850
</taxonomicName>
, male from Atol das Rocas, Brazil (MNRJ 20755): a, major cheliped, lateral; b, same, distal portion of chela, mesial; c, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="597CA920FFACFFAD83D7F9A2FC20C637" blockId="7.[151,1437,1583,2009]" pageId="7" pageNumber="8">
<collectingCountry id="21D4E9B0FFACFFAD83D7F9A2FE9BC747" box="[198,258,1583,1609]" name="Malta" pageId="7" pageNumber="8">Male</collectingCountry>
minor cheliped (
<figureCitation id="C1F8B5A5FFACFFAD82C0F9A2FD80C747" box="[465,537,1583,1609]" captionStart="FIGURE 3" captionStartId="9.[151,255,1899,1923]" captionTargetBox="[198,1376,158,1870]" captionTargetId="figure@9.[191,1397,145,1883]" captionTargetPageId="9" captionText="FIGURE 3. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180925/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
) with short, stout ischium; merus about three times as long as wide proximally; ventrolateral margin straight; ventromesial margin somewhat rugose, usually with spinules; distally with acute tooth (
<figureCitation id="C1F8B5A5FFACFFAD8272F9F2FE31C797" box="[355,424,1663,1689]" captionStart="FIGURE 3" captionStartId="9.[151,255,1899,1923]" captionTargetBox="[198,1376,158,1870]" captionTargetId="figure@9.[191,1397,145,1883]" captionTargetPageId="9" captionText="FIGURE 3. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180925/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
d); carpus cup-shaped; chela smooth, ovate in cross-section, somewhat setose mesially (
<figureCitation id="C1F8B5A5FFACFFAD838EF92AFF7EC7CF" box="[159,231,1703,1729]" captionStart="FIGURE 3" captionStartId="9.[151,255,1899,1923]" captionTargetBox="[198,1376,158,1870]" captionTargetId="figure@9.[191,1397,145,1883]" captionTargetPageId="9" captionText="FIGURE 3. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180925/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
b); distomesial margin of palm with strong acute tooth (
<figureCitation id="C1F8B5A5FFACFFAD8096F92AFC57C7CF" box="[903,974,1703,1729]" captionStart="FIGURE 3" captionStartId="9.[151,255,1899,1923]" captionTargetBox="[198,1376,158,1870]" captionTargetId="figure@9.[191,1397,145,1883]" captionTargetPageId="9" captionText="FIGURE 3. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180925/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
a, b); linea impressa conspicuous (
<figureCitation id="C1F8B5A5FFACFFAD867CF92AFF3FC7E7" captionStart="FIGURE 3" captionStartId="9.[151,255,1899,1923]" captionTargetBox="[198,1376,158,1870]" captionTargetId="figure@9.[191,1397,145,1883]" captionTargetPageId="9" captionText="FIGURE 3. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180925/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
a); adhesive discs small (
<figureCitation id="C1F8B5A5FFACFFAD82C0F942FD81C7E7" box="[465,536,1743,1769]" captionStart="FIGURE 3" captionStartId="9.[151,255,1899,1923]" captionTargetBox="[198,1376,158,1870]" captionTargetId="figure@9.[191,1397,145,1883]" captionTargetPageId="9" captionText="FIGURE 3. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180925/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
a); fingers about as long as palm, with well developed rows of balaeniceps setae (
<figureCitation id="C1F8B5A5FFACFFAD83F3F97AFEB1C61F" box="[226,296,1783,1809]" captionStart="FIGURE 3" captionStartId="9.[151,255,1899,1923]" captionTargetBox="[198,1376,158,1870]" captionTargetId="figure@9.[191,1397,145,1883]" captionTargetPageId="9" captionText="FIGURE 3. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm." httpUri="https://zenodo.org/record/180925/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
ac), tips strongly curved, crossing when chela closed. Female minor cheliped similar to that of male; chela with well developed rows of balaeniceps setae (
<figureCitation id="C1F8B5A5FFACFFAD8045F892FC02C637" box="[852,923,1823,1849]" captionStart="FIGURE 4" captionStartId="10.[151,255,1608,1632]" captionTargetBox="[177,1430,752,1577]" captionTargetId="figure@10.[151,1436,742,1584]" captionTargetPageId="10" captionText="FIGURE 4. Alpheus formosus Gibbes, 1850, neotype female from Big Pine Key, Florida Keys (USNM 1100697): a, frontal region, dorsal; b, major cheliped, mesial; c, same, detail of distomesial tooth; d, minor chela and carpus, mesial; e, detail of distolateral spine of uropod, dorsal. Scale bars = 1 mm." httpUri="https://zenodo.org/record/180926/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
d).
</paragraph>
<paragraph id="597CA920FFACFFA283D7F8CAFE58C05C" blockId="7.[151,1437,1583,2009]" lastBlockId="8.[151,1437,152,2018]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Second pereiopod with ischium subequal to merus (
<figureCitation id="C1F8B5A5FFACFFAD8021F8CAFCEDC66F" box="[816,884,1863,1889]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
d); carpus five-segmented, ratio of carpal segments (from proximal to distal) approximately: 6/3/1.5/1.5/3 (
<figureCitation id="C1F8B5A5FFACFFAD8098F8E2FC56C687" box="[905,975,1903,1929]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
d); chela simple, with fingers slightly longer than palm. Third pereiopod with ischium bearing small ventrolateral spine (
<figureCitation id="C1F8B5A5FFACFFAD8760F81AFB2CC6BF" box="[1137,1205,1943,1969]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
e); merus unarmed, about six times as long as wide; carpus unarmed; propodus with stout spines or pairs of spines along ventral margin and two spines distoventrally near articulation with dactylus (
<figureCitation id="C1F8B5A5FFA3FFA280C8FF15FB87C1BC" box="[985,1054,152,178]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
e); dactylus about 1/3 propodus length, simple, conical, gradually curved towards acute tip, with some setae (
<figureCitation id="C1F8B5A5FFA3FFA2870DFF4DFBC6C1D4" box="[1052,1119,192,218]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
e). Fourth pereiopod similar to third pereiopod. Fifth pereiopod smaller and slenderer than third and fourth pereiopods; ischium unarmed; propodus with row of spines and at least eight rows of grooming setae distolaterally; dactylus conical, as in third pereiopod.
</paragraph>
<paragraph id="597CA920FFA3FFA283D7FEEDFBD0C204" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="9">
Abdominal pleura with posteroventral margins broadly rounded; sixth somite without articulated flap, dorsolateral projections on each side of telson bluntly rounded.
<collectingCountry id="21D4E9B0FFA3FFA28083FE05FC57C0AC" box="[914,974,392,418]" name="Malta" pageId="8" pageNumber="9">Male</collectingCountry>
second pleopod with appendix masculina reaching somewhat beyond appendix interna, with numerous slender apical and subapical setae. Uropod with protopod bearing elongate acute distolateral tooth (
<figureCitation id="C1F8B5A5FFA3FFA28022FE55FCE1C0FC" box="[819,888,472,498]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
f); exopod with strong acute distolateral tooth adjacent to distolateral spine; diaeresis bearing strong lateral tooth, mesial to distolateral spine (
<figureCitation id="C1F8B5A5FFA3FFA28609FD8DFAC7C314" box="[1304,1374,512,538]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
f, g); distolateral spine stout, usually black to tan-brown (
<figureCitation id="C1F8B5A5FFA3FFA281E7FDA5FCD1C34C" box="[758,840,552,578]" captionStart="FIGURE 4" captionStartId="10.[151,255,1608,1632]" captionTargetBox="[177,1430,752,1577]" captionTargetId="figure@10.[151,1436,742,1584]" captionTargetPageId="10" captionText="FIGURE 4. Alpheus formosus Gibbes, 1850, neotype female from Big Pine Key, Florida Keys (USNM 1100697): a, frontal region, dorsal; b, major cheliped, mesial; c, same, detail of distomesial tooth; d, minor chela and carpus, mesial; e, detail of distolateral spine of uropod, dorsal. Scale bars = 1 mm." httpUri="https://zenodo.org/record/180926/files/figure.png" pageId="8" pageNumber="9">Figs. 4</figureCitation>
e, 7e), sometimes pale in alcohol (
<figureCitation id="C1F8B5A5FFA3FFA287C7FDA5FA81C34C" box="[1238,1304,552,578]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
f, g); distal margins of endopod with row of spinules (
<figureCitation id="C1F8B5A5FFA3FFA281A3FDDDFD62C364" box="[690,763,592,618]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">
Fig.
<quantity id="9E3B04C5FFA3FFA281FEFDDDFD62C364" box="[751,763,592,618]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="8" pageNumber="9" unit="g" value="1.0">1</quantity>
</figureCitation>
g). Telson moderately broad, subrectangular, tapering towards posterior margin (
<figureCitation id="C1F8B5A5FFA3FFA282C9FDF5FD84C39C" box="[472,541,632,658]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
f); dorsal surface without median groove, with two pairs of strong dorsal spines, situated at some distance from lateral margins, both pairs situated in posterior 2/3 length of telson (
<figureCitation id="C1F8B5A5FFA3FFA2867DFD2DFF3BC3EC" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
f); posterior margin convex centrally, with two pairs of posterolateral spines, mesial being much longer than lateral (
<figureCitation id="C1F8B5A5FFA3FFA283E1FD7DFEABC204" box="[240,306,752,778]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
f); anal tubercles well developed. Gill formula typical for
<taxonomicName id="9EC3D2A3FFA3FFA280F2FD7DFBD8C207" box="[995,1089,752,777]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="6BB77532FFA3FFA280F2FD7DFBD8C207" box="[995,1089,752,777]" italics="true" pageId="8" pageNumber="9">Alpheus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="597CA920FFA3FFA283D7FC9AFF5BC2DC" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="9">
<emphasis id="6BB77532FFA3FFA283D7FC9AFF60C23F" bold="true" box="[198,249,791,817]" pageId="8" pageNumber="9">Size</emphasis>
.—The largest Caribbean specimen is a male from Cahuita,
<collectingCountry id="21D4E9B0FFA3FFA280DFFC95FBCDC23C" box="[974,1108,792,818]" name="Costa Rica" pageId="8" pageNumber="9">Costa Rica</collectingCountry>
, with CL
<quantity id="9E3B04C5FFA3FFA287DEFC95FAA3C23C" box="[1231,1338,792,818]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="9" unit="mm" value="10.3">10.3 mm</quantity>
and TL
<quantity id="9E3B04C5FFA3FFA28386FCCDFE9EC254" box="[151,263,832,858]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" pageId="8" pageNumber="9" unit="mm" value="33.5">33.5 mm</quantity>
; the CL of remaining adult specimens range from
<quantity id="9E3B04C5FFA3FFA2806BFCCDFC41C254" box="[890,984,832,858]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="8" pageNumber="9" unit="mm" value="5.3">5.3 mm</quantity>
to
<quantity id="9E3B04C5FFA3FFA28717FCCDFBF1C254" box="[1030,1128,832,858]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="8" pageNumber="9" unit="mm" value="7.8">7.8 mm</quantity>
. The CL of the
<typeStatus id="86781782FFA3FFA2862EFCCDFA05C254" box="[1343,1436,832,858]" pageId="8" pageNumber="9" type="neotype">neotype</typeStatus>
(ovigerous female) from Florida Keys is
<quantity id="9E3B04C5FFA3FFA28166FCE5FD4DC28C" box="[631,724,872,898]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="8" pageNumber="9" unit="mm" value="9.7">9.7 mm</quantity>
; the maximum CL of Florida specimens was
<quantity id="9E3B04C5FFA3FFA287F9FCE5FAC9C28C" box="[1256,1360,872,898]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="8" pageNumber="9" unit="mm" value="14.7">14.7 mm</quantity>
(
<bibRefCitation id="3D52D4D1FFA3FFA28671FCE5FE94C2A4" author="Hendrix" pageId="8" pageNumber="20" refString="Hendrix, G. Y. (1971) A systematic study of the genus Alpheus (Crustacea: Decapoda: Alpheidae) in South Florida. PhD Dissertation, University of Miami, Coral Gables, Florida, USA. 184 pp., 24 pls." type="book" year="1971">Hendrix 1971</bibRefCitation>
). The largest specimen from
<collectingCountry id="21D4E9B0FFA3FFA2817DFC1DFD2DC2A4" box="[620,692,912,938]" name="Brazil" pageId="8" pageNumber="9">Brazil</collectingCountry>
is a male from Atol das Rocas with 11.0 mm CL and
<quantity id="9E3B04C5FFA3FFA28622FC1DFA05C2A4" box="[1331,1436,912,938]" metricMagnitude="-2" metricUnit="m" metricValue="3.57" pageId="8" pageNumber="9" unit="mm" value="35.7">35.7 mm</quantity>
TL.
</paragraph>
</subSubSection>
<subSubSection id="11D9FAABFFA3FFA283D7FC52FC7DC7AC" pageId="8" pageNumber="9" type="description">
<paragraph id="597CA920FFA3FFA283D7FC52FD2DC46C" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="9">
<emphasis id="6BB77532FFA3FFA283D7FC52FEEDC2F7" bold="true" box="[198,372,991,1017]" pageId="8" pageNumber="9">Color pattern</emphasis>
.—Body red-brown to purple-brown, with more or less intense yellow mediodorsal band extending from rostrum tip to posterior margin of sixth abdominal somite; carapace with white lateral band extending from anterolateral margin to cardiac notch, continued by zigzag white band ending at sixth abdominal somite; rostrum entirely yellow, usually not bordered by reddish brown on margins; carapace flanks below white band pale; telson and uropods red-brown with large yellow patches and spots; walking legs and second pereiopods bluish, sometimes intense magenta blue; antennular peduncles yellow-brown; antennal scaphocerite brown with white areas, flagella pale bluish; chelipeds pale brown to intense orange brown, major chela with pale patch near articulation with dactylus; dactylus and pollex distally pinkish; minor chela mostly pale brown to orange brown (
<figureCitation id="C1F8B5A5FFA3FFA28159FAADFD17C434" box="[584,654,1312,1338]" captionStart="FIGURE 7" captionStartId="14.[151,255,1823,1847]" captionTargetBox="[151,1436,317,1795]" captionTargetId="figure@14.[151,1436,317,1797]" captionTargetPageId="14" captionText="FIGURE 7. Color patterns of species of the Alpheus formosus Gibbes, 1850 complex: Alpheus panamensis Kingsley, 1878 [a, b]; A. formosus Gibbes, 1850 [c e]; A. paraformosus n. sp. [f h]: a, male from Río Mar, Panama (USNM 1100693); b, ovigerous female from La Chorrera, Panama (not preserved); c, male from Isla Grande, Panama (USNM 1100700); d, ovigerous female from Cahuita, Costa Rica (USNM 1100698); e, neotype female from Big Pine Key, Florida Keys (USNM 1100697); f, g, paratype ovigerous female from Bocas del Toro, Panama (USNM 1100705); h, holotype male from Bocas del Toro, Panama (USNM 1100703)." httpUri="https://zenodo.org/record/180929/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
ce); distolateral spine on uropdal exopod usually brown to almost black (
<figureCitation id="C1F8B5A5FFA3FFA283F7FAC5FEB4C46C" box="[230,301,1352,1378]" captionStart="FIGURE 7" captionStartId="14.[151,255,1823,1847]" captionTargetBox="[151,1436,317,1795]" captionTargetId="figure@14.[151,1436,317,1797]" captionTargetPageId="14" captionText="FIGURE 7. Color patterns of species of the Alpheus formosus Gibbes, 1850 complex: Alpheus panamensis Kingsley, 1878 [a, b]; A. formosus Gibbes, 1850 [c e]; A. paraformosus n. sp. [f h]: a, male from Río Mar, Panama (USNM 1100693); b, ovigerous female from La Chorrera, Panama (not preserved); c, male from Isla Grande, Panama (USNM 1100700); d, ovigerous female from Cahuita, Costa Rica (USNM 1100698); e, neotype female from Big Pine Key, Florida Keys (USNM 1100697); f, g, paratype ovigerous female from Bocas del Toro, Panama (USNM 1100705); h, holotype male from Bocas del Toro, Panama (USNM 1100703)." httpUri="https://zenodo.org/record/180929/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
e); eggs orange-reddish (
<figureCitation id="C1F8B5A5FFA3FFA2815EFAC5FD0CC46C" box="[591,661,1352,1378]" captionStart="FIGURE 7" captionStartId="14.[151,255,1823,1847]" captionTargetBox="[151,1436,317,1795]" captionTargetId="figure@14.[151,1436,317,1797]" captionTargetPageId="14" captionText="FIGURE 7. Color patterns of species of the Alpheus formosus Gibbes, 1850 complex: Alpheus panamensis Kingsley, 1878 [a, b]; A. formosus Gibbes, 1850 [c e]; A. paraformosus n. sp. [f h]: a, male from Río Mar, Panama (USNM 1100693); b, ovigerous female from La Chorrera, Panama (not preserved); c, male from Isla Grande, Panama (USNM 1100700); d, ovigerous female from Cahuita, Costa Rica (USNM 1100698); e, neotype female from Big Pine Key, Florida Keys (USNM 1100697); f, g, paratype ovigerous female from Bocas del Toro, Panama (USNM 1100705); h, holotype male from Bocas del Toro, Panama (USNM 1100703)." httpUri="https://zenodo.org/record/180929/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
d).
</paragraph>
<paragraph id="597CA920FFA3FFA283D7FAE2FC7DC7AC" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="9">
<emphasis id="6BB77532FFA3FFA283D7FAE2FEB1C487" bold="true" box="[198,296,1391,1417]" pageId="8" pageNumber="9">Ecology</emphasis>
.—Intertidal to about
<quantity id="9E3B04C5FFA3FFA2813FFAFDFDF4C484" box="[558,621,1392,1418]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="m" value="40.0">40 m</quantity>
(
<bibRefCitation id="3D52D4D1FFA3FFA2816FFAFDFC80C484" author="Sterrer" box="[638,793,1392,1418]" pageId="8" pageNumber="21" refString="Sterrer, W. (1986) Marine fauna and flora of Bermuda. A systematic guide to the identification of marine organisms. John Wiley &amp; Sons, 742 pp., 16 pls." type="book" year="1986">Sterrer 1986</bibRefCitation>
), commonly associated with reef-fringing habitats, in crevices of rocks, dead and living corals and coral rubble (
<bibRefCitation id="3D52D4D1FFA3FFA2802AFA15FBE1C4BC" author="Verrill" box="[827,1144,1432,1458]" pageId="8" pageNumber="21" refString="Verrill, A. E. (1922) Decapod Crustacea of Bermuda. Part II. Macrura. Transactions of the Connecticut Academy of Arts and Sciences, 26, 1 - 179, pls. 1 - 48." type="journal article" year="1922">Verrill 1922; present study</bibRefCitation>
); mixed sand-rubble bottoms among turtle grass (
<taxonomicName id="9EC3D2A3FFA3FFA282C1FA4DFD57C4D7" box="[464,718,1472,1497]" class="Liliopsida" family="Hydrocharitaceae" genus="Thalassia" kingdom="Animalia" order="Alismatales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="testudinum">
<emphasis id="6BB77532FFA3FFA282C1FA4DFD57C4D7" box="[464,718,1472,1497]" italics="true" pageId="8" pageNumber="9">Thalassia testudinum</emphasis>
</taxonomicName>
), under rocks and coral rubble; in crevices on reefs of the sabellariid polychaete
<taxonomicName id="9EC3D2A3FFA3FFA282B8FA65FDF7C70F" box="[425,622,1512,1537]" class="Polychaeta" family="Sabellariidae" genus="Phragmatopoma" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">
<emphasis id="6BB77532FFA3FFA282B8FA65FDF7C70F" box="[425,622,1512,1537]" italics="true" pageId="8" pageNumber="9">Phragmatopoma</emphasis>
</taxonomicName>
sp. (
<bibRefCitation id="3D52D4D1FFA3FFA281A1FA65FC16C70C" author="Camp" box="[688,911,1512,1538]" pageId="8" pageNumber="18" refString="Camp, D. K., Whiting, N. H. &amp; Martin, R. E. (1977) Nearshore marine ecology at Hutchinson Island, Florida: 1971 - 1974. V. Arthropods. Florida Marine Research Publication, 25, 1 - 63." type="journal article" year="1977">
Camp
<emphasis id="6BB77532FFA3FFA28014FA65FCA7C70F" box="[773,830,1512,1537]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1977
</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA280B0FA65FBEAC70C" author="Gore" box="[929,1139,1512,1538]" pageId="8" pageNumber="19" refString="Gore, R. H., Scotto, L. E. &amp; Becker, L. J. (1978) Community composition, stability, and trophic partitioning in decapod crustaceans inhabiting some subtropical sabellariid worm reefs. Studies on decapod Crustacea from the Indian River region of Florida. IV. Bulletin of Marine Science, 28, 221 - 248." type="journal article" year="1978">
Gore
<emphasis id="6BB77532FFA3FFA280F9FA65FBB8C70F" box="[1000,1057,1512,1537]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1978
</bibRefCitation>
); in empty conch shells (
<taxonomicName id="9EC3D2A3FFA3FFA283B0F99DFE95C727" box="[161,268,1552,1577]" class="Gastropoda" family="Strombidae" genus="Strombus" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="6BB77532FFA3FFA283B0F99DFE95C727" box="[161,268,1552,1577]" italics="true" pageId="8" pageNumber="9">Strombus</emphasis>
</taxonomicName>
) on the outer fringing reefs near
<taxonomicName id="9EC3D2A3FFA3FFA281BCF99DFC86C727" box="[685,799,1552,1577]" class="Liliopsida" family="Hydrocharitaceae" genus="Thalassia" kingdom="Animalia" order="Alismatales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="6BB77532FFA3FFA281BCF99DFC86C727" box="[685,799,1552,1577]" italics="true" pageId="8" pageNumber="9">Thalassia</emphasis>
</taxonomicName>
beds (
<bibRefCitation id="3D52D4D1FFA3FFA28065F99DFBB9C724" author="Hendrix" box="[884,1056,1552,1578]" pageId="8" pageNumber="20" refString="Hendrix, G. Y. (1971) A systematic study of the genus Alpheus (Crustacea: Decapoda: Alpheidae) in South Florida. PhD Dissertation, University of Miami, Coral Gables, Florida, USA. 184 pp., 24 pls." type="book" year="1971">Hendrix 1971</bibRefCitation>
); occasionally in sponges, e.g.,
<taxonomicName id="9EC3D2A3FFA3FFA28386F9B5FEAAC75F" box="[151,307,1592,1617]" class="Demospongiae" family="Callyspongiidae" genus="Callyspongia" kingdom="Animalia" order="Haplosclerida" pageId="8" pageNumber="9" phylum="Porifera" rank="genus">
<emphasis id="6BB77532FFA3FFA28386F9B5FEAAC75F" box="[151,307,1592,1617]" italics="true" pageId="8" pageNumber="9">Callyspongia</emphasis>
</taxonomicName>
,
<taxonomicName id="9EC3D2A3FFA3FFA28250F9B5FE09C75F" box="[321,400,1592,1617]" class="Demospongiae" family="Irciniidae" genus="Ircinia" kingdom="Animalia" order="Dictyoceratida" pageId="8" pageNumber="9" phylum="Porifera" rank="genus">
<emphasis id="6BB77532FFA3FFA28250F9B5FE09C75F" box="[321,400,1592,1617]" italics="true" pageId="8" pageNumber="9">Ircinia</emphasis>
</taxonomicName>
(
<bibRefCitation id="3D52D4D1FFA3FFA2828EF9B5FDABC75C" author="Pearse" box="[415,562,1592,1618]" pageId="8" pageNumber="21" refString="Pearse, A. S. (1950) Notes on the inhabitants of certain sponges at Bimini. Ecology, 31, 149 - 151." type="journal article" year="1950">Pearse 1950</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA2812FF9B5FD54C75C" author="Chace" box="[574,717,1592,1618]" pageId="8" pageNumber="19" refString="Chace, F. A. Jr. (1956) Crustaceos decapodos y estomatopodos del Archipielago de los Roques y Isla de la Orchila. In: Mendez, A. et al. (eds.), Archipielago de Los Roques y la Orchila, pp. 145 - 168, 4 pls. Sociedad de Ciencias Naturales La Salle, Editorial Sucre, Caracas." type="journal article" year="1956">Chace 1956</bibRefCitation>
), among bryozoan colonies,
<taxonomicName id="9EC3D2A3FFA3FFA2870DF9B5FB25C75F" box="[1052,1212,1592,1617]" class="Gymnolaemata" family="Schizoporellidae" genus="Schizoporella" kingdom="Animalia" order="Cheilostomatida" pageId="8" pageNumber="9" phylum="Bryozoa" rank="genus">
<emphasis id="6BB77532FFA3FFA2870DF9B5FB25C75F" box="[1052,1212,1592,1617]" italics="true" pageId="8" pageNumber="9">Schizoporella</emphasis>
</taxonomicName>
sp. (
<bibRefCitation id="3D52D4D1FFA3FFA287E2F9B5FF4CC774" author="Christoffersen" pageId="8" pageNumber="19" refString="Christoffersen, M. L. (1979) Decapod Crustacea: Alpheoidea. Campagnes de la Calypso au large des cotes atlantiques de l'Amerique du Sud (1961 - 1962). I. Resultats Scientifiques de la Campagne de la Calypso, 11, 297 - 377." type="journal article" year="1979">Christoffersen 1979</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA283F5F9EDFD80C774" author="Morgado" box="[228,537,1632,1658]" pageId="8" pageNumber="21" refString="Morgado, E. H. &amp; Tanaka, N. O. (2001) The macrofauna associated with the bryozoan Schizoporella errata (Walters) in southeastern Brazil. Scientia Marina, 65, 173 - 181." type="journal article" year="2001">Morgado &amp; Tanaka 2001</bibRefCitation>
), and in living corals, e.g.,
<taxonomicName id="9EC3D2A3FFA3FFA2807CF9EDFC52C777" box="[877,971,1632,1657]" class="Anthozoa" family="Oculinidae" genus="Oculina" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="6BB77532FFA3FFA2807CF9EDFC52C777" box="[877,971,1632,1657]" italics="true" pageId="8" pageNumber="9">Oculina</emphasis>
</taxonomicName>
and
<taxonomicName id="9EC3D2A3FFA3FFA2871AF9EDFBC6C777" box="[1035,1119,1632,1657]" class="Anthozoa" family="Poritidae" genus="Porites" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="6BB77532FFA3FFA2871AF9EDFBC6C777" box="[1035,1119,1632,1657]" italics="true" pageId="8" pageNumber="9">Porites</emphasis>
</taxonomicName>
(
<bibRefCitation id="3D52D4D1FFA3FFA28762F9EDFA9EC774" author="Chace" box="[1139,1287,1632,1658]" pageId="8" pageNumber="19" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, i - x + 1 - 179." type="journal article" year="1972">Chace 1972</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA28607F9EDFF4FC7AC" author="Reed" pageId="8" pageNumber="21" refString="Reed, J. K., Gore, R. H., Scotto L. E. &amp; Wilson, K. A. (1982) Community composition, structure, areal and trophic relationships of decapods associated with shallow and deep-water Oculina varicosa coral reefs. Studies on decapod crustacea from the Indian River Region of Florida, XXIV. Bulletin of Marine Science, 32, 761 - 786." type="journal article" year="1982">
Reed
<emphasis id="6BB77532FFA3FFA2864FF9EDFA0CC777" box="[1374,1429,1632,1657]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1982
</bibRefCitation>
); occasionally entering sea-connected cave systems (
<bibRefCitation id="3D52D4D1FFA3FFA28053F905FC4EC7AC" author="Hobbs" box="[834,983,1672,1698]" pageId="8" pageNumber="20" refString="Hobbs, H. H. III. (1998) Decapoda (Caridea, Astacidea, Anomura): In Juberthie, C. and Decu, V. (eds.), Encyclopdia Biospeologica, Vol. II, pp. 891 - 911. Societe de Biospeologie, Paris." type="book chapter" year="1998">Hobbs 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11D9FAABFFA3FFA283D7F922FC8CC7C4" box="[198,789,1711,1738]" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="597CA920FFA3FFA283D7F922FC8CC7C4" blockId="8.[151,1437,152,2018]" box="[198,789,1711,1738]" pageId="8" pageNumber="9">
<emphasis id="6BB77532FFA3FFA283D7F922FEFDC7C7" bold="true" box="[198,356,1711,1737]" pageId="8" pageNumber="9">
<typeStatus id="86781782FFA3FFA283D7F922FE9DC7C7" box="[198,260,1711,1737]" pageId="8" pageNumber="9">Type</typeStatus>
locality
</emphasis>
.—Key West, Florida (
<bibRefCitation id="3D52D4D1FFA3FFA2817CF93DFC91C7C4" author="Gibbes" box="[621,776,1712,1738]" pageId="8" pageNumber="19" refString="Gibbes, L. R. (1850) On the carcinological collections of the cabinets of natural history in the United States: with an enumeration of the species contained therein and descriptions of new species. Proceedings of third meeting of American Association for Advancement of Science, 3, 165 - 201." type="journal article" year="1850">Gibbes 1850</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11D9FAABFFA3FFA083D7F95AFE7EC3B4" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="distribution">
<paragraph id="597CA920FFA3FFA083D7F95AFE7EC3B4" blockId="8.[151,1437,152,2018]" lastBlockId="10.[151,1437,152,698]" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<emphasis id="6BB77532FFA3FFA283D7F95AFEC6C7FF" bold="true" box="[198,351,1751,1777]" pageId="8" pageNumber="9">Distribution</emphasis>
.—Western Atlantic: North Carolina (
<bibRefCitation id="3D52D4D1FFA3FFA2800AF955FC67C7FC" author="Hay" box="[795,1022,1752,1778]" pageId="8" pageNumber="19" refString="Hay, W. P. &amp; Shore, C. A. (1918) The decapod crustaceans of Beaufort, North Carolina, and surrounding regions. Bulletin of Bureau of Fisheries, Washington, 35, 371 - 475, pls. 25 - 39." type="journal article" year="1918">Hay &amp; Shore 1918</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA2871AF955FB23C7FC" author="Williams" box="[1035,1210,1752,1778]" pageId="8" pageNumber="21" refString="Williams, A. B. (1965) Marine decapod crustaceans of the Carolinas. Fishery Bulletin of the U. S. Fish and Wildlife Service, 65, 1 - 298." type="journal article" year="1965">Williams 1965</bibRefCitation>
,
<bibRefCitation id="3D52D4D1FFA3FFA287D6F955FA9CC7FC" author="Williams" box="[1223,1285,1752,1778]" pageId="8" pageNumber="21" refString="Williams, A. B. (1984) Shrimps, lobsters and crabs of the Atlantic Coast of the eastern United States, Maine to Florida. Smithsonian Institution Press, Washington DC, 550 pp." type="book" year="1984">1984</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA28603F955FE9FC614" author="Christoffersen" pageId="8" pageNumber="19" refString="Christoffersen, M. L. (1979) Decapod Crustacea: Alpheoidea. Campagnes de la Calypso au large des cotes atlantiques de l'Amerique du Sud (1961 - 1962). I. Resultats Scientifiques de la Campagne de la Calypso, 11, 297 - 377." type="journal article" year="1979">Christoffersen 1979</bibRefCitation>
); reefs of South Carolina (
<bibRefCitation id="3D52D4D1FFA3FFA28157F88DFCEBC614" author="Pearse" box="[582,882,1792,1818]" pageId="8" pageNumber="21" refString="Pearse, A. S. &amp; Williams, L. G. (1951) The biota of the reefs off the Carolinas. Journal of Elisha Mitchell Scientific Society, 67, 133 - 161." type="journal article" year="1951">Pearse &amp; Williams 1951</bibRefCitation>
); Florida: southern and central-eastern Florida (e.g., Fort Pierce,
<collectingCountry id="21D4E9B0FFA3FFA28261F8A5FD90C64C" box="[368,521,1832,1858]" name="Saint Lucia" pageId="8" pageNumber="9">Sainte-Lucie</collectingCountry>
, Hutchinson
<collectingCountry id="21D4E9B0FFA3FFA281B7F8A5FD69C64C" box="[678,752,1832,1858]" name="Iceland" pageId="8" pageNumber="9">Island</collectingCountry>
, Sebastian Inlet), Florida Keys (e.g., Key West, Big Pine Key, Cedar Key), Dry
<collectingCountry id="21D4E9B0FFA3FFA282B1F8DDFD9FC664" box="[416,518,1872,1898]" name="Haiti" pageId="8" pageNumber="9">Tortugas</collectingCountry>
(
<bibRefCitation id="3D52D4D1FFA3FFA28104F8DDFD36C664" author="Gibbes" box="[533,687,1872,1898]" pageId="8" pageNumber="19" refString="Gibbes, L. R. (1850) On the carcinological collections of the cabinets of natural history in the United States: with an enumeration of the species contained therein and descriptions of new species. Proceedings of third meeting of American Association for Advancement of Science, 3, 165 - 201." type="journal article" year="1850">Gibbes 1850</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA281AAF8DDFCFCC664" author="Coutiere" box="[699,869,1872,1898]" pageId="8" pageNumber="19" refString="Coutiere, H. (1910) The snapping shrimps (Alpheidae) of the Dry Tortugas, Florida. Proceedings of the United States National Museum, 37, 485 - 487." type="journal article" year="1910">Coutière 1910</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA28060F8DDFB8CC664" author="Zimmer" box="[881,1045,1872,1898]" pageId="8" pageNumber="22" refString="Zimmer, C. (1913) Westindische Decapoden, 1: Die Familie Alpheidae. Zoologische Jahrbucher, Supplement 11, 381 - 412." type="journal article" year="1913">Zimmer 1913</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA28730F8DDFB3FC664" author="Balss" box="[1057,1190,1872,1898]" pageId="8" pageNumber="18" refString="Balss, H. (1924) Westindische Decapoden. Zoologischer Anzeiger, 61, 177 - 182." type="journal article" year="1924">Balss 1924</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA287A3F8DDFADDC664" author="Boone" box="[1202,1348,1872,1898]" pageId="8" pageNumber="18" refString="Boone, L. (1930) Crustacea: Anomura, Macrura, Schizopoda, Isopoda, Amphipoda, Mysidacea, Cirripedia, and Copepoda. In: Scientific Results of Cruises of the yachts &quot; Eagle &quot; and &quot; Ara &quot;, 1921 - 1928, William K. Vanderbilt Commanding. Bulletin Vanderbilt Marine Museum, 3, 1 - 221, pls. 1 - 83." type="journal article" year="1930">Boone 1930</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA2865EF8DDFF4CC69C" author="Pearse" pageId="8" pageNumber="21" refString="Pearse, A. S. (1932) Observations on the parasites and commensals found associated with crustaceans and fishes at Dry Tortugas, Florida. Papers from the Tortugas Laboratory, Carnegie Institution of Washington, 28, 103 - 115." type="journal article" year="1932">Pearse 1932</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA283F3F8F5FE1EC69C" author="Hendrix" box="[226,391,1912,1938]" pageId="8" pageNumber="20" refString="Hendrix, G. Y. (1971) A systematic study of the genus Alpheus (Crustacea: Decapoda: Alpheidae) in South Florida. PhD Dissertation, University of Miami, Coral Gables, Florida, USA. 184 pp., 24 pls." type="book" year="1971">Hendrix 1971</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA28285F8F5FDFAC69C" author="Camp" box="[404,611,1912,1938]" pageId="8" pageNumber="18" refString="Camp, D. K., Whiting, N. H. &amp; Martin, R. E. (1977) Nearshore marine ecology at Hutchinson Island, Florida: 1971 - 1974. V. Arthropods. Florida Marine Research Publication, 25, 1 - 63." type="journal article" year="1977">
Camp
<emphasis id="6BB77532FFA3FFA282F3F8F5FD8FC69F" box="[482,534,1912,1937]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1977
</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA28161F8F5FCAAC69C" author="Gore" box="[624,819,1912,1938]" pageId="8" pageNumber="19" refString="Gore, R. H., Scotto, L. E. &amp; Becker, L. J. (1978) Community composition, stability, and trophic partitioning in decapod crustaceans inhabiting some subtropical sabellariid worm reefs. Studies on decapod Crustacea from the Indian River region of Florida. IV. Bulletin of Marine Science, 28, 221 - 248." type="journal article" year="1978">
Gore
<emphasis id="6BB77532FFA3FFA281A3F8F5FD7FC69F" box="[690,742,1912,1937]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1978
</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA28051F8F5FB9CC69C" author="Reed" box="[832,1029,1912,1938]" pageId="8" pageNumber="21" refString="Reed, J. K., Gore, R. H., Scotto L. E. &amp; Wilson, K. A. (1982) Community composition, structure, areal and trophic relationships of decapods associated with shallow and deep-water Oculina varicosa coral reefs. Studies on decapod crustacea from the Indian River Region of Florida, XXIV. Bulletin of Marine Science, 32, 761 - 786." type="journal article" year="1982">
Reed
<emphasis id="6BB77532FFA3FFA28095F8F5FC21C69F" box="[900,952,1912,1937]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1982
</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA28703F8F5FB78C69C" author="Posey" box="[1042,1249,1912,1938]" pageId="8" pageNumber="21" refString="Posey, M. H., Troy, D. A., Banner, S., Vose, F. &amp; Lindberg, W. (1998) Temporal variability, diversity and guild structure of a benthic community in the notheastern Gulf of Mexico. Bulletin of Marine Science, 63, 143 - 155." type="journal article" year="1998">
Posey
<emphasis id="6BB77532FFA3FFA28771F8F5FB0DC69F" box="[1120,1172,1912,1937]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1998
</bibRefCitation>
; present study);
<collectingCountry id="21D4E9B0FFA3FFA28386F82DFE9AC6B4" box="[151,259,1952,1978]" name="Bermuda" pageId="8" pageNumber="9">Bermuda</collectingCountry>
(
<bibRefCitation id="3D52D4D1FFA3FFA28205F82DFE26C6B4" author="Heilprin" box="[276,447,1952,1978]" lastPageId="8" lastPageNumber="20" pageId="8" pageNumber="19" refString="Heilprin, A. (1888) Contributions to the natural history of the Bermuda Islands (incl. Crustacea). Proceedings of the Academy of Natural Sciences of Philadelphia, 1888, 302 - 328." type="proceedings paper" year="1888">Heilprin 1888</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA282DFF82DFDF8C6B4" author="Verrill" box="[462,609,1952,1978]" pageId="8" pageNumber="21" refString="Verrill, A. E. (1922) Decapod Crustacea of Bermuda. Part II. Macrura. Transactions of the Connecticut Academy of Arts and Sciences, 26, 1 - 179, pls. 1 - 48." type="journal article" year="1922">Verrill 1922</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA2817EF82DFC9EC6B4" author="Sterrer" box="[623,775,1952,1978]" pageId="8" pageNumber="21" refString="Sterrer, W. (1986) Marine fauna and flora of Bermuda. A systematic guide to the identification of marine organisms. John Wiley &amp; Sons, 742 pp., 16 pls." type="book" year="1986">Sterrer 1986</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA3FFA2800EF82DFCDCC6B4" box="[799,837,1952,1978]" name="United States of America" pageId="8" pageNumber="9">US</collectingCountry>
<collectingCountry id="21D4E9B0FFA3FFA2805FF82DFC6CC6B4" box="[846,1013,1952,1978]" name="Virgin Islands" pageId="8" pageNumber="9">Virgin Islands</collectingCountry>
(
<bibRefCitation id="3D52D4D1FFA3FFA28716F82DFB2BC6B4" author="Hendrix" box="[1031,1202,1952,1978]" pageId="8" pageNumber="20" refString="Hendrix, G. Y. (1971) A systematic study of the genus Alpheus (Crustacea: Decapoda: Alpheidae) in South Florida. PhD Dissertation, University of Miami, Coral Gables, Florida, USA. 184 pp., 24 pls." type="book" year="1971">Hendrix 1971</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA3FFA287D8F82DFAB3C6B4" box="[1225,1322,1952,1978]" name="Virgin Islands" pageId="8" pageNumber="9">St. John</collectingCountry>
(
<bibRefCitation id="3D52D4D1FFA3FFA2862DF82DFF4FC6EC" author="Zimmer" pageId="8" pageNumber="22" refString="Zimmer, C. (1913) Westindische Decapoden, 1: Die Familie Alpheidae. Zoologische Jahrbucher, Supplement 11, 381 - 412." type="journal article" year="1913">Zimmer 1913</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA3FFA283FBF845FE70C6EC" box="[234,489,1992,2018]" name="British Virgin Islands" pageId="8" pageNumber="9">British Virgin Islands</collectingCountry>
: Guana (present study);
<collectingCountry id="21D4E9B0FFA3FFA28015F845FCEDC6EC" box="[772,884,1992,2018]" name="Bahamas" pageId="8" pageNumber="9">Bahamas</collectingCountry>
: Bimini, Cay Sal Bank (
<bibRefCitation id="3D52D4D1FFA3FFA2879CF845FAB9C6EC" author="Pearse" box="[1165,1312,1992,2018]" pageId="8" pageNumber="21" refString="Pearse, A. S. (1950) Notes on the inhabitants of certain sponges at Bimini. Ecology, 31, 149 - 151." type="journal article" year="1950">Pearse 1950</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA3FFA0863AF845FF4FC1BC" author="Lemaitre" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="20" refString="Lemaitre, R. (1984) Decapod crustaceans from Cay Sal Bank, Bahamas, with notes on their zoogeographic affinities. Journal of Crustacean Biology, 4, 425 - 447." type="journal article" year="1984">Lemaitre 1984</bibRefCitation>
), Blue Holes Cave System (
<bibRefCitation id="3D52D4D1FFA1FFA08122FF15FD55C1BC" author="Hobbs" box="[563,716,152,178]" pageId="10" pageNumber="20" refString="Hobbs, H. H. III. (1998) Decapoda (Caridea, Astacidea, Anomura): In Juberthie, C. and Decu, V. (eds.), Encyclopdia Biospeologica, Vol. II, pp. 891 - 911. Societe de Biospeologie, Paris." type="book chapter" year="1998">Hobbs 1998</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA081F4FF15FCE1C1BC" box="[741,888,152,178]" name="Puerto Rico" pageId="10" pageNumber="11">Puerto Rico</collectingCountry>
; Culebra (
<bibRefCitation id="3D52D4D1FFA1FFA080E9FF15FB3EC1BC" author="Rathbun" box="[1016,1191,152,178]" pageId="10" pageNumber="21" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Puerto Rico. United States Fish Commission Bulletin, 20, 1 - 137, pls. 1, 2." type="journal article" year="1901">Rathbun 1901</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA087A6FF15FAC5C1BC" author="Schmitt" box="[1207,1372,152,178]" pageId="10" pageNumber="21" refString="Schmitt, W. L. (1935) Crustacea Macrura and Anomura of Porto Rico and the Virgin Islands. Science Survey of Porto Rico (New York Academy of Sciences), 15, 125 - 227." type="journal article" year="1935">Schmitt 1935</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA0867CFF15FF4FC1D4" author="Ray" pageId="10" pageNumber="21" refString="Ray, J. P. (1974) A study of the coral reef crustaceans (Decapoda and Stomatopoda) of two Gulf of Mexico reef systems: West Flower Garden, Texas and Isla de Lobos, Veracruz, Mexico. PhD Dissertation. Texas A &amp; M University, College Station, Texas. 323 pp." type="book" year="1974">Ray 1974</bibRefCitation>
); French Antilles:
<collectingCountry id="21D4E9B0FFA1FFA082A0FF4DFDAEC1D4" box="[433,567,192,218]" name="Martinique" pageId="10" pageNumber="11">Martinique</collectingCountry>
,
<collectingCountry id="21D4E9B0FFA1FFA08153FF4DFD49C1D4" box="[578,720,192,218]" name="Guadeloupe" pageId="10" pageNumber="11">Guadeloupe</collectingCountry>
(
<bibRefCitation id="3D52D4D1FFA1FFA081F1FF4DFBDCC1D4" author="Carvacho" box="[736,1093,192,218]" pageId="10" pageNumber="18" refString="Carvacho, A. (1982) Sur une petite collection de crevettes de la Martinique. Caribbean Journal of Science, 17, 15 - 20." type="journal article" year="1982">Carvacho 1982; present study</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA0874BFF4DFB28C1D4" box="[1114,1201,192,218]" name="Trinidad and Tobago" pageId="10" pageNumber="11">Tobago</collectingCountry>
(
<bibRefCitation id="3D52D4D1FFA1FFA087D0FF4DFACAC1D4" author="Chace" box="[1217,1363,192,218]" pageId="10" pageNumber="19" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, i - x + 1 - 179." type="journal article" year="1972">Chace 1972</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA08679FF4DFF42C00C" name="Barbados" pageId="10" pageNumber="11">Barbados</collectingCountry>
(
<bibRefCitation id="3D52D4D1FFA1FFA083FBFF65FE16C00C" author="Zimmer" box="[234,399,232,258]" pageId="10" pageNumber="22" refString="Zimmer, C. (1913) Westindische Decapoden, 1: Die Familie Alpheidae. Zoologische Jahrbucher, Supplement 11, 381 - 412." type="journal article" year="1913">Zimmer 1913</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA0828AFF65FDD5C00C" author="Schmitt" box="[411,588,232,258]" pageId="10" pageNumber="21" refString="Schmitt, W. L. (1924 b) Report on Macrura, Anomura and Stomatopoda collected by the Barbados-Antigua Expedition from the University of Iowa in 1918. Studies of Natural History, Iowa University, 10, 65 - 99, pls. 1 - 5." type="journal article" year="1924" yearSuffix="b">Schmitt 1924b</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA08171FF65FCCFC00C" box="[608,854,232,258]" name="Curaçao" pageId="10" pageNumber="11">Netherlands Antilles</collectingCountry>
:
<collectingCountry id="21D4E9B0FFA1FFA08070FF65FC34C00C" box="[865,941,232,258]" name="Aruba" pageId="10" pageNumber="11">Aruba</collectingCountry>
,
<collectingCountry id="21D4E9B0FFA1FFA080A8FF65FB87C00C" box="[953,1054,232,258]" name="Curaçao" pageId="10" pageNumber="11">Curaçao</collectingCountry>
,
<collectingCountry id="21D4E9B0FFA1FFA08738FF65FB1FC00C" box="[1065,1158,232,258]" name="Bonaire, Sint Eustatius and Saba" pageId="10" pageNumber="11">Bonaire</collectingCountry>
(
<bibRefCitation id="3D52D4D1FFA1FFA08784FF65FADAC00C" author="Schmitt" box="[1173,1347,232,258]" pageId="10" pageNumber="21" refString="Schmitt, W. L. (1924 b) Report on Macrura, Anomura and Stomatopoda collected by the Barbados-Antigua Expedition from the University of Iowa in 1918. Studies of Natural History, Iowa University, 10, 65 - 99, pls. 1 - 5." type="journal article" year="1924" yearSuffix="a">Schmitt 1924a</bibRefCitation>
,
<bibRefCitation id="3D52D4D1FFA1FFA0865EFF65FA17C00C" author="Schmitt" box="[1359,1422,232,258]" pageId="10" pageNumber="21" refString="Schmitt, W. L. (1936) Macruran and anomuran Crustacea from Bonaire, Curacao and Aruba. Zoologische Ergebnisse einer Reise nach Bonaire, Curacao und Aruba im Jahre 1930. Nr 16. Zoologische Jahrbucher, Abteilung fur Systematik, Oekologie und Geographie der Tiere, 67, 363 - 378, pls. 11 - 13." type="journal article" year="1936">1936</bibRefCitation>
); present study);
<collectingCountry id="21D4E9B0FFA1FFA08242FE9DFE49C024" box="[339,464,272,298]" name="Venezuela" pageId="10" pageNumber="11">Venezuela</collectingCountry>
: Los Roques, Tucacas,
<collectingCountry id="21D4E9B0FFA1FFA081E2FE9DFC84C024" box="[755,797,272,298]" name="Iceland" pageId="10" pageNumber="11">Isla</collectingCountry>
Margarita, Arrecifes (
<bibRefCitation id="3D52D4D1FFA1FFA0873EFE9DFB5DC024" author="Chace" box="[1071,1220,272,298]" pageId="10" pageNumber="19" refString="Chace, F. A. Jr. (1956) Crustaceos decapodos y estomatopodos del Archipielago de los Roques y Isla de la Orchila. In: Mendez, A. et al. (eds.), Archipielago de Los Roques y la Orchila, pp. 145 - 168, 4 pls. Sociedad de Ciencias Naturales La Salle, Editorial Sucre, Caracas." type="journal article" year="1956">Chace 1956</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA087C5FE9DFA0EC024" author="Rodriguez" box="[1236,1431,272,298]" pageId="10" pageNumber="21" refString="Rodriguez, G. (1980) Crustaceos decapodos de Venezuela. Instituto Venezolano de Investigaciones Cientificas, Caracas, 494 pp." type="book" year="1980">Rodríguez 1980</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA08386FEB5FECEC05C" author="Rodriguez" box="[151,343,312,338]" pageId="10" pageNumber="21" refString="Rodriguez, B. (1986) Los camarones (Crustacea; Decapoda; Natantia) del Parque Nacional Archipielago de los Roques. Trabajo especial de grado para optar al titulo de Licenciado en Biologia, Universidad Central de Venezuela, Caracas. 350 pp." type="book" year="1986">Rodríguez 1986</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA0827AFEB5FE7AC05C" box="[363,483,312,338]" name="Colombia" pageId="10" pageNumber="11">Colombia</collectingCountry>
: Providencia (
<bibRefCitation id="3D52D4D1FFA1FFA08199FEB5FCB2C05C" author="Schmitt" box="[648,811,312,338]" pageId="10" pageNumber="21" refString="Schmitt, W. L. (1939) Decapod and other Crustacea collected on the Presidential Cruise of 1938 (with introduction and station data). Smithsonian Miscellaneous Collections, 98, 1 - 29, 3 pls." type="journal article" year="1939">Schmitt 1939</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA0802EFEB5FC39C05C" box="[831,928,312,338]" name="Panama" pageId="10" pageNumber="11">Panama</collectingCountry>
:
<collectingCountry id="21D4E9B0FFA1FFA080BAFEB5FC4CC05C" box="[939,981,312,338]" name="Iceland" pageId="10" pageNumber="11">Isla</collectingCountry>
Grande, Colón, Bocas del Toro (Abele 1976;
<bibRefCitation id="3D52D4D1FFA1FFA083F2FEEDFD98C074" author="Cubit" box="[227,513,352,378]" pageId="10" pageNumber="19" refString="Cubit, J. &amp; Williams, S. (1983) The invertebrates of Galeta Reef (Caribbean Panama): a species list and bibliography. Atoll Research Bulletin, 269, 1 - 45." type="journal article" year="1983">Cubit &amp; Williams 1983</bibRefCitation>
; present study);
<collectingCountry id="21D4E9B0FFA1FFA081D8FEEDFCD6C074" box="[713,847,352,378]" name="Costa Rica" pageId="10" pageNumber="11">Costa Rica</collectingCountry>
: Limón, Cahuita (
<bibRefCitation id="3D52D4D1FFA1FFA08738FEEDFF42C0AC" author="Vargas" pageId="10" pageNumber="21" refString="Vargas, R. &amp; Cortes, J. (1999) Biodiversidad marina de Costa Rica: Crustacea: Decapoda (Penaeoidea, Sergestoidea, Stenopodidea, Caridea, Thalassinidea, Palinura) del Caribe. Revista de Biologia Tropical, 47, 887 - 911." type="journal article" year="1999">Vargas &amp; Cortés 1999; present study</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA083E3FE05FEF0C0AC" box="[242,361,392,418]" name="Honduras" pageId="10" pageNumber="11">Honduras</collectingCountry>
: Utila (A. Anker, pers. obs.);
<collectingCountry id="21D4E9B0FFA1FFA081C7FE05FCADC0AC" box="[726,820,392,418]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
: Quintana Roo (
<bibRefCitation id="3D52D4D1FFA1FFA080EAFE05FB17C0AC" author="Chace" box="[1019,1166,392,418]" pageId="10" pageNumber="19" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, i - x + 1 - 179." type="journal article" year="1972">Chace 1972</bibRefCitation>
), Veracruz, Campeche (Hernández-Aguilera
<emphasis id="6BB77532FFA1FFA08289FE3DFE55C0C7" box="[408,460,432,457]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. 1996), Puerto Morelos, Punta Estrella (
<bibRefCitation id="3D52D4D1FFA1FFA080BDFE3DFB32C0C4" author="Markham" box="[940,1195,432,458]" pageId="10" pageNumber="20" refString="Markham, J. C., Donath-Hernandez, F. E., Villalobos-Hiriart, J. L. &amp; Diaz-Barriga, A. C. (1990) Notes on the shallow water marine Crustacea of the Caribbean coast of Quintana Roo, Mexico. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, zoologia, 61, 405 - 446 [not seen]." type="journal article" year="1990">
Markham
<emphasis id="6BB77532FFA1FFA08739FE3DFBC4C0C7" box="[1064,1117,432,457]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. 1990
</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA087D0FE3DFABDC0C4" box="[1217,1316,432,458]" name="Jamaica" pageId="10" pageNumber="11">Jamaica</collectingCountry>
: Kingston (
<bibRefCitation id="3D52D4D1FFA1FFA0838FFE55FEDEC0FC" author="Zimmer" box="[158,327,472,498]" pageId="10" pageNumber="22" refString="Zimmer, C. (1913) Westindische Decapoden, 1: Die Familie Alpheidae. Zoologische Jahrbucher, Supplement 11, 381 - 412." type="journal article" year="1913">Zimmer 1913</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA0824DFE55FE06C0FC" box="[348,415,472,498]" name="Cuba" pageId="10" pageNumber="11">Cuba</collectingCountry>
: Gulf of Batabano (
<bibRefCitation id="3D52D4D1FFA1FFA0819BFE55FC7BC0FC" author="Martinez-Iglesias" box="[650,994,472,498]" pageId="10" pageNumber="20" refString="Martinez-Iglesias, J. C., Carvacho, A. &amp; Rios, R. (1996) Catalogo de los carideos marinos (Crustacea, Decapoda, Caridea) de las aguas someras de Cuba. Avicennia, 4 / 5, 27 - 40." type="journal article" year="1996">
Martínez-Iglesias
<emphasis id="6BB77532FFA1FFA08070FE55FC0FC0FF" box="[865,918,472,497]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. 1996
</bibRefCitation>
,
<bibRefCitation id="3D52D4D1FFA1FFA080E1FE55FBB6C0FC" author="Martinez-Iglesias" box="[1008,1071,472,498]" pageId="10" pageNumber="20" refString="Martinez-Iglesias, J. C., Rios, R. &amp; Carvacho, A. (1997) Las especies del genero Alpheus (Decapoda: Alpheidae) de Cuba. Revista de Biologia Tropical, 44 / 45, 401 - 429." type="journal article" year="1997">1997</bibRefCitation>
);
<collectingCountry id="21D4E9B0FFA1FFA08754FE55FB08C0FC" box="[1093,1169,472,498]" name="Brazil" pageId="10" pageNumber="11">Brazil</collectingCountry>
:
<collectingCountry id="21D4E9B0FFA1FFA0878CFE55FB46C0FC" box="[1181,1247,472,498]" name="South Africa" pageId="10" pageNumber="11">Natal</collectingCountry>
, Rio Grande do Norte, Pernambuco, Alagoas, Bahia (including Abrolhos Islands), Ceara, Espirito Santo, Rio de Janeiro, São Paulo (
<bibRefCitation id="3D52D4D1FFA1FFA083FFFDA5FE07C34C" author="Rathbun" box="[238,414,552,578]" pageId="10" pageNumber="21" refString="Rathbun, M. J. (1900) The decapod and stomatopod Crustacea, Part I. In: Results of the Branner-Agassiz Expedition to Brazil. Proceedings of the Washington Academy of Sciences, 2, 133 - 156, pl. 8." type="journal article" year="1900">Rathbun 1900</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA082A1FDA5FD08C34C" author="Fausto-Filho" box="[432,657,552,578]" pageId="10" pageNumber="19" refString="Fausto-Filho, J. (1970) Quarta contribuicao ao inventario dos crustaceos decapodos do nordeste brasileiro. Arquivos de Ciencias do Mar, 10, 55 - 60." type="journal article" year="1970">Fausto-Filho 1970</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA081B2FDA5FCA4C34C" author="Correa" box="[675,829,552,578]" pageId="10" pageNumber="19" refString="Correa, M. M. G. (1972) Contribuicao ao conhecimento da fauna do Arquipelago de Abrolhos, Bahia, Brasil. 2. Lista preliminar dos crustaceos decapodos. Boletim de Museu de Historia Natural, Universidade Federal de Minas Gerais, zoologia, 15, 1 - 19 [not seen]." type="journal article" year="1972">Corrêa 1972</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA0805EFDA5FBD9C34C" author="Christoffersen" box="[847,1088,552,578]" pageId="10" pageNumber="19" refString="Christoffersen, M. L. (1979) Decapod Crustacea: Alpheoidea. Campagnes de la Calypso au large des cotes atlantiques de l'Amerique du Sud (1961 - 1962). I. Resultats Scientifiques de la Campagne de la Calypso, 11, 297 - 377." type="journal article" year="1979">Christoffersen 1979</bibRefCitation>
,
<bibRefCitation id="3D52D4D1FFA1FFA08743FDA5FB09C34C" author="Christoffersen" box="[1106,1168,552,578]" pageId="10" pageNumber="19" refString="Christoffersen, M. L. (1998) Malacostraca. Eucarida. Caridea, Crangonoidea and Alpheoidea (except Glyphocrangonidae and Crangonidae). In Young, P. S. (ed.), Catalogue of Crustacea of Brazil, pp. 351 - 372. Museu Nacional, Rio de Janeiro." type="book chapter" year="1998">1998</bibRefCitation>
;
<bibRefCitation id="3D52D4D1FFA1FFA087B3FDA5FA16C34C" author="Coelho" box="[1186,1423,552,578]" pageId="10" pageNumber="19" refString="Coelho, P. A., de Almeida, A. O., de Souza-Filho, J. F., Bezerra, L. E. A. &amp; Giraldes, B. W. (2006) Diversity and distribution of the marine and estuarine shrimps (Dendrobranchiata, Stenopodidea and Caridea) from North and Northeast Brazil. Zootaxa, 1221, 41 - 62." type="journal article" year="2006">
Coelho
<emphasis id="6BB77532FFA1FFA08612FDA5FAA5C34F" box="[1283,1340,552,577]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. 2006
</bibRefCitation>
), Fernando do Noronha (
<bibRefCitation id="3D52D4D1FFA1FFA082A1FDDDFDCBC364" author="Pocock" box="[432,594,592,618]" pageId="10" pageNumber="21" refString="Pocock, R. I. (1890) Crustacea. In: Ridley, H. N. (ed.), Notes on the zoology of Fernando Noronha. Journal of the Linnean Society, Zoology, 20, 506 - 526." type="journal article" year="1890">Pocock 1890</bibRefCitation>
); Atol das Rocas (present study). Some of the above-listed literature records may actually refer to the below-described new species, and not to
<taxonomicName id="9EC3D2A3FFA1FFA080E2FDF5FBE4C39F" box="[1011,1149,632,657]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFA1FFA080E2FDF5FBE4C39F" box="[1011,1149,632,657]" italics="true" pageId="10" pageNumber="11">A. formosus</emphasis>
</taxonomicName>
, as the two species were not distinguished in the past.
</paragraph>
</subSubSection>
<caption id="0DBCF9A8FFA2FFA38386F8E6FE83C6C1" httpUri="https://zenodo.org/record/180925/files/figure.png" pageId="9" pageNumber="10" targetBox="[198,1376,158,1870]" targetPageId="9">
<paragraph id="597CA920FFA2FFA38386F8E6FE83C6C1" blockId="9.[151,1436,1899,1999]" pageId="9" pageNumber="10">
<emphasis id="6BB77532FFA2FFA38386F8E6FE83C68D" bold="true" box="[151,282,1899,1923]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
<taxonomicName id="9EC3D2A3FFA2FFA38233F8E1FDE5C68D" authority="Gibbes, 1850" authorityName="Gibbes" authorityYear="1850" box="[290,636,1900,1923]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFA2FFA38233F8E1FE7BC68D" box="[290,482,1900,1923]" italics="true" pageId="9" pageNumber="10">Alpheus formosus</emphasis>
Gibbes, 1850
</taxonomicName>
, male from Atol das Rocas, Brazil (MNRJ 20755): a, minor cheliped, lateral; b, same, chela, mesial; c, distal portion of chela, lateral; d, same, detail of ischium, merus and carpus, mesial. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="0DBCF9A8FFA1FFA08386F9C5FCD2C7A5" httpUri="https://zenodo.org/record/180926/files/figure.png" pageId="10" pageNumber="11" targetBox="[177,1430,752,1577]" targetPageId="10">
<paragraph id="597CA920FFA1FFA08386F9C5FCD2C7A5" blockId="10.[151,1436,1608,1707]" pageId="10" pageNumber="11">
<emphasis id="6BB77532FFA1FFA08386F9C5FE82C76E" bold="true" box="[151,283,1608,1632]" pageId="10" pageNumber="11">FIGURE 4.</emphasis>
<taxonomicName id="9EC3D2A3FFA1FFA08234F9C4FD1BC751" authority="Gibbes, 1850" authorityName="Gibbes" authorityYear="1850" box="[293,642,1608,1632]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFA1FFA08234F9C4FE7FC76E" box="[293,486,1609,1632]" italics="true" pageId="10" pageNumber="11">Alpheus formosus</emphasis>
Gibbes, 1850
</taxonomicName>
, neotype female from Big Pine Key, Florida Keys (USNM 1100697): a, frontal region, dorsal; b, major cheliped, mesial; c, same, detail of distomesial tooth; d, minor chela and carpus, mesial; e, detail of distolateral spine of uropod, dorsal. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="11D9FAABFFA1FFA183D7F96FFE85C0AC" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
<paragraph id="597CA920FFA1FFA183D7F96FFE77C024" blockId="10.[151,1436,1762,2029]" lastBlockId="11.[151,1436,152,418]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="6BB77532FFA1FFA083D7F96FFEA1C7F2" bold="true" box="[198,312,1762,1788]" pageId="10" pageNumber="11">Remarks</emphasis>
.—
<taxonomicName id="9EC3D2A3FFA1FFA0824CF96EFDB7C7F2" box="[349,558,1763,1788]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFA1FFA0824CF96EFDB7C7F2" box="[349,558,1763,1788]" italics="true" pageId="10" pageNumber="11">Alpheus formosus</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="9EC3D2A3FFA1FFA0800EF96EFC50C7F2" box="[799,969,1763,1788]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="6BB77532FFA1FFA0800EF96EFC50C7F2" box="[799,969,1763,1788]" italics="true" pageId="10" pageNumber="11">A. panamensis</emphasis>
</taxonomicName>
but can be distinguished from the latter by the orbitorostral grooves extending to the level of eye base posteriorly (vs. to the base of the orbital hoods in
<taxonomicName id="9EC3D2A3FFA1FFA083A7F8BEFEFBC642" box="[182,354,1843,1868]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="6BB77532FFA1FFA083A7F8BEFEFBC642" box="[182,354,1843,1868]" italics="true" pageId="10" pageNumber="11">A. panamensis</emphasis>
</taxonomicName>
); the rostral plate being slightly narrower at base; and the usually much darker distolateral spine on the uropod (black in
<taxonomicName id="9EC3D2A3FFA1FFA08119F8D6FD0EC67A" box="[520,663,1883,1908]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFA1FFA08119F8D6FD0EC67A" box="[520,663,1883,1908]" italics="true" pageId="10" pageNumber="11">A. formosus</emphasis>
</taxonomicName>
vs. pale, distally tan in
<taxonomicName id="9EC3D2A3FFA1FFA080D1F8D6FBF7C67A" box="[960,1134,1883,1908]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="6BB77532FFA1FFA080D1F8D6FBF7C67A" box="[960,1134,1883,1908]" italics="true" pageId="10" pageNumber="11">A. panamensis</emphasis>
</taxonomicName>
) (see also
<bibRefCitation id="3D52D4D1FFA1FFA087E7F8D6FF4FC693" author="Kim" pageId="10" pageNumber="20" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
). The color of the uropodal spine may be lighter if the animal has recently molted. For instance, some alcohol-preserved specimens from Atol das Rocas, including the illustrated male specimen, have a pale spine, without trace of black pigment (
<figureCitation id="C1F8B5A5FFA1FFA08135F85EFDF2C6E3" box="[548,619,2003,2029]" captionStart="FIGURE 1" captionStartId="5.[151,255,1497,1521]" captionTargetBox="[172,1407,211,1464]" captionTargetId="figure@5.[172,1416,194,1473]" captionTargetPageId="5" captionText="FIGURE 1. Alpheus formosus Gibbes, 1850, male from Atol das Rocas, Brazil (MNRJ 20755): a, frontal region, dorsal; b, same, lateral; c, same, detail of orbital hoods, rostrum and first segment of antennular peduncle; d, second pereiopod, lateral; e, third pereiopod, lateral; f, telson and uropod, dorsal (fringing setae on uropod omitted); g, uropod, distal region of exopod and endopod, dorsal (setae omitted). Scale bars = 1 mm." httpUri="https://zenodo.org/record/180923/files/figure.png" pageId="10" pageNumber="11">Fig. 1</figureCitation>
f). Most individuals, however, including the
<typeStatus id="86781782FFA1FFA0878CF85EFB64C6E3" box="[1181,1277,2003,2029]" pageId="10" pageNumber="11" type="neotype">neotype</typeStatus>
, have a dark brown to blackish spine in life and in alcohol (
<figureCitation id="C1F8B5A5FFA0FFA181CCFF15FCB0C1BC" box="[733,809,152,178]" captionStart="FIGURE 4" captionStartId="10.[151,255,1608,1632]" captionTargetBox="[177,1430,752,1577]" captionTargetId="figure@10.[151,1436,742,1584]" captionTargetPageId="10" captionText="FIGURE 4. Alpheus formosus Gibbes, 1850, neotype female from Big Pine Key, Florida Keys (USNM 1100697): a, frontal region, dorsal; b, major cheliped, mesial; c, same, detail of distomesial tooth; d, minor chela and carpus, mesial; e, detail of distolateral spine of uropod, dorsal. Scale bars = 1 mm." httpUri="https://zenodo.org/record/180926/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
e).
<taxonomicName id="9EC3D2A3FFA0FFA18041FF15FBBCC1BF" box="[848,1061,152,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="6BB77532FFA0FFA18041FF15FBBCC1BF" box="[848,1061,152,177]" italics="true" pageId="11" pageNumber="12">Alpheus formosus</emphasis>
</taxonomicName>
can be distinguished from the below-described new species by the presence of balaeniceps setae on the fingers of the minor chela in both sexes (more developed in males), the posteriorly more extending orbitorostral grooves and several features of the color pattern (see below).
</paragraph>
<paragraph id="597CA920FFA0FFA183D7FEBAFE85C0AC" blockId="11.[151,1436,152,418]" pageId="11" pageNumber="12">
<emphasis id="6BB77532FFA0FFA183D7FEBAFE2AC05F" bold="true" box="[198,435,311,337]" pageId="11" pageNumber="12">GenBank number</emphasis>
.—
<accessionNumber id="469034C3FFA0FFA182C5FEB5FDC8C05C" box="[468,593,312,338]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF532605" pageId="11" pageNumber="12">EF532605</accessionNumber>
(fcn 98-163),
<accessionNumber id="469034C3FFA0FFA1800EFEB5FC00C05C" box="[799,921,312,338]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF532606" pageId="11" pageNumber="12">EF532606</accessionNumber>
(fcn 98-189),
<accessionNumber id="469034C3FFA0FFA18776FEB5FB7BC05C" box="[1127,1250,312,338]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF532607" pageId="11" pageNumber="12">EF532607</accessionNumber>
(fcn 98-190),
<accessionNumber id="469034C3FFA0FFA18386FEEDFE88C074" box="[151,273,352,378]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF532608" pageId="11" pageNumber="12">EF532608</accessionNumber>
(fcn 05-101, USNM 1100701),
<accessionNumber id="469034C3FFA0FFA181D5FEEDFCA7C074" box="[708,830,352,378]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF532609" pageId="11" pageNumber="12">EF532609</accessionNumber>
(fcn 06-414),
<accessionNumber id="469034C3FFA0FFA1871BFEEDFB1DC074" box="[1034,1156,352,378]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF532610" pageId="11" pageNumber="12">EF532610</accessionNumber>
(06-499),
<accessionNumber id="469034C3FFA0FFA18609FEEDFA0EC074" box="[1304,1431,352,378]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF309903" pageId="11" pageNumber="12">AF309903</accessionNumber>
,
<accessionNumber id="469034C3FFA0FFA18386FE05FE80C0AC" box="[151,281,392,418]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF309904" pageId="11" pageNumber="12">AF309904</accessionNumber>
.
</paragraph>
</subSubSection>
</treatment>
</document>