408 lines
42 KiB
XML
408 lines
42 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.412.7619" ID-GBIF-Dataset="4a1d248c-92c6-404e-aaa6-da0703439376" ID-PMC="PMC4042698" ID-Pensoft-Pub="1313-2970-412-103" ID-PubMed="24899864" ID-ZBK="12259F296B7743E9ACB2748E7D79E3C7" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-412-103" ModsDocOrigin="ZooKeys 412" ModsDocTitle="A new species of the genus Diplocentrus Peters, 1861 (Scorpiones, Diplocentridae) from Oaxaca, Mexico" checkinTime="1451245838245" checkinUser="pensoft" docAuthor="Santibanez-Lopez, Carlos Eduardo" docDate="2014" docId="D0FE028941ECF545CBAC7FE644D43510" docLanguage="en" docName="ZooKeys 412: 103-116" docOrigin="ZooKeys 412" docSource="http://dx.doi.org/10.3897/zookeys.412.7619" docTitle="Diplocentrus franckei Santibanez-Lopez, 2014, sp. n." docType="treatment" docUuid="B219152E-7FEA-4EDC-985C-592C5C5B0422" docUuidSource="ZooBank" docVersion="4" lastPageNumber="112" masterDocId="8356C345FFD09F47FF972151FFBA7F79" masterDocTitle="A new species of the genus Diplocentrus Peters, 1861 (Scorpiones, Diplocentridae) from Oaxaca, Mexico" masterLastPageNumber="116" masterPageNumber="103" pageNumber="105" updateTime="1668158548050" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of the genus Diplocentrus Peters, 1861 (Scorpiones, Diplocentridae) from Oaxaca, Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santibanez-Lopez, Carlos Eduardo</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>412</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>103</mods:start>
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<mods:end>116</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.412.7619</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.412.7619</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-412-103</mods:identifier>
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<mods:identifier type="ZBK">12259F296B7743E9ACB2748E7D79E3C7</mods:identifier>
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<mods:identifier type="ZooBank">12259F296B7743E9ACB2748E7D79E3C7</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152053276" LSID="urn:lsid:zoobank.org:act:B219152E-7FEA-4EDC-985C-592C5C5B0422" httpUri="http://treatment.plazi.org/id/D0FE028941ECF545CBAC7FE644D43510" lastPageId="9" lastPageNumber="112" pageId="2" pageNumber="105">
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<subSubSection pageId="2" pageNumber="105" type="nomenclature">
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<paragraph pageId="2" pageNumber="105">
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<taxonomicName LSID="http://zoobank.org/B219152E-7FEA-4EDC-985C-592C5C5B0422" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="105">sp. n.</taxonomicNameLabel>
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Figures 1-6
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="105" type="type material">
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<paragraph pageId="2" pageNumber="105">Type material.</paragraph>
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<paragraph pageId="2" pageNumber="105">
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Holotype: Male from MEXICO. OAXACA. Distrito de Villa Alta. Municipio de San Melchor Betaza (CNAN-T0668), km 101 road to Villa Alta
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.224382">17°13.463'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.15207">96°09.124'W</geoCoordinate>
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, 992 m., 21 June 2007, C.
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<normalizedToken originalValue="Santibáñez">Santibanez</normalizedToken>
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and A. Valdez. Paratypes one adult female, one subadult male, one juvenil male and one juvenil female (CNAN- T0669) (same data as holotype). One adult male, three adult females (CNAN-T0670) from San Melchor Betaza
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.251017">17°15.061'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.15313">96°09.188'W</geoCoordinate>
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, 1415 m., 1 June 2007, C.
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<normalizedToken originalValue="Santibáñez">Santibanez</normalizedToken>
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and H. Jara. One adult female, one adult male (AMNH), two adult males, six juvenil male, three adult females and six juvenile female (CNAN) from Municipio de San
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<normalizedToken originalValue="Andrés">Andres</normalizedToken>
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Zoolaga, 6 km south
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.25787">17°15.4722'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.23988">96°14.3928'W</geoCoordinate>
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, 1119 m., 21 June 2007, O. Francke, A. Ballesteros, H.
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<normalizedToken originalValue="Montaño">Montano</normalizedToken>
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, C.
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<normalizedToken originalValue="Santibáñez">Santibanez</normalizedToken>
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and A. Valdez.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="106" pageId="2" pageNumber="105" type="additional material">
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<paragraph pageId="2" pageNumber="105">Additional material.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="106" pageId="2" pageNumber="105">
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MEXICO. OAXACA. Distrito de VILLA ALTA: Municipio de San Juan Tabaa, one adult female (CALA) [
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.304867">17°18.292'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.2065">96°12.390'W</geoCoordinate>
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, 1280 m.], 10 June 2004, T.
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<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
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. One adult female (CALA) same data, 12 June 2004, T.
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<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
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.
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<pageBreakToken pageId="3" pageNumber="106" start="start">One</pageBreakToken>
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adult male (CALA) from Municipio de San Francisco Yovego [
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.55808">17°33.4848'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.22585">96°13.551'W</geoCoordinate>
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, 589 m.], 1 April 2005, T.
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<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
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. Two adult males (CNAN) from Municipio de San Melchor Betaza [
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.251017">17°15.061'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.15313">96°09.188'W</geoCoordinate>
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, 1415 m.], June 2008, R.
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<normalizedToken originalValue="Mejía">Mejia</normalizedToken>
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. One adult female (CNAN) from 9 km from San Andres Yaa on the road to San Juan Tabaa
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.33552">17°20.1312'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.18698">96°11.2188'W</geoCoordinate>
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, 787 m., 3. April 2007, C.
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<normalizedToken originalValue="Santibáñez">Santibanez</normalizedToken>
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and H. Jara. One adult female (CAIMsc-01136) from Municipio San Juan
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<normalizedToken originalValue="Yaeé">Yaee</normalizedToken>
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, Santiago Yagallo [
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.42361">17°25.4166'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.29722">96°17.833'W</geoCoordinate>
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, 1200 m.], 12 March 1997, P. Ruiz Figueroa.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="112" pageId="3" pageNumber="106" type="diagnosis">
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<paragraph pageId="3" pageNumber="106">Diagnosis.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="107" pageId="3" pageNumber="106">
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The following character combination is diagnostic for
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<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">
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<pageBreakToken pageId="4" pageNumber="107" start="start">Diplocentrus</pageBreakToken>
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franckei
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</taxonomicName>
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, sp. n. Total length (adult), 55 to 60 mm. Base coloration (adult) brown to dark brown. Carapace anteromedian notch moderately deep, U-shaped (Fig. 1A). Pedipalp femur, dorsal surface sparsely and finely granular (Fig. 2A). Pedipalp patella, dorsomedian carina moderately developed, crenulate to feebly granular (male); dorsoexternal strongly developed, crenulate to feebly granular (male); externomedian carina moderately developed, crenulate (male; Fig. 2B); ventromedian carina weakly developed to faint, granular (male, female). Pedipalp chela manus, dorsal surface markedly reticulate (male, Fig. 3A) or weakly reticulate (female, Fig. 3B); digital carina strongly developed, smooth to crenulate (male) or weakly to moderately developed, smooth (female); dorsal secondary carina weakly developed to faint, coarsely granular (male) or faint, smooth (female); dorsal external carina weakly developed to faint, smooth to crenulate (male), or faint, smooth (female). Legs I-IV telotarsi, counts of spiniform macrosetae in pro- and retroventral rows, 4-5/5:5/6:6/6:6/6-7 (variation in Table 1); basitarsi spiniform macrosetae pattern: leg I pst, rst, pm, rm; leg II pt, rt, pst, rst, pm, rm, Rm; legs III-IV pt, rt, vt, rst, vst, vm (variation in Table 2). Pectinal tooth count, 12-15, mode = 13 (male) or 11-13, mode = 12 (female) (variation in Table 3).
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</paragraph>
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<caption pageId="4" pageNumber="107">
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<paragraph pageId="4" pageNumber="107">
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Figure 1.
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<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
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sp. n., A Carapace, dorsal aspect B sternum, genital operculum and pectines, ventral aspect.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="107">
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<paragraph pageId="4" pageNumber="107">
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Figure 2.
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<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
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sp. n., A dextral pedipalp femur, dorsal aspect B dextral pedipalp patella, external aspect.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="107">
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<paragraph pageId="4" pageNumber="107">
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Figure 3.
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<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
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sp. n., dextral pedipalp chela, dorsoexternal aspect. A Holotype ♂ (CNAN) B Paratype ♀ (CNAN). Scale bar = 1 mm.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="107">
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<paragraph pageId="4" pageNumber="107">
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Table 1. Telotarsal spiniform macrosetal count (number of macrosetae in pro- and retroventral rows of telotarsi on legs
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<normalizedToken originalValue="I–IV">I-IV</normalizedToken>
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) in
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<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
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sp. n., given as number of legs observed with corresponding proventral (p) and retroventral (r) setal count.
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</paragraph>
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</caption>
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<paragraph pageId="4" pageNumber="107">
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<table pageId="4" pageNumber="107">
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<tr pageId="4" pageNumber="107">
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<th colspan="4" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">Leg I</th>
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<th colspan="4" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">Leg II</th>
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<th colspan="4" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">Leg III</th>
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<th colspan="5" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">Leg IV</th>
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</tr>
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<tr pageId="4" pageNumber="107">
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<th colspan="2" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">p</th>
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<th colspan="2" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">R</th>
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<th colspan="2" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">p</th>
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<th colspan="2" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">r</th>
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<th colspan="2" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">p</th>
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<th colspan="3" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">r</th>
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<th colspan="3" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">p</th>
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<th colspan="2" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">r</th>
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</tr>
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<tr pageId="4" pageNumber="107">
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">4</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">5</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">4</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">5</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">5</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">6</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">5</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">6</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">5</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">6</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">5</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">6</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">7</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">5</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">6</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">7</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">6</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">7</th>
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</tr>
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<tr pageId="4" pageNumber="107">
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<td colspan="1" pageId="4" pageNumber="107" rowspan="1">
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<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
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</td>
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</tr>
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</table>
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</paragraph>
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<caption ID-Table-UUID="3EED9EFDA37731BFDCF8732D11330E00" httpUri="http://table.plazi.org/id/3EED9EFDA37731BFDCF8732D11330E00" pageId="4" pageNumber="107">
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<paragraph pageId="4" pageNumber="107">
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Table 2. Basitarsal spiniform macrosetal count (number of macrosetae of basitarsi on legs I-IV) in
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<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
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sp. n., given as numbers of legs observed with corresponding setae count.
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</paragraph>
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</caption>
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<paragraph pageId="4" pageNumber="107">
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<table pageId="4" pageNumber="107">
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<tr pageId="4" pageNumber="107">
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:left">Leg</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">n</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">pt</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">rt</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">vt</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">pst</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">rst</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">vst</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">pm</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">rm</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">vm</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">Rm</th>
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</tr>
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</table>
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</paragraph>
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<caption pageId="4" pageNumber="107">
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<paragraph pageId="4" pageNumber="107">
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Table 3. Pectinal tooth count (number of teeth per pecten) in
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<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
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sp. n., given as number of male and female pectines observed with corresponding tooth count.
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</paragraph>
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</caption>
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<paragraph pageId="4" pageNumber="107">
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<table pageId="4" pageNumber="107">
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<tr pageId="4" pageNumber="107">
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">11</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">12</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">13</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">14</th>
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<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">15</th>
|
||
<th colspan="1" pageId="4" pageNumber="107" rowspan="1" style="text-align:center">16</th>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="107">
|
||
<td colspan="1" pageId="4" pageNumber="107" rowspan="2">
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="4" pageNumber="107" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="5" pageNumber="108">
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">
|
||
<pageBreakToken pageId="5" pageNumber="108" start="start">Diplocentrus</pageBreakToken>
|
||
franckei
|
||
</taxonomicName>
|
||
sp. n. resembles
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus mexicanus" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="mexicanus">Diplocentrus mexicanus</taxonomicName>
|
||
Peters, 1861,
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus melici" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="melici">Diplocentrus melici</taxonomicName>
|
||
Armas et al., 2004 and
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus jaca" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="jaca">Diplocentrus jaca</taxonomicName>
|
||
Armas and
|
||
<normalizedToken originalValue="Martín-Frías">Martin-Frias</normalizedToken>
|
||
, 2000 in adult size and coloration, but can be distinguished as follows. The leg III telotarsi counts of spiniform macrosetae is higher in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus mexicanus" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="mexicanus">Diplocentrus mexicanus</taxonomicName>
|
||
(7/8), while is lower in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
, sp. n. (6/6). The carapace and mesosoma surfaces are strongly granular (male) in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus mexicanus" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="mexicanus">Diplocentrus mexicanus</taxonomicName>
|
||
, but weakly granulose to shagreened in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
(male). Metasomal and pedipalp carination is slightly granular to crenulate in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus mexicanus" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="mexicanus">Diplocentrus mexicanus</taxonomicName>
|
||
(male), and crenulate to smooth in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
(male). The rt spiniform macroseta in leg I is present in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus mexicanus" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="mexicanus">Diplocentrus mexicanus</taxonomicName>
|
||
, while it is absent in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
; also rsm in leg II is present in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus mexicanus" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="mexicanus">Diplocentrus mexicanus</taxonomicName>
|
||
and absent in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="5" pageNumber="108">
|
||
Pedipalp surfaces in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus melici" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="melici">Diplocentrus melici</taxonomicName>
|
||
are punctuate, and minutely granular or smooth in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
. Carapace anteromargin notch U-shaped in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
, while in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus melici" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="melici">Diplocentrus melici</taxonomicName>
|
||
is V-shaped. Pedipalp patella dorsoexternal and externomedian carinae are strongly developed in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
(male), but both are obsolete in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus melici" order="Scorpiones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="species" species="melici">Diplocentrus melici</taxonomicName>
|
||
(male).
|
||
</paragraph>
|
||
<paragraph lastPageId="6" lastPageNumber="109" pageId="5" pageNumber="108">
|
||
Adults of
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="franckei">
|
||
<pageBreakToken pageId="6" pageNumber="109" start="start">Diplocentrus</pageBreakToken>
|
||
franckei
|
||
</taxonomicName>
|
||
(55-60 mm) are smaller than adults of
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus jaca" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="jaca">Diplocentrus jaca</taxonomicName>
|
||
(75-90 mm). Metasomal segment V ventral carinae are strongly serrate in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus jaca" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="jaca">Diplocentrus jaca</taxonomicName>
|
||
, while in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
are granular. Carapace anteromargin notch is strongly deep (reaching the second pair of lateral ocelli level) in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus jaca" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="jaca">Diplocentrus jaca</taxonomicName>
|
||
, while it is moderately deep (reaching the first pair of lateral ocelli level) in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
. Pedipalp chela is slender in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus jaca" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="jaca">Diplocentrus jaca</taxonomicName>
|
||
(male, chela length:ratio= 4.97), and rounded in
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
(male, chela length:ratio= 2.32).
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="109">Description. Based on holotype (male) and paratype (male) (Fig. 4A, B) with differences in paratype (female) (Figs 4C, D) noted. Measurements in Table 4.</paragraph>
|
||
<caption pageId="6" pageNumber="109">
|
||
<paragraph pageId="6" pageNumber="109">
|
||
Figure 4.
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
sp. n., habitus, dorsal (A, C) and ventral (B, D) aspect. A, B. Holotype ♂ (CNAN) C, D Paratype ♀ (CNAN). Scale bars = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-Table-UUID="9E21FC8EBCEC779D00925CF2BEF279E0" httpUri="http://table.plazi.org/id/9E21FC8EBCEC779D00925CF2BEF279E0" pageId="6" pageNumber="109">
|
||
<paragraph pageId="6" pageNumber="109">
|
||
Table 4. Measurements (mm) of male and female type specimens of
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="6" pageNumber="109" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
, sp. n., in the CNAN.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="6" pageNumber="109">
|
||
<table pageId="6" pageNumber="109">
|
||
<tr pageId="6" pageNumber="109">
|
||
<th colspan="1" pageId="6" pageNumber="109" rowspan="1" style="text-align:center">♂ Holotype</th>
|
||
<th colspan="1" pageId="6" pageNumber="109" rowspan="1" style="text-align:center">♀ Paratype</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="109">Coloration: Carapace brown (male) or darker brown (female), moderately infuscate throughout, uniformly around median ocelli, variegated elsewhere. Coxosternum pale yellow to light brown. Pedipalps brown to dark brown, carinae darker. Legs brown to reddish brown, uniformly and faintly infuscate. Mesosoma brown (male) to dark brown (female), tergites moderately (male) to weakly (female) infuscate; sternites pale brown to pale yellow. Metasoma reddish brown (male) to dark brown (female). Telson brown, uniformly infuscate.</paragraph>
|
||
<paragraph pageId="6" pageNumber="109">Carapace: Anterior margin weakly setose; anteromedian notch weakly to moderately deep, U-shaped (Fig. 1A). Frontal lobes and interocular surface moderately granular; surfaces around median ocular tubercle shagreened; other surfaces minutely, sparsely and finely granular. Three pairs of subequal lateral ocelli.</paragraph>
|
||
<paragraph pageId="7" pageNumber="110">
|
||
<pageBreakToken pageId="7" pageNumber="110" start="start">Pedipalps</pageBreakToken>
|
||
: Orthobothriotaxic, Type C. Femur width greater than height (Fig. 2A); dorsal intercarinal surface flat, sparsely granular; external intercarinal surface smooth; ventral intercarinal surface flat, shagreened to minutely, finely granular; internal intercarinal surface coarsely and densely granular; dorsointernal carina strongly developed, comprising several large granules; dorsoexternal carina moderately to weakly developed, comprising few large granules; ventroexternal carina obsolete; ventrointernal carina moderately developed, comprising large granules. Patella, dorsal and external intercarinal surfaces slightly, minutely and finely granular-reticulate (Fig. 2B); ventral intercarinal surface flat, minutely granular between ventrointernal and ventromedian carinae, smooth to slightly reticulate between ventromedian and ventroexternal carinae (male) or smooth (female); internal intercarinal surface sparsely granular; proximal tubercle strongly developed, comprising two large granules; dorsointernal carina obsolete; dorsomedian carina moderately developed, crenulate to feebly granular; dorsoexternal carina strongly developed, crenuate to feebly granular (male) or weakly developed to faint, smooth (female); externomedian and ventroexternal carinae moderately developed, smooth to slightly crenulate; ventromedian carina weakly to moderately developed, granular (male) or weakly developed, smooth to slightly granular (female); ventrointernal carina weakly to moderately developed, granular. Chela manus rounded (male, female) height greater than width, moderately (male) to sparsely setose (female); dorsal intercarinal surface granular-reticulate (male, female); external intercarinal surface granular-reticulate (male) or reticulate (female) (Fig. 3A, B); dorsal margin strongly developed, granular; digital carina strongly developed, crenulate to smooth (male, female); dorsal secondary carina weakly developed, granular; external secondary carina weakly developed, smooth to slightly granular (male) or weakly developed, smooth (female); ventroexternal carina weakly developed, crenulate; ventromedian carina moderately to strongly developed, coarsely granular to crenulate (male) or crenulate to smooth (female), directed towards midpoint of movable finger articulation; ventrointernal carina weakly developed, smooth to slightly crenulate; internoventral, internomedian and internodorsal carinae weakly developed, slightly granular; internal surface with shallow longitudinal depression where chela rest against patella. Chela fixed finger slightly curved, length equals femur length; dorsal surface smooth and densely setose proximally, external surface flat, internal surface shallowly concave.
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="111">
|
||
<pageBreakToken pageId="8" pageNumber="111" start="start">Legs</pageBreakToken>
|
||
: Legs I-IV femora and tibiae, prolateral surfaces shagreened; telotarsi, counts of spiniform macrosetae in pro- and retroventral rows (dextral/sinistral): 4/5 5/5:5/5 5/6:6/6 6/6:6/6 6/6 (holotype); basitarsi, spiniform macrosetae pattern: Leg I pst. rst, pm, rm; leg II pt, rt, pst, rst, pm, rm, Rm; legs III-IV pt, rt, vt, rst, vst, vm (holotype).
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="111">Pectines: Tooth count: 13-14 (male; Fig. 1B) or 11-11 (female)</paragraph>
|
||
<paragraph pageId="8" pageNumber="111">Mesosoma: Tergites I-VI, pre-tergites smooth, post-tergites minutely granular; VII granular. Sternites smooth; dorsosubmedian and dorsolateral carinae weakly to moderately developed, crenulate.</paragraph>
|
||
<paragraph pageId="9" pageNumber="112">
|
||
<pageBreakToken pageId="9" pageNumber="112" start="start">Metasoma</pageBreakToken>
|
||
: Metasomal segments I-V, dorsal intercarinal surfaces shagreened on segments I-IV, smooth on V; lateral intercarinal surfaces shagreened on segments I-IV, smooth on V; ventral intercarinal surfaces smooth on I-V. Segments I-IV, dorsolateral carinae weakly developed, granular; lateral surpamedian carinae moderately developed, granular on I, moderately developed, granular to crenulate on II-III; moderately developed, granular to serrate on IV; lateral inframedian carinae strongly developed, crenulae on I, moderately developed, crenulate to granular on II-IV; ventrolateral carinae strongly developed, crenulate on I, moderately developed, crenulate to slightly granular on II-IV; ventrosubmedian carinae strongly developed, smooth to crenulate on I, moderately developed, smooth to crenulate on II-IV. Segment V length: pedipalp femur length ratio, 1.31 (male), 1.09 (female); dorsolateral carinae strongly developed, crenulate to feebly granular; lateral inframedian carinae moderately to weakly developed, feebly granular on I-III, weakly developed, sparsely granular on IV; ventrolateral carinae moderately developed, granular to feebly serrate, with subspiniform granules; ventromedian carina moderately to strongly developed, granular, with subspiniform granules; ventral transverse carina moderately developed, comprising four subspiniform granules posteriorly; anal arch semicircular; anal subterminal carina moderately developed, comprising twelve subspiniform granules; anal terminal carina vestigial, weakly granular.
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="112">Telson: Telson, width: length ratio, 0.46 (male), 0.58 (female). Vesicle, lateral surface smooth; ventral surface granular anteriorly. Subaculear tubercle stout, subconical. Aculeus length 1.5.</paragraph>
|
||
<paragraph pageId="9" pageNumber="112">Hemispermatophore: Lamelliform, weakly sclerotized (Fig. 5); total length 8.9 mm; distal lamella, length 4.8 mm, capsular region width, 1.8 mm; median lobe narrow, margin with small crenulations.</paragraph>
|
||
<caption pageId="9" pageNumber="112">
|
||
<paragraph pageId="9" pageNumber="112">
|
||
Figure 5.
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="9" pageNumber="112" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
sp. n., paratype ♂ (CNAN) sinistral hemispermatophore. A dorsal aspect B ental view C Capsular region. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="112" type="etymology">
|
||
<paragraph pageId="9" pageNumber="112">Etymology.</paragraph>
|
||
<paragraph pageId="9" pageNumber="112">
|
||
This species is dedicated to Dr. Oscar Francke, for his enormous contribution to the taxonomy and systematics of the genus
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus" order="Scorpiones" pageId="9" pageNumber="112" phylum="Arthropoda" rank="genus">Diplocentrus</taxonomicName>
|
||
, and his guidance through the years in my education.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="112" type="distribution">
|
||
<paragraph pageId="9" pageNumber="112">Distribution.</paragraph>
|
||
<paragraph pageId="9" pageNumber="112">
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="9" pageNumber="112" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
, sp. n. is known from the Villa Alta district within the Northern mountain range in Oaxaca; in the municipalities of San Andres Yaa, San Andres Zoolaga, San Juan Tabaa, San Melchor Betaza (Fig. 6).
|
||
</paragraph>
|
||
<caption pageId="9" pageNumber="112">
|
||
<paragraph pageId="9" pageNumber="112">
|
||
Figure 6.
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="9" pageNumber="112" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
sp. n., known records in Oaxaca (in circles).
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus mexicanus" order="Scorpiones" pageId="9" pageNumber="112" phylum="Arthropoda" rank="species" species="mexicanus">Diplocentrus mexicanus</taxonomicName>
|
||
(type locality in triangle).
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus jaca" order="Scorpiones" pageId="9" pageNumber="112" phylum="Arthropoda" rank="species" species="jaca">Diplocentrus jaca</taxonomicName>
|
||
(type locality in square).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="112" type="ecology">
|
||
<paragraph pageId="9" pageNumber="112">Ecology.</paragraph>
|
||
<paragraph pageId="9" pageNumber="112">
|
||
This species was observed first on the soil surface, walking at night with UV detection; later it was collected inside houses, under stones and also in the crevices of rock walls. It was also found doorkeeping at burrow entrances in walls of road cuts. The burrows were constructed at an angle of ca. 30° to the wall, ca. 40-50 cm long and mostly straight with some turns around stones in the soil matrix. The dominant vegetation was the transition between dry tropical forest and pine-oak forest at 1500 m.
|
||
<taxonomicName class="Arachnida" family="Buthidae" genus="Centruroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centruroides serrano" order="Scorpiones" pageId="9" pageNumber="112" phylum="Arthropoda" rank="species" species="serrano">Centruroides serrano</taxonomicName>
|
||
<normalizedToken originalValue="Santibáñez-López">Santibanez-Lopez</normalizedToken>
|
||
& Ponce-Saavedra, 2009 was collected in sympatry. The habitat and habitus of
|
||
<taxonomicName class="Arachnida" family="Diplocentridae" genus="Diplocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocentrus franckei" order="Scorpiones" pageId="9" pageNumber="112" phylum="Arthropoda" rank="species" species="franckei">Diplocentrus franckei</taxonomicName>
|
||
, sp. n. are consistent with the pelophilous ecomorphotype (
|
||
<bibRefCitation author="Prendini, L" editor="Fet, V" journalOrPublisher="Burnham Beeches, British Arachnological Society, UK" pageId="10" pageNumber="113" pagination="113 - 118" title="Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revised." volumeTitle="Scorpions 2001. In Memoriam Gary A. Polis" year="2001">Prendini 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |