275 lines
34 KiB
XML
275 lines
34 KiB
XML
<document id="6949493034D508FA9375576F583EC14F" ID-CLB-Dataset="39225" ID-DOI="10.11646/zootaxa.4059.2.2" ID-GBIF-Dataset="65f3dd28-09dd-4739-99cc-0031cbd72b0f" ID-ISSN="1175-5326" ID-Zenodo-Dep="253812" ID-ZooBank="5D7E8244-3A68-40A7-ABCF-8932C6492948" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461361456702" checkinUser="plazi" docAuthor="Steinhoff, Philip O. M. & Uhl, Gabriele" docDate="2015" docId="D04487FBA670FFB1FF78F8DAFD2517AC" docLanguage="en" docName="zt04059p276.pdf" docOrigin="Zootaxa 4059 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeliccia pyriformis Laidlaw 1932" docType="treatment" docVersion="8" lastPageNumber="267" masterDocId="2C7DFF83A678FFBBFFEFFF94FFB81013" masterDocTitle="Taxonomy and nomenclature of some mainland SE-Asian Coeliccia species (Odonata, Platycnemididae) using micro-CT analysis" masterLastPageNumber="276" masterPageNumber="257" pageNumber="265" updateTime="1698661837837" updateUser="ExternalLinkService">
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<mods:titleInfo id="3C4D4DBE7B3E1E5573A0A26E74EBF736">
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<mods:title id="77BAE331EC57DA96511CD223492A5AAE">Taxonomy and nomenclature of some mainland SE-Asian Coeliccia species (Odonata, Platycnemididae) using micro-CT analysis</mods:title>
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<mods:namePart id="780BA395CE95572B4ED1B9A94C251E87">Steinhoff, Philip O. M.</mods:namePart>
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<mods:name id="EFEF9AD75F8945F0F3A2D79951C9ACA6" type="personal">
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<mods:roleTerm id="2A8ED7637FD2729AFCB3B832B42904E9">Author</mods:roleTerm>
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<mods:namePart id="507463DDA4B94A1FAE83D70F86E0347C">Uhl, Gabriele</mods:namePart>
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<mods:typeOfResource id="9C1464C41464B782C4945F443D7733BE">text</mods:typeOfResource>
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<mods:title id="958CA9BD59F1B179F5B5212930901C63">Zootaxa</mods:title>
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<mods:part id="D999D8D1F9B3E19AFCDBD0BE068C5D04">
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<mods:date id="C05B997832A152021069042AB4C81919">2015</mods:date>
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<mods:number id="5563D797EB4DD6664AF0DDA393DEAE64">4059</mods:number>
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<mods:detail id="93981B577E39A955E668F294B46F7362" type="issue">
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<mods:number id="BEA52F9BAE4D0DA5D00C8B6BD09E3615">2</mods:number>
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<mods:identifier id="3F82B87DBF2880F4C72F2EC2B1703E83" type="DOI">10.11646/zootaxa.4059.2.2</mods:identifier>
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<treatment id="D04487FBA670FFB1FF78F8DAFD2517AC" ID-DOI="http://doi.org/10.5281/zenodo.6100443" ID-GBIF-Taxon="119648610" ID-Zenodo-Dep="6100443" LSID="urn:lsid:plazi:treatment:D04487FBA670FFB1FF78F8DAFD2517AC" httpUri="http://treatment.plazi.org/id/D04487FBA670FFB1FF78F8DAFD2517AC" lastPageId="10" lastPageNumber="267" pageId="8" pageNumber="265">
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<subSubSection id="10F76566A670FFB3FF78F8DAFAA217E5" pageId="8" pageNumber="265" type="nomenclature">
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<paragraph id="585236EDA670FFB3FF78F8DAFDFA177A" blockId="8.[151,578,1870,1897]" box="[151,578,1870,1897]" pageId="8" pageNumber="265">
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<heading id="031A8181A670FFB3FF78F8DAFDFA177A" bold="true" box="[151,578,1870,1897]" fontSize="11" level="1" pageId="8" pageNumber="265" reason="1">
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<taxonomicName id="9FED4D6EA670FFB3FF78F8DAFDFA177A" ID-CoL="WQD3" authority="Laidlaw, 1932" authorityName="Laidlaw" authorityYear="1932" box="[151,578,1870,1897]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="265" phylum="Arthropoda" rank="species" species="pyriformis">
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<emphasis id="6A99EAFFA670FFB3FF78F8DAFDFA177A" bold="true" box="[151,578,1870,1897]" pageId="8" pageNumber="265">
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<emphasis id="6A99EAFFA670FFB3FF78F8DAFE33177B" bold="true" box="[151,395,1870,1896]" italics="true" pageId="8" pageNumber="265">Coeliccia pyriformis</emphasis>
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<bibRefCitation id="3C7C4B1CA670FFB3FE7CF8DBFDFA177A" author="Laidlaw" box="[403,578,1871,1897]" pageId="8" pageNumber="265" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw, 1932</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="585236EDA670FFB3FF78F802FAA217E5" blockId="8.[151,1436,1941,2038]" pageId="8" pageNumber="265">
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<collectingCountry id="20FA767DA670FFB3FF78F802FF0E17BD" box="[151,182,1942,1966]" name="American Samoa" pageId="8" pageNumber="265">As</collectingCountry>
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pointed out by
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<bibRefCitation id="3C7C4B1CA670FFB3FE89F802FE4317BD" author="Dow" box="[358,507,1942,1966]" pageId="8" pageNumber="265" refString="Dow, R. A. (2011 a) Coeliccia pyriformis. In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.2. Available from: http: // www. iucnredlist. org / details / 167436 / 0 (accessed 11 April 2013)" type="journal article" year="2011" yearSuffix="a">Dow (2011a)</bibRefCitation>
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, the original description of
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<taxonomicName id="9FED4D6EA670FFB3FCDFF801FBAF17BD" box="[816,1047,1941,1966]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="265" phylum="Arthropoda" rank="species" species="pyriformis">
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<emphasis id="6A99EAFFA670FFB3FCDFF801FBAF17BD" box="[816,1047,1941,1966]" italics="true" pageId="8" pageNumber="265">Coeliccia pyriformis</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="3C7C4B1CA670FFB3FBAFF801FB5517BD" author="Laidlaw" box="[1088,1261,1941,1966]" pageId="8" pageNumber="265" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw (1932)</bibRefCitation>
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lacks drawings as well as a sufficiently detailed description. Here, we redescribe the
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<typeStatus id="8756884FA670FFB3FC27F82DFB9217C1" box="[968,1066,1977,2002]" pageId="8" pageNumber="265" type="holotype">holotype</typeStatus>
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and discuss inaccuracies in the original description. The potential synonymy of
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<taxonomicName id="9FED4D6EA670FFB3FD5AF84AFCF217E5" box="[693,842,2014,2038]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="265" phylum="Arthropoda" rank="species" species="pyriformis">
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<emphasis id="6A99EAFFA670FFB3FD5AF84AFCF217E5" box="[693,842,2014,2038]" italics="true" pageId="8" pageNumber="265">C. pyriformis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="9FED4D6EA670FFB3FC6EF84AFB3A17E5" authority="Asahina, 1997" authorityName="Asahina" authorityYear="1997" box="[897,1154,2013,2038]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="265" phylum="Arthropoda" rank="species" species="acco">
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<emphasis id="6A99EAFFA670FFB3FC6EF84AFC6E17E6" box="[897,982,2014,2037]" italics="true" pageId="8" pageNumber="265">C. acco</emphasis>
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<bibRefCitation id="3C7C4B1CA670FFB3FC32F849FB3A17E5" author="Asahina" box="[989,1154,2013,2038]" pageId="8" pageNumber="265" refString="Asahina, S. (1997) Records of the northernVietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 5: Coenagrionidae, Protoneuridae, Platycnemididae. Bulletin of the National Science Museum, Tokyo (A), 23, 17 - 34." type="journal article" year="1997">Asahina, 1997</bibRefCitation>
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</taxonomicName>
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, is discussed.
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</paragraph>
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</subSubSection>
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<subSubSection id="10F76566A671FFB2FF28FF03FE9E10C6" pageId="9" pageNumber="266" type="materials_examined">
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<paragraph id="585236EDA671FFB2FF28FF03FE9E10C6" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="266">
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<emphasis id="6A99EAFFA671FFB2FF28FF03FE0D10A3" bold="true" box="[199,437,151,176]" pageId="9" pageNumber="266">Material examined.</emphasis>
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<specimenCount id="4EEBFD64A671FFB2FE2FFF0CFE5B10A3" box="[448,483,152,176]" pageId="9" pageNumber="266" type="generic">1♂</specimenCount>
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<typeStatus id="8756884FA671FFB2FE00FF03FDE310A3" box="[495,603,151,176]" pageId="9" pageNumber="266" type="holotype">Holotype</typeStatus>
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: “Bao Ha,
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<collectingCountry id="20FA767DA671FFB2FD0CFF03FC8F10A3" box="[739,823,151,176]" name="Vietnam" pageId="9" pageNumber="266">Tonkin</collectingCountry>
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”,
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<date id="2C53102DA671FFB2FCBEFF0CFC4D10A3" box="[849,1013,151,176]" pageId="9" pageNumber="266" value="1924-07-21">21. July 1924</date>
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, leg. H. Stevens [
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<collectionCode id="3EFCAE28A671FFB2FB26FF0CFA9A10BC" box="[1225,1314,152,175]" pageId="9" pageNumber="266">BMNH</collectionCode>
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,
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<collectionCode id="3EFCAE28A671FFB2FADDFF0CFA2410BC" LSID="urn:lsid:biocol.org:col:34665" box="[1330,1436,152,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="266">NHMUK</collectionCode>
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010266914].
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</paragraph>
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</subSubSection>
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<subSubSection id="10F76566A671FFB1FF28FF4BFBD01524" lastPageId="10" lastPageNumber="267" pageId="9" pageNumber="266" type="description">
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<paragraph id="585236EDA671FFB2FF28FF4BFEBF119B" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="266">
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<emphasis id="6A99EAFFA671FFB2FF28FF4BFEE110EB" bold="true" box="[199,345,223,248]" pageId="9" pageNumber="266">Description.</emphasis>
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Head and prothorax (
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<figureCitation id="C0D62A68A671FFB2FDBEFF4BFD2D10E4" box="[593,661,223,248]" captionStart="FIGURE 7" captionStartId="9.[151,250,1807,1829]" captionTargetBox="[317,1269,445,1770]" captionTargetId="figure@9.[317,1269,445,1786]" captionTargetPageId="9" captionText="FIGURE 7. Coeliccia pyriformis Holotype. A head dorsolateral, B prothorax and thorax dorsolateral, C prothorax and thorax dorsal, D anal appendages dorsolateral, E anal appendages lateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253819/files/figure.png" pageId="9" pageNumber="266">Fig. 7</figureCitation>
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A): Head black, anteclypeus, antefrons and genae cream white; two pale spots between lateral ocelli and base of antennae. Prothorax entirely black, hindlobe slightly raised. Synthorax (
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<figureCitation id="C0D62A68A671FFB2FEA2FEB3FE2A112C" box="[333,402,295,320]" captionStart="FIGURE 7" captionStartId="9.[151,250,1807,1829]" captionTargetBox="[317,1269,445,1770]" captionTargetId="figure@9.[317,1269,445,1786]" captionTargetPageId="9" captionText="FIGURE 7. Coeliccia pyriformis Holotype. A head dorsolateral, B prothorax and thorax dorsolateral, C prothorax and thorax dorsal, D anal appendages dorsolateral, E anal appendages lateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253819/files/figure.png" pageId="9" pageNumber="266">Fig. 7</figureCitation>
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B–C): Black with grey-purplish markings that are nearly faded away. Dorsally with two inward curved stripes (
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<figureCitation id="C0D62A68A671FFB2FE4FFED8FE671177" box="[416,479,332,357]" captionStart="FIGURE 7" captionStartId="9.[151,250,1807,1829]" captionTargetBox="[317,1269,445,1770]" captionTargetId="figure@9.[317,1269,445,1786]" captionTargetPageId="9" captionText="FIGURE 7. Coeliccia pyriformis Holotype. A head dorsolateral, B prothorax and thorax dorsolateral, C prothorax and thorax dorsal, D anal appendages dorsolateral, E anal appendages lateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253819/files/figure.png" pageId="9" pageNumber="266">Fig 7</figureCitation>
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C); mesepimeron mostly black, metepisternum and metepimeron covered by large markings.
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</paragraph>
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<caption id="0C926665A671FFB2FF78F89BFF5E1771" httpUri="https://zenodo.org/record/253819/files/figure.png" pageId="9" pageNumber="266" targetBox="[317,1269,445,1770]" targetPageId="9">
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<paragraph id="585236EDA671FFB2FF78F89BFF5E1771" blockId="9.[151,1436,1807,1890]" pageId="9" pageNumber="266">
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<emphasis id="6A99EAFFA671FFB2FF78F89BFEAB1736" bold="true" box="[151,275,1807,1829]" pageId="9" pageNumber="266">FIGURE 7.</emphasis>
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<taxonomicName id="9FED4D6EA671FFB2FEF5F89BFE541736" box="[282,492,1807,1829]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="266" phylum="Arthropoda" rank="species" species="pyriformis">
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<emphasis id="6A99EAFFA671FFB2FEF5F89BFE541736" box="[282,492,1807,1829]" italics="true" pageId="9" pageNumber="266">Coeliccia pyriformis</emphasis>
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</taxonomicName>
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Holotype.
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<emphasis id="6A99EAFFA671FFB2FDB1F89BFDC81737" bold="true" box="[606,624,1807,1828]" pageId="9" pageNumber="266">A</emphasis>
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head dorsolateral,
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<emphasis id="6A99EAFFA671FFB2FCDCF884FCFC1736" bold="true" box="[819,836,1808,1829]" pageId="9" pageNumber="266">B</emphasis>
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prothorax and thorax dorsolateral,
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<emphasis id="6A99EAFFA671FFB2FB42F89BFB071736" bold="true" box="[1197,1215,1807,1829]" pageId="9" pageNumber="266">C</emphasis>
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prothorax and thorax dorsal,
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<emphasis id="6A99EAFFA671FFB2FF0EF8BAFF4B1750" bold="true" box="[225,243,1838,1859]" pageId="9" pageNumber="266">D</emphasis>
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anal appendages dorsolateral,
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<emphasis id="6A99EAFFA671FFB2FDC3F8BAFD851750" bold="true" box="[556,573,1838,1859]" pageId="9" pageNumber="266">E</emphasis>
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anal appendages lateral,
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<emphasis id="6A99EAFFA671FFB2FCD3F8BAFCF31750" bold="true" box="[828,843,1838,1859]" pageId="9" pageNumber="266">F</emphasis>
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anal appendages dorsal,
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<emphasis id="6A99EAFFA671FFB2FBA6F8BAFBE51757" bold="true" box="[1097,1117,1838,1860]" pageId="9" pageNumber="266">G</emphasis>
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left Fw,
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<emphasis id="6A99EAFFA671FFB2FB58F8BAFB731750" bold="true" box="[1207,1227,1838,1859]" pageId="9" pageNumber="266">H</emphasis>
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left Hw.
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<emphasis id="6A99EAFFA671FFB2FAC6F8BAFA241757" box="[1321,1436,1838,1860]" italics="true" pageId="9" pageNumber="266">Scale bars:</emphasis>
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500Μm.
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</paragraph>
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</caption>
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<caption id="0C926665A672FFB1FF78FC87FBC21354" httpUri="https://zenodo.org/record/253820/files/figure.png" pageId="10" pageNumber="267" targetBox="[205,1383,199,758]" targetPageId="10">
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<paragraph id="585236EDA672FFB1FF78FC87FBC21354" blockId="10.[151,1436,787,839]" pageId="10" pageNumber="267">
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<emphasis id="6A99EAFFA672FFB1FF78FC87FEAA133B" bold="true" box="[151,274,787,809]" pageId="10" pageNumber="267">FIGURE 8.</emphasis>
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Genital ligulae as reconstructed from micro-CT (
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<emphasis id="6A99EAFFA672FFB1FCEAFC80FC09133A" box="[773,945,787,809]" italics="true" pageId="10" pageNumber="267">orientation bars:</emphasis>
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dl dorsolateral, l lateral, p posterior, v ventral, vl ventrolateral).
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<emphasis id="6A99EAFFA672FFB1FEC3FCA5FEE41355" bold="true" box="[300,348,817,839]" pageId="10" pageNumber="267">A–B</emphasis>
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Genital ligula of
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<taxonomicName id="9FED4D6EA672FFB1FDE1FCA6FD2D1354" box="[526,661,817,839]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="pyriformis">
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<emphasis id="6A99EAFFA672FFB1FDE1FCA6FD2D1354" box="[526,661,817,839]" italics="true" pageId="10" pageNumber="267">C. pyriformis</emphasis>
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</taxonomicName>
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holotype,
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<emphasis id="6A99EAFFA672FFB1FCEEFCA5FC881354" bold="true" box="[769,816,817,839]" pageId="10" pageNumber="267">B–C</emphasis>
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genital ligula of
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<taxonomicName id="9FED4D6EA672FFB1FC32FCA6FBCE1354" box="[989,1142,817,839]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="cyanomelas">C. cyanomelas</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph id="585236EDA672FFB1FF28FCFAFE8313E0" blockId="10.[151,1437,878,1875]" pageId="10" pageNumber="267">
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Wings (
|
||
<figureCitation id="C0D62A68A672FFB1FEF0FCFAFEDA1394" box="[287,354,878,903]" captionStart="FIGURE 7" captionStartId="9.[151,250,1807,1829]" captionTargetBox="[317,1269,445,1770]" captionTargetId="figure@9.[317,1269,445,1786]" captionTargetPageId="9" captionText="FIGURE 7. Coeliccia pyriformis Holotype. A head dorsolateral, B prothorax and thorax dorsolateral, C prothorax and thorax dorsal, D anal appendages dorsolateral, E anal appendages lateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253819/files/figure.png" pageId="10" pageNumber="267">Fig. 7</figureCitation>
|
||
G–H): Hyaline, with black Pt, covering about 1¼ cells. 16 Px in both Hw, 17 Px in left Fw and 18 Px in right Fw.
|
||
<collectingCountry id="20FA767DA672FFB1FE81FC07FE2913B8" box="[366,401,915,939]" name="Poland" pageId="10" pageNumber="267">RP</collectingCountry>
|
||
2 arising at Px
|
||
<quantity id="9F159B08A672FFB1FDD0FC06FDD413B8" box="[575,620,914,939]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="267" unit="in" value="6.0">6 in</quantity>
|
||
both Hw, at Px
|
||
<quantity id="9F159B08A672FFB1FCC8FC07FCEC13B8" box="[807,852,914,939]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="10" pageNumber="267" unit="in" value="7.0">7 in</quantity>
|
||
left Fw and at Px
|
||
<quantity id="9F159B08A672FFB1FBC5FC07FBEE13B8" box="[1066,1110,914,939]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="10" pageNumber="267" unit="in" value="8.0">8 in</quantity>
|
||
right Fw. IR1 arises slightly distal to Px
|
||
<quantity id="9F159B08A672FFB1FECEFC23FEE313DC" box="[289,347,950,975]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="267" unit="in" value="10.0">10 in</quantity>
|
||
both Fw and slightly distal to Px
|
||
<quantity id="9F159B08A672FFB1FD0CFC23FCB713DC" box="[739,783,950,975]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="10" pageNumber="267" unit="in" value="9.0">9 in</quantity>
|
||
both Hw. IR2 arises distal to subnodus and is inserted at subnodal vein.
|
||
</paragraph>
|
||
<paragraph id="585236EDA672FFB1FF28FC6AFC3B144C" blockId="10.[151,1437,878,1875]" pageId="10" pageNumber="267">Abdomen: Black, long and slender; ventral part of the abdomen cream white, the white extending upwards at the end of the segments, forming lateral spots that are visible from S3–7, the spot at S7 being notably smaller (and not visible dorsally) than those on S3–6. Segments 9 and 10 ochre.</paragraph>
|
||
<paragraph id="585236EDA672FFB1FF28FBFEFB3414B4" blockId="10.[151,1437,878,1875]" pageId="10" pageNumber="267">
|
||
Anal appendages (
|
||
<figureCitation id="C0D62A68A672FFB1FE78FBFEFE611491" box="[407,473,1130,1155]" captionStart="FIGURE 7" captionStartId="9.[151,250,1807,1829]" captionTargetBox="[317,1269,445,1770]" captionTargetId="figure@9.[317,1269,445,1786]" captionTargetPageId="9" captionText="FIGURE 7. Coeliccia pyriformis Holotype. A head dorsolateral, B prothorax and thorax dorsolateral, C prothorax and thorax dorsal, D anal appendages dorsolateral, E anal appendages lateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253819/files/figure.png" pageId="10" pageNumber="267">Fig. 7</figureCitation>
|
||
D–F): Ochreous, cerci slightly shorter than paraprocts. Cerci with ventromedial tooth that is rather difficult to see from lateral view, obvious from dorsolateral and dorsal views.
|
||
</paragraph>
|
||
<paragraph id="585236EDA672FFB1FF28FB26FDF81500" blockId="10.[151,1437,878,1875]" pageId="10" pageNumber="267">
|
||
Genital ligula (
|
||
<figureCitation id="C0D62A68A672FFB1FE80FB26FE0A14D8" box="[367,434,1202,1227]" captionStart="FIGURE 8" captionStartId="10.[151,250,787,809]" captionTargetBox="[205,1383,199,758]" captionTargetId="figure@10.[198,1388,193,765]" captionTargetPageId="10" captionText="FIGURE 8. Genital ligulae as reconstructed from micro-CT (orientation bars: dl dorsolateral, l lateral, p posterior, v ventral, vl ventrolateral). A – B Genital ligula of C. pyriformis holotype, B – C genital ligula of C. cyanomelas." httpUri="https://zenodo.org/record/253820/files/figure.png" pageId="10" pageNumber="267">Fig. 8</figureCitation>
|
||
A–B, Supplementary
|
||
<figureCitation id="C0D62A68A672FFB1FD43FB26FD4814D8" box="[684,752,1202,1227]" captionStart="FIGURE 4" captionStartId="6.[151,250,1562,1584]" captionTargetBox="[317,1270,193,1519]" captionTargetId="figure@6.[316,1270,193,1541]" captionTargetPageId="6" captionText="FIGURE 4. Coeliccia hainanense Lectotype. A head dorsolateral, B prothorax and thorax dorsolateral, C anal appendages dorsal, D prothorax and thorax dorsal, E anal appendages dorsolateral, F appendages lateral, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253816/files/figure.png" pageId="10" pageNumber="267">Fig. 4</figureCitation>
|
||
): First segment simple, distally curved; second segment with internal, but without terminal fold; third segment with two laterally placed flagella. Ends of the flagella turned inwards and thickened in lateral view.
|
||
</paragraph>
|
||
<paragraph id="585236EDA672FFB1FF28FAB4FBD01524" blockId="10.[151,1437,878,1875]" box="[199,1128,1310,1335]" pageId="10" pageNumber="267">
|
||
<emphasis id="6A99EAFFA672FFB1FF28FAB4FE0D1525" box="[199,437,1311,1335]" italics="true" pageId="10" pageNumber="267">Measurements (mm):</emphasis>
|
||
Total length: 42, Abd.: 36, Wingspan: 43, Fw/Hw (right): 21.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="10F76566A672FFB1FF28FAD6FEAC1740" pageId="10" pageNumber="267" type="discussion">
|
||
<paragraph id="585236EDA672FFB1FF28FAD6FCD31620" blockId="10.[151,1437,878,1875]" pageId="10" pageNumber="267">
|
||
<emphasis id="6A99EAFFA672FFB1FF28FAD6FE831548" bold="true" box="[199,315,1346,1371]" pageId="10" pageNumber="267">Remarks.</emphasis>
|
||
The original description of the
|
||
<typeStatus id="8756884FA672FFB1FD4FFAD6FCBA1548" box="[672,770,1346,1371]" pageId="10" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="9FED4D6EA672FFB1FCC7FAD7FBCB1548" authority="Laidlaw 1932" authorityName="Laidlaw" authorityYear="1932" box="[808,1139,1346,1371]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="6A99EAFFA672FFB1FCC7FAD7FC061548" box="[808,958,1347,1371]" italics="true" pageId="10" pageNumber="267">C. pyriformis</emphasis>
|
||
(
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FC22FAD6FBD31548" author="Laidlaw" box="[973,1131,1346,1371]" pageId="10" pageNumber="267" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw 1932</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
does not mention the pale spots between lateral ocelli and base of antennae. Additionally,
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FC6DFAF2FBD9156C" author="Laidlaw" box="[898,1121,1382,1407]" pageId="10" pageNumber="267" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw (1932: 26)</bibRefCitation>
|
||
described two “...pyriform marks of pale blue…” dorsally on each side of the synthorax. He noted that these markings “...faded...the whole dorsum being uniformly black...” (
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FDDFFA3AFCBB15D4" author="Laidlaw" box="[560,771,1454,1479]" pageId="10" pageNumber="267" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw 1932: 26</bibRefCitation>
|
||
) after a while. However, during the examination of the
|
||
<typeStatus id="8756884FA672FFB1FF78FA46FF4415F8" box="[151,252,1490,1515]" pageId="10" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
, we noted that the markings are still clearly visible, when the specimen is viewed under good lighting. Furthermore, the markings on the synthorax do not have a pyriform shape but are rather simple inwardly curved stripes (cf.
|
||
<figureCitation id="C0D62A68A672FFB1FEFBF98EFEDA1620" box="[276,354,1562,1587]" captionStart="FIGURE 11" captionStartId="13.[151,250,1631,1653]" captionTargetBox="[316,1270,409,1608]" captionTargetId="figure@13.[316,1270,409,1609]" captionTargetPageId="13" captionText="FIGURE 11. Photos in life of Coeliccia pyriformis and Coeliccia cyanomelas. A Coeliccia pyriformis, male, Tam Đ ảo, northern Vietnam. B Coeliccia pyriformis, tandem, Tam Đ ảo, northern Vietnam. C Coeliccia pyriformis, male thorax and head dorsal, Tam Đ ảo, northern Vietnam. D Coeliccia cyanomelas, male head and thorax dorsolateral, Tam Đ ảo, northern Vietnam. E Coeliccia cyanomelas, male with four dorsal synthorax markings, Tam Đ ảo, northern Vietnam. F Coeliccia cyanomelas, male with two dorsal synthorax markings, Tam Đ ảo, northern Vietnam. All photos by Sebastién Delonglee." httpUri="https://zenodo.org/record/253823/files/figure.png" pageId="10" pageNumber="267">Fig. 11</figureCitation>
|
||
A–C for photographs in life of
|
||
<taxonomicName id="9FED4D6EA672FFB1FD2AF98FFCE31620" box="[709,859,1563,1587]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="6A99EAFFA672FFB1FD2AF98FFCE31620" box="[709,859,1563,1587]" italics="true" pageId="10" pageNumber="267">C. pyriformis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="585236EDA672FFB1FF28F9AAFDAE168C" blockId="10.[151,1437,878,1875]" pageId="10" pageNumber="267">
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FF28F9AAFE1A1644" author="Laidlaw" box="[199,418,1598,1623]" pageId="10" pageNumber="267" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw (1932: 26)</bibRefCitation>
|
||
declared that the superior appendages did not have any “...marked projection...”. This is, however, not the case. There is a ventromedial tooth that is rather difficult to see from lateral view, but obvious from dorsolateral and dorsal view.
|
||
</paragraph>
|
||
<paragraph id="585236EDA672FFB1FF28F93EFEAC1740" blockId="10.[151,1437,878,1875]" pageId="10" pageNumber="267">
|
||
Comparison of the
|
||
<typeStatus id="8756884FA672FFB1FE4CF93EFDBD16D0" box="[419,517,1706,1731]" pageId="10" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="9FED4D6EA672FFB1FDC1F93FFD7D16D0" box="[558,709,1707,1731]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="6A99EAFFA672FFB1FDC1F93FFD7D16D0" box="[558,709,1707,1731]" italics="true" pageId="10" pageNumber="267">C. pyriformis</emphasis>
|
||
</taxonomicName>
|
||
with the original description of
|
||
<taxonomicName id="9FED4D6EA672FFB1FBD8F93FFB3716D1" box="[1079,1167,1707,1730]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="acco">
|
||
<emphasis id="6A99EAFFA672FFB1FBD8F93FFB3716D1" box="[1079,1167,1707,1730]" italics="true" pageId="10" pageNumber="267">C. acco</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FB53F93EFAD316D0" author="Asahina" box="[1212,1387,1706,1731]" pageId="10" pageNumber="267" refString="Asahina, S. (1997) Records of the northernVietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 5: Coenagrionidae, Protoneuridae, Platycnemididae. Bulletin of the National Science Museum, Tokyo (A), 23, 17 - 34." type="journal article" year="1997">Asahina (1997)</bibRefCitation>
|
||
and the recent redescription of this species, based on new material, by
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FC4CF95AFB1716F4" author="Steinhoff" box="[931,1199,1742,1767]" pageId="10" pageNumber="267" refString="Steinhoff, P. O. M. & Do, M. C. (2013) Notes on some Coeliccia - species from Vietnam. Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
, shows that
|
||
<taxonomicName id="9FED4D6EA672FFB1FAACF95BFA2416F4" box="[1347,1436,1743,1767]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="acco">
|
||
<emphasis id="6A99EAFFA672FFB1FAACF95BFA2416F4" box="[1347,1436,1743,1767]" italics="true" pageId="10" pageNumber="267">C. acco</emphasis>
|
||
</taxonomicName>
|
||
agrees with
|
||
<taxonomicName id="9FED4D6EA672FFB1FEF0F967FE0D1718" box="[287,437,1779,1803]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="6A99EAFFA672FFB1FEF0F967FE0D1718" box="[287,437,1779,1803]" italics="true" pageId="10" pageNumber="267">C. pyriformis</emphasis>
|
||
</taxonomicName>
|
||
in all diagnostic characters. Only the number of Px given for
|
||
<taxonomicName id="9FED4D6EA672FFB1FB82F967FB7B1719" box="[1133,1219,1779,1802]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="acco">
|
||
<emphasis id="6A99EAFFA672FFB1FB82F967FB7B1719" box="[1133,1219,1779,1802]" italics="true" pageId="10" pageNumber="267">C. acco</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FB01F966FF59173C" author="Steinhoff" pageId="10" pageNumber="267" refString="Steinhoff, P. O. M. & Do, M. C. (2013) Notes on some Coeliccia - species from Vietnam. Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
differ from the
|
||
<typeStatus id="8756884FA672FFB1FE7DF882FE4C173C" box="[402,500,1814,1839]" pageId="10" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
<taxonomicName id="9FED4D6EA672FFB1FE15F883FD37173C" box="[506,655,1815,1839]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="6A99EAFFA672FFB1FE15F883FD37173C" box="[506,655,1815,1839]" italics="true" pageId="10" pageNumber="267">C. pyriformis</emphasis>
|
||
</taxonomicName>
|
||
, which has 3 Px less in all wings. Therefore we propose the following synonymy:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="10F76566A672FFB1FF78F817FD2517AC" pageId="10" pageNumber="267" type="reference_group">
|
||
<paragraph id="585236EDA672FFB1FF78F817FD9D1788" blockId="10.[151,669,1922,1983]" box="[151,549,1922,1947]" pageId="10" pageNumber="267">
|
||
<taxonomicName id="9FED4D6EA672FFB1FF78F817FD9D1788" ID-CoL="WQD3" authority="Laidlaw, 1932" authorityName="Laidlaw" authorityYear="1932" box="[151,549,1922,1947]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="6A99EAFFA672FFB1FF78F817FEC61788" box="[151,382,1923,1947]" italics="true" pageId="10" pageNumber="267">Coeliccia pyriformis</emphasis>
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FE6AF816FD9D1788" author="Laidlaw" box="[389,549,1922,1947]" pageId="10" pageNumber="267" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw, 1932</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="585236EDA672FFB1FF71F833FD2517AC" blockId="10.[151,669,1922,1983]" box="[158,669,1958,1983]" pageId="10" pageNumber="267">
|
||
Syn.
|
||
<taxonomicName id="9FED4D6EA672FFB1FF38F833FD9117AC" ID-CoL="WQBM" authority="Asahina, 1997" authorityName="Asahina" authorityYear="1997" box="[215,553,1958,1983]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="267" phylum="Arthropoda" rank="species" species="acco">
|
||
<emphasis id="6A99EAFFA672FFB1FF38F833FEC717AC" box="[215,383,1959,1983]" italics="true" pageId="10" pageNumber="267">Coeliccia acco</emphasis>
|
||
<bibRefCitation id="3C7C4B1CA672FFB1FE6AF832FD9117AC" author="Asahina" box="[389,553,1958,1983]" pageId="10" pageNumber="267" refString="Asahina, S. (1997) Records of the northernVietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 5: Coenagrionidae, Protoneuridae, Platycnemididae. Bulletin of the National Science Museum, Tokyo (A), 23, 17 - 34." type="journal article" year="1997">Asahina, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="71AA5784A672FFB1FDDAF832FD2517AC" box="[565,669,1958,1983]" pageId="10" pageNumber="267">
|
||
<emphasis id="6A99EAFFA672FFB1FDDAF832FD2517AC" bold="true" box="[565,669,1958,1983]" pageId="10" pageNumber="267">syn. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |