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<document id="49FFA2FACCDC46968C0344DC20CB58B2" ID-DOI="10.37828/em.2024.80.7" ID-ISSN="2336-9744" ID-Zenodo-Dep="14653158" ID-ZooBank="7F46AFC2-5105-4E02-AAB5-A625E070BE72" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736951107901" checkinUser="felipe" docAuthor="Legalov, Andrei A., Vasilenko, Dmitry V. &amp; Perkovsky, Evgeny E." docDate="2024" docId="D1130013FFA11E72FF7BF8D8FB98E2BE" docLanguage="en" docName="EcolMontenegrina.80.78-85.pdf" docOrigin="Ecologica Montenegrina 80" docSource="https://doi.org/10.37828/em.2024.80.7" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.3:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="3" docTitle="Orthocis vilhelmseni Legalov et Perkovsky 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="82" masterDocId="2D2A786BFFA31E76FFC6FFF1FFD8E65A" masterDocTitle="Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber" masterLastPageNumber="85" masterPageNumber="78" pageNumber="80" updateTime="1736951822116" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D5A5FBE57CAEEAD32DF5C553124EBBFE">Three new species of the genus Orthocis Casey, 1898 (Coleoptera: Ciidae) from Eocene Danish amber</mods:title>
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<mods:namePart id="79A1A077C58265C903872D9FCF137E45">Legalov, Andrei A.</mods:namePart>
<mods:affiliation id="8084D7253DB946C446F2ABC880166C92">Institute of Systematics and Ecology of Animals, SB RAS, Frunze Street 11, Novosibirsk, 630091, Russia; &amp; Altai State University, Lenina Street 61, Barnaul, 656049, Russia. &amp; Tomsk State University, Lenina Prospekt 36, 634050 Tomsk, Russia;</mods:affiliation>
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<mods:namePart id="19378F2EE0A92C899D748303F08F8998">Vasilenko, Dmitry V.</mods:namePart>
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<mods:affiliation id="FEC32378D349C1FA9FAB64D9138C0D70">Paleontological Laboratory, Cherepovets State University, Cherepovets 162600, Russia. &amp; Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia. E-mail: vasilenko @ paleo. ru; https: // orcid. org / 0000 - 0002 - 4827 - 7290</mods:affiliation>
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<mods:namePart id="3A6B16B7753BF0F2648C63E898465230">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="03F08C67A8465D6131FB2734ED0A53DA">Natural History Museum of Denmark, Universitetsparken 15, Copenhagen, Denmark; E-mail: perkovsk @ gmail. com; https: // orcid. org / 0000 - 0002 - 7959 - 4379 Corresponding author. E-mail: fossilweevils @ gmail. com; https: // orcid. org / 0000 - 0001 - 7347 - 8169</mods:affiliation>
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<mods:date id="AF6265B0C06A97131736AAE9C7B01662">2024</mods:date>
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<subSubSection id="11A0E28EFFA11E74FF7BF8D8FB49E138" pageId="2" pageNumber="80" type="nomenclature">
<paragraph id="5905B105FFA11E74FF7BF8D8FCE2E119" blockId="2.[189,1169,1833,1924]" box="[189,826,1833,1859]" pageId="2" pageNumber="80">
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<taxonomicName id="9EBACA86FFA11E74FF7BF8D8FD16E119" authority="Legalov et Perkovsky" authorityName="Legalov et Perkovsky" authorityYear="2024" box="[189,718,1833,1859]" class="Insecta" family="Ciidae" genus="Orthocis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="80" phylum="Arthropoda" rank="species" species="vilhelmseni" status="sp. nov.">
<emphasis id="6BCE6D17FFA11E74FF7BF8D8FE6FE119" bold="true" box="[189,439,1833,1859]" italics="true" pageId="2" pageNumber="80">Orthocis vilhelmseni</emphasis>
Legalov et Perkovsky
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,
<taxonomicNameLabel id="70FDD06CFFA11E74FD1CF8D8FCE2E119" box="[730,826,1833,1859]" pageId="2" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
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https://zoobank.org/
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<subSubSection id="11A0E28EFFA11E74FF7BF898FEF6E1DE" box="[189,302,1897,1924]" pageId="2" pageNumber="80" type="description">
<paragraph id="5905B105FFA11E74FF7BF898FEF6E1DE" blockId="2.[189,1169,1833,1924]" box="[189,302,1897,1924]" pageId="2" pageNumber="80">
(
<figureCitation id="C181AD80FFA11E74FF00F898FEFCE1DE" box="[198,292,1897,1924]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStartId-0="3.[189,264,1971,1995]" captionStartId-1="4.[189,264,1094,1118]" captionTargetBox-0="[220,1370,848,1948]" captionTargetBox-1="[228,1359,189,1074]" captionTargetId-0="figure-487@3.[216,1370,842,1951]" captionTargetId-1="figure-461@4.[228,1359,189,1074]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figure 2. Orthocis vilhelmseni sp. nov., holotype, NHMD 153508: a habitus, dorsal view; b habitus, lateral view, right; c habitus, ventral view; d anterior half of body, ventral view; e anterior half of body, dorso-lateral view, right; f anterior half of body, frontal view." captionText-1="Figure 3. Orthocis vilhelmseni sp. nov., holotype, NHMD 153508, anterior half of body, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.14653164" figureDoi-1="http://doi.org/10.5281/zenodo.14653166" httpUri-0="https://zenodo.org/record/14653164/files/figure.png" httpUri-1="https://zenodo.org/record/14653166/files/figure.png" pageId="2" pageNumber="80">Fig. 2-3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="11A0E28EFFA11E74FF7BF85CFE7DE1B3" pageId="2" pageNumber="80" type="materials_examined">
<paragraph id="5905B105FFA11E74FF7BF85CFE7DE1B3" blockId="2.[189,1399,1964,2025]" pageId="2" pageNumber="80">
<emphasis id="6BCE6D17FFA11E74FF7BF85CFEA1E19D" bold="true" box="[189,377,1965,1991]" pageId="2" pageNumber="80">Type material.</emphasis>
<materialsCitation id="E9D2BB58FFA11E74FE42F85DFE7DE1B3" collectingDate="1959-11-14" collectionCode="NHMD" pageId="2" pageNumber="80" specimenCode="NHMD 153508" specimenCount="1" typeStatus="holotype">
<typeStatus id="86010FA7FFA11E74FE42F85DFE2EE19D" box="[388,502,1964,1991]" pageId="2" pageNumber="80" type="holotype">Holotype</typeStatus>
:
<specimenCode id="091C197EFFA11E74FDC5F85CFD18E19D" box="[515,704,1964,1991]" collectionCode="NHMD" pageId="2" pageNumber="80">NHMD 153508</specimenCode>
with label '
<taxonomicName id="9EBACA86FFA11E74FC96F85DFBA5E19D" authority=", C. V. Hennigsen" authorityName="C. V. Hennigsen" box="[848,1149,1964,1991]" class="Insecta" family="Ciidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="80" phylum="Arthropoda" rank="family">Ciidae, C. V. Hennigsen</taxonomicName>
/
<date id="2D0497C5FFA11E74FB40F85DFACAE19D" box="[1158,1298,1964,1991]" pageId="2" pageNumber="80" value="1959-11-14">
<collectingDate id="3D406E2DFFA11E74FB40F85DFACAE19D" box="[1158,1298,1964,1991]" pageId="2" pageNumber="80" value="1959-11-14">14-11-1959</collectingDate>
</date>
', Danish amber, late Eocene.
</materialsCitation>
</paragraph>
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<subSubSection id="11A0E28EFFA01E75FEDAFF35FD2DE75B" pageId="3" pageNumber="81" type="etymology">
<paragraph id="5905B105FFA01E75FEDAFF35FD2DE75B" blockId="3.[189,1399,196,830]" pageId="3" pageNumber="81">
<emphasis id="6BCE6D17FFA01E75FEDAFF35FE72E684" bold="true" box="[284,426,196,222]" pageId="3" pageNumber="81">Etymology.</emphasis>
The epithet of this new species is dedicated to outstanding hymenopterist Lars Vilhelmsen, keeper of NHMD amber collection.
</paragraph>
</subSubSection>
<subSubSection id="11A0E28EFFA01E72FEDAFEF9FB98E2BE" lastPageId="4" lastPageNumber="82" pageId="3" pageNumber="81" type="description">
<paragraph id="5905B105FFA01E75FEDAFEF9FE67E564" blockId="3.[189,1399,196,830]" pageId="3" pageNumber="81">
<emphasis id="6BCE6D17FFA01E75FEDAFEF9FE6CE778" bold="true" box="[284,436,264,290]" pageId="3" pageNumber="81">Description.</emphasis>
Body black, oblong-oval, convex dorsally, with bristles. Clypeus without teeth. Forehead convex, wide, finely punctate. Eyes well protruded, coarsely faceted, about 0.7 times as long as wide. Antennae clavate, extend beyond middle of pronotum, ten-segmented. Antennomeres first and second suboval. Third antennomere long-oval. Fourth antennomere oval, about 0.8 times as long as antennomere three. Antennomeres four-nine suboval, transverse. Antennal club consists of antennomeres eightten. Tenth antennomere about 1.5 times as long as antennomere nine. Pronotum almost trapezoid, of same length as wide across apex, about 0.8 times as long as wide across middle and about 0.7 times as long as wide across base. Lateral margins narrow, slightly explanate, visible for their entire lengths from above, lacking bristles. Disc with large and small punctures. Scutellum small, subpentagonal. Elytra oval, coarsely punctate, with lateral margins subparallel at basal three-fourths, then converging and forming moderately acute apex, widest at middle, about 2.2 times as long as wide across base, about 1.5 times as long as wide across middle, about 2.4 times as long as wide across apical fourth, about 3.3 times as long as pronotum, with indistinct humeri. Elytral striae absent. Metaventrite weakly convex, about 2.5 times as long as length of mesocoxal cavity, sparsely punctate. Metepisternum narrow, with row of punctures. Abdomen convex. Legs long. Femora clavate. Protibiae untoothed on the outer edge. Apex of protibiae simply rounded. Tarsi long, four-segmented. Last tarsomere very long. Body length:
<quantity id="9E421CE0FFA01E75FE9BFCD2FE63E564" box="[349,443,803,830]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="81" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
<caption id="0DC5E18DFFA01E75FF7BF842FD0DEE52" ID-DOI="http://doi.org/10.5281/zenodo.14653164" ID-Zenodo-Dep="14653164" httpUri="https://zenodo.org/record/14653164/files/figure.png" pageId="3" pageNumber="81" startId="3.[189,264,1971,1995]" targetBox="[220,1370,848,1948]" targetPageId="3" targetType="figure">
<paragraph id="5905B105FFA01E75FF7BF842FD0DEE52" blockId="3.[189,1398,1970,2056]" pageId="3" pageNumber="81">
<emphasis id="6BCE6D17FFA01E75FF7BF842FEFCE191" bold="true" box="[189,292,1971,1995]" pageId="3" pageNumber="81">Figure 2.</emphasis>
<taxonomicName id="9EBACA86FFA01E75FEEBF843FDD4E190" authorityName="Legalov &amp; Vasilenko &amp; Perkovsky" authorityYear="2024" box="[301,524,1970,1994]" class="Insecta" family="Ciidae" genus="Orthocis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="81" phylum="Arthropoda" rank="species" species="vilhelmseni" status="sp. nov.">
<emphasis id="6BCE6D17FFA01E75FEEBF843FDD4E190" box="[301,524,1970,1994]" italics="true" pageId="3" pageNumber="81">Orthocis vilhelmseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70FDD06CFFA01E75FDD2F842FDB4E190" box="[532,620,1971,1994]" pageId="3" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NHMD 153508: a habitus, dorsal view; b habitus, lateral view, right; c habitus, ventral view; d anterior half of body, ventral view; e anterior half of body, dorso-lateral view, right; f anterior half of body, frontal view.
</paragraph>
</caption>
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<paragraph id="5905B105FFA71E72FF7BFBB7FADDE207" blockId="4.[189,1285,1093,1118]" box="[189,1285,1093,1118]" pageId="4" pageNumber="82">
<emphasis id="6BCE6D17FFA71E72FF7BFBB7FEC4E204" bold="true" box="[189,284,1094,1118]" pageId="4" pageNumber="82">Figure 3</emphasis>
.
<taxonomicName id="9EBACA86FFA71E72FEECFBB4FDDFE207" authorityName="Legalov &amp; Vasilenko &amp; Perkovsky" authorityYear="2024" box="[298,519,1093,1117]" class="Insecta" family="Ciidae" genus="Orthocis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="82" phylum="Arthropoda" rank="species" species="vilhelmseni" status="sp. nov.">
<emphasis id="6BCE6D17FFA71E72FEECFBB4FDDFE207" box="[298,519,1093,1117]" italics="true" pageId="4" pageNumber="82">Orthocis vilhelmseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70FDD06CFFA71E72FDC8FBB7FDBDE207" box="[526,613,1094,1117]" pageId="4" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NHMD 153508, anterior half of body, ventral view.
</paragraph>
</caption>
<paragraph id="5905B105FFA71E72FEDAFB59FB98E2BE" blockId="4.[189,1398,1192,1252]" pageId="4" pageNumber="82">
<emphasis id="6BCE6D17FFA71E72FEDAFB59FE66E298" bold="true" box="[284,446,1192,1218]" pageId="4" pageNumber="82">Comparison.</emphasis>
The new species differs from
<taxonomicName id="9EBACA86FFA71E72FCEAFB59FBF6E299" authorityName="Legalov &amp; Vasilenko &amp; Perkovsky" authorityYear="2024" box="[812,1070,1192,1219]" class="Insecta" family="Ciidae" genus="Orthocis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="82" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
<emphasis id="6BCE6D17FFA71E72FCEAFB59FBF6E299" box="[812,1070,1192,1219]" italics="true" pageId="4" pageNumber="82">Orthocis kompantsevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70FDD06CFFA71E72FBF1FB58FB4DE299" box="[1079,1173,1193,1219]" pageId="4" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
in having a bristly body, a densely punctate pronotum and elytra, and longer antennomere nine.
</paragraph>
</subSubSection>
</treatment>
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